taxonID	type	description	language	source
DE6D87F0465A8376089DFF22C1980F4B.taxon	description	Pileus (20 –) 40 – 60 (– 95) mm, dark greyish brown, then melleous to pale deer-brown. Lamellae off-white to pale beige or pale grey. Stipe (20 –) 30 – 60 (– 70) × 2 – 6 (– 7) mm, pale yellow-brown, darker towards base with greyish blue or steel blue tones. Context with greyish blue or steel blue tinge at stipe base. Basidiospores (7.0 –) 8.0 – 10 (– 12) × 5.0 – 7.0 μm, (broadly) ellipsoid or ellipsoidfusoid, subovoid, verruculose, rarely with small ridges. Cheilocystidia urticoid, 28 – 49 × (5.0 –) 6.0 – 11 μm. Pileipellis an ixocutis. Caulohymenium developed. Holotype: — HUNGARY, Budapest, RÁkospalota, 29 November 2013 I. Rimóczi (BP 104371; Isotype BRNM 751957). Etymology: — Dedicated to the collector’s mother, who supported his first steps in mycology. Pileus (20 –) 40 – 60 (– 95) mm broad, slightly convex with small central umbo and involute margin when young, then applanate to slightly infundibuliform with indistinct to rather distinct umbo and straight or revolute margin, dull, smooth, rarely slightly felty at centre, otherwise innately fibrillose, especially when old. Margin non-grooved or slightly grooved in old specimens, dark greyish brown when young (and also in wet older basidiocarps), then melleous to pale deer-brown with darker centre. Lamellae medium to almost crowded, 2 – 6 (– 8) mm wide, narrowly to broadly adnate, off-white to pale beige or pale grey (but may have slight blue tinge when old), with even concolorous edge. Stipe (20 –) 30 – 60 (– 70) × 2 – 6 (– 7) mm, straight, cylindrical, sometimes slightly broadened at apex and / or at base, hard in touch, floccose-felty at apex, longitudinally fibrillose to fibrillose-squamulose otherwise, pale yellow-brown, darker towards base with ± distinct greyish blue or steel blue tinge, sometimes blue colour ascending towards stipe apex, at base with poor pallid tomentum and covered with distinct sandy cover. Context very thin and soft in pileus, fibrillose-stuffed, then hollow in stipe, off-white when young, then becoming pale greyish beige under pileipellis and near lamellae, silky white in stipe in very young specimens, then quickly turning pale brownish grey in upper part and darker greyish brown towards base; at base, the colour becomes gradually violaceous-blue and then almost blackish blue at very base; dark blue colour of stipitipellis often runs upwards; bluish colour of the stipe context is also distinct in exsiccates. Smell and taste indistinct. Basidiospores (7.0 –) 8.0 – 10 (– 12) × 5.0 – 7.0 μm, average = 9.0 × 6.0 μm, E = 1.30 – 1.90, Q = (1.43 –) 1.57 – 1.64, (broadly) ellipsoid, ellipsoid-fusoid, or subovoid, verruculose, rarely with small ridges, warts regular or irregular in shape and variable in size, up to 0.5 – 0.75 μm broad and 0.25 – 0.34 μm high, amyloid. Basidia 32 – 41 (– 48) × 9.0 – 13 μm, 4 - spored, clavate. Basidioles 15 – 40 × 4.5 – 13 μm, clavate, cylindrical, rarely subfusoid. Cheilocystidia urticoid, 28 – 49 × (5.0 –) 6.0 – 11 μm, of two types: the exscissa - type more common, basal part often (sub) fusoid; the brevipes - type less frequent, basal part clavate to subvesiculose; apical part of both types 2.0 – 5.0 μm wide at base, subulate to cylindrical, thin-walled, mostly not muricate. Marginal cells 19 – 31 (– 44) × 8.0 – 11 (– 17) μm, clavate, fusoid, subpyriform, utriform, ± thin-walled, regular