taxonID	type	description	language	source
E33C87BDC633FFEEF4FA64F870017472.taxon	description	= Ino Hinds, 1843, non Ino Schrank, 1803 Type species: Triforis (Ino) gigas Hinds, 1843. Subsequent designation by Jousseaume (1884). Recent, southwestern Pacific. Diagnosis: paucispiral protoconch with very reduced nucleus and typically two spiral cords on the following whorls; teleoconch elongated, with three main spiral cords, the adapical one usually initially narrower but gradually developing.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC633FFECF4FA66A8751773A5.taxon	description	Figures 3 Y, 4	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC633FFECF4FA66A8751773A5.taxon	materials_examined	Type material. Holotype: ANSP 367841 (Fig. 3 Y). Paratypes: ANSP 367840, type locality [4]. Type locality. 26 º 29 ʹ 45 ʺ N, 78 º 37 ʹ 15 ʺ W, 300 m, Lucaya, Grand Bahama Island, Bahamas. Material examined. The type material and: Brazil: Espírito Santo state: IBUFRJ 19619, REVIZEE-Central C 1 - VV 38 [1]. Almirante Saldanha Seamount: MNRJ 33780, REVIZEE-Central C 1 - D 1 [1]. Rio de Janeiro state: MNRJ 18302, HAB 13 - H 4 [2]; MNRJ 18412, HAB 16 - H 4 [1]; MNRJ 18300, HAB 11 - G 4 [1]; MNRJ 18402, HAB 16 - G 4 [1]; MNRJ 18301, HAB 13 - D 3 [1]; MNRJ 31124, 22 º 42 ʹ S, 40 º 40 ʹ W, 110 – 120 m, xii / 2001 [2]; MNRJ 32629, 22 º 42 ʹ S, 40 º 40 ʹ W, 110 – 120 m, viii / 2002 [2]; MNRJ 30868, 22 º 42 ʹ S, 40 º 40 ʹ W, 110 – 120 m, 19 / ix / 2003 [3]; MNRJ 31108, 22 º 42 ʹ S, 40 º 40 ʹ W, 110 – 120 m, xi / 2007 [2]; MNRJ 18379, HAB 16 - C 3 [1]; IBUFRJ 19577, 22 º 48 ʹ S, 40 º 45 ʹ W, 106 – 110 m, 27 / i / 1998 [28]; IBUFRJ 19533, REVIZEE-Central C 1 - D 1 - 2 [2]; MNRJ 18423, HAB 16 - C 4 [2]; MNRJ 33806, 23 º 03 ʹ 00 ʺ S, 41 º 04 ʹ 06 ʺ W, 98 m, 12 / iv / 1995 [2]; MNRJ 32043, 23 º 03 ʹ 18 ʺ S, 41 º 02 ʹ 06 ʺ W, 97 m, x / 2008 [3]; MNRJ 18965 [20], MORG 50111 [46], 23 º 04 ʹ 14 ʺ S, 40 º 59 ʹ 31 ʺ W, 100 m, 17 / xii / 2004; MNRJ 31115, 23 º 04 ʹ 14 ʺ S, 40 º 59 ʹ 31 ʺ W, 100 m, iii / 2005 [4]; MNRJ 32354, 23 º 04 ʹ 14 ʺ S, 40 º 59 ʹ 31 ʺ W, 100 m, vii / 2005 [3]; MNRJ 18948 [6], MNRJ 26603 [16], MNRJ 33075 [11], MNRJ 33076 [5], MORG 47452 [7], MORG 47495 [5], MORG 48069 [5], MORG 48131 [9], MORG 48182 [2], MORG 48195 [5], MORG 48220 [2], MORG 48264 [5], MORG 48430 [7], MORG 48556 [2], MORG 48573 [5], MORG 48610 [6], MORG 48614 [1], 23 º 05 ʹ S, 40 º 58 ʹ W, 100 m; MNRJ 32391, 23 º 05 ʹ S, 40 º 59 ʹ W, 100 m, v / 2009 [12]; MNRJ 30867, 23 º 05 ʹ 23 ʺ S, 40 º 58 ʹ 55 ʺ W, 100 m, 17 / i / 2004 [5]; MNRJ 32072, 23 º 09 ʹ 00 ʺ S, 41 º 02 ʹ 06 ʺ W, 100 m, x / 2008 [5]; MNRJ 18368, HAB 16 - B 4 [2]; MNRJ 18299 [5], MNRJ 18703 [1], HAB 11 - B 5; MNRJ 18426, HAB 16 - B 5 [3]; MNRJ 32061, 23 º 13 ʹ 30 ʺ S, 41 º 21 ʹ 00 ʺ W, 93 m, x / 2008 [6]; MNRJ 32403, 23 º 18 ʹ 35 ʺ S, 41 º 14 ʹ 03 ʺ W, 100 m [1]; MNRJ 32048, 23 º 39 ʹ 00 ʺ S, 41 º 24 ʹ 00 ʺ W, 154 m, x / 2008 [2]; MNRJ 32038, 24 º 07 ʹ 44 ʺ S, 44 º 15 ʹ 12 ʺ W, 142 m, x / 2008 [1]; MNRJ 32040, 24 º 12 ʹ 20 ʺ S, 44 º 16 ʹ 34 ʺ W, 145 m, x / 2008 [2]; IBUFRJ 8992, Campos Basin, Oc. Ship Astro Garoupa coll., 1984 [1]; MORG 39386, Campos Basin, 1997 [1]; MNRJ 27846, REVIZEE-Sul sta. 6666 [2]. São Paulo state: MNRJ 27847, PADCT sta. 6573 [5]; MNRJ 27853, REVIZEE-Sul sta. 6676 [4]; MNRJ 60188, PADCT sta. 6577 [1].	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC633FFECF4FA66A8751773A5.taxon	description	Description. Shell sinistral, elongated, conical / fusiform, with rectilinear profile, reaching 11.21 mm in length, 1.80 mm in width, ratio length / width 4.0 to 6.1. Protoconch subcylindrical / columnar, 0.63 – 0.71 mm in length, 0.54 – 0.62 mm in width in its last whorl; 2.25 to 3.25 convex protoconch whorls, weak distinction between protoconch and teleoconch; nucleus very small, slightly elevated, in an oblique descent; two main strong spiral cords present from the beginning of the protoconch, with nearly the same size, situated at 34 % and 72 % of last whorl height, in addition to a small subsutural cord, more evident in the last whorl; axial sculpture entirely absent. Teleoconch with up to 16.5 whorls; three spiral cords in the beginning, all of them continuous with those of the protoconch; adapical spiral cord very close to the median cord and narrower than other cords through first whorls of teleoconch, but ac- quiring nearly the same size as others on the eighth / twelfth whorl; on the body whorl, distance between spiral cords is equal or as much as twice the width of cords; 13 to 16 nearly orthocline axial ribs, rarely slightly opisthocline; rounded (especially the adapical and median cords) to slightly elliptical (especially the abapical cord) nodules of a medium to moderately large size; distinct but little developed suture, with a small sutural cord; thick, smooth subperipheral cord, two to three smooth basal cords, the adapical one thick and very close to the subperipheral cord, the other two (or one) usually thin; no supranumerical cords; ovate aperture, 0.61 – 0.95 mm long, 0.47 – 0.82 mm wide, ratio length / width 1.0 – 1.3; anterior canal very short, directed downwards / backwards, and open, sometimes partially crossed in its base by the projection of the outer lip, 0.20 – 0.29 mm long, 0.22 – 0.29 mm wide, ratio length / width 0.9 – 1.2; posterior canal as a small notch, 0.11 – 0.34 mm long. Light brown / beige shell, apex (protoconch and one / two initial whorls of teleoconch) slightly lighter; nodules of teleoconch usually somewhat lighter than internodular spaces.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC633FFECF4FA66A8751773A5.taxon	discussion	Remarks. This is by far the most common species of Inella in Campos Basin, southeastern Brazil. The single shell from Espírito Santo state is of a very worn juvenile, thus it is uncertain if living populations of I. apexbilirata indeed occur in Brazil north of Campos Basin. The type material of I. apexbilirata (Fig. 3 Y) is represented by five shells, at least some of which are adult (Rolán & Fernández-Garcés 2008), but the maximum shell length is only 3.5 mm for about nine whorls of teleoconch, in contrast to 11.21 mm for 16.5 whorls in shells from Brazil (those between 8.5 and 9.5 whorls of teleoconch reach from 4.1 to 4.8 mm long). It remains unclear if Caribbean specimens can reach larger dimensions, owing to the scarcity of available material. The most remarkable difference between shells from the Caribbean and Brazil is related to shell color: white in the former, light brown in the latter (Fig. 4 A – E). A few Triphoridae species present polychromatism in the shell along their geographical distribution, like the usually white shells in Caribbean but usually light brown shells in Brazil of Metaxia rugulosa (C. B. Adams, 1850) (Fernandes & Pimenta 2011). In spite of it, no available shell of Caribbean I. apexbilirata is light brown, nor is any shell from Brazil white. Another difference occurs in their bathymetric range, as shells from Brazil do not seem to reach the depth of 300 m (type locality), but some juveniles were sampled down to 163 m. The record of Triphora compsa Dall, 1927 from Brazil by Absalão (1989) actually consists of I. apexbilirata (Fig. 4 E). Strobiligera compsa (Fig. 3 R) is presumably restricted to the southeastern U. S. A., possessing a wider shell, larger teleoconch whorls, and a deeper bathymetric record (805 m). Geographical distribution. Bahamas (type locality); Brazil: Espírito Santo (pending confirmation), Almirante Saldanha Seamount, Rio de Janeiro, São Paulo (this study).	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC633FFECF4FA66A8751773A5.taxon	description	Bathymetric distribution. 69 – 163 m (this study) to 300 m (type locality).	