taxonID	type	description	language	source
0B140524D2EA514CA2C74AD27D77DB1C.taxon	description	Figs 1 – 2, 3 – 6, 77	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
0B140524D2EA514CA2C74AD27D77DB1C.taxon	distribution	General distribution. Species endemic to northern Korea.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
786F2F84D4CA5C51A119879215F3539B.taxon	description	Figs 7, 8, 78	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
786F2F84D4CA5C51A119879215F3539B.taxon	distribution	General distribution. Species occurs both in Eastern and Western Palaearctic, it is recorded from Armenia, Finland, eastern Kazakhstan, Norway, Sweden, and Tajikistan. Recorded from Korean Peninsula for the first time.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
E14982E580FE58EDA1A0CF82CFF0F5F7.taxon	description	Figs 9 – 15, 79	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
E14982E580FE58EDA1A0CF82CFF0F5F7.taxon	distribution	General distribution. Wide-spread throughout the whole Palaearctic.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
CAA400C2B20D57F3BE3B467F0DF65595.taxon	description	Figs 16 – 18, 80	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
CAA400C2B20D57F3BE3B467F0DF65595.taxon	distribution	General distribution. Species was recorded from Hokkaido Island (Japan), Sakhalin and Kuril Islands (Russia).	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
89F480D088345A6AAE3597C9ABF11941.taxon	type_taxon	Type species. Dicranota guerini Zetterstedt, 1838 (by monotypy) (Western and Eastern Palaearctic).	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
89F480D088345A6AAE3597C9ABF11941.taxon	description	Description. Medium-sized to largest Dicranota crane flies with body length 6.3 – 9.0 mm and wing length 7.0 – 11.5 mm. Body colouration varies from brownish yellow to dark brown or dark grey. Head. Antenna 12 - or 13 - segmented, longer than in most Dicranota, reaching at least to approximately middle of presutural scutum, sometimes well beyond base of abdomen, if bent backwards. Male antenna often comparatively longer than that of female. Flagellomeres elongate, verticils short, not exceeding length of respective segments. Apical antennomere usually small, shorter than preceding segment. Thorax. Presutural scutum with three or four dark longitudinal stripes, medial stripe could have paler narrow line along middle. Wing with comparatively short radial sector, it is only ~ 2.3 × as long as vein m-cu. Cell r 2 with supernumerary cross-vein. Discal cell usually open due to atrophy of m-m, sometimes closed. Cell m 1 short to very short, distinctly shorter than its stem, sometimes missing. Abdomen. Epandrium of male terminalia comparatively simple, posterior margin nearly straight or concave, without additional lobes at the middle, lateral lobe, if present, small and simple. Gonocoxite with blunt, densely setose dorso-apical lobe. Interbase elongate, species specific, often used for species delimitation. Two pairs of gonostyli. Outer gonostylus elongate, fleshy and setose, inner gonostylus pale, elongate, usually covered with small spines and often bearing subbasal lobe. Ovipositor with long cercus and hypogynial valve, tip of cercus raised upwards, dorsal margin of hypogynial valve setose at base, setae comparatively short.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
510603B8E44B51599D6904211BA43700.taxon	description	Figs 19 – 20, 21 – 24, 81	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
510603B8E44B51599D6904211BA43700.taxon	diagnosis	Diagnosis. Pale yellow species with slightly darkened basal abdominal tergites. Wing translucent, milky with indistinct darker areas surrounding cross-veins, stigma missing. Gonocoxite of male terminalia with narrow subapical cone-shaped lobe on medio-dorsal surface, small subapical setose bump on dorso-lateral surface, small gonostylus and distinct strong darkened horn-shaped paramere.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
510603B8E44B51599D6904211BA43700.taxon	etymology	Etymology. Species is named after that to which it is most closely related, Dicranota perdistincta, and because the most reliable character for its discrimination is the distinct horn- or spine-shaped paramere.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
