identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
E2083F003B0CFFAEFF74FD81FA5D60AE.text	E2083F003B0CFFAEFF74FD81FA5D60AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyclocephala bicolor	<div><p>Key for identification of the species of the Cyclocephala bicolor group of the genus Cyclocephala</p><p>1. Anterior tarsi enlarged (males) ..................................................................................................................................... 2</p><p>- Anterior tarsi slender (females) .................................................................................................................................... 6</p><p>2. Second ventrite with a triangular, flat projection on posterior half; basal tooth of protibiae (Fig. 29) distinctly directed forward, separated from middle tooth by a distance about 2.0 times longer than that separating middle from anterior tooth; clypeus on apical half slightly convex in lateral view (Fig. 12) ....................... C. dichroa Dechambre.</p><p>- Second ventrite without triangular projection; basal tooth of protibiae (Figs. 24, 26, 28, 31) with anterior border usually perpendicular to longitudinal axis of tibiae and separated from the middle tooth by a distance 1.5 times or less than that separating middle from anterior tooth; clypeus on apical half more or less flat to slightly concave in lateral</p><p>view (Figs. 4, 8, 16) .................................................................................................................................................... 3 3. Frontoclypeal suture (Fig. 15) posteriorly projecting angularly; punctation on scutellum very fine, much more so than on middle of base of pronotum; punctation on apical region of pygidium very fine, distinctly finer than that of basal region, which is moderately fine ............................................................................................... C. anibali, sp. n.</p><p>- Frontoclypeal suture (Figs. 3, 7, 11) posteriorly projecting arcuately or almost straight; punctation of scutellum fine, scarcely finer than that on center of pronotal base; punctation on apical region of pygidium fine, scarcely smaller than that of basal region, which is strong to moderately strong .................................................................................. 4</p><p>4. Punctures on base of clypeus not discrete, closely set, mostly confluent, appearing as irregular and fine, more or less transversely striate; frontoclypeal suture posteriorly projecting arcuately (Fig. 3); surface of pygidium smooth (50X) ...................................................................................................................................................................................... 5</p><p>- Punctures on base of clypeus moderately strong, discrete, somewhat transverse, the distance between most punctures smaller than the diameter of puncture; frontoclypeal suture very slightly arcuately projecting posteriorly (Fig. 7); surface of pygidium finely alutaceous (50X) ........................................................................ C. bicolorata Endrödi</p><p>5. Labium (Fig. 5) with strong, dense piliferous punctures on basal two-thirds; metatibiae (Figs. 49–50) with piliferous punctures almost restricted to those of normal longitudinal series close to posterior border; distances between most punctures on apex of pygidium usually smaller than 2.0 times the diameter of punctures; apex of pygidium (in posterior view) feebly convex-truncate. .................................................................................................... C . bicolor Laporte</p><p>- Labium with one longitudinal series of long bristles on each side, converging towards base; mesotibia and especially metatibiae with numerous piliferous punctures over all surface, in addition to those of normal longitudinal series; distances between most punctures on apex of pygidium usually 2.0 times or more larger than the diameter of punctures; apex of pygidium (in posterior view) rounded..................................................................... C. sarahae Ratcliffe</p><p>6. Basal tooth of protibiae distinctly anteriorly projected, separated from the middle tooth by a distance about 2.0 times larger than that separating the middle from the anterior (Fig. 30); clypeus convex in lateral view (Fig. 12) ................ .................................................................................................................................................... C . dichroa Dechambre</p><p>- Basal tooth of protibiae more diagonal and/or its anterior border more or less perpendicular to longitudinal axis of tibiae, separated from the middle tooth by a distance 1.5 or less than that separating the middle from the anterior (Figs. 25, 27, 29, 32); clypeus more or less flat in lateral view (Figs. 4, 8, 16) .......................................................... 7</p><p>7. Punctures on apical region of pygidium distinctly smaller than those of basal region. ...................... C. anibali sp. n.</p><p>- Punctures on apical region of pygidium scarcely smaller than those of basal region, which are strong to moderately strong ........................................................................................................................................................................... 8</p><p>8. Pygidium feebly convex in lateral view, surface smooth, punctures more or less strong almost to apex, on apical region somewhat smaller, usually dense, sometimes scattered ....................................................... C. bicolor Laporte</p><p>- Pygidium strongly convex in lateral view, surface finely alutaceous (50X), punctures more or less strong at base, smaller on apex, most distances between punctures 2.0 or more times larger than diameter of puncture. .................... .................................................................................................................................................... C. bicolorata Endrödi</p></div>	https://treatment.plazi.org/id/E2083F003B0CFFAEFF74FD81FA5D60AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Joly, Luis J.	Joly, Luis J. (2009): Review of the species in the Cyclocephala bicolor Laporte species group (Coleoptera: Scarabaeidae: Dynastinae). Zootaxa 2048: 47-64, DOI: 10.5281/zenodo.186485