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC634FFF7F4FA60447353737E.taxon	description	Figure 8 A – J	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC634FFF7F4FA60447353737E.taxon	materials_examined	Material examined. Brazil: Bahia state: MZSP 133326, Parcel Paredes, Alcobaça, 2 – 3 m, P. J. Souza & E. P. Gonçalves coll., 15 / i / 1970 [1]. Rio de Janeiro state: MNRJ 18414, HAB 16 - H 3 [1]; MNRJ 18297, HAB 11 - C 3 [1]; IBUFRJ 19557, REVIZEE-Central C 1 - D 3 [1]; IBUFRJ 11698, 22 º 20 ʹ S, 40 º 32 ʺ W, iv / 1998 [1]; MNRJ 29509, 22 º 40 ʹ 35 ʺ S, 41 º 43 ʹ 21 ʺ W, 47 m [1]; MNRJ 32624 [1], MNRJ 32644 [2], 23 º 04 ʹ 14 ʺ S, 40 º 59 ʹ 31 ʺ W, 100 m; MNRJ 18961 [1], MORG 52216 [1], 23 º 04 ʹ 14 ʺ S, 40 º 59 ʹ 31 ʺ W, 100 m, 17 / xii / 2004; MNRJ 31117, 23 º 04 ʹ 14 ʺ S, 40 º 59 ʹ 31 » W, 100 m, iii / 2005 [1]; MNRJ 26606, 23 º 04 ʹ 14 ʺ S, 40 º 59 ʹ 31 ʺ W, 100 m, vii / 2005 [1]; MNRJ 33592, 23 º 05 ʹ S, 40 º 58 ʹ W, 100 m [1]; MNRJ 32054, 23 º 06 ʹ 00 ʺ S, 41 º 06 ʹ 00 ʺ W, 89 m, x / 2008 [1]; MNRJ 32068, 23 º 09 ʹ 00 ʺ S, 41 º 06 ʹ 00 ʺ W, 99 m, x / 2008 [1]; MNRJ 32046, 23 º 15 ʹ 00 ʺ S, 41 º 06 ʹ 00 ʺ W, 112 m, x / 2008 [1]. São Paulo state: MNRJ 32918, 25 º 13 ʹ 06 ʺ S, 44 º 59 ʹ 31 ʺ W, 160 m, viii / 2008 [1].	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC634FFF7F4FA60447353737E.taxon	description	Description. Shell sinistral, elongated, conical, nearly rectilinear profile, reaching 7.79 mm in length, 1.85 mm in width, ratio length / width 4.0 to 4.2. Protoconch subcylindrical, 0.85 – 1.08 mm in length, 0.64 – 0.72 mm in width of its last whorl; 4.25 to 4.75 convex protoconch whorls, weak distinction between protoconch and teleoconch; initial whorl dome-shaped, slightly globose in some cases, smooth; remaining whorls with two spiral cords, situated at 33 % and 63 % of last whorl height, the abapical one slightly more prominent especially at the end of protoconch, as the adapical one vanishes before the transition to the teleoconch; a subsutural small cord is also apparent on the last whorl; very small axial wrinkles above the adapical cord, being nearly orthocline to oblique. Teleoconch with up to 11 whorls; two spiral cords (adapical and abapical) in the beginning, the abapical one continuous with that of the protoconch; median spiral cord emerges narrowly at the very end of the fourth to the beginning of the sixth whorl, reaching nearly the same size as the abapical cord (the adapical one is more prominent on late whorls) after 3.5 to five whorls; on the body whorl, distance between spiral cords is 1.1 times the width of cords; 17 to 20 opisthocline axial ribs; rounded (especially in the adapical cord) to nearly elliptical (median and abapical cords) nodules of a medium size; distinct and well-developed suture, with a small sutural cord; weakly nodulose to slightly wavy subperipheral cord, two to three slightly wavy to smooth basal cords; no supranumerical cords are present, but in one shell there are two very fine threads behind the peristome; ovate aperture, 0.90 – 1.12 mm long, 0.87 – 0.89 mm wide, ratio length / width 1.0 – 1.1; anterior canal curved almost downwards, medium to moderately large, mainly closed, 0.62 – 0.78 mm long, 0.27 – 0.29 mm wide, ratio length / width 2.3 to 2.7; posterior canal as a wide sinus, 0.10 – 0.20 mm long, not detached from the aperture. Light brown to cream shell, with the base slightly darker.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC634FFF7F4FA60447353737E.taxon	discussion	Remarks. The single remarkable difference between “ Inella ” sp. 2 and shells of “ I ”. faberi from Brazil is related to the size of protoconch, as it varies from 0.85 – 1.08 mm in length and 0.64 – 0.72 mm in width (4.25 to 4.75 whorls) in “ Inella ” sp. 2, but from 0.55 – 0.79 mm in length and 0.51 – 0.59 mm in width (3 to 3.5 whorls) in “ I ”. faberi. This could represent the existence of very similar sympatric species or a single species with a considerable variation in protoconch size, as it occurs with other taxa in the present study. However, no shell was observed to possess an intermediate number of protoconch whorls (3.5 to 4.25 whorls) in the material examined. The single shell, immature and worn, sampled from Bahia (Fig. 8 C) cannot be confidently regarded as indicative that “ Inella ” sp. 2 truly occurs north of Campos Basin. It is probably a subfossil / fossil form, and its shallow bathymetric record is also doubtful. A large shell from Espírito Santo state (MNRJ 30892, Fig. 8 K – L), 8.30 mm in length and 2.20 mm in width, has the adapical spiral cord of protoconch disappearing much before the transition to the teleoconch, the median spiral cord of teleoconch emerging yet at the end of the second whorl (Fig. 8 L), and the three spiral cords of similar strength. Because of the similarity of its protoconch with “ Inella ” sp. 2 and the lack of more available material, this shell is herein illustrated without being designated as a new morphospecies. Geographical distribution. Brazil: Rio de Janeiro and São Paulo, immature / worn shell northern up to Bahia.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC634FFF7F4FA60447353737E.taxon	description	Bathymetric distribution. 47 – 160 m. The record of 2 – 3 m from Bahia must be disregarded, owing to the immature / worn condition of the material.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC628FFF2F4FA676C7373760A.taxon	description	urn: lsid: zoobank. org: act: AAAA 9787 - 50 A 1 - 4917 - B 07 D- 08 D 301 EBF 7 D 3	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC628FFF2F4FA676C7373760A.taxon	materials_examined	Type material. Holotype: MNRJ 29365, P. Gonçalves coll., iii / 2003. Paratypes: Brazil: Rio de Janeiro state: MNRJ 29410, 22 º 24 ʹ 05 ʺ S, 41 º 05 ʹ 08 ʺ W, 48 m, Campos Basin [2]; MNRJ 29409, 22 º 40 ʹ 35 ʺ S, 41 º 43 ʹ 21 ʺ W, 47 m, Campos Basin [1]; MZSP 39584 [5], MORG 47194 [1], Arraial do Cabo, 25 – 30 m, P. Gonçalves coll., viii / 2003. Type locality. Arraial do Cabo, 25 – 30 m, Rio de Janeiro state, Brazil.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC628FFF2F4FA676C7373760A.taxon	etymology	Etymology. The specific epithet alludes to the citizens of Arraial do Cabo (locally known as “ galo ”, in Portuguese), the region where this species was exclusively found so far. Epithet as a noun in apposition.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC628FFF2F4FA676C7373760A.taxon	diagnosis	Diagnosis. Robust shell, usually light brown / golden, with a large aperture; median spiral cord emerges very narrowly on the fourth or fifth teleoconch whorl, eventually on the sixth whorl; two supranumerical cords may develop.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC628FFF2F4FA676C7373760A.taxon	description	Description. Shell sinistral, elongated, conical, broad at the base, nearly rectilinear profile, reaching 9.90 mm in length, 2.64 mm in width, length / width ratio 3.4 to 4.1. Protoconch subcylindrical, 0.62 – 0.89 mm in length, 0.53 – 0.66 mm in width of its last whorl; 3.25 to 4.25 convex whorls, clear distinction between protoconch and teleoconch; initial whorl dome-shaped, mainly smooth; remaining whorls with two spiral cords, situated at 33 – 36 % and 61 – 73 % of last whorl height, the adapical one vanishing and disappearing before the transition to the teleoconch; very small, almost indistinct, axial wrinkles above the adapical cord, being nearly orthocline. Teleoconch with up to 13 whorls; two spiral cords (adapical and abapical) in the beginning, the abapical one continuous with that of the protoconch; median spiral cord emerges very narrowly on the fourth or fifth whorl, eventually on the sixth whorl, reaching the same size of the abapical cord (the adapical one is more prominent) after 3.5 to six whorls; on the body whorl, distance between spiral cords is 0.8 to 1.5 times the width of cords; 21 to 26 orthocline to moderately opisthocline axial ribs; rounded nodules of a moderately large size, positioned very close to each other in the spiral cords; distinct and well-developed suture, with a small sutural cord; weakly nodulose to wavy subperipheral cord, one or two very narrow, smooth basal cords; two supranumerical cords may develop, one between median and abapical cords, the other between abapical and subperipheral cords; ovate, large aperture, very projected, 1.39 – 1.51 mm long, 1.10 – 1.24 mm wide, length / width ratio 1.1 – 1.3; anterior canal curved downwards / backwards, moderately short, open or partly closed by the projection of the outer lip, 0.47 – 0.66 m long, 0.34 – 0.56 mm wide, length / width ratio 1.1 – 1.6; posterior canal as a wide notch, 0.18 – 0.25 mm long, not detached from the aperture. Light brown / golden shell, but the peristome and the first whorl of protoconch can be whitish; teleoconch rarely whitish.