510603B8E44B51599D6904211BA43700.taxon	description	Description. General body colouration brownish to pale yellow (Figs 19, 20). Body length of male 5.8 mm, of female ~ 7.3 – 7.5 mm. Wing length of male 6.1 – 7.3 mm, of female 6.6 – 7.1 mm. Head. Grey dorsally, paler grey along eye margin, yellowish grey posteriorly, obscure yellow ventrally, covered with sparse yellowish setae. Eyes widely separated, distance between them at base of antenna exceeds length of scape in both sexes. Antenna 0.9 mm long in male, reaching slightly beyond frontal margin of prescutum if bent backwards, 1.1 mm in female. Scape yellowish grey, elongate, pedicel oval, yellowish grey, covered with dark brown setae. Flagellum 12 - segmented, light grey, slightly darker at distal end, basal flagellomere oval, 2 – 5 flagellomeres subglobular, remaining segments oval, decreasing in length towards apex, apical flagellomere approximately as long as preceding. Longest verticils slightly exceed length of respective flagellomeres. Rostrum obscure yellow, palpus greyish yellow covered with short dark brown setae. Labellum pale greyish yellow. Thorax. Pale brownish yellow, covered with sparse greyish pruinosity. Cervical sclerites pale yellow. Pronotum pale, covered with few erect whitish setae. Presutural scutum pale greyish yellow with three indistinct stripes. Stripes not much darker than surrounding area, but they are semi-polished and not covered with pruinosity. Area around stripes covered with bluish grey pruinosity. Tubercular pit missing, pseudosutural fovea indistinct. Prothoracic spiracle surrounded by whitish membrane. Scutal lobe with pale yellow semi-polished area in the middle, margins covered with greyish pruinosity, area between scutal lobes and scutellum whitish. Mediotergite pale. Pleuron pale brownish yellow dorsally, turning totally pale ventrally. Wing (Figs 21, 22) elongate, length / width ratio 3.5, widest slightly before tip of vein CuP, milky, iridescent. Stigma missing. Narrow darker brown areas surround sc-r, tip of Sc, base of Rs, R 2 and supernumerary cross-vein in cell r 1, cross-veins r-m and m-cu, distal margin of discal cell. Veins greyish brown to slightly grey and pale at wing base. Venation: Sc long, reaching wing margin slightly before level of supernumerary cross-vein in cell r 1, sc-r far before the level of Rs base or tip of anal vein, approximately at the middle between humeral vein and base of radial sector. Rs long, slightly> 4 × as long as cross-vein m-cu, usually arcuate at base, but could be also angulate and short spurred. Free end of R 1 very short or missing, reaching wing margin together with R 2. Vein R 2 nearly transverse, supernumerary cross-vein in cell r 1 beyond or at the same level as branching point of R 4 + 5. R 3, R 4, and R 5 nearly parallel to each other. Cell r 3 without stem, R 2 + 3 starting slightly before or at same point as r-m, cell r 4 with short stem, cell itself 3.5 – 4.6 × as long as its stem. Cross-vein r-m distinct, discal cell large, ~ 2.4 × as long as wide, in rare occasions open by atrophy of vein m-m. Cell m 1 short, its stem ≥ 1.3 × as long as cell itself. Cross-vein m-cu at or slightly beyond branching point of M. Veins CuP and A 1 nearly straight just slightly arched before wing margin. Anal angle widely rounded. No brachypterous females were observed. Halter long, its length nearly reaches that of thorax. Length of male halter 1.1 – 1.2 mm, of female 1.0 – 1.2 mm. Stem of halter pale with yellowish base, basal half of knob milky, distal part slightly infuscate. Coxae obscure yellow to pale yellow, depending on specimen. Trochanters of fore and middle legs pale greyish yellow, trochanter of hind leg pale or pale yellow. Femora pale greyish yellow with slightly infuscate distal part. Tibiae greyish yellow with widely darkened distal margin. Basitarsi brownish with darker brown distal end and pale base, remaining tarsomeres dark brown. Male femur I: 3.3 – 3.8 mm long, II: 4.0 mm, III: 4.0 – 4.4 mm, tibia I: 3.4 – 3.8 mm, II: 3.2 – 3.4 mm, III: 3.7 – 4.1 mm, tarsus I: 4.2 – 5.2 mm, II: 3.6 – 3.9 mm, III: 3.8 – 4.6 mm. Female femur I: 3.3 – 4.0 mm long, II: 3.8 mm, III: 4.0 – 4.5 mm, tibia I: 3.0 mm, II: 2.9 mm, III: 2.9 – 3.8 mm, tarsus