taxonID	type	description	language	source
E31187EDFFB6FFF6D1F0FA60FCC1FE94.taxon	description	(Figs. 2 – 12) Material examined: Panamá: Chiriquí, Jaramillo Abajo, Caldera River, 1065 masl, 8 ° 44 ’ 46.97 ” N and 82 ° 25 ’ 8.77 ” O. 1 n, 04. ii. 2015, # 17; ibid., 1 n, 04. ii. 2015, # 19; ibid., 1 n, 04. ii. 2015, # 22; ibid., 1 n, 21. ii. 2015, # 23; 1 n, 21. ii. 2015, # 24; 1 n, 21. ii. 2015, # 25; 1 n, 21. ii. 2015, # 26; 1 n, 21. ii. 2015, # 27; 1 n, 21. ii. 2015, # 28; 1 n, 21. iii. 2015, # 48; ibid., 1 n, 21. iii. 2015, # 49; ibid., 1 n, 21. iii. 2015, # 50; ibid., 1 n, 21. iii. 2015, # 51; ibid., 1 n, 21. iii. 2015, # 52; ibid., 1 n, 21. iii. 2015, # 53; ibid., 1 n, 21. iii. 2015, # 54; ibid., 1 n, 25. iv. 2015, # 99; ibid., 1 n, 25. iv. 2015, # 100; ibid., 1 n, 24. viii. 2015, # 190; 1 n, 17. ix. 2015, # 196; ibid., K. Castillo, MUPADI (nymphs: m # 1, f # 1, adults: m # 2, f # 16). Adult: Description by Stark (1998). Description of the last nymphal instar. Male (n = 3): Body brown and stout, body length 12.3 ± 0.23 mm (not including cerci) (Fig. 2). Head (length = 1.9 ± 0.17 mm, width = 2.9 ± 0.26 mm): mostly brown, with a dark spot in dorsal view that extends from the back of the ocelli to the middle part of the head. Light colored ocelli with dark margins and a clear area extended laterally, front with a clear and discontinuous M-shape spot, brown labrum (Fig. 3), filamentous antennae (length: 7.5 ± 1.10 mm) with dark hairs in the proximal region of the scape (Fig. 4). Thorax (n = 3): Pronotum (length = 1.6 ± 0.11 mm, width = 2.9 ± 0.21 mm) with band pattern (dark, light, dark, light) from the midline to the lateral margin of the pronotum, pronotal margin clear with spines (Fig. 3). Mesonotum (length = 2.6 ± 0.73 mm, width = 32 ± 0.21 mm) and metanotum (length = 2.6 ± 0.17 mm, width = 2.7 ± 0.23 mm) with pattern of clear, diffuse drawings, wing pads dark in nymphs closed to emerge, with spines in the margins (Fig. 9). Thoracic gills: sc 1 present, sc 2 and sc 3 absent; pl 1, pl 2 and pl 3 present (Fig. 10). Legs mostly brown, covered with dark hairs and outer side margins with few fringes of bristles. Foreleg (Figs 5 – 6): coxa with numerous dorsal spines in the distal region and without ventral spines (Fig. 8). Femur covered with dark hairs, with a transverse row of spines (7 to 8) in the middle part and a patch of small spines in the proximal lower part, presence of a dark spot at the distal end, without ventral distal spines (Fig. 7). Abdomen: Brown tergites with long, strong spines, presence of a circular brown spot in the ninth sternite (Fig. 11), anal gills absent, cerci with few fine bristles at the distal end. Variations Female (n = 11): Body characteristics similar to those of the male. Body length 16.1 ± 0.82 mm (does not include cerci, n = 3), antennas: n = 10, long = 7.55 ± 2.69 mm, head: long = 2.42 ± 0.17 mm, width = 3.71 ± 0.21 mm, pronotum: long = 2.15 ± 0.08 mm, width = 3.77 ± 0.28 mm, mesonotum: long = 3.23 ± 0.24 mm, width = 4.08 ± 0.32 mm, metanotum: long = 3.31 ± 0.16 mm, width = 3.68 ± 0.25 mm, inverted V-shaped notch in the eighth sternite (Fig. 12).	en	Castillo Sánchez, Kayla N., Aguirre, Yusseff P., Ríos González, Tomás A., Bernal Vega, Juan A. (2019): The Plecoptera of Panama. IV. Description of the last nymphal instar for two species of Anacroneuria Klapálek (Plecoptera: Perlidae) from the Caldera River, Chiriquí, Panamá. Zootaxa 4711 (3): 446-458, DOI: 10.11646/zootaxa.4711.3.2
