taxonID	type	description	language	source
E35A87B9871AFF81C5C7FECCFB95FCF4.taxon	description	Figs 1 – 12	en	Nabozhenko, M. V. (2025): COMPOSITION AND DISTRIBUTION OF THE GENUS ALLECULA FABRICIUS, 1801 (COLEOPTERA: TENEBRIONIDAE) IN RUSSIA WITH DESCRIPTION OF A NEW SPECIES. Far Eastern Entomologist 518: 1-13, DOI: 10.25221/fee.518.1, URL: https://doi.org/10.25221/fee.518.1
E35A87B9871AFF81C5C7FECCFB95FCF4.taxon	materials_examined	MATERIAL EXAMINED (only for the Russian Far East). Russia: Primorsky Krai; Shkotovsky Distr., 30 km W of Partizansk, Anisimovka village, light trap, 24. VIII 2022, 1 ♂ (I. D. Solodkiy) (ZIN); same locality and collector, 13. VIII 2023, 2 ♂, 1 ♀ (ZIN); Partizansky Distr., Vasil’evka village, 7. VIII 2023 1 ♂ (I. D. Solodkiy) (PCMN); Vladivostok, Okeansky ridge, light trap, 11. VIII 2023, 2 ♂, 1 ♀, (S. Veriga) (PCMN); Krasnoarmeysky Distr., Dersu env., 45 ° 45 ʹN, 135 ° 19 ʹE, 9 – 15. VIII 2023, 3 ♂, 2 ♀, (M. E. Sergeev) (PCMN). NOTES. We studied specimens of A. ussuriensis from the type locality and several other localities and didn’t find any differences between one male from Crimea (listed as Allecula morio in Nabozhenko & Gadaborsheva (2023 )) and Far Eastern specimens. They have the same OI, measurements of antennomeres, the shape and puncturation of the pronotum, dull pronotum and elytra and shiny head. The shape of the male VIII inner sternite, spiculum gastrale and the aedeagus are identical in both species. The shape of the aedeagus in lateral view is slightly different in specimen from Crimea, due to desiccation of the parameres. There are two explanations for this gap in the range: the introduction of the species from the Far East with timber or historical disjunction. Disjuctions in the nemoral animal ranges in the Palaearctic are well known, including among insects. Gorodkov (1984) called such ranges “ Trans-Eurasian nemoral ”. He associated these disjunctions with the disappearance of broad-leaved forests in Siberia during the glacial periods in Pleistocene. Dubatolov and Kosterin (1998, 2000) discussed in details disjunctions of nemoral ranges on the example of Lepidoptera. They called such ranges as amphipalaearctic and showed that the ranges of elm, linden and oak were restored in Siberia during interglacial periods and were continuous from Atlantic to Pacific oceans.	en	Nabozhenko, M. V. (2025): COMPOSITION AND DISTRIBUTION OF THE GENUS ALLECULA FABRICIUS, 1801 (COLEOPTERA: TENEBRIONIDAE) IN RUSSIA WITH DESCRIPTION OF A NEW SPECIES. Far Eastern Entomologist 518: 1-13, DOI: 10.25221/fee.518.1, URL: https://doi.org/10.25221/fee.518.1
E35A87B9871AFF81C5C7FECCFB95FCF4.taxon	distribution	DISTRIBUTION. This species with the amphipalaearctic range is widely distributed in the Russian Far East (south of Khabarovsky Krai and Primorsky Krai), Korean peninsula and Ulleungdo Island (Lee et al., 2024), NE China (Mařan, 1940). One male was collected in Crimea (Nabozhenko & Gadaborsheva, 2023).	en	Nabozhenko, M. V. (2025): COMPOSITION AND DISTRIBUTION OF THE GENUS ALLECULA FABRICIUS, 1801 (COLEOPTERA: TENEBRIONIDAE) IN RUSSIA WITH DESCRIPTION OF A NEW SPECIES. Far Eastern Entomologist 518: 1-13, DOI: 10.25221/fee.518.1, URL: https://doi.org/10.25221/fee.518.1
E35A87B9871DFF81C5C7FC05FC39FA1D.taxon	description	Figs 13 – 19	en	Nabozhenko, M. V. (2025): COMPOSITION AND DISTRIBUTION OF THE GENUS ALLECULA FABRICIUS, 1801 (COLEOPTERA: TENEBRIONIDAE) IN RUSSIA WITH DESCRIPTION OF A NEW SPECIES. Far Eastern Entomologist 518: 1-13, DOI: 10.25221/fee.518.1, URL: https://doi.org/10.25221/fee.518.1