I: 3.3 mm, II: 2.8 mm, III: 2.9 – 3.7 mm. Claw small and simple, without spines. Abdomen. Obscure yellow. Three basal tergites dark grey medially, margins of darkening vague, lateral margins widely yellow. Remaining tergites and sternites pale yellow, but sclerites are so thin that blackish inner content of guts is visible through and gives dark appearance of the whole abdomen in museum specimens. Abdomen covered with sparse semierect yellowish setae. Male terminalia (Fig. 23) pale yellow, sometimes with weak greyish shade. Posterior margin of epandrium protuberant postero-medially, postero-lateral angle with long blade-shaped lobe, tip of which extended into narrow pale slightly curved caudal lobule. Whole posterior margin of epandrium widely covered with long setae. Gonocoxite elongate, slightly arched, ~ 2.4 × as long as wide, with narrow conical subapical lobe on medio-dorsal surface and small subapical rounded setose bump on dorso-lateral surface. One pair of gonostyli. Gonostylus subglobular with indistinctly protuberant dorsal part (rostrum) covered with long setae. Distal (caudal) part of gonostylus covered with numerous black peg-like spines. Aedeagus short, not visible in dorsal view. Paramere strong, horn-shaped, slightly curved, wider basally, brown to dark brown. Posterior segments of female abdomen generally yellow, concolourous with the rest of abdomen. Tenth tergite pale, cercus greyish yellow with pale yellow distal part, apex acute, slightly raised upwards. Hypogynial valve straight, pale yellow with brownish yellow base. Three spermathecae small, drop-shaped (Fig. 24).	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
510603B8E44B51599D6904211BA43700.taxon	distribution	Distribution. Currently known only from South Korea.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
BF48623D0EC658799C022B942F2E35F6.taxon	description	Figs 25 – 28, 82	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
BF48623D0EC658799C022B942F2E35F6.taxon	distribution	General distribution. Species is endemic to Korean Peninsula.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
B84A68095AA35056A4A5D7DB7CDF03B7.taxon	description	Figs 29 – 33, 83	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
B84A68095AA35056A4A5D7DB7CDF03B7.taxon	distribution	General distribution. Species was recorded from the eastern Kazakhstan, Honshu Island of Japan, and the Far East of Russia.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
A5D968F07E4B5A0A9AE2F2981E95E17B.taxon	description	Figs 34 – 37, 84	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
A5D968F07E4B5A0A9AE2F2981E95E17B.taxon	distribution	General distribution. Species was recorded from northern Korea and the Far East of Russia close to the border with North Korea.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
510A7F361A385E33A41A50352033EEB8.taxon	type_taxon	Type species. Tricyphona lucidipennis Edwards 1921 (original designation) (West Palaearctic).	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
3BD80680773050AB89098145FCA9B0F8.taxon	description	Figs 38 – 43, 85	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
3BD80680773050AB89098145FCA9B0F8.taxon	distribution	General distribution. Japan, Far East of Russia, South Korea, Taiwan.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
ADDAF999BA645354A80556BA6C03D2E9.taxon	description	Figs 44 – 48, 86	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
ADDAF999BA645354A80556BA6C03D2E9.taxon	distribution	General distribution. North Korea and Far East of Russia.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
70315E3BB5C15F16AF0C4817F7907895.taxon	description	Figs 49 – 52, 87	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
70315E3BB5C15F16AF0C4817F7907895.taxon	distribution	General distribution. Korean Peninsula and Far East of Russia.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
DE51D7F2BAEA54B48045AD4A63735AFE.taxon	description	Figs 53 – 57, 88	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
DE51D7F2BAEA54B48045AD4A63735AFE.taxon	distribution	General distribution. North and South Koreas, Far East of Russia.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
326BA982B5E15DD18275889B66148B3B.taxon	description	Figs 58 – 61, 89	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
