taxonID	type	description	language	source
E245A643FF91FFFC73AFFE42FE3AD24E.taxon	materials_examined	Type species: Mucomyia emersa Kvifte and Curler, sp. nov., by present designation. Other included species: Mucomyia browni Kvifte, sp. nov.	en	Kvifte, Gunnar Mikalsen, Curler, Gregory R., Marshall, Stephen A. (2018): Aquatic insects in the forest canopy: a new genus of moth flies (Diptera: Psychodidae) developing in slime on aerial roots. Journal of Natural History 52 (3 - 4): 137-153, DOI: 10.1080/00222933.2017.1410590, URL: http://dx.doi.org/10.1080/00222933.2017.1410590
E245A643FF91FFFC73AFFE42FE3AD24E.taxon	diagnosis	Diagnosis Adult male. Eyes with very dense row of supraocular setae, single row at eyebridge, several rows laterally; labellum bulbous, fleshy, diminutive; palp shorter than head length, with 4 th palpomere fully sclerotized; antennae with 14 flagellomeres, flagellomeres 1 – 11 carrying one pair of Y-shaped ascoids and two or three pairs of additional smaller sensillae; distal three flagellomeres diminutive, not fused; wing membrane with setae; gonostylus without subapical bristle, gonocoxal condyles dorsally with wingshaped sclerites connected to the base of parameres and presumably the subepandrial plate; aedeagus of two phallomeres that either are separate and asymmetrical (M. emersa) or fused and symmetrical (M. browni); hypoproct oval, broad, with non-sclerotized fleshy lateral projections dorsal to bases of surstyli. Pupa. Mesonotum with conspicuous transverse bands of microtrichia inserted anteriorly, dorsally; posterior margins of abdominal segments 1 – 8 each with fringe of prominent microtrichia. 4 th instar larva. Antenna with single prominent mushroom element; postmentum truncate, without teeth; trunk hirsute; tergal plates each with row of prominent microtrichia along posterior margin.	en	Kvifte, Gunnar Mikalsen, Curler, Gregory R., Marshall, Stephen A. (2018): Aquatic insects in the forest canopy: a new genus of moth flies (Diptera: Psychodidae) developing in slime on aerial roots. Journal of Natural History 52 (3 - 4): 137-153, DOI: 10.1080/00222933.2017.1410590, URL: http://dx.doi.org/10.1080/00222933.2017.1410590
E245A643FF91FFFC73AFFE42FE3AD24E.taxon	etymology	Etymology From Latin mucus, slime and Greek myia, fly, referring to the larval habitat of the type species. The grammatical gender of the genus is feminine.	en	Kvifte, Gunnar Mikalsen, Curler, Gregory R., Marshall, Stephen A. (2018): Aquatic insects in the forest canopy: a new genus of moth flies (Diptera: Psychodidae) developing in slime on aerial roots. Journal of Natural History 52 (3 - 4): 137-153, DOI: 10.1080/00222933.2017.1410590, URL: http://dx.doi.org/10.1080/00222933.2017.1410590
E245A643FF91FFFC73AFFE42FE3AD24E.taxon	discussion	Remarks In Psychodinae, the mushroom element of the larval antennae is unique to Psychodini (Duckhouse 1985). This structure could be derived from an enlarged and / or fused series of sensilla basiconica. Nonetheless, for larval Psychodini it is a diagnostic character that can easily be recognized. The head capsule of Mucomyia, particularly the postmentum, is similar to that of Philosepedon humeralis (Meigen, 1818) (Vaillant 1974, p. 113, figure 247). Diagnostic characters given above for the larva and pupa are unique relative to descriptions that have been published to date. Adults of Mucomyia key to Philosepedon Eaton in keys to Neotropical genera of Psychodidae (Quate 1996; Wagner and Ibañez-Bernal 2009). Philosepedon was, however, recently redefined and restricted by Omelková and Ježek (2012) and later Cordeiro et al. (2015), and in the strict sense it currently only comprises species in the Afrotropical and Palearctic regions. The generic