E2083F003B0FFFABFF74FE66FC4D65A6.text	E2083F003B0FFFABFF74FE66FC4D65A6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyclocephala bicolor Laporte 1840	<div><p>Cyclocephala bicolor Laporte, 1840</p><p>(Figs. 1, 3–6, 19, 24–25, 33–35, 41–42, 49–50, 58, 69–70)</p><p>Bibliography. Cyclocephala bicolor Laporte 1840: 124 (description). Type (sex?) MNHN. Type locality: Cayenne. Burmeister 1847: 60 (description), Cayenne. Prudhomme 1906: 13, French Guiana. Bodkin 1919: 215, British Guiana. Höhne 1923: 359 (compared with C. vestita). Arrow 1937: 8, Guiana. Blackwelder 1944: 251, Guiana. Endrödi 1964: 442 (compared with C. bicolorata), Fig. 11 (genitalia). Endrödi 1966: 91 (key, male), 140, 143 (key, female), 156 (description male, female), Figs. 386–397 (genitalia), Venezuela, British Guiana, French Guiana, Suriname, Brazil. Gruner 1971: 845, French Guiana. Dechambre 1979: 160, Venezuela, British Guiana, French Guiana, Suriname, Brazil. Endrödi 1985: 109 (key, male), 146 (key, female), Figs. 648–649 (genitalia), French Guiana, Suriname, Guyana, Brazil. Ratcliffe 1992: 187 (compared with C. sarahae). Lachaume 1992: 16, plate 1, Fig. 28, Guyana, French Guiana, Suriname, Brazil. Dechambre 1992: 67 (compared with C. dichroa), Fig. 32 (genitalia), plate 1, Fig. 28.</p><p>Description. Short, oval, moderately shiny. Brownish red, frons and vertex black, elytra testaceous. Dorsal surface and pygidium glabrous, ventral pubescence sparse. Length: 9.0–13.0 mm.</p><p>MALE: Clypeus (Fig. 3) about 2.3 times as wide across base as long; sides straight, strongly convergent to truncated and feebly reflexed apex; shallowly sulcate longitudinally along middle and shallowly depressed laterally before apex, in lateral view flat to slightly concave on apical half; surface finely alutaceous, with fine, shallow and widely annulate punctures, annulation wide, not well defined posteriorly, usually confluent, appearing as irregular and fine, more or less transverse striae; punctures simple toward apex and anterior half of sides. Frontoclypeal suture thin, arcuately projecting posteriorly. Frons width about 2.5 times transverse diameter of eye, finely alutaceous; punctures similar to those of clypeus, but less dense, somewhat thinner and better delimited.</p><p>Mandibles with external border sinuate, apex projecting externally. Maxillae (Fig. 6) with galea moderately robust, with two apical, one median, and three basal teeth; basal teeth very small and arising from common base; stipes projected anteriorly on inner side and with a brush of fine, black setae on apex of projection; maxillary palpi (Fig. 19) moderately robust, second palpomere longer than third, weakly widened towards apex, fourth palpomere fusiform, 2.5 times longer than preceding. Labium (Fig. 5) with sides almost parallel on basal two-thirds, constricted at middle of apical third; apex truncate on middle and fimbriate with long, fine yellowish-golden setae; surface almost flat, with some very small punctures on disc and one longitudinal series of long bristles on each side, converging towards base.</p><p>Antennae with 10 antennomeres, club more than 2.0 times the length of antennomeres 2–7 together.</p><p>Pronotum 1.5 times as wide as long, widest near base, sides more strongly convergent to apex on anterior half; anterior border widely and arcuately projected at middle; basal angles widely rounded; basal border without marginal bead, almost uniformly arcuate, slightly more projected in central third; surface smooth (40X); punctures annulate, moderately fine and dense, the distance between most punctures on disc, as long as or longer than diameter of punctures, punctures gradually larger and denser towards sides, the diameter of a puncture at basal angle at least 2.0 times the diameter of those on center of disc, more dense on basal angles, where the distance between most punctures is as large or smaller than the diameter of a puncture; all surface also covered with numerous minute punctures (50X or more). Scutellum triangular, sides feebly arcuate, apex briefly rounded; punctures similar to those of pronotal disc.</p><p>Elytra 1.1 times as long as wide, arcuately widened from first quarter to middle, then rounded to apex, sutural angle well defined, but rounded; humeral and apical umbones weak, with a feeble, short, elongate tumescence behind humeral umbones on external border; surface smooth, with large, shallow, annulate punctures, much smaller on sides and apex, double rows not evident, additional numerous minute punctures (50X or more) scattered over the surface; sutural interstria flat, slightly elevated on apical half. Epipleura narrow, more so apically.</p><p>Pygidium 2.0 times as wide as long; in posterior view the inferior borders slightly convex, converging to feebly convexly truncated apex at an angle distinctly greater than 90°; in lateral view (Fig. 62) uniformly convex; surface smooth, shining, with large annulate punctures, punctures more dense on lateral angles; on apex more shining, the punctures smaller and less dense, the distances between most punctures less than 2.0 times their own diameter.