326BA982B5E15DD18275889B66148B3B.taxon	distribution	General distribution. North and South Koreas.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
7A22AE3DFB6257CF83328330BE6BAA87.taxon	type_taxon	Type species. Rhaphidolabis tenuipes Osten Sacken, 1869 (original designation) (Nearctic).	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
D4CD0FF4EE6A5006AFEBA17423D9BE93.taxon	description	Figs 62 – 65, 90	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
D4CD0FF4EE6A5006AFEBA17423D9BE93.taxon	distribution	General distribution. South Korea, Kyushu and Shikoku islands of Japan.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
3D14342586D95A478D01BB43890E5053.taxon	description	Figs 66 – 68, 91	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
3D14342586D95A478D01BB43890E5053.taxon	diagnosis	Diagnosis. Medium-sized dark brown species. Wing semi-translucent, milky without darker areas except brownish stigma. Antenna 14 - segmented with large apical segment. Epandrium of male genitalia with straight caudal margin bordered with small lateral lobe. Gonocoxite with rounded dorso-apical lobe. Both gonostyli elongate. Gonostyli and gonocoxal lobe covered with short pale spines. Interbase large, flat, with strong spine dorso-caudally. Aedeagus short with blunt apex. Ovipositor yellow with arched cercus and straight hypogynial valve.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
3D14342586D95A478D01BB43890E5053.taxon	etymology	Etymology. The species is named after the Korean entomologist Dr Hong-Yul Seo, who not only studied Korean aphids, but did a lot for other groups of insects. He is a very warm person, our friend and colleague, and with whom we collected together on that cold October day when the holotype of this species was found.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
3D14342586D95A478D01BB43890E5053.taxon	description	Description. General body colouration dark brown. Male body length 10.0 mm, wing length 9.2 mm. Head. Dark greyish brown, narrowly paler along eye margin, covered with sparse semi-erect dark setae dorsally, longer frontally, shorter caudally. Eyes widely separated, distance between them at base of antenna nearly equals length of scape. Male antenna 1.5 mm long, reaching to approx. frontal margin of prescutum if bent backwards, female antenna not reaching frontal margin of prescutum if bent backwards. Scape dark brown, subcylindrical, bearing few erect setae dorsally. Pedicel somewhat paler, slightly widening towards distal end. Flagellum 12 - segmented in both sexes, brown, basal flagellomere slightly elongate, similar to pedicel, succeeding flagellomeres short, subglobular, distal flagellomeres slightly elongate, apical flagellomere large, distinctly longer than preceding segment, especially in female. Verticils short, not reaching length of respective flagellomeres. Rostrum and palpus brown, labellum pale. Thorax. Uniformly dark brown. Cervical sclerites paler. Pronotum with yellowish lateral margin, bearing few erect whitish setae dorsally. Presutural scutum dark brown with three longitudinal stripes. Medial stripe very wide, not reaching suture caudally, area between caudal margin of stripe and suture yellowish. Stripes separated by grey area, covered with erect pale setae. Pseudosutural fovea indistinct. Prothoracic spiracle surrounded by yellowish membrane. Scutal lobe dark brown, area between lobes yellowish. Scutellum yellowish brown. Mediotergite dark brown. Pleuron uniformly dark brown, just ventral margin of katepisternum narrowly blackish. Wing (Fig. 66) elongate, length / width ratio 3.8, widest slightly before tip of vein CuP, semi-translucent, milky, yellowish at base, without any darker spots besides brownish stigma. Veins greyish to brownish, yellowish at wing base. Venation: Sc long, reaching wing margin slightly beyond frontal margin of stigma, far beyond of R 2 + 3 + 4 branching point, sc-r closer to origin of radial sector than to humeral vein. Rs short, 2.3 × as long as cross-vein m-cu, slightly arched. Free end of R 1 short, approximately as long as R 2. Vein R 2 nearly transverse. R 3, R 4, and R 5 nearly straight and parallel to each other. Cell r 3 with distinct stem, which is as long as cross-vein