classification of Psychodini is currently being reassessed, with new genera regularly being described and ‘ traditional’ genera being redefined (Curler and Moulton 2008; Omelková and Jezek 2012; Cordeiro et al. 2015; Kvifte 2015; Ježek and Le Pont 2016), and this process is likely to continue given the high diversity particularly in the Neotropical and Oriental regions. Since many taxa still are insufficiently characterized or undescribed, it is premature to offer a detailed differential diagnosis against other taxa; however, the combination of characters given in the diagnosis above separates Mucomyia from all other described world genera of Psychodini.	en	Kvifte, Gunnar Mikalsen, Curler, Gregory R., Marshall, Stephen A. (2018): Aquatic insects in the forest canopy: a new genus of moth flies (Diptera: Psychodidae) developing in slime on aerial roots. Journal of Natural History 52 (3 - 4): 137-153, DOI: 10.1080/00222933.2017.1410590, URL: http://dx.doi.org/10.1080/00222933.2017.1410590
E245A643FF96FFF27387FDEDFE41D636.taxon	description	(Figures 1 – 6)	en	Kvifte, Gunnar Mikalsen, Curler, Gregory R., Marshall, Stephen A. (2018): Aquatic insects in the forest canopy: a new genus of moth flies (Diptera: Psychodidae) developing in slime on aerial roots. Journal of Natural History 52 (3 - 4): 137-153, DOI: 10.1080/00222933.2017.1410590, URL: http://dx.doi.org/10.1080/00222933.2017.1410590
E245A643FF96FFF27387FDEDFE41D636.taxon	materials_examined	Type material Holotype male. COSTA RICA, Heredia Province: Braulio Carrillo National Park, Quebrada Gonzalez loop trail, (WGS 84) 10 ° 9 ʹ 39.23 ʺN 83 ° 56 ʹ 18.41 ʺW, 527 m, 15. viii. 2013, leg. S. A. Marshall; deposited in MRCA. Specimen dissected, mounted on micro-slide. Other material Same data as holotype, 1 instar IV larval exuvia, 1 pupal exuvia (mounted in Canada Balsam on one slide); deposited in LACM.	en	Kvifte, Gunnar Mikalsen, Curler, Gregory R., Marshall, Stephen A. (2018): Aquatic insects in the forest canopy: a new genus of moth flies (Diptera: Psychodidae) developing in slime on aerial roots. Journal of Natural History 52 (3 - 4): 137-153, DOI: 10.1080/00222933.2017.1410590, URL: http://dx.doi.org/10.1080/00222933.2017.1410590
E245A643FF96FFF27387FDEDFE41D636.taxon	diagnosis	Diagnosis Frontal scar patch without dorsal extension leading to eyebridge. Parabasal processes parallel to upper margins of parameres, lateral base less than twice as broad as (medial) apex. Parameres not distinctly hooked apicolaterally. Aedeagus with two distal phallomeres of slightly different length. Surstylus with single apical tenaculum.	en	Kvifte, Gunnar Mikalsen, Curler, Gregory R., Marshall, Stephen A. (2018): Aquatic insects in the forest canopy: a new genus of moth flies (Diptera: Psychodidae) developing in slime on aerial roots. Journal of Natural History 52 (3 - 4): 137-153, DOI: 10.1080/00222933.2017.1410590, URL: http://dx.doi.org/10.1080/00222933.2017.1410590
E245A643FF96FFF27387FDEDFE41D636.taxon	description	Description Adult male (n = 1). Head (Figure 5 (a )) broadly oval; vertex a third of head length, hind margin of head nearly straight except median thoracic joint; eyebridge of four facet rows, separated by 1.5 facet diameters; interocular suture broadly V-shaped; more than 20 supraocular setae present, reaching stems of interocular suture, uniseriate dorsal to eyebridges, biseriate dorsal to eye, triseriate laterally; frontal scar patch broadly oval with median posterior extension reaching level of anterior margin of eyebridge; clypeus trapezoid, anterior margin about half as broad as posterior margin, sides tapering evenly; frontoclypeal suture complete, broadly U-shaped; labrum, maxillae and labium as long and wide as anterior