</p><p>Protibiae (fig. 24) tridentate, small basal tooth separated from the intermediate tooth by a distance 1.5 times as long as that between the intermediate and the anterior teeth; intermediate and anterior teeth well developed, basal tooth with its anterior border more or less perpendicular to long axis of tibiae; apical spur almost straight, surpassing the apex of the first tarsomere. Protarsi (Fig. 34–35) enlarged; tarsomere 5 shorter than 1–4 together, without ventrointernal carina, ventral side with short and fine longitudinal striae on inner side of apex and sometimes also on inner side of base; tarsomere 4 longitudinally striated on about apical half, tarsomere 3 on apical third; largest claw (Fig. 41–42) wide, flattened, moderately widely cleft at apex, both branches feebly divergent (external border of small branch not continuous with internal curvature of the claw). Metafemur (Fig. 49) about 2.3 times as long as its maximum width. Metatibiae (Fig. 49) robust, widened at apex, bristles on diagonal carina and apex strong. Metatarsi (Fig. 33) subequal in length to tibiae, with first tarsomere widened towards apex, with apical bristles similar to those of apex of tibiae.</p><p>Prosternal projection high, apex alutaceous, anterior part transversely oval, externally pointed; posterior part shorter but wider than anterior, posterior border fimbriate with long bristles.</p><p>Pronotal hypomera alutaceous, with sparse, erect setae. Mesopleura, metapleura, and sides of metasternum alutaceous, with large and very dense confluent, annulate punctures; disc of metasternum smooth, with small, scattered punctures; metacoxae with punctures similar to those of pleurae but smaller and more disperse. Ventrites with annulate, moderately dense punctures, punctures smaller and sparser on apical ventrites.</p><p>Aedeagus: Figs. 69–70.</p><p>FEMALE: differs from the male in the following characters: punctation generally stronger, that of pygidium of the same size; protibiae Fig. 25; protarsi slender; posterior femora (Fig. 50) somewhat stronger; epipleura (Fig. 58) strongly narrowed at level of metacoxa, then widened up to level of middle of fourth ventrite.</p><p>Distribution: Venezuela, Colombia, Guyana, Suriname, French Guiana, Brazil, Peru, and Bolivia. Colombia, Peru, and Bolivia are new country records.</p><p>Specimens studied: VENEZUELA: Amazonas: Coromoto, Dto. Atures, VI-1979, 1 F (HAHC). Culebra, Caño Negro, P. N. Duida-Marahuaca, 3º 33’ N, 65º 55’ W, 6-16-X-1983, Exped. Terramar, 4 F (MIZA). La Esmeralda, LF Station, 01-X-1997, leg. Krimse, 1 F (MIZA). Pto. Ayacucho, 22-IV-1967, P. J. Anduze, 1 M (HAHC); 4-15-XII-1992, G. Romero, 1 F (MIZA). San Carlos de Río Negro, 7-13-1982, A.</p><p>Chacón-G. Yépez, 1 M, 15 F (MIZA); idem, 1º 55’ N, 67º 01’ W, 4-14-III-1984, J. Clavijo-J. De Marmels, 1 F (MIZA). San Fernando de Atabapo, 17-VIII-1982, CEUM Fac. Agronomía, 1 F (MIZA). Santa Lucía, 15-21- XI-1982, A. Chacón-G. Yépez, 1 F (MIZA). Talud Cerro Aracamuni, 1 º 29’ N, 65º 38’ W, 600 m, 18-24-X- 1987, Exped. Terramar, 3 M, 4 F (MIZA). Bolívar: Anacoco, 60 m, 6º 5’ N, 61º 8’ O, 10-30-VIII-1979, Expedición La Salle, 3 M, 2 F (MIZA). Canaima, Río Carrao, (Campamento Yuri), 9-15-II-1972, J. M. Ayala, 1 F (JMAL). El Paují, 100 Km O Sta. Elena, 1100 m, 5-IV-80, 2 M, 2 F (CCBM). El Dorado-Sta. Elena, Km 88, 160 m, 28-IX-1967, C. J. Rosales-M. Gelbez-L. Rodríguez V., 2 F (MIZA); 27-IX-1967, L. J. Joly, 1 F (MHNLS). El Dorado-Sta. Elena, Km 109, 460 m, 6-IX-1964, P. J. Salinas, 1 F (MIZA). Carret. El Dorado- Sta. Elena, Km 88, 100 m, 27-IX-1967, L. J. Joly, 1 F (MHNLS); idem, 200 m, 7-IV-80, C. Bordón, 1 F (CCBM). 5 Km E Caicara, 11-VI-1996, B. D. Gill, 1 M (BDGC). Carret. Caicara-San Juan de Manapiare, Km 170, 300 m, 21-30-XII-1973, J. L. García, 1 M (HAHC). Cerro Guaiquinima, 1000 m, 5º 53‘ N, 63º 30’ O, 3- 4-II-1990, Exped. FUDECI, J. De Marmels-A. Chacón, 2 F (MIZA). Guri, Río Caroní, 100 m, 16-XI-1966, J. &amp; B. Bechyne-E. Osuna, 3 F (MIZA); 11-IV-1968, J. Salcedo, 1 M, 1 F (MIZA). Kamarata, 450 m, 24-II- 1978, L. J. Joly, 2 M, 5 F (MIZA). Kanarakuni, Alto Caura, 450 m, 10-13-IX-1964, F. Fernández Y.-J. Bechyne, 5 F (MIZA). 35 Km N Las Trincheras, 17-VI-1987, S. &amp; J. Peck, woodland, forest edge, UV trap, 15 Km N Corocito, 2 M (HAHC). Los Pijiguaos, 190 m, Exped. MIZA, UCV, 3-13-VI-1992, 4 M, 4 F (MIZA). Pendare, Río Parguaza, 6º 6’ N, 67º 5’ W, 6-7-VI-1997, E. Osuna-A. Chacón-F. Rojas, 1 M, 38 F (MIZA). Río Caroní, Salto Las Babas, 8-10-IV-1983, Exped. Inst. Zool. Agric., 5 M, 5 F (MIZA). Río Caura, El Playón, 23-XI-1978, 3 F (MIZA). Río Caura, Salto Pará, 250 m, 21-XI- 178, B. Bechyné, 3 F; idem, 20-22- XI-1978, A. Chacón, 1 F (MIZA). Río Caura, Jabillal, 100 m, 25-XI-1978, A. Chacón, 2 F (MIZA). Río Guaniamo, 6º 45’ N, 66º 01’ O, 160 m, 25-28-V-1979, J. Clavijo-A. Chacón-G. Yépez, 3 M, 2 F (MIZA). Río Paragua, E Río Cachicamo, 425 m, 11-IV-1983, Exped. Inst. Zool. Agric., 1 F (MIZA). San Ignacio de Yuruaní, Gran Sabana, 830 m, 19-IV-1988, A. Chacón, 1 M (MIZA). Suapure, Caura River, V-27-1899, E. A. Klages, 1 M (HAHC). Uonquen, 850 m, 4-III-1966, A. Pérez, 1 F (MIZA). Táchira: Navay, 200 m, 30-X-78, C. Bordón, 1 F (CCBM). COLOMBIA: Caquetá, Río Caquetá at Tres Esquinas, 18-I-1969, R. E. Dietz IV, 1 M (NMNH). GUYANA: British Guiana, 4 M, 1 F (AMNH), 1 M (MIZA). Demerara (Bartica), R. J. Crew, 1 M, 3 F (NMNH). Kamakusa, Jan, 1923, 1 M, 1 F, (AMNH). Kartabo, Bartica District, 7-IX-1920, Wm. L. Beebe collection, 1 F (AMNH); 11-X-1920, Gift