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC628FFF2F4FA676C7373760A.taxon	discussion	Remarks. This species shares several similarities with “ I ”. faberi from Brazil, such as the shell coloration, the protoconch sculpture and the whorl of emergence of the median spiral cord on the teleoconch. However, a typical shell of “ Inella ” galo sp. nov. is thicker and broader, with a very projected aperture (Fig. 10 A), narrower basal cord (s) (usually numbering only one, very close to the subperipheral cord), base not darker than remaining shell (Fig. 10 A – D), with more axial ribs (21 to 26, instead of 16 to 22 in “ I ”. faberi), and it may have distinct supranumerical cords (Fig. 10 G), contrary to “ I ”. faberi. The two species are partially sympatric, with the deepest records of “ Inella ” sp. 3 coinciding with the shallowest records of “ I ”. faberi and “ Inella ” sp. 2 at a depth zone of 45 – 50 m.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC62EFFF0F4FA65BC736D7656.taxon	description	urn: lsid: zoobank. org: act: 44 EB 94 EF- 5688 - 435 C-A 44 D-A 88 F 82 BA 42 AB	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC62EFFF0F4FA65BC736D7656.taxon	materials_examined	Type material. Holotype: MNRJ 32545. Paratypes: Brazil: Almirante Saldanha Seamount: MNRJ 33763, RE- VIZEE-Central C 1 - D 1 [1]. Rio de Janeiro state: MNRJ 18962 [1], MNRJ 32627 [1], 23 º 04 ʹ 14 ʹʹS, 40 º 59 ʹ 31 ʹʹW, 100 m, 17 / xii / 2004; MNRJ 31113, type locality, 22 / iii / 2005 [2]. Other material examined. Brazil: Rio de Janeiro state: MNRJ 18705, HAB 11 - G 4 [1]; IBUFRJ 19579, 22 º 48 ʹ S, 40 º 45 ʹ W, 106 – 110 m, 27 / i / 1998 [1]; MORG 52212 [4], MORG 52240 [1], MORG 52244 [1], type locality; MNRJ 30910, type locality, 17 / ix / 2004 [1]; MNRJ 32401, 23 º 15 ʹ 00 ʺ S, 41 º 06 ʹ 00 ʺ W, 112 m, x / 2008 [1]; MNRJ 32402, 23 º 18 ʹ 35 ʺ S, 41 º 14 ʹ 03 ʺ W, 100 m, x / 2005 [1]. São Paulo state: MNRJ 32919, 25 º 13 ʹ 06 ʺ S, 44 º 59 ʹ 31 ʺ W, 160 m, viii / 2008 [1]. Type locality. 23 º 05 ʹS, 40 º 58 ʹW, 100 m, Rio de Janeiro state, Brazil.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC62EFFF0F4FA65BC736D7656.taxon	etymology	Etymology. The specific epithet renders a meaning in Portuguese regarding the generic name; translating to English, it would mean “ in her I trust ”. Epithet as a noun in apposition.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC62EFFF0F4FA65BC736D7656.taxon	diagnosis	Diagnosis. Protoconch with 3.5 to 4.25 very convex whorls; median spiral cord emerges at the end of the first to the beginning of the third whorl of teleoconch; light brown to cream shell.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC62EFFF0F4FA65BC736D7656.taxon	description	Description. Shell sinistral, elongated, conical, rectilinear profile, reaching 14.33 mm in length, 3.15 mm in width, length / width ratio 3.8 to 4.6. Protoconch slightly globose, subcylindrical, 0.65 – 0.89 mm in length, 0.57 – 0.66 mm in width of its last whorl; 3.5 to 4.25 very convex whorls, clear distinction between protoconch and teleoconch; initial whorl dome-shaped but slightly elevated, covered by small, spirally-disposed vesicles; remaining whorls with two spiral cords, situated at 28 % and 57 % of last whorl height, the abapical one becoming more prominent and assuming a keeled shape on the last whorl, the adapical one sometimes disappearing in the last whorl; numerous small axial wrinkles above the adapical cord, being nearly orthocline to very prosocline, sparsely present also below the abapical cord. Teleoconch with up to 15 whorls; two spiral cords (adapical and abapical) in the beginning, the abapical one continuous with that of the protoconch; median spiral cord emerges at the end of the first whorl to the beginning of the third whorl, reaching the same size of other cords after 2 to 2.5 whorls; on the body whorl, distance between spiral cords is 1.1 to 1.7 times the width of cords; 18 to 20 nearly orthocline to opisthocline axial ribs; rounded nodules of a medium size; distinct and well-developed suture, with a small sutural cord; moderately to weakly nodulose subperipheral cord, two weakly nodulose to wavy basal cords, the adapical one situated closer to the subperipheral cord than to the thick abapical basal cord; a very fine supranumerical cord may develop just before the peristome, between the median and the abapical spiral cords; ovate aperture, 1.00 – 1.72 mm long, 0.81 – 1.39 mm wide, length / width ratio 1.2 – 1.3; anterior canal curved almost downwards, being moderately long and partly or almost closed, 0.59 – 0.95 mm long, 0.34 – 0.51 mm wide, length / width ratio 1.7 – 1.9; posterior canal as a small notch, 0.20 – 0.31 mm long, not detached from the aperture. Light brown to cream shell; coloration usually faint in worn shells.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC62EFFF0F4FA65BC736D7656.taxon	discussion	Remarks. The most similar species to “ Inella ” euconfio sp. nov. is “ Inella ” triserialis (Dall, 1881) sensu lectotype (Rolán & Fernández-Garcés 2008), especially considering the early development of the median spiral cord of teleoconch (Fig. 3 C – D). In contrast, “ I ”. triserialis has the median spiral cord already present in the beginning of the teleoconch (but emerging at the end of the first to the beginning of the third whorl in Inella euconfio sp. nov.), a different protoconch, and its bathymetry ranges from 366 m to 1472 m (Rolán & Fernández-Garcés 2008), instead of 91 m to 160 m in “ Inella ” euconfio sp. nov. The poorly preserved condition of the lectotype of “ I ”. triserialis (Rolán & Fernández-Garcés 2008: fig. 14 C – E), without a complete base and formation of anterior and posterior canals, precludes further comparisons. Relative to species from Brazil, “ Inella ” euconfio sp. nov. has a similar protoconch to “ I ”. faberi and “ Inella ” sp. 2, but in these species the median spiral cord usually emerges at the fifth or sixth whorl of teleoconch, instead of the first to the beginning of the third whorl in “ Inella ” euconfio sp. nov. (Fig. 11 E – F). Also, the three main spiral cords on the teleoconch of “ Inella ” euconfio sp. nov. have similar thickness, although the adapical cord of “ I ”. faberi and “ Inella ” sp. 2 is more prominent than other cords at the body whorl.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC62CFFF1F4FA65BC737370C0.taxon	description	Figure 12 urn: lsid: zoobank. org: act: 4 D 9 E 7 E 84 - 49 AD- 4793 - B 8 D 9 - 849 D 0 FEB 39 B 9	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC62CFFF1F4FA65BC737370C0.taxon	materials_examined	Type material. Holotype: MNRJ 18637. Paratypes: Brazil: Espírito Santo state: MNRJ 33816, 18 º 46 ʹ 48 ʺ S, 38 º 31 ʹ 24 ʺ W, 65 m, 20 / iv / 1985 [1]; MNRJ 32658 [1], MNRJ 32661 [1], 20 º 14 ʹ S, 40 º 12 ʹ W, vi / 2008; MORG 40404, REVIZEE-Central C 1 - VV 22 [1]; MNRJ 34031, Ilha Escalvada, Guarapari, 2012 [1]; MNRJ 31080, 20 º 47 ʹ S, 40 º 34 ʹ W, iii / 2010 [1]; IBUFRJ 19687, Piúma, 1993 [7]. Rio de Janeiro state: MNRJ 18303 [1], MNRJ 18711 [1], HAB 13 - H 2; MNRJ 18642, HAB 16 - H 2 [1]; IBUFRJ 19583, REVIZEE-Central C 1 - D 3 [5]. Other material examined. Brazil: Espírito Santo state: MNRJ 32295, 19 º 24 ʹ 17 ʺ S, 39 º 15 ʹ 33 ʺ W, 45 m [1]; IBUFRJ 19615, REVIZEE-Central C 1 - VV 38 [1]; MNRJ 32651 [1], MNRJ 32652 [2], MNRJ 32655 [1], 20 º 14 ʹ S, 40 º 12 ʹ W, vi / 2008; MNRJ 33797, REVIZEE-Central C 1 - VV 21 [1]; MNRJ 31033 [1], MNRJ 31044 [2], MNRJ 31063 [1], 20 º 47 ʹ S, 40 º 34 ʹ W, iii / 2010. Type locality. HAB 16 - H 3, 21 º 43 ʹ 06 ʺ S, 40 º 11 ʹ 37 ʺ W, 71 m, Rio de Janeiro state, Brazil.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC62CFFF1F4FA65BC737370C0.taxon	etymology	Etymology. leukos, Gr. = white; cephalo, derived from the Gr. kephale = head. It refers to the color of the apex of these shells.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC62CFFF1F4FA65BC737370C0.taxon	diagnosis	Diagnosis. Protoconch with two spiral cords, the adapical one disappearing one to two whorls after its emergence; median spiral cord emerges in the second or third whorl of teleoconch; protoconch and first whorl of teleoconch white, remainder whorls brown.