width of clypeus; labellae fleshy, lateral margins more strongly sclerotized, with six dorsolateral and three ventral setae; palp of four segments, cylindrical except fourth segment spool-shaped; fourth palp segment fully sclerotized; length of palp segments 50: 60: 70: 70; antennae incomplete in type specimen, with only scape, pedicel and first five flagellomeres preserved; scape cylindrical, pedicel globular, flagellomeres nodiform symmetrical; ascoid attachments paired, present on all flagellomeres; flanked by smaller circular or spiniform sensilla (hard to ascertain in specimen); length of scape, pedicel and first five flagellomeres 70: 55: 100: 90: 90: 90: 85; ascoids Y-shaped; thorax (Figure 3) with dorsum, anepisternum, laterotergite, scutellum and ventral half of katepisternum setose; postpronotal lobe, anepimeron, katepisternum and meron naked; prothoracal spiracle naked, metathoracic spiracle densely setose; coxae all with dorsoventral anterior aseriate rows of setae, densest at ventral half of mid coxa; pointed extensions present on anterior side of front tibia and posterior end of mid and hind tibiae; tarsomere 5 without dorsal extension; wing (Figure 3) 1.4 mm long, not fully sclerotized in type specimen, veins generally hard to discern; C with two breaks, Sc terminating in wing membrane but approaching R 1, reaching halfway between base of M 1 + 2 and R 5; wing membrane setose; R 5 terminating in wing apex; radial fork distal to medial fork, complete; medial fork hard to discern; jugum broadly angulate oval, setose; narrow oval patch of setae present dorsal to jugum; male genitalia (Figure 5 (b )) with hypandrium membranous, curved concavely at median, reaching tips of gonocoxites; gonocoxites reniform-cylindrical, 1.3 × length of gonostyles; gonostyles conical with blunt apices, with scattered sensilla; aedeagus with basiphallus narrow at base, distally with median keel; distiphallus apparently spatulate with two phallomeres originating from lateral margins of apex of basiphallus; phallomeres narrow, pointed apically, left phallomere slightly longer than right; gonocoxal condyles wing-shaped, flat, approaching medially but not touching, with margin strip-like, thicker and more strongly sclerotized than rest of condyle; subtriangular parameres present, presumably originating from sclerotized posterior margin of gonocoxite, hooked at medial basal end at junction with subepandrial membrane, distally hooked towards the lateral; parameres as long as gonocoxite; epandrium (Figure 5 (b )) broader than long; base with strip-like apodeme, concave without laterobasal projections; single oval aperture present at median; fleshy setose projections present laterally on distal third; hypoproct oval with slightly pointed lateral edges, fused with subepandrial membrane; subepandrial membrane triangular, tapering towards base; epiproct (not illustrated) present as strip-like sclerite on ventral distal edge of hypoproct; surstyli cigar-shaped, about as long as epandrium; with single distal tenaculum; tenaculum spatulate with longitudinal striations, apex slightly pointed but without fringes. Pupa (Figures 4 (c), 4 (d) and 6 (e – h )). Cuticle glabrous, without pigment. Respiratory organ digitiform; apical 1 / 3 of respiratory organ curved anteriorly, with paired, longitudinal row of pores dorsally, with two pores at mid-length of organ, at apices of bumps. Mesonotum with conspicuous transverse bands of microtrichia inserted anteriorly, dorsally. Abdominal segments 1 – 8 with fringe of prominent microtrichia along posterior margins; segment 9 with four protuberances lateroventrally on each side; lateroventral protuberances each with a seta inserted apically; hook-shaped protuberances posterodorsally, conical protuberances posteroventrally; posteroventral protuberances each with a prominent seta inserted apically; genital lobes each with a spiniform protuberance flanked by setae apically. 