of N. Y. Zool. Soc. 1 F (AMNH). Mazaruni, Potaro District, Takuku Mountains, 6º 15’ N, 59º 5’ W, 7-XII-1983, at blacklight in forest clearing near streams, Earthwatch Research Expedition, P. J. Spangler &amp; W. E. Steiner, 1 M (NMNH). Penal settlement, Bartica District, 1917, Gift of N. Y. Zool. Soc., 1 F (AMNH). Rupununi, XI, 1913, 1 F (AMNH). Rockstone, Essequibo River, 4 F (NMNH). Tumatumari, 1918, 19 F (AMNH), 1 F (MIZA); idem, VIII, 1913, 1 M, 17 F (AMNH). Tumatumari, Potaro R., 29-VI-1927, Cornell Unv. Lot 760 Sub 117, 1 F (HAHC). SURINAME: Aibinia, 29- VII-1975, D. Engleman, at light, 1 F (UNSM). Brokopando District, Brownsberg Natuurpark, Mazaroni Plateau, 400–500 m, 20 August 1982, W. E. Steiner, Earthwatch Suriname Expedition, August 1982, Collins, Early, Oberman, Pollock, Putnem, Steiner, 4 M, 2 F (NMNH). Moengo Boven, Cottica, 16-V-1927, Cornell Univ., Lot 760, Sub. 60, 1 F (NMNH). FRENCH GUIANA: Kaw Rd. PK 37, 19 -VIII-1995, 1 M (UNSM). Maroni River, 1 F (NMNH). Route de Petit Saut PK9 25-VIII-2000, Y. Ponchel, 5 M (HAHC). St. Jean, W. M. Schaus col. 7 F (NMNH). 13 Km W D-5, Risq. Rd. 14-VIII-1995, J. E. Wappes, 3 M, 6 F (UNSM); 22-VIII- 1995, 2 M, 1 F (UNSM), 1 M, 1 F (HAHC). BRAZIL: Robinson bequest, 1929, 1F (NMNH). Amazonas: AM 0 10, Km 244, 20 -I-1977, B. C. Ratcliffe, 7 M, 6 F (UNSM). AM 0 10, Km 268, 18 -I-1977, B. C. Ratcliffe, 3 F (UNSM). Manaus, 30-X-83, C. Bordón, 2 M, 1 F (CCBM); 13-I-1978, B. C. Ratcliffe, 4 M, 2 F (UNSM); 13-XI-1978, B. C. Ratcliffe, 3 M, 3 F (UNSM). CEPLAC, 30 Km NE Manaus, 12-XII-1976, B. C. Ratcliffe, 1 M, 1 F (UNSM). Amapá: Serra Lombard Lomao, 24-VIII-1961, J. &amp; B. Bechyne, 3 F (MIZA). Ceará: Fortaleza, IX-1943, at light, Mac Creary, 1 F (NMNH). Mato Grosso: Reserva Humboldt, 10º 11’ S, 59º 48’W, 13-14-VII-1977, B. C. Ratcliffe, 1 F (UNSM). Pará: Braçança, 20-I-1949, J. M. Díaz, 1 M (HAHC). Rio Negro, Santa Isabel, X-1930, Holt, Blake &amp; Agostini, 5 F (NMNH). Cachimbo, 400 m, 14/ 21-IX-1955, L. Travassos &amp; Oliveira, 5 M, 1 F (HAHC). Rio Xingu Camp., 3º 39’ S, 52º 22’ W, ca 60 Km S Altamira, 1-7-X- 1986, 1F (NMNH). Santarem, IX-1933, L. C. Scaramuzza, 2 F (NMNH). Pernambuco: Recife, 10-X-50, Asua leg, 1 F (HAHC). Río Grande do Norte: Natal, VIII-X-1954, 1 M (HAHC). Rondonia: Porto Velho, Rio Madeira, Mann &amp; Baker, 1918, 1 F (NMNH). Terr. Acre: Río Branco, 15-V-1952, M. Alvarenga col., 1 F (HAHC). Ro. (?): Santa Cruz da Serra, 20-VIII-85, J. P. Caldwell, 3 F (HAHC). I have also seen specimens from BOLIVIA: Beni, Guayamerin (AMNH). BRAZIL: Amazonas: Pará (AMNH). PERU: Tingo María (AMNH).</p><p>Other localities: SURINAME: Pakira-Iméné; Paramaribo; Demerara. FRENCH GUIANA: Acarouany; Ante cume Pata (Haut Kialo); Carbet République; Cayenne; Fleuve Oyapock, du Saut-Maripa à Massikiri; Kourou fôret; Mana; Organabo; route Conte; Saint Élie; Sinnamary. BRAZIL: Gorotire; São Felix do Xingu (Moyen Xingu) (Endrödi 1966, Gruner 1971, Dechambre 1979).</p></div>	https://treatment.plazi.org/id/E2083F003B0FFFABFF74FE66FC4D65A6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Joly, Luis J.	Joly, Luis J. (2009): Review of the species in the Cyclocephala bicolor Laporte species group (Coleoptera: Scarabaeidae: Dynastinae). Zootaxa 2048: 47-64, DOI: 10.5281/zenodo.186485
E2083F003B05FFA0FF74FC3EFF226581.text	E2083F003B05FFA0FF74FC3EFF226581.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyclocephala bicolorata Endrodi 1964	<div><p>Cyclocephala bicolorata Endrödi, 1964</p><p>(Figs. 2, 7–10, 20, 26–27, 36, 44–45, 51 –52, 59, 63–64, 73–74)</p><p>Bibliography. Cyclocephala bicolorata Endrödi 1964: 441 (description, male; compared with C. bicolor), Fig. 10 (genitalia). Holotype male Col. Endrödi. Type locality: Venezuela: Llanos. Endrödi 1966: 91 (key, male), 143 (key, female), 157 (description, male), Fig. 388 (genitalia), Venezuela. Dechambre 1979: 160, Venezuela, Brazil. Endrödi 1985: 110 (key, male), 146 (key, female), Fig. 650 (genitalia), Venezuela. Dechambre 1992: 67 (compared with C. dichroa), Fig. 33 (genitalia).</p><p>Description. Short, oval, moderately shiny. Red to brownish red; frons and vertex black, elytra testaceous. Dorsal surface and pygidium glabrous, ventral pubescence sparse. Length: 11.0–11.8 mm.</p><p>MALE: Clypeus (Fig. 7) about 2.4 times as wide across base as long; sides straight, strongly convergent to truncated and feebly reflexed apex; shallowly sulcate longitudinally along middle and shallowly depressed laterally before apex, in lateral view flat to slightly concave on apical half; surface finely alutaceous, with punctures moderately strong, discrete, somewhat transverse (especially on base), dense, the distance between most punctures smaller than the diameter of the puncture, more dense towards apex. Frontoclypeal suture (Fig. 7) thin, slightly arcuately projecting posteriorly. Frons width about 2.5 times transverse diameter of eye, finely alutaceous; punctures similar to those of clypeus, but somewhat more scattered, except on a narrow, smooth, transverse band behind frontoclypeal suture and another longitudinal band along midline.</p><p>Mandibles with external border sinuate, apex projecting externally. Maxillae (Fig. 10) with galea moderately strong, with two apical, one median and three basal teeth, the basal teeth very small and arising from a common base; stipes projected anteriorly on inner side and with a brush of fine, yellowish brown setae on apex of projection; maxillary palpi (Fig. 20) moderately robust, second palpomere with a small projection at base, longer than third, fourth palpomere fusiform, 3.0 times as long as preceding. Labium (Fig. 9) with sides slightly convex on basal two-thirds, not constricted at the middle of apical half; apex truncate on ventral side, with two small projections at middle on oral side, and fimbriated with long, fine, yellowish brown setae; surface without punctures on disc, with one longitudinal series of long bristles on each side, converging towards base.