or irregular. Pleurocystidia absent. Trama hyphae cylindrical, thinwalled, hyaline, non-dextrinoid, up to 15 μm wide. Pileipellis an ixocutis composed of radially arranged, ± thin-walled, gelatinized, non-dextrinoid, up to 9.0 μm wide hyphae; terminal cells adpressed to erect, cylindrical or subclavate, ± thin-walled, obtuse, sometimes slightly irregular, 5.0 – 15 μm wide, ochraceous brown to ochraceous-grey in KOH. Stipitipellis a cutis of cylindrical, parallel, slightly thick-walled, 3.0 – 6.0 μm wide hyphae, pale yellow-brown in KOH. Caulohymenium of (1) 20 – 40 (– 55) × (6.0 –) 9.0 – 12 μm, clavate, cylindrical, thin-walled cells, (2) 32 – 46 × 8.0 – 13 μm, urticoid cystidia similar to cheilocystidia, and (3) 24 – 32 × 10 – 13 μm, caulobasidia, 4 - spored, cylindrical to clavate; urticoid caulocystidia and caulobasidia may be rare or even absent. Clamp connections absent. Habitat: — Single or in small groups on sandy soil in Festucetum vaginatae stipetosum, Junipero - Populetum albae, and secondary Brometum tectorum associations. All up-to-now known localities belong to the Eupannonicum floristical district of the Pannonian floristical province. Typical for them are loose sandy soils poor in humus (1 – 2 %), with xeric surface soils rich in chalk. Material studied: — HUNGARY. Budapest, RÁkospalota, 29 November 2013, I. Rimóczi (holotype BP 104371, isotype BRNM 751957). Pest County: Kiskunság National Park, Örkény, 30 September 2005, I. Rimóczi (BRNM 751958 and BP 104372). Ibid., 13 November 2012, I. Rimóczi (BP 104373). Bács-Kiskun County: Kiskunság National Park, Orgovány, 14 June 2006, I. Rimóczi (BRNM 751959 and BP 104374). Kiskunság National Park, Fülöpháza, 9 June 2006, I. Rimóczi (BP 104375). Ibid., 9 June 2006, I. Rimóczi and L. Nagy NL- 5202 (Herbarium of the University of Szeged). Remarks: — Melanoleuca juliannae is characterized by a violaceous-blue stipe context, which represents an unique feature in this genus. Additionally it has moderately large (broadly) ellipsoid, ellipsoid-fusoid, subovoid, verruculose basidiospores, urticoid cheilocystidia of two types (exscissa - type most frequent), and a caulohymenium. Is seems to grow exclusively in primary and secondary xerothermic sandy habitats. Because of the urticoid cheilocystidia, Melanoleuca juliannae belongs to subgen. Urticocystis Boekhout (1988: 400). The collection by I. Rimóczi and L. Nagy (NL- 5202) differs from other ones by the presence of both 4 - and 2 - spored basidia and slightly larger basidiospores (8.5 – 10.5 × 6.0 – 7.5 µm, average = 9.4 × 6.7 μm); other characters fully agree with the type specimen.	en	Antonín, Vladimír, Benedek, Imre Rimóczi Lajos, Papp, Viktor, Szarkándi, János Gergő, Dima, Bálint, Nagy, László G., Papp, Tamás, Ďuriška, Ondrej, Tomšovský, Michal (2014): Melanoleuca juliannae (Basidiomycota, Tricholomataceae), a new species from subgen. Urticocystis. Phytotaxa 170 (1): 13-23, DOI: 10.11646/phytotaxa.170.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.170.1.2