4 th instar larva (Figures 4 (a), 4 (b) and 6 (a – d )). Head capsule ovate, brown; antenna composed of single prominent mushroom element, one digitiform, six globular sensilla basiconica, two sensilla trichodea; labrum covered with simple microtrichia ventrally, with one sensilla basiconica inserted anteriorly on each side; maxillary palp cylindrical, rounded apically, with single seta inserted at apex; lacinia composed of spiniform and setiform microtrichia; mandibles each with five dentations apically; prostheca composed of one brush-like seta and one simple seta; postmentum truncate, without teeth. Anterior spiracles digitiform, inserted laterally on posterior annulus of prothorax. Trunk: cylindrical; integument pale, except brown tergal plates; segmentation typical of Psychodinae: pro, meso and mesothorax, abdominal segment 1 with two annuli, abdominal segments 2 – 7 each with three annuli; hirsute, covered with inconspicuous microtrichia; microtrichia more numerous, entangled laterally, ventrally; tergal plates quadrangular, each with multiple, irregular, transverse rows of microtrichia; microtrichia along posterior margins of tergites more prominent; protergites without setae, mesotergites with transverse row of four setae, metatergites with two setae dorsally, flanked by paired setae dorsolaterally; mesosternum of segments 1 and 2 with transverse, clustered rows of prominent microtrichia along anterior and posterior margins; mesosternum of segments 3 – 7 with transverse bands of less prominent microtrichia, not arranged in rows; anal division with dorsal and ventral sclerites fused laterally, encircling anal division basally, conical apically, with inconspicuous microtrichia distributed evenly on surface; flabellar processes not elongate, approximately 1 / 8 the length of anal division, with fringe of spathiform setae; length of ventral flabellar processes approximately two times that of dorsal processes.	en	Kvifte, Gunnar Mikalsen, Curler, Gregory R., Marshall, Stephen A. (2018): Aquatic insects in the forest canopy: a new genus of moth flies (Diptera: Psychodidae) developing in slime on aerial roots. Journal of Natural History 52 (3 - 4): 137-153, DOI: 10.1080/00222933.2017.1410590, URL: http://dx.doi.org/10.1080/00222933.2017.1410590
E245A643FF96FFF27387FDEDFE41D636.taxon	distribution	Distribution Known only from the type locality in Braulio Carillo National Park, Costa Rica.	en	Kvifte, Gunnar Mikalsen, Curler, Gregory R., Marshall, Stephen A. (2018): Aquatic insects in the forest canopy: a new genus of moth flies (Diptera: Psychodidae) developing in slime on aerial roots. Journal of Natural History 52 (3 - 4): 137-153, DOI: 10.1080/00222933.2017.1410590, URL: http://dx.doi.org/10.1080/00222933.2017.1410590
E245A643FF96FFF27387FDEDFE41D636.taxon	biology_ecology	Biology Mucomyia larvae were observed moving freely within the mucilaginous matrix on root tendrils hanging at c. 150 – 200 cm above ground. The same mucilaginous masses also held several dead thrips (Thysanoptera) and some unidentifiable remains of other insects, suggesting that this sticky, viscous material is only hospitable for some insects. Larvae did not make contact with either the root tissue or the embedded, dead insects and during the short (less than one hour) period of observation prior to collection, no obvious feeding behaviour was observed. The plant from which this species was collected could not be identified beyond family (Araceae). In order to identify it to genus