</p><p>Antennae with 10 antennomeres, club about 2.0 times the length of antennomeres 2–7 together.</p><p>Pronotum 1.4 times as wide as long, widest near base, sides more strongly convergent to apex on anterior half; anterior border widely and arcuately projected at middle; basal angles widely rounded; basal border without marginal bead, almost uniformly arcuate, slightly more projected in central third; surface finely alutaceous (40X); punctures annulate, moderately fine and dense, the distance between most punctures on disc as long as or longer than diameter of punctures, punctures gradually larger toward sides, diameter of a puncture about 2.0 times the diameter of those on center of disc, more dense on basal angles, where the distance between most punctures is as long as or shorter than the diameter of a puncture; surface also covered with numerous minute punctures (50X or more). Scutellum triangular, sides feebly arcuate, apex briefly rounded; punctures similar to those on pronotal disc.</p><p>Elytra 1.2 times as long as wide, arcuately widened from first quarter to middle, then rounded to apex, sutural angle well defined, but rounded; humeral and apical umbones weak; with feeble, short, elongate tumescence behind humeral umbones on external border; surface smooth, with large, shallow, annulate punctures, much smaller on sides and apex, double rows not evident; also with numerous minute punctures (50X or more) scattered over all surface; sutural interstria flat, weakly elevated on apical half. Epipleura narrow, more so apically.</p><p>Pygidium 2.0 times as wide as long; in posterior view the inferior borders straight, converging to narrow and feebly convex-truncate apex in an angle distinctly greater than 90°; in lateral view (Fig. 63) slightly convex, more strongly convex on apical half; surface finely alutaceous, with abundant minute punctures and with large annulate punctures, more dense on lateral angles; punctures on apex distinctly smaller and less dense, the distances between most punctures more than 2.0 times its own diameter.</p><p>Protibiae (Fig. 26) tridentate, small basal tooth separated from the intermediate tooth by a distance 1.4 times as long as that between the intermediate and anterior teeth; intermediate and anterior teeth well developed, basal tooth with its anterior border more or less perpendicular to longitudinal axis of tibiae; apical spur almost straight, surpassing apex of first tarsomere. Protarsi (Fig. 36) enlarged; tarsomere 5 shorter than 1–4 together, without ventro-internal carina, ventral side with very short and very fine longitudinal striae on inner side of apex; tarsomere 4 longitudinally striated on less than apical half; tarsomere 3 striated only on inner side of apical half; largest claw (Figs. 44–45) wide, flattened, moderately narrowly cleft at apex, the external border of small branch continuous with internal curvature of the claw. Metafemur (Fig. 51) robust, 2.0 times as long as its maximum width, its anterior border strongly convex. Metatibiae (Fig. 51) robust, widened at apex, bristles on diagonal carina and apex very strong. Metatarsi subequal in length to tibiae, with first tarsomere widened towards apex, with apical bristles similar to those of apex of tibiae.</p><p>Prosternal projection high, apex alutaceous, anterior part transversely oval, externally pointed; posterior part shorter but wider than anterior, posterior border fimbriate with long bristles.</p><p>Pronotal hypomera alutaceous, with sparse erect setae. Mesopleura, metapleura, and sides of metasternum alutaceous, with large, dense confluent, annulate punctures; disc of metasternum smooth, with small, scattered punctures; metacoxae with punctures similar to those of pleurae but smaller and more disperse. Ventrites with annulate, moderately dense punctures, smaller and sparser on apical ventrites.</p><p>Aedeagus: Figs. 73–74.</p><p>FEMALE: differs from the male in the following characters: punctation generally stronger; protibiae Fig.</p><p>27; protarsi slender; metafemora (Fig. 52) somewhat stronger; metatibiae Fig. 52; epipleura (Fig. 59) strongly narrowed at level of anterior border of metacoxa, then widened up to level of middle of second ventrite; pygidium in lateral view Fig. 64.</p><p>Distribution: Venezuela, Brazil.</p><p>Specimens studied: VENEZUELA: Apure: Hato El Frío, Fundo El Ceibote, 100 m, 29-V-1975, C. J. Rosales, 1 F (MIZA). Bolívar: Guri, Río Caroní, 11-IV-1968, J. Salcedo, 1 M (MIZA).</p><p>Other localities: VENEZUELA: Llanos. BRAZIL: São Felix do Xingu (Moyen Xingu) (Dechambre 1979).</p></div>	https://treatment.plazi.org/id/E2083F003B05FFA0FF74FC3EFF226581	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Joly, Luis J.	Joly, Luis J. (2009): Review of the species in the Cyclocephala bicolor Laporte species group (Coleoptera: Scarabaeidae: Dynastinae). Zootaxa 2048: 47-64, DOI: 10.5281/zenodo.186485