r-m. Discal cell missing due to atrophy of vein m-m. Cell m 1 rather long, its stem 1.8 × as long as cell itself. Cross-vein m-cu half of its own length beyond branching point of M. Vein CuP nearly straight, A 1 slightly arched before wing margin. Anal angle wide and rounded. Halter long, uniformly pale, knob not darker than stem in male, slightly infuscate in female. Length of male halter 1.2 mm. Fore coxa dorsally brown, ventrally obscure yellow, brown area approximately equals yellow. Middle coxa brown with yellowish distal part, posterior coxa entirely brown. Trochanters obscure yellow with narrowly blackened distal margin. Femora brown with widely pale base and darker distal part. Tibiae brown with indistinctly darker distal part. Tarsomeres brown, except dark brown last segment. Male femur I: 3.3 – 3.8 mm long, II: 4.0 mm, III: 4.0 – 4.4 mm, tibia I: 3.4 – 3.8 mm, II: 3.2 – 3.4 mm, III: 3.7 – 4.1 mm, tarsus I: 4.2 – 5.2 mm, II: 3.6 – 3.9 mm, III: 3.8 – 4.6 mm. Female femur I: 3.3 – 4.0 mm long, II: 3.8 mm, III: 4.0 – 4.5 mm, tibia I: 3.0 mm, II: 2.9 mm, III: 2.9 – 3.8 mm, tarsus I: 3.3 mm, II: 2.8 mm, III: 2.9 – 3.7 mm. Claw small, brownish, slightly arched, with subbasal spine. Abdomen. Brown, slightly dusted with grey, basal sternite yellowish. Both tergites and sternites covered with very sparse short pale setae. Male terminalia (Fig. 67) brown with yellowish gonocoxites. Posterior margin of epandrium straight with small serration, covered with short pale setae, lateral margin with small simple, triangle-shaped lobe. Gonocoxite elongate, ~ 2 × as long as wide, with rounded lobe dorso-apically. Mesal margin of lobe covered with sparse short, spine-shaped setae. Two pairs of gonostyli. Outer gonostylus elongate, brown, fleshy and setose, round-apexed. Mesal and caudal margins covered with pale short spines. Inner gonostylus long and narrow, wider at base, slightly arched, pale brown, nearly as long as outer gonostylus. Mesal margin with few long pale spines. Interbase large, wide plate-shaped, dorso-caudal angle extended into large strong spine, ventral margin finely serrate. Aedeagus short, blunt-apexed. Paramere elongate, slightly curved. Ovipositor (Fig. 68) brownish yellow with pale distal part of hypogynial valve. Cercus uniformly brownish yellow, just tip paler. Distal part distinctly narrower, raised upwards, apex blunt. Hypogynial valve straight, brownish yellow at base. Dorsal margin with few short subbasal setae, reaching to ~ 1 / 4 of valve. Two spermathecae small, subglobular.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
3D14342586D95A478D01BB43890E5053.taxon	distribution	Distribution. Currently known only from South Korea, but probably has a wider distribution throughout the country, because both localities are far from each other.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
AEB9B234C009598E96862ED1522B4728.taxon	description	Figs 69 – 71, 92	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
AEB9B234C009598E96862ED1522B4728.taxon	distribution	General distribution. North Korea, Sakhalin Island (Russia).	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
4718333F740A5A669805952302CF19E3.taxon	description	Figs 72 – 76, 93	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
4718333F740A5A669805952302CF19E3.taxon	diagnosis	Diagnosis. Pale yellow species with contrastingly black eyes. Wing semi-translucent, milky without darker areas and without stigma. Male gonostylus with large mesal lobe, posterior margin of epandrium with U-shaped invagination, margins of which usually asymmetrical, interbase large with elongate postero-dorsal and postero-ventral angles, the latter with long curved spine. Female cercus just slightly arched with distinct black base of ventral margin.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
4718333F740A5A669805952302CF19E3.taxon	etymology	Etymology. The species is named after the locality where it was collected, Yeongok-myeon.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
