taxonID	type	description	language	source
0F6484DEEBB85FEAA4E6B11DFA8FD882.taxon	etymology	Etymology. The specific name Occidozyga lingnanica refers to the lingnan region, a geographic area covering Guangdong, Guangxi, and Hainan in southeastern China, where this new frog species occurs in. This specific name is also dedicated to the former Lingnan University (1888 - 1952) that was incorporated into Sun Yat-sen University after 1953.	en	Lyu, Zhi-Tong, Wang, Jian, Zeng, Zhao-Chi, Luo, Lin, Zhang, Yan-Wu, Guo, Chun-Peng, Ren, Jin-Long, Qi, Shuo, Mo, Yun-Ming, Wang, Ying-Yong (2022): Taxonomic clarifications on the floating frogs (Anura: Dicroglossidae: Occidozyga sensu lato) in southeastern China. Vertebrate Zoology 72: 495-512, DOI: http://dx.doi.org/10.3897/vz.72.e80019, URL: http://dx.doi.org/10.3897/vz.72.e80019
0F6484DEEBB85FEAA4E6B11DFA8FD882.taxon	diagnosis	Diagnosis. (1) Body stocky, size small, SVL 19.9 - 22.1 mm in males (n = 8) and 26.8 - 28.8 mm in females (n = 3); (2) snout short triangular shaped; (3) nostrils located laterally; (4) eye orientation laterally; (5) canthus rostralis visible, rounded; (6) loreal region vertical, not concave / convex; (7) tongue wide and swollen, unnotched, rounded distally, lingual papilla absent; (8) vomerine ridge and vomerine teeth absent; (9) supratympanic fold distinct and raised, slightly curved on the temporal region; (10) tympanum hidden, edge invisible; (11) relative finger lengths II = I <IV <III, relative toe lengths I <II <V <III <IV; (12) fingers without webs, toes with two third webs; (13) heels not meeting, tibio-tarsal articulation reaching at the posterior margin of supratympanic fold; (14) tarsal fold absent.	en	Lyu, Zhi-Tong, Wang, Jian, Zeng, Zhao-Chi, Luo, Lin, Zhang, Yan-Wu, Guo, Chun-Peng, Ren, Jin-Long, Qi, Shuo, Mo, Yun-Ming, Wang, Ying-Yong (2022): Taxonomic clarifications on the floating frogs (Anura: Dicroglossidae: Occidozyga sensu lato) in southeastern China. Vertebrate Zoology 72: 495-512, DOI: http://dx.doi.org/10.3897/vz.72.e80019, URL: http://dx.doi.org/10.3897/vz.72.e80019
0F6484DEEBB85FEAA4E6B11DFA8FD882.taxon	description	Description of holotype. SYS a 008846, adult male (Fig. 5). Body stocky, small-sized, SVL 21.4 mm. Head longer than wide (HDW / HDL 0.97), plat above; pineal ocellus distinct; snout short triangular shaped, distinctly protruding beyond lower jaw, tip of snout rounded in dorsal view and profile; canthus rostralis rounded, loreal region oblique, not concave or convex; nostril rounded, laterally, not raised, at the middle between tip of snout and eye; eye orientation laterally, pupil diamond shaped; interorbital space distinctly narrower than internarial distance; tympanum hidden, edge invisible; vomerine ridge and teeth absent; tongue wide and swollen, unnotched, rounded distally, lingual papilla absent. Forelimbs short, lower arm 15 % of SVL and hand 25 % of SVL; fingers relatively thin and long, relative finger lengths II = I <IV <III; tips of fingers rounded, not dilated and without disks; fingers without webs and fringes; subarticular tubercles present at the bases of each finger, prominent and rounded; supernumerary tubercles absent; inner and outer palmar tubercles prominent and rounded. Hind limbs robust, tibia 44 % of SVL and foot 63 % of SVL; heels not meeting when hind limbs flexed at right angles to the axis of the body; tibio-tarsal articulation reaching at the posterior margin of supratympanic fold when hind limb is stretched