E35A87B9871DFF81C5C7FC05FC39FA1D.taxon	materials_examined	MATERIAL EXAMINED. (ZIN), Russia: Stavropol Krai, Lermontov env., Beshtau Mt., light trap, 25 – 27. VII 2015, 1 ♂ (E. N. Terskov); Abkhazia: Tsadripsh near Russian border, 15. VIII 2011, 1 ♂ (E. A. Khachikov). NOTES. The shape of the aedeagus imaged (outline drawing) in Novák et al. (2011) differs from those in our specimen from Russia and similar to A. olexai dorsally and to European specimens of A. rhenana laterally. This difference is probably due to a slight deformation of the parameres during drying. The photo presented here was taken with the newly dissected aedeagus.	en	Nabozhenko, M. V. (2025): COMPOSITION AND DISTRIBUTION OF THE GENUS ALLECULA FABRICIUS, 1801 (COLEOPTERA: TENEBRIONIDAE) IN RUSSIA WITH DESCRIPTION OF A NEW SPECIES. Far Eastern Entomologist 518: 1-13, DOI: 10.25221/fee.518.1, URL: https://doi.org/10.25221/fee.518.1
E35A87B9871DFF81C5C7FC05FC39FA1D.taxon	distribution	DISTRIBUTION. Widespread in Europe east to the Caucasus and Iran (Novák, 2023). This species was listed from Transcaucasia (Dubrovina, 1978; Dubrovina et al., 1979): Abkhazia (Sukhum) and Georgia (Borjomi). In Russia, A. rhenana was known only from Ubinskaya (Dubrovina, 1978) and Guzeripl’ of Krasnodar Krai (Nikitsky et al., 2008; Nabozhenko et al., 2010). Beshtau (Western Ciscaucasia) is the easternmost locality of the species in the North Caucasus and Russia.	en	Nabozhenko, M. V. (2025): COMPOSITION AND DISTRIBUTION OF THE GENUS ALLECULA FABRICIUS, 1801 (COLEOPTERA: TENEBRIONIDAE) IN RUSSIA WITH DESCRIPTION OF A NEW SPECIES. Far Eastern Entomologist 518: 1-13, DOI: 10.25221/fee.518.1, URL: https://doi.org/10.25221/fee.518.1
E35A87B9871DFF80C5C7FA3DFCFEFECE.taxon	distribution	DISTRIBUTION. This species is known only from the type locality by the type series, despite that Dubrovin (1992) indicated in the original description also Khabarovsky Krai of Russia and North-Eastern China.	en	Nabozhenko, M. V. (2025): COMPOSITION AND DISTRIBUTION OF THE GENUS ALLECULA FABRICIUS, 1801 (COLEOPTERA: TENEBRIONIDAE) IN RUSSIA WITH DESCRIPTION OF A NEW SPECIES. Far Eastern Entomologist 518: 1-13, DOI: 10.25221/fee.518.1, URL: https://doi.org/10.25221/fee.518.1
E35A87B9871FFF82C5C7FAECFCFEFECE.taxon	description	Figs 20 – 28	en	Nabozhenko, M. V. (2025): COMPOSITION AND DISTRIBUTION OF THE GENUS ALLECULA FABRICIUS, 1801 (COLEOPTERA: TENEBRIONIDAE) IN RUSSIA WITH DESCRIPTION OF A NEW SPECIES. Far Eastern Entomologist 518: 1-13, DOI: 10.25221/fee.518.1, URL: https://doi.org/10.25221/fee.518.1
E35A87B9871FFF82C5C7FAECFCFEFECE.taxon	materials_examined	TYPE MATERIAL EXAMINED. Holotype – ♂ (ZIN): “ Yakovlevka Spas. u, Ussur kr. 12. VIII 1926, D’yakonov Filip’ev ” (now Russia: Primorsky Krai, Yakovlevka Distr., Yakovlevka village, 44 ° 25 ′ 37 ″ N, 133 ° 28 ′ 47 ″ E). Paratypes, 3 specimens: the same locality and collectors, but different dates: “ 15. VIII 1926 ” (♂), “ 26. VIII 1926 ” (♂), and “ 2. IX 1926 ” (♀).	en	Nabozhenko, M. V. (2025): COMPOSITION AND DISTRIBUTION OF THE GENUS ALLECULA FABRICIUS, 1801 (COLEOPTERA: TENEBRIONIDAE) IN RUSSIA WITH DESCRIPTION OF A NEW SPECIES. Far Eastern Entomologist 518: 1-13, DOI: 10.25221/fee.518.1, URL: https://doi.org/10.25221/fee.518.1
E35A87B9871FFF82C5C7FAECFCFEFECE.taxon	distribution	DISTRIBUTION. This species is known only from the type locality by the type series, despite that Dubrovin (1992) indicated in the original description also Khabarovsky Krai of Russia and North-Eastern China.	en	Nabozhenko, M. V. (2025): COMPOSITION AND DISTRIBUTION OF THE GENUS ALLECULA FABRICIUS, 1801 (COLEOPTERA: TENEBRIONIDAE) IN RUSSIA WITH DESCRIPTION OF A NEW SPECIES. Far Eastern Entomologist 518: 1-13, DOI: 10.25221/fee.518.1, URL: https://doi.org/10.25221/fee.518.1