4718333F740A5A669805952302CF19E3.taxon	description	Description. General body colouration pale yellow. Body length of male 5.0 – 5.7 mm, of female 7.3 mm. Wing length of male 5.0 – 6.2 mm, of female 5.9 – 6.4 mm. Head. Greyish yellow, paler posteriorly, covered with sparse short pale setae. Eyes widely separated, distance between them at base of antenna nearly equals length of both basal antennomeres. Antenna 0.9 – 1.0 mm long in male, reaching slightly beyond frontal margin of prescutum if bent backwards, 1.1 mm in female. Scape pale, subcylindrical, ~ 2 × as long as second antennomere, bearing few setae dorsally. Pedicel pear-shaped, slightly darker than scape. Flagellum 13 – 14 - segmented, pale brown, basal flagellomeres oval, distal flagellomeres slightly elongate, apical flagellomere comparatively large, but shorter than preceding segment. Longest verticils nearly as long as respective flagellomeres. Rostrum and palpus yellow. Labellum pale. Thorax. Pale brownish yellow. Cervical sclerites pale. Pronotum somewhat darker, bearing few erect setae dorsally. Presutural scutum uniformly pale brownish yellow, longitudinal stripes missing (stripes could be faded because of preservation in ethanol). Tubercular pit missing, pseudosutural fovea indistinct. Prothoracic spiracle surrounded by pale membrane. Scutal lobe concolourous with presutural scutum. Scutellum paler. Mediotergite pale brownish yellow. Pleuron paler than dorsum, darker areas missing. Wing (Fig. 72) elongate, length / width ratio 4.2, widest approximately at tip of vein CuP, semi-translucent, milky. Stigma and any darker areas missing. Veins greyish to brownish, paler at wing base. Venation: Sc long, reaching wing margin slightly beyond level of branching point of R 2 + 3 + 4, sc-r slightly closer to humeral vein than to origin of radial sector. Rs short, 2.5 × as long as cross-vein m-cu, slightly arched. Free end of R 1 nearly missing, reaching wing margin together with R 2. Vein R 2 oblique. R 3, R 4, and R 5 parallel to each other. Cell r 3 with distinct stem. Cross-vein r-m also distinct. Discal cell missing due to atrophy of vein m-m. Cell m 1 very short, its stem 4 × as long as cell itself. Cross-vein m-cu less than its own length beyond branching point of M. Vein CuP nearly straight, A 1 slightly sinuous. Anal angle long and narrow. Halter long, uniformly pale yellow. Length of male halter 0.8 – 0.9 mm, of female 0.8 – 0.9 mm. Coxae and trochanters pale yellow, distal margin of trochanter narrowly blackened. Femora pale brownish yellow. Tibiae pale yellow with slightly darkened apices. Basitarsi pale yellow at base, turning darker towards distal end, remaining tarsomeres brownish. Male femur I: 3.2 mm long, II: 3.8 mm, III: 3.6 – 4.0 mm, tibia I: 3.7 mm, II: 3.8 mm, III: 3.9 – 4.5 mm, tarsus II: 4.2 mm, III: 4.7 – 4.8 mm. Female femur I: 2.9 mm long, II: 3.4 mm, III: 3.5 – 3.7 mm, tibia I: 4.2 mm, II: 3.5 mm, III: 4.1 mm, tarsus I: 4.3 mm, II: 3.6 mm, III: 3.0 – 3.7 mm. Claw small and simple, nearly straight, without spines. Abdomen. Pale, pregenital segments slightly infuscate. Male terminalia (Figs 73 – 75) pale brownish yellow. Posterior margin of epandrium with U-shaped concavity at middle, bottom of which with few setae starting from small bumps. Setae often not symmetrically arranged on both sides. Margin of concavity extended into larger setose lobe, two or three much smaller lobules could be present further laterally. Often size of larger lobe is different on both sides, number of smaller lobules often varies too, and sometimes they are missing completely. Thus posterior margin of epandrium often, but not always, is asymmetrical. Gonocoxite slightly elongate, 1.4 × as long as wide, approximately egg-shaped, without additional spiny lobe distally, but with few short spine-shaped setae subapically. One pair of gonostyli. Outer part of gonostylus darkened, elongate, basal half of mesal surface strongly swollen, rounded, with darker margin, lateral part elongated, darker brown, covered with small spines on distal two-thirds and apex. Middle of mesal margin with large pale, subglobular lobe bearing small brownish rostral appendage, lower, expanded part with scattered sensilla. Interbase extended into large flattened lobe, postero-dorsal angle of which extended into slightly arched spine. Postero-ventral angle extended into long rostrum, bearing long curved spine, turned backwards. Aedeagus short, not visible in dorsal view, tip shallowly bifid. Paramere elongate, slightly curved. Posterior segments of female abdomen generally yellow, concolourous with the rest of abdomen. Tenth tergite yellow. Cercus (Fig. 76) yellow, slightly arched, tip raised upwards, blunt-apexed with paler distal part and distinctly blackened basal part of ventral margin, dorsal margin slightly infuscate. Hypogynial valve straight, obscure yellow with brownish base and pale distal part. Dorsal margin with few short subbasal setae, reaching to ~ 1 / 4 of valve. Two small subglobular spermathecae.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