along the side of the body; toes distinctly long and thin, relative lengths I <II <V <III <IV; tips of toes rounded, dilated into rounded disks; toes with two third webs, metatarsal web present, distinct lateral fringes on lateral edges of toes I and V; subarticular tubercles rounded, prominent; inner metatarsal tubercle large and long-elliptic, distinctly raised, length triple the width; outer metatarsal tubercle absent; tarsal fold absent. Dorsal surface relatively rough, large tubercles scattering on skin of dorsum, flanks, and dorsal limbs, not bearing spinules on the dorsal skin; a faint fold across head between orbits; supratympanic fold distinct, raised, extending from the posterior corner of the eye, slightly curved on the temporal region, to the previous shoulder; dorsolateral fold absent. Ventral surface with large flattened tubercles; a fold across breast; dense granules on the ventral tarsi.	en	Lyu, Zhi-Tong, Wang, Jian, Zeng, Zhao-Chi, Luo, Lin, Zhang, Yan-Wu, Guo, Chun-Peng, Ren, Jin-Long, Qi, Shuo, Mo, Yun-Ming, Wang, Ying-Yong (2022): Taxonomic clarifications on the floating frogs (Anura: Dicroglossidae: Occidozyga sensu lato) in southeastern China. Vertebrate Zoology 72: 495-512, DOI: http://dx.doi.org/10.3897/vz.72.e80019, URL: http://dx.doi.org/10.3897/vz.72.e80019
0F6484DEEBB85FEAA4E6B11DFA8FD882.taxon	description	Male secondary sexual characteristics. Male with a single subgular vocal sac; in breeding season, a single, light yellow, swollen, and granular nuptial pad on the dorsal surface of finger I, nuptial spinules invisible. Males (SVL 19.9 - 22.1 mm) distinctly smaller than females (SVL 26.8 - 28.8 mm) (Fig. 4 C, D).	en	Lyu, Zhi-Tong, Wang, Jian, Zeng, Zhao-Chi, Luo, Lin, Zhang, Yan-Wu, Guo, Chun-Peng, Ren, Jin-Long, Qi, Shuo, Mo, Yun-Ming, Wang, Ying-Yong (2022): Taxonomic clarifications on the floating frogs (Anura: Dicroglossidae: Occidozyga sensu lato) in southeastern China. Vertebrate Zoology 72: 495-512, DOI: http://dx.doi.org/10.3897/vz.72.e80019, URL: http://dx.doi.org/10.3897/vz.72.e80019
0F6484DEEBB85FEAA4E6B11DFA8FD882.taxon	distribution	Distribution. Occidozyga lingnanica sp. nov. can be recognized from multiple localities in Guangdong and Hainan of southeastern China based on the phylogenetic result in this work (Figs 1, 2). The records of O. martensi from Yunnan, southwestern China require further clarifications with morphological examinations (see Discussion).	en	Lyu, Zhi-Tong, Wang, Jian, Zeng, Zhao-Chi, Luo, Lin, Zhang, Yan-Wu, Guo, Chun-Peng, Ren, Jin-Long, Qi, Shuo, Mo, Yun-Ming, Wang, Ying-Yong (2022): Taxonomic clarifications on the floating frogs (Anura: Dicroglossidae: Occidozyga sensu lato) in southeastern China. Vertebrate Zoology 72: 495-512, DOI: http://dx.doi.org/10.3897/vz.72.e80019, URL: http://dx.doi.org/10.3897/vz.72.e80019
BFD1E4E6F78259D29A9F27662F224A79.taxon	materials_examined	Specimens examined. Seven adult males and three adult females. Males CIB 44475 - 44476, from Guangzhou, Guangdong Province, China; male SYS a 000534, from Mt Danxia (25.0347 N, 113.7407 E), Renhua County, Guangdong; males SYS a 008120 - 8121, 8123, 8125, and females SYS a 008122, 8124, from Shimentai Nature Reserve (24.3818 N, 113.3927 E), Yingde City, Guangdong; female SYS a 000488, from Mt Luoyang, Lingshan County, Guangxi Zhuang Autonomous Region, China.	en	Lyu, Zhi-Tong, Wang, Jian, Zeng, Zhao-Chi, Luo, Lin, Zhang, Yan-Wu, Guo, Chun-Peng, Ren, Jin-Long, Qi, Shuo, Mo, Yun-Ming, Wang, Ying-Yong (2022): Taxonomic clarifications on the floating frogs (Anura: Dicroglossidae: Occidozyga sensu lato) in southeastern China. Vertebrate Zoology 72: 495-512, DOI: http://dx.doi.org/10.3897/vz.72.e80019, URL: http://dx.doi.org/10.3897/vz.72.e80019