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC62CFFF1F4FA65BC737370C0.taxon	description	Description. Shell sinistral, elongated, conical, rectilinear profile, reaching 11.23 mm in length, 2.69 mm in width, length / width ratio 3.4 to 3.9. Protoconch slightly globose, subcylindrical, 0.45 – 0.67 mm in length, 0.44 – 0.54 mm in width in its last whorl; 2.75 to 3.75 convex whorls, weak distinction between protoconch and teleoconch; initial whorl dome-shaped but slightly pointed, covered by small rounded granules; remaining whorls initially with two spiral cords, situated at 32 – 37 % and 58 – 74 % of whorl height, the adapical one disappearing one to two whorls after its emergence, the abapical one readily strengthening and assuming a keeled shape on the last whorl; small or almost indistinct axial wrinkles more evident above the adapical cord, being orthocline to much oblique. Teleoconch with up to 14 whorls; two spiral cords (adapical and abapical) in the beginning, the abapical one continuous with that of the protoconch; median spiral cord emerges in the second or third whorl, reaching the same size of other cords after 1.5 to 2 whorls; on the body whorl, distance between spiral cords is 0.9 to 1.8 times the width of cords; 18 to 21 orthocline to slightly opisthocline axial ribs; rounded (especially in the adapical cord) to elliptical (median and abapical cords) nodules of a medium to small size, becoming less pronounced on late whorls of large shells; distinct and well-developed suture, with a small sutural cord; narrow and weakly nodulose to wavy subperipheral cord, two slightly wavy to smooth basal cords; two very fine threads may appear behind the peristome in some shells, one above, the other below the median spiral cord; ovate aperture, 0.69 – 0.86 mm long, 0.52 – 0.71 mm wide, length / width ratio 1.2 – 1.3; anterior canal curved almost downwards, moderately long and open or partly closed by the projection of the outer lip, 0.28 – 0.50 mm long, 0.24 – 0.30 mm wide, length / width ratio 1.0 – 1.7; posterior canal a small notch, 0.14 – 0.17 mm long, not detached from the aperture. Protoconch and first whorl of teleoconch white, remainder whorls light brown to brown; coloration usually faint in worn shells.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC62CFFF1F4FA65BC737370C0.taxon	discussion	Remarks: There is intraspecific variation in “ Inella ” leucocephala sp. nov. regarding the protoconch (Fig. 12 J – K), as the variable number of protoconch whorls (2.75 to 3.75) is associated with great differences in its size (0.45 – 0.67 mm in length, 0.44 – 0.54 mm in width) and in the persistence of the adapical spiral cord (for one to two whorls). The coloration pattern of “ Inella ” leucocephala sp. nov. (Fig. 12 A – D) is similar to that of the Caribbean species Isotriphora peetersae Moolenbeek & Faber, 1989. A juvenile of “ Inella ” leucocephala sp. nov. (Fig. 12 E) was sampled from Bahia state (MORG 26919, Ilha Guarita, Abrolhos, 5 m). Owing to this unusual shallow bathymetric record and the early developmental stage of the shell, Bahia was not included in the geographical distribution list of this species, requiring more specimens to confirm this northward range. Geographical distribution. Brazil: Espírito Santo and Rio de Janeiro.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC62CFFF1F4FA65BC737370C0.taxon	description	Bathymetric distribution. 33 – 80 m.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC621FFFAF4FA6113736D737E.taxon	description	Figure 14 urn: lsid: zoobank. org: act: ED 2 D 1925 - 84 E 6 - 405 E- 93 BA- 2 AD 917 BEE 15 E	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC621FFFAF4FA6113736D737E.taxon	materials_examined	Type material. Holotype: MNRJ 18712. Paratypes: Brazil: Bahia state: MNRJ 32913, 13 º 25 ʹ 43 ʺ S, 38 º 49 ʹ 30 ʺ W, 32 m, v / 2007 [1]. Rio de Janeiro state: MNRJ 18641, HAB 16 - G 3 [1]; MNRJ 32062, 23 º 13 ʹ 30 ʺ S, 41 º 21 ʹ 00 ʺ W, 93 m, x / 2008 [2]; IBUFRJ 2319, 23 º 42 ʹ 12 ʺ S, 43 º 01 ʹ 06 ʺ W, 129 m, 28 / iii / 1983 [1]. Other material examined. Brazil: Espírito Santo state: IBUFRJ 19503 [1], IBUFRJ 19602 [3], REVIZEE- Central C 1 - VV 38; MORG 52259, REVIZEE-Central C 1 - VV 22 [1]. Rio de Janeiro state: MORG 39415, Campos Basin [2]; MNRJ 18713, HAB 13 - I 4 [1]; MNRJ 32365, 22 º 42 ʹ S, 40 º 40 ʹ W, 110 – 120 m, ix / 2004 [1]; MNRJ 31120, 22 º 42 ʹ S, 40 º 40 ʹ W, 110 – 120 m, xi / 2007 [1]; IBUFRJ 11697 [5], IBUFRJ 11700 [1], IBUFRJ 19578 [3], 22 º 48 ʹ S, 40 º 45 ʹ W, 106 – 110 m, 27 / i / 1998; MNRJ 32045, 23 º 03 ʹ 18 ʺ S, 41 º 02 ʹ 06 ʺ W, 97 m, x / 2008 [1]; MORG 52213, 23 º 04 ʹ 14 ʺ S, 40 º 59 ʹ 31 ʺ W, 100 m, 17 / xii / 2004 [4]; MNRJ 31114, 23 º 04 ʹ 14 ʺ S, 40 º 59 ʹ 31 ʺ W, 100 m, iii / 2005 [1]; MNRJ 32398, 23 º 05 ʹ S, 40 º 59 ʹ W, 100 m, v / 2009 [1]; MNRJ 32543 [3], MORG 52235 [1], MORG 52243 [2], 23 º 05 ʹ S, 40 º 58 ʹ W, 100 m; MNRJ 32070, 23 º 09 ʹ 00 ʺ S, 41 º 02 ʹ 06 ʺ W, 100 m, x / 2008 [1]; MNRJ 60196, PADCT sta. 6627 [1]. São Paulo state: MNRJ 32920, 25 º 13 ʹ 06 ʺ S, 44 º 59 ʹ 31 ʺ W, 160 m, viii / 2008 [1]. Type locality. HAB 13 - H 5, 21 º 42 ʹ 33 ʺ S, 40 º 09 ʹ 06 ʺ W, 147 m, Rio de Janeiro state, Brazil.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC621FFFAF4FA6113736D737E.taxon	etymology	Etymology. facetus, L. = fine, elegant. It alludes to the refined shell sculpture and color pattern of this species.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC621FFFAF4FA6113736D737E.taxon	diagnosis	Diagnosis. Golden protoconch, with two spiral cords very close to each other, the abapical one much more prominent; adapical spiral cord of teleoconch with orange nodules alternating with white nodules; median spiral cord often emerges on the sixth whorl of the teleoconch, usually reaching the same size of the abapical cord after five whorls.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC621FFFAF4FA6113736D737E.taxon	description	Description. Shell sinistral, elongated, conical, rectilinear profile, reaching 8.95 mm in length, 2.07 mm in width, length / width ratio 3.9 to 4.3. Protoconch somewhat tumid, subcylindrical, 0.59 – 0.84 mm in length, 0.58 – 0.65 mm in width in its last whorl; 3.25 to 4 very convex whorls, clear distinction between protoconch and teleoconch; initial whorl dome-shaped, covered by several vesicles and spirally disposed granules; remaining whorls initially with two spiral cords, situated at 41 % and 54 – 67 % of penultimate whorl height, the abapical one much more prominent and gradually assuming a very keeled shape (barely resulting in a pagoda-shape to each protoconch whorl), the adapical one gradually diminishing and becoming much closer to the abapical cord, disappearing just before the transition to the teleoconch or even earlier; small, numerous faint axial riblets, more evident above the adapical cord, being nearly orthocline to much prosocline. Teleoconch with up to 12 whorls; two spiral cords (adapical and abapical) in the beginning, the abapical one continuous with that of the protoconch; median spiral cord emerges narrowly at the end of the fifth to the beginning of the seventh whorl, reaching the same size of the abapical cord (the adapical one is more prominent, especially in late whorls) after 4.5 to 5.5 whorls; on the body whorl, distance between spiral cords is 0.9 to 1.2 times the width of cords; 18 to 21 strongly opisthocline axial ribs; rounded (especially the adapical cord) to slightly elliptical (median and abapical cords) nodules of a medium to moderately large size; distinct and well-developed suture, with a small sutural cord; weakly nodulose to wavy subperipheral cord, one narrow, slightly wavy basal cord, besides a smooth thickening below it, directed downwards, not developing in a true abapical basal cord; no supranumerical cords; ovate aperture, 0.98 – 1.22 mm long, 0.87 – 1.03 mm wide, length / width ratio 1.1 – 1.2; anterior canal curved almost downwards, moderately long, partly or entirely closed, 0.58 – 0.69 mm long, 0.37 – 0.43 mm wide, length / width ratio 1.5 – 1.6; posterior canal a deep, wide sinus, 0.16 – 0.19 mm long, not detached from the aperture. Protoconch golden; cream / whitish background of teleoconch, the adapical spiral cord irregularly tinted with one or two orange nodules usually alternating with one or two (hardly ever three) white nodules, with the orange patches occasionally reaching the median spiral cord; light brown / orange base.