and perhaps species, flowers must be examined; unfortunately, the plant encountered in this study was not flowering. Multiple genera of Araceae are known to have mucilage covering their hanging roots (Paul Hanson, pers. comm.)	en	Kvifte, Gunnar Mikalsen, Curler, Gregory R., Marshall, Stephen A. (2018): Aquatic insects in the forest canopy: a new genus of moth flies (Diptera: Psychodidae) developing in slime on aerial roots. Journal of Natural History 52 (3 - 4): 137-153, DOI: 10.1080/00222933.2017.1410590, URL: http://dx.doi.org/10.1080/00222933.2017.1410590
E245A643FF96FFF27387FDEDFE41D636.taxon	etymology	Etymology Past tense of Latin emertare, ‘ emerge’, referring to the type material being a reared specimen. The epithet is to be treated as a participle and thus follows the grammatical gender of the genus.	en	Kvifte, Gunnar Mikalsen, Curler, Gregory R., Marshall, Stephen A. (2018): Aquatic insects in the forest canopy: a new genus of moth flies (Diptera: Psychodidae) developing in slime on aerial roots. Journal of Natural History 52 (3 - 4): 137-153, DOI: 10.1080/00222933.2017.1410590, URL: http://dx.doi.org/10.1080/00222933.2017.1410590
E245A643FF99FFF673C3FF5EFEB8D17D.taxon	description	(Figure 7)	en	Kvifte, Gunnar Mikalsen, Curler, Gregory R., Marshall, Stephen A. (2018): Aquatic insects in the forest canopy: a new genus of moth flies (Diptera: Psychodidae) developing in slime on aerial roots. Journal of Natural History 52 (3 - 4): 137-153, DOI: 10.1080/00222933.2017.1410590, URL: http://dx.doi.org/10.1080/00222933.2017.1410590
E245A643FF99FFF673C3FF5EFEB8D17D.taxon	materials_examined	Type material Holotype male. COSTA RICA, San José Province: Moravia, Zurquí de Moravia, Tower path, (WGS 84) 10.046806 ° N, 84.008036 ° W, 1600 m, 18 - 25. i. 2013, leg. ZADBI project (Malaise trap); deposited in MRCA. Eight male and 10 female paratypes: COSTA RICA, San José Province: Moravia, Zurquí de Moravia, Creek 2 North, (WGS 84) 10.048127 ° N, 84.008600 ° W, 1600 m, leg. ZADBI project (Malaise trap), 3 females, 18 - 24. ix. 2012 (MRCA, LACM, ZMUB), same but 29. xii. 2012 - 3. i. 2013, 1 male (MRCA), same but 3 – 11. i. 2013, 3 females (MRCA, LACM, ZMUB), same but 11 – 18. i. 2013, 2 males (MRCA, ZMUB), same but 18 – 25. i. 2013, 2 females (MRCA), same but 25. i- 1. ii. 2013, 2 males 1 female (MRCA, LACM), same but 1 – 8. ii. 2013, 1 male (MRCA), same but 22 – 29. iii. 2013, 2 males (MRCA, LACM). COSTA RICA, San José Province: Moravia, Zurquí de Moravia, Creek 2 North, (WGS 84), 10.048127 ° N, 84.008600 ° W, 1600 m, leg. ZADBI project (Aquatic emergence trap), 29. xi- 7. xii. 2012. 1 female. Specimens dissected, mounted on micro-slides.	en	Kvifte, Gunnar Mikalsen, Curler, Gregory R., Marshall, Stephen A. (2018): Aquatic insects in the forest canopy: a new genus of moth flies (Diptera: Psychodidae) developing in slime on aerial roots. Journal of Natural History 52 (3 - 4): 137-153, DOI: 10.1080/00222933.2017.1410590, URL: http://dx.doi.org/10.1080/00222933.2017.1410590
E245A643FF99FFF673C3FF5EFEB8D17D.taxon	diagnosis	Diagnosis Adult male. Narrow strip of setae alveoli reaching from frontal patch to interocular suture. Parabasal processes with lower margin parallel to upper margins of parameres, upper margin parallel to gonocoxal condyles, broadly triangular with lateral base subequal to width of gonocoxite and tapering to medial apex. Parameres distinctly hooked apicolaterally. Aedeagus with distal phallomeres fused. Surstylus with two apical tenacula and one medial subapical tenaculum.	en	Kvifte, Gunnar Mikalsen, Curler, Gregory R., Marshall, Stephen A. (2018): Aquatic insects in the forest canopy: a new genus of moth flies (Diptera: Psychodidae) developing in slime on aerial roots. Journal of Natural History 52 (3 - 4): 137-153, DOI: 10.1080/00222933.2017.1410590, URL: http://dx.doi.org/10.1080/00222933.2017.1410590