BFD1E4E6F78259D29A9F27662F224A79.taxon	etymology	Etymology. The specific name obscura means ' obscure' in Latin.	en	Lyu, Zhi-Tong, Wang, Jian, Zeng, Zhao-Chi, Luo, Lin, Zhang, Yan-Wu, Guo, Chun-Peng, Ren, Jin-Long, Qi, Shuo, Mo, Yun-Ming, Wang, Ying-Yong (2022): Taxonomic clarifications on the floating frogs (Anura: Dicroglossidae: Occidozyga sensu lato) in southeastern China. Vertebrate Zoology 72: 495-512, DOI: http://dx.doi.org/10.3897/vz.72.e80019, URL: http://dx.doi.org/10.3897/vz.72.e80019
BFD1E4E6F78259D29A9F27662F224A79.taxon	diagnosis	Diagnosis. (1) Body stocky, size small, SVL 24.2 - 27.5 mm in adult males (n = 7) and 31.5 - 32.2 mm in adult females (n = 3); (2) snout short triangular shaped; (3) nostrils located dorsally; (4) eye orientation vertically; (5) loreal region oblique, not concave or convex; (6) interorbital space narrower than internarial distance; (7) tongue narrow and slender, unnotched, pointed distally, lingual papilla absent; (8) vomerine ridge and vomerine teeth absent; (9) supratympanic fold distinct, raised, and granulated, curved on the temporal region; (10) tympanum hidden, edge invisible; (11) fingers with rudimentary webs, toes with full webs; (12) heels not meeting, tibio-tarsal articulation reaching between the posterior and anterior of the eye.	en	Lyu, Zhi-Tong, Wang, Jian, Zeng, Zhao-Chi, Luo, Lin, Zhang, Yan-Wu, Guo, Chun-Peng, Ren, Jin-Long, Qi, Shuo, Mo, Yun-Ming, Wang, Ying-Yong (2022): Taxonomic clarifications on the floating frogs (Anura: Dicroglossidae: Occidozyga sensu lato) in southeastern China. Vertebrate Zoology 72: 495-512, DOI: http://dx.doi.org/10.3897/vz.72.e80019, URL: http://dx.doi.org/10.3897/vz.72.e80019
BFD1E4E6F78259D29A9F27662F224A79.taxon	description	Re-description. Based on the examined specimens (n = 10). All specimens were similar in morphology. The measurements are given in Table 3. Body stocky, small-sized, SVL 24.2 - 27.5 mm (n = 7) in males and 31.5 - 32.2 mm (n = 3) in females. Head flat above, almost as wide as long (HDW / HDL 0.97 - 1.09, n = 10); pineal ocellus absent; snout short triangular shaped, distinctly protruding beyond lower jaw, tip of snout rounded in dorsal view and profile; canthus rostralis absent, loreal region oblique, not concave or convex; nostril rounded, located dorsally, distinctly raised, closer to the tip of snout than to the eye; eye orientation vertically, pupil diamond shaped; interorbital space narrower than internarial distance; tympanum hidden, edge invisible; vomerine ridge and teeth absent; tongue narrow and slender, unnotched, pointed distally, lingual papilla absent. Forelimbs short, lower arm 13 - 18 % (n = 10) of SVL and hand 26 - 32 % (n = 10) of SVL; fingers distinctly thin and long, relative finger lengths I <II <IV <III; tips of fingers pointed, not dilated, and without disks; distinct lateral fringes on inner and outer sides of each finger, fingers with rudimentary webs, more distinct between fingers I and II; subarticular tubercles present at the bases of each finger, prominent and rounded; supernumerary tubercles absent; inner and outer palmar tubercles prominent and rounded. Hind limbs robust, tibia 41 - 47 % (n = 8) of SVL and foot 66 - 80 % (n = 8) of SVL; heels not meeting when hind