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC621FFFAF4FA6113736D737E.taxon	discussion	Remarks. This is a fairly common species in Campos Basin, southeastern Brazil, possessing very distinct features, like the protoconch sculpture with two close spiral cords (Fig. 14 I – K), the gradual development of the median spiral cord of the teleoconch (reaching the same size of the abapical cord about five whorls after its emergence) and the notable color pattern of the adapical spiral cord of teleoconch (Fig. 14 A – D). The teleoconch of the western Atlantic species Monophorus olivaceus (Dall, 1889) superficially resembles that of “ Inella ” faceta sp. nov., especially in the irregular brown / orange blotches alternating with white nodules in the adapical spiral cord. Geographical distribution. Brazil: Bahia to São Paulo.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC621FFFAF4FA6113736D737E.taxon	description	Bathymetric distribution. 32 – 160 m.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC627FFF8F4FA61AC736D737E.taxon	description	Figure 15 urn: lsid: zoobank. org: act: 667 B 1 EA 1 - E 5 CF- 42 FC- 85 D 8 - D 0 E 174 AD 95 AA	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC627FFF8F4FA61AC736D737E.taxon	materials_examined	Type material. Holotype: MNRJ 32392, v / 2009. Paratypes: Brazil: Espírito Santo state: IBUFRJ 19502, REVIZEE- Central C 1 - VV 38 [4]. Rio de Janeiro state: MNRJ 18966, 23 º 04 ʹ 14 ʺ S, 40 º 59 ʹ 31 ʺ W, 100 m, 17 / xii / 2004 [2]. Other material examined. Brazil: Espírito Santo state: MNRJ 30698, 19 º 26 ʹ 22 ʺ S, 39 º 15 ʹ 22 ʺ W, 51 m, x / 2003 [1]; IBUFRJ 19604, REVIZEE-Central C 1 - VV 38 [5]; MNRJ 32660, 20 º 14 ʹ S, 40 º 12 ʹ W, vi / 2008 [1]; MNRJ 33798, REVIZEE-Central C 1 - VV 21 [1]; MNRJ 33790, REVIZEE-Central C 6 - Y 7 [1]; IBUFRJ 19695, Piú- ma, 1993 [9]. Almirante Saldanha Seamount: MNRJ 33761, REVIZEE-Central C 1 - D 1 [2]. Rio de Janeiro state: IBUFRJ 7683, 22 º 08 ʹ S, 40 º 31 ʹ W, 55 m, 29 / viii / 1979 [1]; IBUFRJ 19595, REVIZEE-Central C 1 - D 3 [2]; MNRJ 32393, type locality, v / 2009 [1]. Type locality. 23 º 05 ʹ S, 40 º 59 ʹ W, 100 m, Rio de Janeiro state, Brazil.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC627FFF8F4FA61AC736D737E.taxon	etymology	Etymology. maculata, L. = stained. It alludes to the color pattern of this shell.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC627FFF8F4FA61AC736D737E.taxon	diagnosis	Diagnosis. Protoconch with two main spiral cords and a nodulose subsutural spiral thread; median spiral cord usually emerges on the second or third teleoconch whorl; teleoconch with irregular light brown / orange patches often comprising all spiral cords on the same whorl.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC627FFF8F4FA61AC736D737E.taxon	description	Description. Shell sinistral, elongated, conical, rectilinear profile, reaching 9.69 mm in length, 2.07 mm in width, length / width ratio 3.8 to 4.6. Protoconch subcylindrical / columnar, robust, 0.67 – 1.13 mm in length, 0.57 – 0.82 mm in width of its last whorl; three to four slightly convex whorls, weak distinction between protoconch and teleoconch; nucleus very small, slightly elevated, in an oblique descent, mainly smooth; two main spiral cords present from nearly the beginning of the protoconch, the abapical one initially slightly more prominent, but gradually both assume the same size, situated at 29 – 31 % and 63 – 73 % of last whorl height; a small subsutural spiral thread is also present, crossed by small axial wrinkles, resulting in a very nodulose sculpture of this spiral thread; the adapical spiral cord is slightly to moderately nodulose, especially on the last whorl, but the abapical one is smooth. Teleoconch with up to 12 whorls; two spiral cords (adapical and abapical) in the beginning, both continuous with those of the protoconch; median spiral cord emerges narrowly at the end of the first to the beginning of the fourth whorl, but usually on the second / third whorl, reaching the same size of other cords after two to 3.5 whorls; on the body whorl, distance between spiral cords is 1.1 to 1.2 times the width of cords; 19 to 22 nearly orthocline to strongly opisthocline axial ribs; rounded nodules of a medium to moderately large size; distinct, well-developed suture, with a small sutural cord; very nodulose subperipheral cord, two basal cords, the adapical one nodulose and close to the subperipheral cord, the abapical one smooth; a small supranumerical cord may develop between the median and abapical spiral cords; ovate aperture, length / width ratio 1.3; anterior canal curved downwards / backwards, moderately short to medium, and partly closed, length / width ratio 1.4; posterior canal a deep sinus, not detached from the aperture. Protoconch brown; cream / whitish background of teleoconch, with irregular, axially disposed, light brown / orange patches often comprising all spiral cords on the same whorl, and usually not exceeding a width of three axial ribs, but in late whorls of some shells the colored patches can fill the width of several axial ribs; light brown / orange base.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC627FFF8F4FA61AC736D737E.taxon	discussion	Remarks. “ Inella ” maculata sp. nov. presents a considerable range in the dimensions of the protoconch, which can be explained by the number of whorls attained (three to four), and in the emergence and strengthening of the median spiral cord of the teleoconch; notwithstanding, these features represent continuous intraspecific variations. An anomalous shell from Espírito Santo state (Fig. 15 D, J), with smaller dimensions (0.61 mm in length, 0.53 mm in width) of a golden protoconch (not brown as in typical shells), is tentatively referred to this species. The light brown / orange patches on the teleoconch of “ Inella ” maculata sp. nov. (Fig. 15 A – D) resemble those on the shell of “ Inella ” faceta sp. nov. (Fig. 14 A – D). The main differences between them are related to an axial pattern of coloration in “ Inella ” maculata sp. nov. (but spirally oriented on the adapical spiral cord of “ Inella ” faceta sp. nov.), the earlier emergence of the median spiral cord of the teleoconch in “ Inella ” maculata sp. nov. (usually on the second / third whorl, but on the sixth whorl in “ Inella ” faceta sp. nov.) and very different protoconch sculptures. Despite the same pattern of coloration, Inella sp. 1 has the adapical spiral cord of teleoconch initially reduced (Fig. 5 I), instead of the median one emerging later in “ Inella ” maculata sp. nov. (Fig. 15 F), in addition to significant differences in protoconch sculpture. The teleoconch of “ Inella ” maculata sp. nov. is most similar to that of Nototriphora decorata (C. B. Adams, 1850), a very common species in the tropical western Atlantic. The protoconch of “ Inella ” maculata sp. nov. is similar to that of the Australian species Inella intercalaris Marshall, 1983, although the teleoconch of the latter species belongs to the Inella s. s. group (Marshall, 1983: fig. 12 B). Geographical distribution. Brazil: Espírito Santo, Almirante Saldanha Seamount and Rio de Janeiro.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC627FFF8F4FA61AC736D737E.taxon	description	Bathymetric distribution. 51 – 105 m.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61BFFC6F4FA64F873DE752A.taxon	materials_examined	Type species. Triforis ibex Dall, 1881. Original designation. Recent, Gulf of Mexico. Diagnosis. See Fernandes & Pimenta (2014).	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61BFFC4F4FA679072F971B0.taxon	materials_examined	Type material. Holotype: MZSP 78886 (Fig. 17 A – B). Paratypes: MZSP 78890 (Fig. 17 C – F), type locality [4]. Type locality. 02 º 14 ʹ 25 ʺ S, 38 º 22 ʹ 50 ʺ W, 240 – 260 m, Canopus Bank, off Ceará state, Brazil. Material examined. The type material and: Brazil: off Maranhão state: USNM 810450, 0 º 18 ʹ N, 44 º 16 ʹ 48 ʺ W, 274 m, Oregon R / V coll., 09 / iii / 1963 [2]. Off Ceará state: MNRJ 30402 [5], MZSP 66560 [1], MZSP 93891 [4], MZSP 94251 [1], type locality, P. M. S. Costa & J. Coltro coll., xi / 2005; MZSP 53729 [1], MZSP 90270 [2], MZSP 70277 [5], MZSP 70291 [2], MZSP 133330 [8], type locality, J. Coltro & C. M. Cunha coll., viii / 2005.