E245A643FF99FFF673C3FF5EFEB8D17D.taxon	description	Description Adult male. (n = 9 except when stated otherwise) Head (Figure 7 (a )) broadly oval; vertex a third of head length, hind margin of head slightly convex; eyebridge of four facet rows, separated by 2 facet diameters; interocular suture U-shaped with median thickening and in some specimens an indistinct spur; 28 – 50 ocular setae present, reaching second eyebridge facet row from the suture, uniseriate dorsal to eyebridges, biseriate dorsal to eye, triseriate laterally; frontal scar patch broadly oval with median posterior extension reaching interocular suture; clypeus rectangular with two anterior concavities; frontoclypeal suture complete, straight; labrum, maxillae and labium as long as clypeus; labellae fleshy, lateral margins sclerotized, with two dorsolateral setae and spiniform anterior sensillum; palp of four segments, cylindrical except fourth segment spool-shaped; fourth palp segment fully sclerotized; length of palp segments 42: 54: 63: 66; antennae with 14 flagellomeres; scape conical-cylindrical, pedicel globular, flagellomeres 1 – 11 symmetrically nodiform with internodes slightly longer than nodes except flagellomere 11 with internode a fifth as long as node (Figure 7 (c )), flagellomeres 12 – 13 globular, flagellomere 14 dropletshaped (Figure 7 (c )); pair of ascoids present on flagellomeres 1 – 11, flanked by pair of small spiniform sensilla with broad round bases; length of antennal segments 91: 57: 135: 135: 132: 132: 135: 132: 129: 111: 111: 102: 51: 21: 24: 30 (n = 9,9,9,9,9,9,8,7,5,2,1,1,1,1,1,1); ascoids Y-shaped, length of anterior ascoid branches reaching beyond internode of segment; thorax with dorsum, proepisternum, anterior half of anepisternum, laterotergite and dorsal part of scutellum setose; proepisternum with tongue-shaped internal ventral projection; setose anterior half of anepisternum sharply delimited from bare posterior half; prothoracal spiracle naked; metathoracal spiracle densely setose; coxae all with dorsoventral anterior aseriate rows of setae, densest at ventral half of mid coxa; pointed extensions present on anterior side of front tibia and posterior end of mid and hind tibiae; tarsomere 5 without dorsal extension; wing (Figure 7 (b )) 2 mm long (n = 8); C with two breaks, Sc terminating in wing membrane but approaching R 1, reaching level of base of M 1 + 2; wing membrane setose; area dorsal of R 1 slightly darker than rest of membrane; R 5 terminating in wing apex; radial fork distal to medial fork, both complete; jugum obtusely V-shaped, setose; narrow oval patch of setae present dorsal to jugum; male genitalia (Figure 7 (d )) with hypandrium reduced; gonocoxites reniform with broad triangular parabasal processes, 1.5 × length of gonostyles; parabasal processes with lower margin parallel to upper margins of parameres, upper margin parallel to gonocoxal condyles, broadly triangular with lateral base subequal to width of gonocoxite and tapering to medial apex; gonostyles conical with blunt apices, with scattered sensilla; aedeagus with basiphallus narrow at base, distally with median keel; distiphallus apparently spatulate with two phallomeres originating from lateral margins of apex of basiphallus; phallomeres narrow, pointed apically, of equal length; gonocoxal condyles angularly arched, approaching medially but not touching, with lateral margin thicker and more strongly sclerotized than rest of condyle; parameres oblong-shaped, hooked at medial basal end at junction with subepandrial membrane, distally hooked towards the