E35A87B9871EFF82C5C7FE0BFE54FD86.taxon	description	Figs 29 – 35 NOTES. This species is known in Russia from Kaliningrad Region (Alekseev & Bukejs, 2010).	en	Nabozhenko, M. V. (2025): COMPOSITION AND DISTRIBUTION OF THE GENUS ALLECULA FABRICIUS, 1801 (COLEOPTERA: TENEBRIONIDAE) IN RUSSIA WITH DESCRIPTION OF A NEW SPECIES. Far Eastern Entomologist 518: 1-13, DOI: 10.25221/fee.518.1, URL: https://doi.org/10.25221/fee.518.1
E35A87B98711FF8FC5C7FE8EFC4EFCF9.taxon	description	Figs 36 – 42	en	Nabozhenko, M. V. (2025): COMPOSITION AND DISTRIBUTION OF THE GENUS ALLECULA FABRICIUS, 1801 (COLEOPTERA: TENEBRIONIDAE) IN RUSSIA WITH DESCRIPTION OF A NEW SPECIES. Far Eastern Entomologist 518: 1-13, DOI: 10.25221/fee.518.1, URL: https://doi.org/10.25221/fee.518.1
E35A87B98711FF8FC5C7FE8EFC4EFCF9.taxon	materials_examined	TYPE MATERIAL. Holotype – ♂ (ZIN): Russia: Khabarovsky Krai, Anyuysky National Park, mouth of Bogbasu River, 49 ° 22 ʹ 32.1 ʺN, 137 ° 43 ʹ 15.3 ʺE, 20. VII 2024 (Yu. N. Sundukov, L. A. Sundukova).	en	Nabozhenko, M. V. (2025): COMPOSITION AND DISTRIBUTION OF THE GENUS ALLECULA FABRICIUS, 1801 (COLEOPTERA: TENEBRIONIDAE) IN RUSSIA WITH DESCRIPTION OF A NEW SPECIES. Far Eastern Entomologist 518: 1-13, DOI: 10.25221/fee.518.1, URL: https://doi.org/10.25221/fee.518.1
E35A87B98711FF8FC5C7FE8EFC4EFCF9.taxon	description	DESCRIPTION. Body large (length 11 mm, width 3.5 mm), slender, black and shiny, gently convex, densely covered by dark-brown recumbent and subrecumbent setae. Anterior margin of epistome slightly rounded. Lateral margin of genae straightly converging to epistome, shortly subparallel at base; lateral margin of head between gena and epistome with obtuse emargination. Eyes large and convex, OI = 30.5. Puncturation of head dense and coarse, puncture diameter near two times as long as interpuncture space; middle of frons with small unpunctured area. Ventral side of head smooth and shiny, only temples with coarse and dense puncturation. Maxillary palpomeres 1 – 2 asymmetrical, 1 st palpomere 2 times as long as 2 nd one; apical palpomere strongly asymmetrical, securiform, 1.46 times as wide as interocular distance. Antennae thin, filiform, reaching elytral middle, when directed backward; antennomeres 3 and 4 long, subequal in length; antennomeres 5 – 11 slightly shorter. Prothorax. PI = 73.9. Pronotum narrow, trapezoid, strongly converging from base to anterior margin, 1.28 times as wide as head; lateral margins almost straight, only near middle slightly shortly rounded; anterior margin straight, base slightly trisinuate. Anterior margin and base finely emarginated; laterial margins emarginated only from base to middle. Anterior and posterior angles obtuse (posterior ones obtuse, because base rounded near angles). Disc flattened in middle and strongly convex laterally, with two median round depressions in middle and one transverse median depression in basal third. Puncturation of disc coarse and dense, puncture diameter 2 – 3 times as long as interpuncture space; punctures slightly sparser on sides from middle. Prohypomera and prothoracic sternite coarsely and very densely punctured, but punctures smaller than on pronotum. Prosternal process protruded in apical portion. Pterothorax. Scutellum triangle, with acute apex, densely punctured. Elytra 