E2083F003B01FFA1FF74F939FDB76121.text	E2083F003B01FFA1FF74F939FDB76121.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyclocephala sarahae Ratcliffe 1992	<div><p>Cyclocephala sarahae Ratcliffe, 1992</p><p>(Figs. 1, 21, 28, 37–38, 43, 57, 75)</p><p>Bibliography. Cyclocephala sarahae Ratcliffe 1992: 187 (description male, female; compared with C. bicolor, C. testacea ab. ovulum), Fig. 6 (photograph), Figs. 27–28 (genitalia). Holotype male UNSM. Type locality: Brazil (Amazonas, Rio Demeni, nr. equator).</p><p>Description. Short, oval, moderately shiny. Brownish red, frons and vertex black, elytra testaceous. Dorsal surface and pygidium glabrous, ventral pubescence sparse. Length: 11.4 mm.</p><p>MALE: Clypeus about 2.8 times as wide across base as long; sides straight, strongly convergent to truncate and feebly reflexed apex; shallowly sulcate longitudinally along middle and shallowly depressed laterally before apex, in lateral view flat to slightly concave on apical half; surface finely alutaceous, with shallow and widely annulate punctures, the annulation wide, not well defined posteriorly, most punctures confluent, appearing as irregular and fine, more or less transverse striae. Frontoclypeal suture thin, bisinuate, not pointed posteriorly at middle. Frons: width about 2.8 times transverse diameter of eye, finely alutaceous; punctures smaller than those of clypeus, somewhat transverse by confluence of punctures, most punctures separated by distances more or less their own diameter.</p><p>Mandibles with external border sinuate, apex projecting externally. Maxillae not visible; maxillary palpi (Fig. 21) moderately robust, second palpomere longer than third, scarcely widened towards apex, fourth fusiform, 3.5 times longer than preceding. Labium narrow, lateral borders slightly arched on basal two-thirds, slightly constricted at middle of apical third; apex truncate on middle of ventral side, more arcuately produced and with small emargination at middle on oral side, and fimbriate with long, fine blackish brown setae; surface convex on anterior part, depressed on center of base, basal two-thirds with strong, dense piliferous punctures, the setae long; anterior third with minute, scattered punctures.</p><p>Antennae with 10 antennomeres, club about 2.0 times the length of antennomeres 2–7 together.</p><p>Pronotum 1.5 times as wide as long, widest near base, sides more strongly convergent to apex on anterior half; anterior border widely and arcuately projected at middle; basal angles widely rounded; basal border without marginal bead, almost uniformly arcuate, slightly more projected in central third; surface finely alutaceous (40X); punctures annulate, moderately fine and dense, the distance between most punctures on disc as long as or longer than diameter of punctures, punctures gradually larger towards sides, the diameter of a puncture at basal angle at least 2.0 times the diameter of those on center of disc, more dense on basal angles, where the distance between most punctures is as large or smaller than the diameter of a puncture; surface also covered with numerous minute punctures (50X or more). Scutellum triangular, sides feebly arcuate, apex briefly rounded; punctures similar to those of pronotal disc.</p><p>Elytra 1.2 times as long as wide, arcuately widened from first quarter to middle, then rounded to apex, sutural angle well defined, but rounded; humeral and apical umbones weak; a feeble, short, elongate tumescence behind humeral umbones on external border; surface smooth, with large, shallow, annulate punctures, much smaller on sides and apex, double rows not evident, also with numerous minute punctures (50X or more) scattered over all the surface; sutural interstria flat, weakly elevated on apical half. Epipleura narrow, more so apically.</p><p>Pygidium 2.2 times as wide as long; in posterior view the inferior borders slightly convex, converging to widely rounded apex in an angle distinctly greater than 90°, but less than in C. bicolor; apex not truncate; in lateral view uniformly convex; surface smooth, shining, with large annulate punctures, more dense on lateral angles; shiner apically, the punctures smaller and less dense, distances between most punctures more than 2.0 times their own diameter.</p><p>Protibiae (Fig. 28) tridentate, small basal tooth separated from intermediate tooth by a distance 1.5 times as long as that between the intermediate and the anterior teeth; intermediate and anterior teeth well developed, basal tooth with its anterior border more or less perpendicular to longitudinal axis of tibiae; apical spur almost straight, slightly surpassing apex of first tarsomere. Protarsi (Fig. 37–38) enlarged; tarsomere 5 shorter than 1–4 together, without ventro–internal carina, ventral side with very short and very fine longitudinal striae on inner side of apex; tarsomere 4 longitudinally striate on apical half, tarsomere 3 striated on apical third; largest claw (Fig. 43) wide, flattened, narrowly cleft at apex, the external border of small branch continuous with internal curvature of the claw. All femora almost uniformly, not densely pilose, especially the mesofemora, on which, in addition to the normal longitudinal series of piliferous punctures close to anterior and posterior borders (seen in natural position of the leg), with numerous piliferous punctures over all the surface. Metafemur (Fig. 57) 2.3 times as long as its maximum width; metatibiae (Fig. 57) robust, widened at apex, bristles on diagonal carina broken in the holotype, and those of apex very strong (mostly missing from the holotype). Metatarsi subequal in length to tibiae, with first tarsomere widened towards apex, with apical bristles similar to those of apex of tibiae.</p><p>Prosternal projection high, apex alutaceous, anterior part transversely oval, externally pointed; posterior part shorter but wider than anterior, posterior border fimbriate with long bristles.</p><p>Pronotal hypomera alutaceous, with sparse erect setae. Mesopleura, metapleura, and sides of metasternum alutaceous, with large, dense, confluent, annulate punctures; disc of metasternum smooth, with small, scattered punctures; metacoxae with punctures similar to pleurae but smaller and more disperse. Ventrites with annulate, moderately dense punctures, punctures smaller and sparser on apical ventrites.