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61BFFC4F4FA679072F971B0.taxon	description	Description. See Simone (2006) for the description of the teleoconch. The protoconch is herein described, based on lots MZSP 53729, MZSP 66560, MZSP 70277 and MZSP 90270, all from the type locality: protoconch globose, much to little inflated, 0.69 – 0.76 mm in length, 0.48 – 0.63 mm in width of its first whorl, 0.52 – 0.62 mm in width of its last whorl; 2.75 to 3.0 whorls, unclear distinction between protoconch and teleoconch, transition defined by a fine, sinuous / opisthocline axial mark; initial 1.5 or 1.75 whorls dome-shaped, broad, white, mostly smooth and translucent; remaining 1.25 whorls with a distinct or faint light brown color, with two main spiral cords, the abapical one initially slightly more prominent, in addition to two minute subsutural cords and one small cord situated right above the suture; two main spiral cords respectively situated at 40 – 46 % and 67 – 75 % of last whorl height; axial sculpture absent except by fine axial marks present above the adapical spiral cord.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61BFFC4F4FA679072F971B0.taxon	discussion	Remarks. This is one of the largest triphorids of the world, reaching 49.7 mm in shell length (Simone 2006). The main problem related to the original description of S. unicornium is that its single paratype lot is a mix of three species (Fig. 17), one of them being Strobiligera gaesona, which is recorded in Brazil in the present study (see below). The protoconch illustrated by Simone (2006) as being S. unicornium actually belongs to S. gaesona, and the real protoconch of S. unicornium is herein described and illustrated (Fig. 19), fitting the globose paucispiral protoconch of Strobiligera and rendering the new generic allocation. Indeed, worn shells of S. unicornium and S. gaesona may be mistaken, but they differ on the broader profile of S. unicornium, its expanded outer lip, the reduced adapical spiral cord through most whorls of the teleoconch, a stronger spiral sculpture on the teleoconch (also developing small elliptical granules, absent in S. gaesona), and protoconch with a faint or distinct light brown color (but entirely white in S. gaesona) and discrete axial marks above its adapical spiral cord. Simone (2006) differentiated S. unicornium from Inella colon (Dall, 1881) (Fig. 3 W), from Cuba and Florida (U. S. A.), by having a larger shell size, proportionally longer whorls, taller spiral cords and a projected outer lip. Two shells identified by Rolán & Fernández-Garcés (2008: figs. 16 E – F) as I. colon, although they noted differences compared to the type material of this species, are very similar to S. unicornium; one shell is from Yucatan (Mexico), the other from Florida (U. S. A.). These shells seem to present a much reduced adapical spiral cord on the teleoconch, which is somewhat similar to the holotype of S. unicornium, but not to the majority of shells of this species (with a gradually developing adapical spiral cord). In addition, their bathymetric range (236 – 366 m) is more similar to that of S. unicornium (240 – 274 m) than to the type material of I. colon (823 – 1834 m); further comparison is required to evaluate if S. unicornium has a wider geographical range. Despite the broken apex, the two shells of USNM 810450 (Fig. 18 G – H) are herein referred to S. unicornium owing to a very large size, similar teleoconch sculpture (although the adapical spiral cord is slightly more developed), identical bathymetric records and close geographical distribution, thus extending the known range of this species to Maranhão state. Geographical distribution. Brazil: off Maranhão (this study), off Ceará (type locality).	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61BFFC4F4FA679072F971B0.taxon	description	Bathymetric distribution. 240 m (type locality) to 274 m (this study).	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC619FFC2F4FA626F72F97235.taxon	description	Figures 3 S – T, 20	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC619FFC2F4FA626F72F97235.taxon	materials_examined	Material examined. The type material and: Brazil: off Ceará state: MNRJ 30401 [6], MZSP 70305 [6, all with dried soft parts], MZSP 78890 [4, but two shells worn / unidentifiable], MZSP 131829 [8], Canopus Bank, 02 º 14 ʹ 25 ʺ S, 38 º 22 ʹ 50 ʺ W, 240 – 260 m, P. M. S. Costa & J. Coltro coll., xi / 2005; MZSP 37077 [2], MZSP 53685 [2], MZSP 53697 [2], MZSP 133332 [1], MZSP 66850 [1], MZSP 70300 [6], MZSP 133341 [4], MZSP 133331 [6], Canopus Bank, 02 º 14 ʹ 25 ʺ S, 38 º 22 ʹ 50 ʺ W, 240 – 60 m, J. Coltro & C. M. Cunha coll., viii / 2005.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC619FFC2F4FA626F72F97235.taxon	description	Description. Shell sinistral, elongated, conical-fusiform, rectilinear profile, reaching 28.56 mm in length, 4.03 mm in width, length / width ratio 7.5 to 7.7. Protoconch globose, much to little inflated, 0.66 – 0.86 mm in length, 0.41 – 0.68 mm in width of its first whorl, 0.45 – 0.61 mm in width of its last whorl; 2.75 to 3.5 slightly convex whorls (except first whorl), unclear distinction between protoconch and teleoconch, transition defined by a very fine, sinuous / opisthocline axial mark; initial whorl dome-shaped, broad, smooth; remaining whorls with two main spiral cords, situated at 45 – 52 % and 66 – 76 % of whorl height, with nearly the same strength or with the adapical spiral slightly more prominent, in addition to a small subsutural cord, rapidly enlarging to the end of the protoconch; axial sculpture absent. Teleoconch with up to 24 whorls; three spiral cords in the beginning, all of them continuous with those of the protoconch, the adapical one initially smallest but soon reaching the same size of others; very weak spiral sculpture, almost indistinct even in initial whorls, with only the micro spiral sculpture (composed of several threads) being evident in late whorls, but some shells have a strengthening of the spiral sculpture of median and abapical cords on late whorls; complete absence of axial sculpture over most of the shell, despite sinuous growth lines or discrete threads in some shells; absence of nodules, whorls totally flat, without any convexity; very indistinct suture, only marked by a small sutural cord; narrow, smooth subperipheral cord, two smooth basal cords; no supranumerical cords are distinguishable on the inconspicuous end of the body whorl; ovate aperture, with a moderately expanded outer lip, 1.09 – 1.32 mm long, 0.96 – 1.01 mm wide, length / width ratio 1.1 – 1.4; anterior canal very short, directed downwards, and totally open, 0.23 – 0.29 mm long, 0.36 – 0.43 mm wide, length / width ratio 0.6 – 0.8; posterior canal a very small notch, almost indistinct, 0.15 – 0.17 mm long. White shell, with a translucent initial protoconch whorl.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC619FFC2F4FA626F72F97235.taxon	discussion	Remarks. As mentioned above, the single lot of paratypes of Strobiligera unicornium is a mix of three species, including Strobiligera gaesona (Fig. 17 E), whose protoconch was illustrated by Simone (2006: figs. 3 – 4). Shells from Brazil (Fig. 20) are remarkably similar to the lectotype of S. gaesona (Fig. 3 S), without any clear difference, except the bathymetric range of this species is shallower in Brazil (240 – 260 m) than in the type locality (805 m). Some types also show a moderately strong spiral sculpture on earlier whorls (Fig. 3 T), fading out later, which is slightly different from the very weak spiral sculpture since the beginning of the teleoconch in Brazilian shells; as mentioned above, shells of Strobiligera dinea (Dall, 1927) may be mixed in that lot. Several shells of S. gaesona from Brazil had drill holes indicative of predation by gastropods. Their smooth shells, without nodules, seem to be prone to suffer attacks. Geographical distribution. U. S. A.: off Georgia (type locality); Brazil: off Ceará (this study).	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC619FFC2F4FA626F72F97235.taxon	description	Bathymetric distribution. 240 m (this study) to 805 m (type locality).	