E31187EDFFB6FFF6D1F0FA60FCC1FE94.taxon	diagnosis	Diagnosis: The nymphs of A. planicollis can be distinguished from A. quetzali, by the presence of a dark spot on the head in dorsal view that extends from the back of the ocelli to the middle part of the head, presence of pattern of bands from the midline towards the lateral margin of the pronotum, coxa without ventral spines, and femur with a transversal row of spines (7 to 8) medially.	en	Castillo Sánchez, Kayla N., Aguirre, Yusseff P., Ríos González, Tomás A., Bernal Vega, Juan A. (2019): The Plecoptera of Panama. IV. Description of the last nymphal instar for two species of Anacroneuria Klapálek (Plecoptera: Perlidae) from the Caldera River, Chiriquí, Panamá. Zootaxa 4711 (3): 446-458, DOI: 10.11646/zootaxa.4711.3.2
E31187EDFFB6FFF6D1F0FA60FCC1FE94.taxon	distribution	Distribution: Belize, Costa Rica, Guatemala, México, Nicaragua, and Panamá.	en	Castillo Sánchez, Kayla N., Aguirre, Yusseff P., Ríos González, Tomás A., Bernal Vega, Juan A. (2019): The Plecoptera of Panama. IV. Description of the last nymphal instar for two species of Anacroneuria Klapálek (Plecoptera: Perlidae) from the Caldera River, Chiriquí, Panamá. Zootaxa 4711 (3): 446-458, DOI: 10.11646/zootaxa.4711.3.2
E31187EDFFB6FFF6D1F0FA60FCC1FE94.taxon	biology_ecology	Ecology: The nymphs of A. planicollis were collected in a shallow area of the Caldera River with bedrock substrate, submerged leaf litter, and slow current. The nymphs of A. planicollis were positively correlated with temperature and conductivity (Castillo Sánchez et al., 2018).	en	Castillo Sánchez, Kayla N., Aguirre, Yusseff P., Ríos González, Tomás A., Bernal Vega, Juan A. (2019): The Plecoptera of Panama. IV. Description of the last nymphal instar for two species of Anacroneuria Klapálek (Plecoptera: Perlidae) from the Caldera River, Chiriquí, Panamá. Zootaxa 4711 (3): 446-458, DOI: 10.11646/zootaxa.4711.3.2
E31187EDFFB2FFFBD1F0FC50FC6AFE94.taxon	description	(Figs. 13 – 27)	en	Castillo Sánchez, Kayla N., Aguirre, Yusseff P., Ríos González, Tomás A., Bernal Vega, Juan A. (2019): The Plecoptera of Panama. IV. Description of the last nymphal instar for two species of Anacroneuria Klapálek (Plecoptera: Perlidae) from the Caldera River, Chiriquí, Panamá. Zootaxa 4711 (3): 446-458, DOI: 10.11646/zootaxa.4711.3.2
E31187EDFFB2FFFBD1F0FC50FC6AFE94.taxon	materials_examined	Material examined: Panamá: Chiriquí, Volcán Barú National Park (PNVB), Caldera River, 1148 ± 12 masl (( 8 ° 51 ’ 06.9 ’’ N and 82 ° 29 ’ 37.4 ’’ W), 4 n, 22. iii. 2015, # 58; ibid., 1 n, 26. iv. 2015, # 108; ibid., 1 n, 26. iv. 2015, # 109; ibid., 1 n, 26. iv. 2015, # 110; ibid., 1 n, 17. v. 2015, # 119; ibid., 1 n, 23. viii. 2015, # 182; ibid., 1493 ± 29 masl (8 ° 51 ’ 02.2 ’’ N and 82 ° 29 ’ 38.4 ’’ W), 1 n, 26. iv. 2015, # 107; ibid., 1 n, 17. v. 2015, # 118; ibid., 1848 ± 8 masl, (8 ° 50 ’ 58.7 ” N and 82 ° 29 ’ 36.2 ” W), 3 n, 22. iii. 2015, # 60; ibid., 6 n, 26. iv. 2015, # 112; 5 n, 17. v. 2015, # 120; ibid., 5 n, 17. v. 2015, # 122; ibid., 1 n, 17. v. 2015, # 123; ibid., 1 n, 17. v. 2015, # 124; ibid., 4 n, 21. vi. 2015, # 128; 7 n, 19. vii. 2015, # 142; ibid., 6 n, 23. viii. 2015, # 184; ibid., 2 n, 27. ix. 2015, # 207; ibid., 1 n, 27. ix. 2015, # 208; ibid., 1 n, 18. x. 2015, # 214; 1 n, 15. xi. 2015, # 222. 