DE6D87F0465F8378089DFEA0C4530FB6.taxon	description	It differs from the typical variety by the absence of the blue-violaceous colour in the stipe base. Holotype: — ITALY. Monti Sibillini National Park, Altino di Montemonaco, 20 November 2012, V. Antonín 10.438 (HOLOTYPE, BRNM 751960). Etymology: — Lacking the blue colour in the stipe base. Pileus 25 – 35 mm broad, plano-convex, with obtuse, conical, rather broad but ± distinct central umbo within sometimes depressed centre, margin reflexed and involute, hygrophanous, not translucently striate, smooth, glabrous, entirely dark brown (6 F 5) when young, pallescent to (grey-) brown (6 D 5 - 7 E 5, 6 E 4 - 5, 6 - 7 D 3) from margin. Lamellae moderately close, L = c. (30 –) 55 – 60, l = 3 – 4, shortly emarginate and attached with a short tooth, sinuate, cream coloured with beige reflex, with concolorous, finely pubescent edge. Stipe 28 – 45 × 3 – 6 mm, cylindrical, slightly broadened at apex, cylindrical to slightly clavate-bulbose (up to 7 mm) at base, longitudinally fibrillose, finely pruinose-floccose at apex, lustrous, dark grey-brown (6 E 3 - 6, 7 E-F 3), with brown (6 D 4) apex; basal tomentum white. Context watery offwhite, hollow in stipe, brown to dark brown in stipe base, in stipitipellis colour in stipe cortex, with indistinct or slight fungoid smell and mild taste. Basidiospores (7.0 –) 8.0 – 10 × (5.0 –) 5.5 – 6.5 μm, average = 8.5 × 5.7 μm, E = 1.27 – 1.73, Q = 1.42 – 1.55, (broadly) ellipsoid, ovoid, verruculose with ridges, amyloid. Basidia 28 – 42 × 10.5 – 13 μm, 4 - spored, clavate. Basidioles 15 – 45 × 4.0 – 12 μm, cylindrical, clavate. Cheilocystidia 35 – 50 × 4.5 – 12 μm, urticoid of two types, the brevipes - type more frequent than the exscissa - type, basal part (sub) cylindrical or fusoid, sometimes irregular, apical part 2.5 – 4.0 μm wide, almost cylindrical to subulate, thin-walled. Pleurocystidia not seen. Trama hyphae cylindrical to subinflated, thin-walled, smooth or minutely incrusted, non-dextrinoid, up to 20 μm wide. Pileipellis an ixocutis, transient to ± ixotrichoderm at centre, composed of ± interwoven, cylindrical, thin-walled, non-dextrinoid, up to 7.0 (– 9.0) μm wide hyphae; terminal cells adpressed to (sub) erect, cylindrical, (narrowly) clavate, subfusoid, obtuse, thin-walled, up to 9.0 μm wide. Stipitipellis a cutis of cylindrical, parallel, ± thin-walled, non-dextrinoid, up to 7.0 μm wide hyphae. Caulohymenium of (1) 25 – 55 × 8.0 – 9.0 μm, clavate, subcylindrical, thin-walled cells, and (2) 68 – 71 × 8.0 μm, urticoid cystidia similar to cheilocystidia, sometimes rare or absent (?). Clamp connections absent. Habitat: — On xeric alcaline (calcareous and serpentinite) soils in a montane pasture with Juniperus communis and J. oxycedrus, in grass in a pasture, and in a grassy xerothermic steppe. Material studied: — CZECH REPUBLIC. Hrubšice, Nad řekami Nature Reserve, elev. 240 – 260 m, 6 November 2010, V. Antonín 10.377 (BRNM 751961). Brno-Slatina, Stránská skála National Nature Reserve, elev. 290 m, 4 December 2006, R. Schles (BRNM 751962). ITALY, Monti Sibillini National Park, Altino di Montemonaco, elev. 1220 – 1320 m, 20 November 2012, V. Antonín 10.438 (holotype BRNM 751960). Remarks: — Melanoleuca juliannae var. decolorans is both macro- and microscopically similar to the type variety. The main differential character is the absence on the blue colour in the stipe base.	en	Antonín, Vladimír, Benedek, Imre Rimóczi Lajos, Papp, Viktor, Szarkándi, János Gergő, Dima, Bálint, Nagy, László G., Papp, Tamás, Ďuriška, Ondrej, Tomšovský, Michal (2014): Melanoleuca juliannae (Basidiomycota, Tricholomataceae), a new species from subgen. Urticocystis. Phytotaxa 170 (1): 13-23, DOI: 10.11646/phytotaxa.170.1.2, URL: http://dx.doi.org/10.11646/phytotaxa.170.1.2