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61DFFC1F4FA65BC739F762E.taxon	description	Figure 22	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61DFFC1F4FA65BC739F762E.taxon	materials_examined	Material examined. Brazil: off Espírito Santo state: MNHN, MD 55 41 - DC 73 [1].	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61DFFC1F4FA65BC739F762E.taxon	description	Description. Shell sinistral, elongated, very thin, almost fusiform, rectilinear profile, reaching 3.19 mm in length, 0.77 mm in width, length / width ratio 4.1. Protoconch slightly globose, subcylindrical, 0.47 mm in length, 0.43 mm in width of its last whorl; 2.75 convex protoconch whorls, weak distinction between protoconch and teleoconch; initial whorl somewhat pointed, mostly smooth, but with several minute, irregular granules especially concentrated on its abapical portion; remaining whorls initially with a strong spiral cord in a median / adapical position of the whorl, assuming an abapical position and a keeled shape; an adapical cord appears soon after, much narrower than the abapical one until the end of protoconch, in addition to a small subsutural thread; the two main cords are situated at 26 % and 60 % of last whorl height. Teleoconch with up to ~ 8.5 whorls; three spiral cords in the beginning, continuous to those of protoconch, the adapical one narrower than other cords through the entire teleoconch; on the body whorl, distance between spiral cords is 2.1 times the width of cords; 15 opisthocline axial ribs; rounded nodules of a medium size; distinct suture, with a small sutural cord; narrow, slightly wavy subperipheral cord, one smooth, very narrow basal cord, close to the subperipheral cord; shortened base; no supranumerical cords; ovate aperture, length / width ratio 1.3; anterior canal partly broken; posterior canal not formed. White shell.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61DFFC1F4FA65BC739F762E.taxon	discussion	Remarks. Strobiligera sp. 1 is similar to shells of S. dinea from Brazil, the differences between them consisting of the width of the teleoconch (1.20 mm in a shell of of S. dinea 3.83 mm in length, 0.77 mm in a shell of Strobiligera sp. 1 3.19 mm in length), the dimensions of the protoconch (first whorl much inflated in S. dinea) and the abapical cord of the protoconch being more prominent in Strobiligera sp. 1 (but the adapical one in S. dinea). The shell of Strobiligera sp. 1 is also apparently narrower than those of Strobiligera enopla (Dall, 1927) (Fig. 3 X) and Strobiligera meteora (Dall, 1927), both from the northwestern Atlantic (Rolán & Fernández-Garcés 2008), in addition to the adapical spiral cord of the teleoconch of Strobiligera sp. 1 being broader than that of S. enopla but narrower than that of S. meteora. Geographical distribution. Brazil: off Espírito Santo.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61DFFC1F4FA65BC739F762E.taxon	description	Bathymetric distribution. 607 – 620 m.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61CFFC1F4FA649C739F71F8.taxon	description	urn: lsid: zoobank. org: act: FBE 0 B 96 F- 3327 - 4 D 19 - BDA 0 - B 4 EDA 1 FA 970 A	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61CFFC1F4FA649C739F71F8.taxon	materials_examined	Type material. Holotype: MNRJ 32350, iii / 2007. Paratypes: Brazil: Rio de Janeiro state: MNRJ 18640, HAB 16 - G 5 [1]; MNRJ 19481, type locality, 11 / iv / 2003 [5]; MNRJ 32349, type locality, ix / 2004 [1]; MNRJ 18950, type locality, iii / 2007 [1]; MNRJ 31122, type locality, xi / 2007 [1]; IBUFRJ 11701 [1], IBUFRJ 19580 [5], 22 º 48 ʹ S, 40 º 45 ʹ W, 106 – 110 m, 27 / i / 1998; MNRJ 32546, 23 º 0 5 ʹ S, 40 º 58 ʹ W, 100 m [1]. Type locality. 22 º 42 ʹ S, 40 º 40 ʹ W, 110 – 120 m, Rio de Janeiro state, Brazil.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61CFFC1F4FA649C739F71F8.taxon	etymology	Etymology. This specific epithet refers to the citizens of Campos dos Goytacazes, in Rio de Janeiro state, because this species is apparently restricted to the Campos Basin (a sedimentary basin named for the above-mentioned city). Epithet as a noun in apposition.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61CFFC1F4FA649C739F71F8.taxon	diagnosis	Diagnosis. Light brown protoconch with 2.75 to 3 whorls, clearly distinct from the cream teleoconch; adapical spiral cord reaches nearly the same size as other spiral cords in the tenth / twelfth whorl of teleoconch; suture very shallow.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61CFFC1F4FA649C739F71F8.taxon	description	Description. Shell sinistral, elongated, conical, rectilinear profile, reaching 12.32 mm in length, 2.34 mm in width, length / width ratio 4.6 to 4.9. Protoconch globose, inflated, 0.51 – 0.65 mm in length, 0.45 – 0.53 mm in width of its last whorl; 2.75 to 3 convex protoconch whorls, clear distinction between protoconch and teleoconch; initial whorl dome-shaped, broad, smooth; remaining whorls with two spiral cords, with nearly the same strength, situated at 41 % and 76 % of last whorl height; micro-sculpture of subsutural axial wrinkles, more evident on the last whorl. Teleoconch with up to ~ 18 whorls; three spiral cords in the beginning, the median and abapical cords continuous with those of the protoconch; adapical cord narrower than other cords through most of teleoconch, but acquiring nearly the same size as other cords on the tenth / twelfth whorl; on the body whorl, distance between spiral cords is 1.2 times the width of cords; 14 to 16 nearly orthocline to opisthocline axial ribs; rounded (especially the adapical and median cords) to slightly elliptical (especially the abapical cord) nodules of a medium to moderately large size; distinct but shallow suture, with a small sutural cord; thick, smooth subperipheral cord, three smooth basal cords, the median one the narrowest of them; no supranumerical cords; ovate aperture, length / width ratio 1.2; anterior canal moderately short, directed downwards / backwards, and open, length / width ratio 0.9; posterior canal a small notch, not detached from the aperture. Light brown protoconch (first whorl sometimes translucent), cream teleoconch, spiral cords with nodules lighter than internodular spaces.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61CFFC1F4FA649C739F71F8.taxon	discussion	Remarks. Strobiligera campista sp. nov. mainly differs from S. dinea by shell color and by having a wider shell, but a smaller, less inflated protoconch (Fig. 23 J). Strobiligera enopla (Fig. 3 X) has a completely white shell and a much reduced adapical spiral cord on the teleoconch, in addition to a bathymetric range (538 – 823 m; Rolán & Fernández-Garcés 2008) deeper than that of Strobiligera campista sp. nov. (100 – 149 m). Geographical distribution. Brazil: Rio de Janeiro.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61CFFC1F4FA649C739F71F8.taxon	description	Bathymetric distribution. 100 – 149 m.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61CFFCCF4FA6227739F745E.taxon	description	Figure 24 urn: lsid: zoobank. org: act: 68 B 24 A 96 - B 28 A- 46 FC-A 366 - F 4 D 5 C 888 FB 4 B	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61CFFCCF4FA6227739F745E.taxon	materials_examined	Type material. Holotype: MZSP 32618. Paratypes: Brazil: São Paulo state: MNRJ 29382, PADCT sta. 6573 [2]; MNRJ 29374, REVIZEE-Sul sta. 6676 [1]; MNRJ 32917, 25 º 13 ʹ 0 6 ʺ S, 44 º 59 ʹ 31 ʺ W, 160 m, viii / 2008 [1].	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61CFFCCF4FA6227739F745E.taxon	description	Description. Shell sinistral, elongated, conical-fusiform, rectilinear profile, reaching 13.89 mm in length, 2.09 mm in width, length / width ratio 6.6. Protoconch globose, inflated, 0.63 – 0.78 mm in length, 0.45 – 0.56 mm in width of its first whorl, 0.49 – 0.58 mm in width of its last whorl; 2.75 convex whorls, clear distinction between protoconch and teleoconch; initial 1.5 whorl dome-shaped, broad, smooth; remaining 1.25 whorl with two main spiral cords, situated at 34 – 44 % and 61 – 69 % of whorl height, the abapical one slightly more pronounced; minute, very faint, axial marks across the entire whorl length. Teleoconch with up to 20 whorls; three spiral cords in the beginning, the median and abapical cords continuous with those of the protoconch; adapical cord narrower than other cords through most of teleoconch, but slowly strengthening and acquiring the same size of the abapical one (the median cord is more pronounced than others in mid-late whorls) only on the body whorl, in which the distance between spiral cords is 1.2 times the width of cords; 14 to 15 orthocline to slightly opisthocline axial ribs; rounded nodules of a medium size; well-developed suture, with a distinct sutural cord; moderately thick, smooth subperipheral cord, only one smooth, thin basal cord; no supranumerical cords; elliptical aperture, 1.35 mm long, 0.99 mm wide, ratio length / width 1.4; posterior and anterior canals not fully formed, with the latter being directed downwards. Light brown protoconch (first whorl usually translucent white), cream teleoconch, with nodules clearer than internodular spaces in the spiral cords.