</p><p>Aedeagus: Fig. 75.</p><p>FEMALE: differs from the male in the following characters: punctation generally stronger; protarsi slender; metafemora more robust; epipleura differences with C. bicolor not observed when studying the allotype.</p><p>Distribution: Brazil: Amazonas (only known from type locality).</p><p>Specimens studied: BRAZIL: Amazonas: Rio Demeni, N. R. Equator, IV-10-78, L. A. Lacey, 1 male (holotype), 1 female (allotype) (UNSM).</p></div>	https://treatment.plazi.org/id/E2083F003B01FFA1FF74F939FDB76121	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Joly, Luis J.	Joly, Luis J. (2009): Review of the species in the Cyclocephala bicolor Laporte species group (Coleoptera: Scarabaeidae: Dynastinae). Zootaxa 2048: 47-64, DOI: 10.5281/zenodo.186485
E2083F003B00FFA3FF74FE99FB3F6702.text	E2083F003B00FFA3FF74FE99FB3F6702.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyclocephala dichroa Dechambre 1992	<div><p>Cyclocephala dichroa Dechambre, 1992</p><p>(Figs. 2, 11–14, 22, 29–30, 39, 46–47, 53 –54, 60, 65–66, 71–72)</p><p>Bibliography. Cyclocephala dichroa Dechambre 1992: 67 (description of male; compared with C. bicolor, C. bicolorata), Fig. 31 (genitalia), French Guiana. Holotype male, MNHN. Type locality: French Guiana: Route du Kaw, PK12.</p><p>Description. Short, oval, moderately shiny. Brownish red, posterior part of clypeus, frons brownish red or black, vertex black, elytra testaceous, sometimes with base and a reddish brown posteriorly narrowed sutural band; dorsal surface and pygidium glabrous, ventral pubescence sparse. Length: 10.0–13.0 mm.</p><p>MALE: Clypeus (Fig. 11) about 2.4 times as wide across base as long; sides straight, strongly convergent to truncate and not reflexed apex, shallowly sulcate longitudinally along middle and shallowly depressed laterally before apex, in lateral view slightly convex on apical half; surface finely alutaceous, with fine, shallow and widely annulate punctures, annulation wide, not well defined posteriorly, sometimes confluent, in which case appearing as irregular and fine, more or less transverse striae. Frontoclypeal suture (Fig. 11) thin, not well defined, feebly bisinuate, but not pointed posteriorly at middle. Frons: width about 2.4 times transverse diameter of eye, finely alutaceous; punctures stronger and more discrete than those on clypeus.</p><p>Mandibles with external border sinuate, apex projecting externally. Maxillae (Fig. 14) with galea moderately robust, with two apical, one median and three basal teeth, the basal teeth very small and arising from a common base; stipes projected anteriorly on inner side and with a brush of fine yellowish brown setae on apex of projection; maxillary palpi (Fig. 22) moderately robust, second palpomere widened towards apex, longer than third, fourth fusiform, more than 3.0 times as long as preceding. Labium (Fig. 13) with sides slightly convex on basal two-thirds, not constricted at middle of apical third; apex uniformly convex on ventral side, more arcuately produced, with small emargination at middle on oral side, fimbriate with long, fine, yellowish brown setae; surface almost flat, impunctate on disc, with one longitudinal series of long bristles on each side converging towards base.</p><p>Antennae with 10 antennomeres, club almost 2.0 times the length of antennomeres 2–7 together.</p><p>Pronotum 1.5 times as wide as long, widest near base, sides more strongly convergent to apex on anterior half; anterior border widely and arcuately projected at middle; basal angles widely rounded; basal border without marginal bead, almost uniformly arcuate, slightly more projected in central third; surface finely alutaceous (40X); punctures annulate, moderately fine and dense, the distance between most punctures on disc as long as or longer than diameter of punctures, punctures gradually larger toward sides, the diameter of a puncture about 2.0 times the diameter of those on center of disc, more dense on basal angles where the distance between most punctures is as long as or shorter than diameter of a puncture; all surface also covered with numerous minute punctures (50X or more). Scutellum triangular, sides feebly arcuate, apex briefly rounded; punctures similar to those on pronotal disc.</p><p>Elytra 1.1 times as long as wide, arcuately widened from first quarter to middle, then rounded to apex, sutural angle well defined, but rounded; humeral and apical umbones weak; a feeble, short, elongate tumescence behind humeral umbones on external border; surface smooth, with large, shallow, annulate punctures, much smaller on sides and apex, double rows not evident, also with numerous micropunctures (50X or more) scattered all over the surface; sutural interstria flat, a little elevated on apical half. Epipleura narrow, more so apically.</p><p>Pygidium almost 2.0 times as wide as long; in posterior view the inferior borders almost straight, converging to feebly convex-truncate apex in an angle distinctly greater than 90°; in lateral view (Fig. 65) convex, more strongly on apical half; surface smooth, shiny, with large annulate punctures, more dense on lateral angles; on apex more shining, the punctures somewhat smaller and less dense, the distances between most punctures 1.0–2.0 times their own diameter.