4718333F740A5A669805952302CF19E3.taxon	distribution	Distribution. Currently known only from Odaesan National Park, South Korea.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
2C645F50FFE0580E965FD1BDDF698789.taxon	type_taxon	Type species. Dicranota guerini Zetterstedt, 1838 (by monotypy) (Western and Eastern Palaearctic).	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
2C645F50FFE0580E965FD1BDDF698789.taxon	description	Description. Small to medium-sized Pediciidae crane flies with body length 4.3 – 9.0 mm and wing length 5.0 – 11.5 mm. Body colouration varies from pale yellow to dark brown, dark grey, or even black. Head. Rounded posteriorly, vertex with low but wide tubercle. Eyes with short erect setae between ommatidia. Antenna usually short, reaching approximately to frontal margin of prescutum, if bent backwards, sometimes longer in males and reaching posterior margin of first abdominal segment if bent backwards. Number of flagellomeres varies from 10 to 15, most usual number being 11 – 13. Flagellomeres slightly elongate or oval, covered with short pubescence, verticils variable, but usually not exceeding length of respective segments. Apical antennomere usually large, as long or longer than preceding segment. Thorax. Pronotum rather big, covered with erect setae medially. Mesonotal prescutum without tubercular pits, pseudosutural fovea indistinct. Katepisternum often bare, bearing few setae in some species. Meron small. Middle and posterior coxae close to each other. Wing usually long and narrow, often ~ 4 × as long as wide, similar in both sexes, sometimes dimorphic: posterior margin of male wing extended, widest at tip of anal vein, or wing reduced, with some brachypterous females, sometimes brachypterous individuals occurring among both sexes. Some species, especially living in cold climate or at higher altitudes are all brachypterous. Wings of most species without any pattern, even stigma often is missing. Wings of only few species with smaller or larger dark spots, mostly surrounding cross-veins, base of Rs and cord. Venation: arculus present, vein Sc very long, reaching wing margin far beyond branching point of Rs, sc-r long distance before base of Rs, sometimes close to the middle between base of Rs and humeral vein, R 1 short, usually not exceeding R 2 in length. Radial sector short, ≤ 3 × as long as m-cu, often shorter than that. Cell r 3 long with short stem or stem is missing completely. Cell m 1 usually short, much shorter than its stem. Discal cell missing due to reduction of vein m-m, rarely present, but even when present, m-m is weak. Cross-vein m-cu beyond branching point of M. Tip of anal vein far not reaches level of Rs base. Anal angle usually wide. Tibial spurs present on all legs. Claw usually simple, without spines. Abdomen. Male terminalia not enlarged, approximately as wide as rest abdominal segments. Epandrium often with medial and lateral lobes, lateral lobes could be elongate and complicated. Gonocoxite simple or with larger or smaller dorsal lobe at apex. Usually this lobe covered with small dark spines. One or two pairs of gonostyli, outer gonostylus usually bearing lots of small spines, interbase large, often complicate. Aedeagus short and simple, paramere elongate. Ovipositor with long cercus and hypogynial valve, tip of cercus raised upwards, dorsal margin of hypogynial valve usually covered with long and strong setae. Most females with two spermathecae, some with three.	en	Podenas, Sigitas, Yum, Jin Whoa, Ahn, Neung-Ho, Kim, Soen Yi, Kim, Jisoo, Podeniene, Virginija (2025): Dicranota Zetterstedt, 1838 crane flies (Diptera, Pediciidae) of Korea. ZooKeys 1253: 1-72, DOI: 10.3897/zookeys.1253.146576