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61CFFCCF4FA6227739F745E.taxon	discussion	Remarks. Strobiligera santista sp. nov. is similar to Strobiligera campista sp. nov. both in teleoconch and protoconch sculpture, in addition to shell coloration. Their main differences comprise the nearly fusiform and thin shell shape of Strobiligera santista sp. nov. (more conical and broader in Strobiligera campista sp. nov.), the more gradual strengthening of the adapical spiral cord and presence of smaller nodules of teleoconch (making internodular spaces more evident in Strobiligera santista sp. nov.), the number of basal cords (one in Strobiligera santista sp. nov., three in Strobiligera campista sp. nov.), and the slightly larger dimensions of the protoconch in Strobiligera santista sp. nov. Preserved shells of Strobiligera campista sp. nov. are recorded as deep as ~ 150 m and are apparently restricted to the Campos Basin, thus showing a shallower and more northerly distribution than Strobiligera santista sp. nov., which is apparently restricted to the Santos Basin, between 153 m and 258 m. Strobiligera santista sp. nov. is distinguished from S. dinea by shell color (mainly cream teleoconch and light brown protoconch, instead of the white shell of S. dinea) and by having reduced dimensions of the protoconch, which is much inflated in S. dinea. Strobiligera santista sp. nov. also inhabits the transition between the continental shelf and slope (153 m to 258 m), contrary to the strictly bathyal records of S. dinea (607 m to 940 m).	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC61CFFCCF4FA6227739F745E.taxon	description	Geographical distribution. Brazil: São Paulo. Bathymetric distribution. 153 – 258 m.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC611FFCCF4FA6656739F705A.taxon	materials_examined	Material examined. Brazil: Rio Grande do Norte state: MNRJ 35113, 04 º 44 ʹ 53 ʺ S, 36 º 25 ʹ 27 ʺ W, 102 – 108 m, 23 / v / 2011 [5].	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC611FFCCF4FA6656739F705A.taxon	description	Description. Shell sinistral, elongated, conical, rectilinear profile, reaching 7.12 mm in length, 1.84 mm in width. Protoconch globose, inflated, 0.55 – 0.60 mm in length, 0.46 – 0.50 mm in width of its last whorl; 2.5 to 2.7 convex whorls, weak distinction between protoconch and teleoconch; first initial whorl dome-shaped, broad, mainly smooth but covered by minute, irregular-shaped granules; remaining whorls with two spiral cords, equal in strength, situated at 36 – 43 % and 66 – 71 % of last whorl height; intense micro-sculpture of axial wrinkles just below the suture. Teleoconch with up to 13.5 whorls; three spiral cords in the beginning, median and abapical ones continuous with those of protoconch; adapical cord narrower than other cords over the entire teleoconch; on the 13 th whorl, distance between spiral cords is 1.3 times the width of cords; 13 to 15 strongly opisthocline axial ribs; elliptical nodules, considerably pointed and partially cut in the middle, of a medium size; distinct suture, with a small sutural cord; base not formed. White shell, internodular spaces slightly darker than nodules of teleoconch.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC611FFCCF4FA6656739F705A.taxon	discussion	Remarks. Strobiligera sp. 2 is distinguished from S. dinea by possessing a smaller, less inflated protoconch (Fig. 25 E, G), minute granules on the first whorl of protoconch (Fig. 25 F; not entirely smooth as in S. dinea: Fig. 21 E), a conical shell (Fig. 25 A – D; not fusiform as in S. dinea: Fig. 21 A – B), and internodular spaces slightly darker than nodules of teleoconch. Strobiligera sp. 2 has similar teleoconch coloration to that of Inella pseudotorticula (Fig. 3 O – P), with internodular spaces darker than nodules, and a similar protoconch sculpture. Slight differences such as the more conical shell shape of Strobiligera sp. 2, its smaller adult shell size (although no complete adult shell was yet sampled) and the wide geographical isolation of both populations (I. pseudotorticula is only known from Bermuda) suggest that they are different species. Complete adult shells are necessary for the description of this species.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC611FFCCF4FA6656739F705A.taxon	description	Geographical distribution. Brazil: Rio Grande do Norte. Bathymetric distribution. 102 – 108 m.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC616FFC9F4FA60187033769A.taxon	description	Figure 27	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC616FFC9F4FA60187033769A.taxon	materials_examined	Material examined. Brazil: off Rio de Janeiro state: MNRJ 26140, REVIZEE-Sul sta. 6753 [approximately 100 larvae and post-larvae and two pieces of biogenic substrata].	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
E33C87BDC616FFC9F4FA60187033769A.taxon	discussion	Remarks. An egg laying of a Strobiligera species was sampled by the Expedition ‘ REVIZEE-Sulʹ in southeast- ern Brazil. This spawn includes nearly 100 larvae and post-larvae associated with two fragments of hard biogenic substrata (Fig. 27). The major fragment measures 18.3 mm in length and is mainly dominated by an encrusted sponge. The minor fragment measures 5.6 mm x 3.2 mm and is composed of a bryozoan colony partly covered by encrusted sponges, especially on its underside. A few post-larvae were still associated with the substrata when photographed (Fig. 27 C – D, F), but the majority were detached and many exhibited distinct spicules adhering to the shell, indicating a moderately loose attachment (i. e., post-larvae were not embedded in the sponge). The light brown protoconch has usually 2.5 whorls, and the white teleoconch reaches one complete whorl. The protoconch is 0.61 – 0.70 mm long, 0.42 – 0.48 mm wide on the first, smooth and inflated whorl, 0.51 – 0.56 mm wide on the last whorl. Two smooth spiral cords are present on the last protoconch whorl, the adapical one situated at 41 – 44 % of whorl height, the abapical one at 68 – 79 %; minute axial marks are present below the suture. The spiral cords of protoconch originate the nodulose median and abapical cords of teleoconch, with the adapical cord of teleoconch emerging immediately after metamorphosis. The most probable identity of this spawn is Strobiligera campista sp. nov., because of protoconch color, similar dimensions and positioning of spiral cords at the larval shell, in addition to being a moderately common species in the lower continental shelf of the sampled locality. The present findings shed light on the larval development of the genus, in this case not a lecithotrophic development with a short demersal stage, but a probable example of intracapsular metamorphosis, which suggests a restricted geographical range of this species. The egg laying in sponges is similar to previous observations in Cerithiopsidae (Nützel 1998; Collin 2004) and Newtoniellidae (Schiaparelli et al. 2003), constituting the second observed spawning of a Triphoridae species, after Pelseneer (1926): he described the spawning of a species related to the “ Triphora perversa ” complex (Bouchet & Guillemot 1978) in a bivalve shell from France.	en	Fernandes, Maurício Romulo, Pimenta, Alexandre Dias (2019): Taxonomic review of Inella and Strobiligera (Gastropoda: Triphoridae) from Brazil. Zootaxa 4613 (1): 1-52, DOI: 10.11646/zootaxa.4613.1.1