1882 ± 6 masl (8 ° 51 ’ 08.2 ” N and 82 ° 29 ’ 44.0 ” O), 2 n, 30. i. 2015, # 1; ibid., 5 n, 22. ii. 2015, # 29; ibid., 3 n, 22. iii. 2015, # 56; ibid., 1 n, 26. iv. 2015, # 102; ibid., 1 n, 26. iv. 2015, # 103; ibid., 4 n, 26. iv. 2015, # 104; ibid., 1 n, 26. iv. 2015, # 106; ibid., 2 n, 17. v. 2015, # 117; ibid., 4 n, 21. vi. 2015, # 126; 1 n, 19. vii. 2015, # 140; ibid., 2 n, 23. viii. 2015, # 181; ibid., 1 n, 27. ix. 2015, # 200; ibid., 1 n, 27. ix. 2015, # 201; ibid., 1 n, 27. ix. 2015, # 202; ibid., 1 n, 27. ix. 2015, # 203; ibid., 1 n, 18. x. 2015, # 210; ibid., 1 n, 15. xi. 2015, # 217; ibid., 1 n, 13. xii. 2015, # 230; ibid., 10 n, 13. xii. 2015, # 231. K. Castillo, MUPADI (nymphs: m # 5, f # 80, adults: m # 5, f # 7). Adults: Male description by Gutiérrez-Fonseca and Springer (2015). Female: Habitus similar to male (Figs. 25 – 26). Mesothoracic wing length 30 ± 2.19 mm (n = 7). Subgenital plate of the eighth sternite bilobed with a V-shaped notch, sinuous margins and with bristles grouped in the upper margin of the ninth sternite (Fig. 27).	en	Castillo Sánchez, Kayla N., Aguirre, Yusseff P., Ríos González, Tomás A., Bernal Vega, Juan A. (2019): The Plecoptera of Panama. IV. Description of the last nymphal instar for two species of Anacroneuria Klapálek (Plecoptera: Perlidae) from the Caldera River, Chiriquí, Panamá. Zootaxa 4711 (3): 446-458, DOI: 10.11646/zootaxa.4711.3.2
E31187EDFFB2FFFBD1F0FC50FC6AFE94.taxon	description	Description of the last nymphal instar. Male (n = 8): Body brown and stout, body length 21.65 ± 1.64 mm (not including cerci) (Fig. 13). Head (length: 3.05 ± 0.29 mm, width: 4.75 ± 0.35 mm), mostly brown covered with scattered dark hairs, anterior region with a lighter area. Labrum with a dark brown spot on the middle, front with a M-shaped mark, clear or sometimes diffuse, black ocelli with laterally expanded lighter areas, black eyes with notable spines and elongated at their margins and a clear area around, region near the antennae with a dark brown spot (Fig. 14), filamentous antennae (n = 7, length: 8.48 ± 0.71 mm) with small and few dark hairs (Fig. 15). Thorax: Pronotum (length: 2.5 ± 0.15 mm, width: 4.6 ± 0.24 mm) with clear and diffuse patterns, mostly covered with dark hairs, dark brown pronotal margin with long spines (Fig. 14). Mesonotum (length: 4.53 ± 0.14 mm, width: 5.88 ± 0.53 mm) and metanotum (length: 4.65 ± 0.27 mm, width: 5.48 ± 0.36 mm) with clear and diffuse patterns, mostly covered with dark hairs, with short spines on their margins (Fig. 20). Legs mostly brown, robust covered with dark hairs and lower lateral margins with numerous fringes of bristles. Foreleg (Figs 16 – 17): Coxa covered with numerous dark hairs, row of numerous small dorsal spines at the distal end and without ventral spines (Fig. 19). Femur covered by numerous dark hairs, leaving a clear fringe free of them and like the area the upper left margin, a transversal row of spines (4 to 5) spines, without ventral spines, dark band at the distal end in nymphs closed to emerge (Fig. 18). Thoracic gills: sc 1, sc 3, sc 2 present; pl 1, pl 2 and pl 3 present (Fig. 21). Brown abdomen with long and noticeable spines, presence of a circular brown spot in the ninth sternite, sometimes be diffuse (it can be observed clearly using a brighter light) (Fig. 22). Brown cerci with evident spines arranged in rows and covered with numerous bristles from the middle towards the distal