2.15 times as long as wide, widest slightly behind the middle, 2.1 times as wide as head, 1.6 times as wide and 4.75 times as long as pronotum. Lateral margins of elytra slightly widely emarginated in basal half. Striae consist of coarse and dense round punctures, in basal half merging in entire furrow; interstriae with moderately dense and coarse puncturation, punctures only slightly smaller than strial ones. Epipleura coarsely and densely puncturated, almost reaching suture angle. Mesoventrite with very dense and coarse rugose puncturation; intercoxal process strongly longitudinally impressed in middle. Metaventrite moderately punctured, punctures small, raduliform, diameter slightly lesser than interpuncture space. Sclerites of meso- and metathorax coarsely and densely punctured by simple rounded punctures. Legs long and slender, mesotibiae slightly curved. Tarsi narrow, pro-and mesotarsomeres 3 – 4 lobed, slightly widened, metatarsomer 4 also lobate, but longer and narrower. Ultimate protarsomere slightly longer than first one. First tarsomere longest in meso- and metatarsi. Each tarsal claw with six teeth. Abdomen. Intercoxal process of the first abdominal ventrite strongly depressed and located below the level of the rest surface. Puncturation of ventrites fine and sparse; ventrite 5 without depressions. Inner sternite VIII with very wide and deep emargination medially, large and wide rounded apices, folded on inner side, covered with short and sparse recumbent setae. Spiculum gastrale with securiform blades (outer portion slightly larger than inner) and with long and straight common stem. Aedeagus wide, parameres comparatively wide, triangle in dorso-ventral position and thickened in lateral position. COMPARATIVE DIAGNOSIS. The new species differs from all known Old world Allecula (s. str.) spp. in the unusual shape of the trapezoid pronotum, very similar to those in Lagria Fabricius, 1775, having lateral edges margined only from the base to the middle; coarsely and densely punctured interstriae also distinguish this species from other congeners. The new species is also different in the structure of the male inner sternite VIII, with very wide, deep emargination and very wide, rounded lateral apices with short sparse recumbent setation (other Allecula spp. have male inner sternite VIII with triangle emargination and acute, pointed lateral angulations with dense erected long hairs). The aedeagus and spiculum gastrale is also different from those in all known species. The new species is one of the largest representative of the genus (s. str.) together with several taxa from Taiwan (Masumoto et al., 2019), having body length 11 mm and more. See additional differences in a key below.	en	Nabozhenko, M. V. (2025): COMPOSITION AND DISTRIBUTION OF THE GENUS ALLECULA FABRICIUS, 1801 (COLEOPTERA: TENEBRIONIDAE) IN RUSSIA WITH DESCRIPTION OF A NEW SPECIES. Far Eastern Entomologist 518: 1-13, DOI: 10.25221/fee.518.1, URL: https://doi.org/10.25221/fee.518.1
E35A87B98711FF8FC5C7FE8EFC4EFCF9.taxon	etymology	ETYMOLOGY. The species is named in honour of Yuri Sundukov (Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, Vladivostok, Russia), known specialist on the Far Eastern Carabidae and one of the collectors of a new species.	en	Nabozhenko, M. V. (2025): COMPOSITION AND DISTRIBUTION OF THE GENUS ALLECULA FABRICIUS, 1801 (COLEOPTERA: TENEBRIONIDAE) IN RUSSIA WITH DESCRIPTION OF A NEW SPECIES. Far Eastern Entomologist 518: 1-13, DOI: 10.25221/fee.518.1, URL: https://doi.org/10.25221/fee.518.1