lateral; parameres slightly longer than gonocoxite; triangular sclerotization present dorsolaterally on parameres; epandrium (Figure 7 (d )) broader than long; base with strip-like apodeme, concave with indistinct projection laterally; aperture indistinct; fleshy setose projections present laterally on distal third; subepandrial membrane indistinct; hypoproct oval with slightly pointed lateral edges, with V-shaped concavity anteriorly and weak concavity posteriorly; epiproct present as strip-like sclerite on ventral distal edge of hypoproct; surstyli cylindrical, about as long as epandrium; with two distal and one subapical mesal tenaculum; tenacula with indistinct longitudinal striations, apex rounded. Adult female (n = 10) similar to male except with length of palps 45: 60: 66: 72 (n = 10,10,10,9), length of antennal segments 87: 54: 123: 111: 111: 111: 108: 108: 102: 99: 93: 51: 24: 27: 30 (n = 10,10,9,9,9,9,9,9,9,6,6,6,6,6,6,6), wing length 2.39. Female terminalia (Figure 7 (e )) with subgenital plate rectangular with two rounded subtriangular distal lobes, pseudospermatheca with curved F-shaped inner sclerotized mesal margins, distally with two triangular sclerites which have their lower margins expanded and slightly produced posteriorly, laterally with hyaline sclerotized margins; elipsoid transverse sclerite present, flanked by pair of comma-shaped sclerites; cerci 1.3 × length of subepandrial plate.	en	Kvifte, Gunnar Mikalsen, Curler, Gregory R., Marshall, Stephen A. (2018): Aquatic insects in the forest canopy: a new genus of moth flies (Diptera: Psychodidae) developing in slime on aerial roots. Journal of Natural History 52 (3 - 4): 137-153, DOI: 10.1080/00222933.2017.1410590, URL: http://dx.doi.org/10.1080/00222933.2017.1410590
E245A643FF99FFF673C3FF5EFEB8D17D.taxon	distribution	Distribution Known only from the type locality at Zurquí de Moravia, Costa Rica.	en	Kvifte, Gunnar Mikalsen, Curler, Gregory R., Marshall, Stephen A. (2018): Aquatic insects in the forest canopy: a new genus of moth flies (Diptera: Psychodidae) developing in slime on aerial roots. Journal of Natural History 52 (3 - 4): 137-153, DOI: 10.1080/00222933.2017.1410590, URL: http://dx.doi.org/10.1080/00222933.2017.1410590
E245A643FF99FFF673C3FF5EFEB8D17D.taxon	biology_ecology	Biology Most specimens have been collected in Malaise traps in tropical cloud forest. One female specimen was collected from an aquatic emergence trap, however this is not necessarily a reliable indicator of its larval habitat. In European studies of psychodids in emergence traps, there very often is a proportion of specimens of species with known non-aquatic larval stages occurring (Wagner 1973, G. M. Kvifte and M. Ivkovic, in prep.).	en	Kvifte, Gunnar Mikalsen, Curler, Gregory R., Marshall, Stephen A. (2018): Aquatic insects in the forest canopy: a new genus of moth flies (Diptera: Psychodidae) developing in slime on aerial roots. Journal of Natural History 52 (3 - 4): 137-153, DOI: 10.1080/00222933.2017.1410590, URL: http://dx.doi.org/10.1080/00222933.2017.1410590
E245A643FF99FFF673C3FF5EFEB8D17D.taxon	etymology	Etymology Named with admiration and appreciation for Brian V. Brown, Head of the Entomology Department and Curator of Entomology at the Natural History Museum of Los Angeles County and co-coordinator of the Zurquí All Diptera Biodiversity Inventory (see Borkent and Brown 2015).	en	Kvifte, Gunnar Mikalsen, Curler, Gregory R., Marshall, Stephen A. (2018): Aquatic insects in the forest canopy: a new genus of moth flies (Diptera: Psychodidae) developing in slime on aerial roots. Journal of Natural History 52 (3 - 4): 137-153, DOI: 10.1080/00222933.2017.1410590, URL: http://dx.doi.org/10.1080/00222933.2017.1410590