end of the cerci (length: 9.62 ± 2.69 mm) (Fig. 27). Variations Female (n = 34): Body characteristics similar to those of the male. Body length 25.33 ± 3.22 mm (does not include cerci), antennas: n = 26, long = 8.76 ± 0.93 mm, head: long = 3.23 ± 0.42 mm, width = 5.53 ± 0.51 mm, pronotum: long = 2.83 ± 0.26, width = 5.08 ± 0.39 mm, mesonotum: long = 4.88 ± 0.77 mm, width = 6.28 ± 0.75 mm, metanotum: long = 4.91 ± 0.85 mm, width = 5.85 ± 0.68 mm, cercus: n = 32, length: 9.78 ± 1.02 mm, inverted V-shaped notch in the eighth sternite (Fig. 23).	en	Castillo Sánchez, Kayla N., Aguirre, Yusseff P., Ríos González, Tomás A., Bernal Vega, Juan A. (2019): The Plecoptera of Panama. IV. Description of the last nymphal instar for two species of Anacroneuria Klapálek (Plecoptera: Perlidae) from the Caldera River, Chiriquí, Panamá. Zootaxa 4711 (3): 446-458, DOI: 10.11646/zootaxa.4711.3.2
E31187EDFFB2FFFBD1F0FC50FC6AFE94.taxon	diagnosis	Diagnosis: The nymphs of A. quetzali can be distinguished from A. planicollis, by the mostly brown head covered with scattered dark hairs, labrum with a dark brown spot in the middle part, supracoxal gills (sc) in the second and third pair legs, coxa without ventral spines, femur with a transversal row of spines (4 to 5) dorsal midsoles, and cerci covered with numerous bristles medially to the distal end.	en	Castillo Sánchez, Kayla N., Aguirre, Yusseff P., Ríos González, Tomás A., Bernal Vega, Juan A. (2019): The Plecoptera of Panama. IV. Description of the last nymphal instar for two species of Anacroneuria Klapálek (Plecoptera: Perlidae) from the Caldera River, Chiriquí, Panamá. Zootaxa 4711 (3): 446-458, DOI: 10.11646/zootaxa.4711.3.2
E31187EDFFB2FFFBD1F0FC50FC6AFE94.taxon	discussion	Comment: The presence of supracoxal gills in the second and third pair of legs was evidenced in the nymphs of A. quetzali. This characteristic has been mentioned previously by Gutiérrez-Fonseca (2009) for Morphospecies 1.	en	Castillo Sánchez, Kayla N., Aguirre, Yusseff P., Ríos González, Tomás A., Bernal Vega, Juan A. (2019): The Plecoptera of Panama. IV. Description of the last nymphal instar for two species of Anacroneuria Klapálek (Plecoptera: Perlidae) from the Caldera River, Chiriquí, Panamá. Zootaxa 4711 (3): 446-458, DOI: 10.11646/zootaxa.4711.3.2
E31187EDFFB2FFFBD1F0FC50FC6AFE94.taxon	distribution	Distribution: Costa Rica and Panamá.	en	Castillo Sánchez, Kayla N., Aguirre, Yusseff P., Ríos González, Tomás A., Bernal Vega, Juan A. (2019): The Plecoptera of Panama. IV. Description of the last nymphal instar for two species of Anacroneuria Klapálek (Plecoptera: Perlidae) from the Caldera River, Chiriquí, Panamá. Zootaxa 4711 (3): 446-458, DOI: 10.11646/zootaxa.4711.3.2
E31187EDFFB2FFFBD1F0FC50FC6AFE94.taxon	biology_ecology	Ecology: The nymphs of A. quetzali were collected more frequently in shallow reaches of Caldera River with lower water temperatures, bedrock substrate, with little current. Physical and chemical parameters for the above collection sites of nymphs were presented by Castillo Sánchez et al. (2018).	en	Castillo Sánchez, Kayla N., Aguirre, Yusseff P., Ríos González, Tomás A., Bernal Vega, Juan A. (2019): The Plecoptera of Panama. IV. Description of the last nymphal instar for two species of Anacroneuria Klapálek (Plecoptera: Perlidae) from the Caldera River, Chiriquí, Panamá. Zootaxa 4711 (3): 446-458, DOI: 10.11646/zootaxa.4711.3.2