limbs flexed at right angles to the axis of the body; tibio-tarsal articulation reaching between the posterior and anterior of the eye when hind limb is stretched along the side of the body; toes distinctly long and thin, relative lengths I <II <III <V <IV; tips of toes pointed, dilated into pear-shaped disks; toes with full webs, metatarsal web present, distinct lateral fringes on lateral edges of toes I and V; subarticular tubercles rounded, prominent; inner metatarsal tubercle large and long-elliptic, slightly raised, length twice the width; outer metatarsal tubercle relatively smaller than the inner metatarsal tubercle, distinctly raised and pointed, length slightly larger than width; inner tarsal fold relatively flat, in contact with the inner metatarsal tubercle; tarsal tubercle large and distinctly raised, close to the tibio-tarsal articulation. Dorsal surface relatively rough, transverse wrinkles and dense tubercles on the dorsum, head, flanks, and limbs; small granules on the dorsal rears of hands and tarsi; not bearing spinules on the dorsal skin; supratympanic fold distinct, raised and granulated, extending from the posterior corner of the eye, curved on the temporal region, to the previous shoulder; dorsolateral fold absent. Ventral surface with large flattened tubercles, denser on the throat and thighs; dense granules on the ventral feet and tarsi.	en	Lyu, Zhi-Tong, Wang, Jian, Zeng, Zhao-Chi, Luo, Lin, Zhang, Yan-Wu, Guo, Chun-Peng, Ren, Jin-Long, Qi, Shuo, Mo, Yun-Ming, Wang, Ying-Yong (2022): Taxonomic clarifications on the floating frogs (Anura: Dicroglossidae: Occidozyga sensu lato) in southeastern China. Vertebrate Zoology 72: 495-512, DOI: http://dx.doi.org/10.3897/vz.72.e80019, URL: http://dx.doi.org/10.3897/vz.72.e80019
BFD1E4E6F78259D29A9F27662F224A79.taxon	description	Male secondary sexual characteristics. Male with a single subgular vocal sac; in breeding season, a single, light grey nuptial pad on the dorsal surface of finger I, nuptial spinules invisible. Males (SVL 24.2 - 27.5 mm) distinctly smaller than females (SVL 31.5 - 32.2 mm) (Fig. 4 A, B).	en	Lyu, Zhi-Tong, Wang, Jian, Zeng, Zhao-Chi, Luo, Lin, Zhang, Yan-Wu, Guo, Chun-Peng, Ren, Jin-Long, Qi, Shuo, Mo, Yun-Ming, Wang, Ying-Yong (2022): Taxonomic clarifications on the floating frogs (Anura: Dicroglossidae: Occidozyga sensu lato) in southeastern China. Vertebrate Zoology 72: 495-512, DOI: http://dx.doi.org/10.3897/vz.72.e80019, URL: http://dx.doi.org/10.3897/vz.72.e80019
BFD1E4E6F78259D29A9F27662F224A79.taxon	distribution	Distribution. Occidozyga obscura comb. nov. can be recognized from several localities of Guangdong and Guangxi, southeastern China (Figs 1, 2), based on the analyzed vouchers in this work. The previous records of O. lima from Guangdong, Guangxi, Hong Kong, Macao, and Fujian, southeastern China are reassigned to O. obscura currently (see Remarks below). The records of O. lima from Yunnan require further clarifications with molecular and morphological vouchers (which might be close to the lineage of O. cf. lima based on the biogeographical similarity).	en	Lyu, Zhi-Tong, Wang, Jian, Zeng, Zhao-Chi, Luo, Lin, Zhang, Yan-Wu, Guo, Chun-Peng, Ren, Jin-Long, Qi, Shuo, Mo, Yun-Ming, Wang, Ying-Yong (2022): Taxonomic clarifications on the floating frogs (Anura: Dicroglossidae: Occidozyga sensu lato) in southeastern China. Vertebrate Zoology 72: 495-512, DOI: http://dx.doi.org/10.3897/vz.72.e80019, URL: http://dx.doi.org/10.3897/vz.72.e80019