</p><p>Protibiae (Fig. 29) tridentate, small basal tooth separated from the intermediate tooth by a distance 2.0 times as long as that between intermediate and anterior teeth; intermediate and anterior teeth well developed, basal tooth acute and anteriorly projected; apical spur almost straight, surpassing apex of first tarsomere. Protarsi (Fig. 39) enlarged; tarsomere 5 shorter than 1–4 together, without ventro-internal carina, ventral side with inconspicuous longitudinal striae on inner side of apex; tarsomere 4 longitudinally striated on about apical half, tarsomere 3 usually without apical striae, sometimes (2 out of 6 males) with 1–2 striae on inner side of apex; largest claw (Figs. 46–47) wide, flattened, moderately widely cleft at apex, the external border of small branch continuous with internal curvature of the claw. Metafemur (Fig. 53) 2.0 times as long as its maximum width. Metatibiae (Fig. 53) robust, widened at apex, bristles on diagonal carina and apex very strong. Metatarsi subequal in length to tibiae, with first tarsomere widened towards apex, with apical bristles similar to those of apex of tibiae.</p><p>Prosternal projection high, apex alutaceous, anterior part transversely oval, externally pointed; posterior part with posterior border fimbriate with long bristles.</p><p>Pronotal hypomera alutaceous, with sparse, erect setae. Mesopleura, metapleura, and sides of metasternum alutaceous, with large, dense, confluent, annulate punctures; disc of metasternum smooth, with small, scattered punctures; metacoxae with small, moderately dense punctures, more dense and confluent on posterior and external borders. Ventrites with moderately dense annulate punctures, some open posteriorly; second ventrite with strong, triangular carina along middle, from basal quarter to apex.</p><p>Aedeagus: Figs. 71–72.</p><p>FEMALE: differs from the male in the following characters: punctation generally stronger, that of pygidium of the same size; protarsi slender; protibiae Fig. 30; metafemora (Fig. 54) somewhat more robust; metatibiae Fig. 54; pygidium (Fig. 66) flattened in lateral view; epipleura (Fig. 60) strongly narrowed before anterior border of metacoxae, then widened up to level of middle of second ventrite.</p><p>Distribution: Venezuela, French Guiana, Brazil, and Peru. Venezuela, Brazil, and Peru are new country records.</p><p>Specimens studied: VENEZUELA: Amazonas: Culebra, P. N. Duida-Marahuaca, 3º 44’ N, 65º 46’ W, 24-26-I-1992, Exped. Terramar, J. Clavijo-A. Chacón, 1 F (MIZA). Pto. Ayacucho, 22-IV-1967, P. J. Salinas, 1 M (MIZA); 4-15-XII-1982, G. Romero, 4 F (MIZA). Rio Baría, 0º 55’ N, 66º 10’ W, 140 m, L. J. Joly-A. Chacón, 1 F (MIZA). Barinas: Reserva Forestal de Ticoporo, 230 m, 10-IV-1968, en la luz, F. Fernández Y.- L. J. Joly, 1 F (MIZA). Bolívar: El Barroso, Río Matú, 7-I-1972, J. Salcedo, 1 F (MIZA). Los Pijiguaos, 600 m, 3-13-VI-1992, Exped. MIZA, 1 M (MIZA). Río Guaniamo, 6º 45’ N, 66º 01’ W, 8-12-V-1979, 1 M (MIZA). Portuguesa: Biscucuy, 13-IV-1981, 1 M (MIZA). Táchira: Nula, 9-III-1967, L. J. Joly-V. Obregón, 2 M, 1 F (MHNLS). BRAZIL: Pará: Cachimbo, 400 m, 14/ 21-IX-1955, L. Travassos &amp; S. Oliveira, 5 M, 1 F (HAHC). PERU: Huánuco: Tingo María, 17-V-1947, 2.200 ft, J. C. Pallister Coll. Donor Frank Johnson, 1 F (AMNH).</p><p>Other localities: FRENCH GUIANA: Route de Kau; Saül; St. Laurent (Dechambre 1992).</p></div>	https://treatment.plazi.org/id/E2083F003B00FFA3FF74FE99FB3F6702	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Joly, Luis J.	Joly, Luis J. (2009): Review of the species in the Cyclocephala bicolor Laporte species group (Coleoptera: Scarabaeidae: Dynastinae). Zootaxa 2048: 47-64, DOI: 10.5281/zenodo.186485
E2083F003B02FFBDFF74F8BEFE5D650D.text	E2083F003B02FFBDFF74F8BEFE5D650D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyclocephala anibali	<div><p>Cyclocephala anibali, new species</p><p>Figs. 1, 15–18, 23, 31–32, 40, 48, 55–56, 61, 67–68, 76–77</p><p>HOLOTYPE male, ALLOTYPE female. VENEZUELA: Apure: Capanaparo, Hato Los Turpiales, 8-V- 1980, E. Osuna (MIZA). PARATYPES. VENEZUELA: Apure: Fundo La Florida [6° 52’ 30” N, 68° 57’ 44” W], cr. río Quitaparo, 7° 05’N, 68° 36’W, 12-13-XI-2001, E. Osuna, 1 M, 1 F (MIZA).</p><p>Description. Holotype male. Short, oval, moderately shiny. Brownish red, frons, except anterior part, and vertex black, elytra testaceous. Dorsal surface and pygidium glabrous; ventral pubescence sparse. Length: 11.0 mm.</p><p>Clypeus (Fig. 15) about 2.3 times as wide across base as long; sides straight, strongly convergent to truncate and arcuately reflexed apex; shallowly sulcate longitudinally along middle and shallowly depressed laterally before apex, in lateral view flat to slightly concave on apical half; surface finely alutaceous, with fine, discrete, shallow, punctures thinner and sparser toward apex and anterior half of sides. Frontoclypeal suture thin, bisinuate and angularly projecting posteriorly at middle. Frons: width about 2.8 times transverse diameter of eye, finely alutaceous; punctures, except on smooth longitudinal middle line, fine, confluent by pairs so they appear as transverse, stronger and not confluent posteriorly.</p><p>Mandibles with external border sinuate, apex projecting externally. Maxillae (Fig. 18) with galea moderately robust, with two apical, one median, and three basal teeth, the basal teeth very small and arising from a common base; stipes anteriorly projected on inner side, and with a brush of fine yellowish brown setae on apex of projection; maxillary palpi (Fig. 23) moderately robust, second palpomere subcylindrical, longer than third, fourth fusiform, more than 3.0 times as long as preceding. Labium (Fig. 17) with sides slightly convex on basal two-thirds, not constricted at middle of apical third; apex convex and widely emarginated at middle, and fimbriate with long, yellowish brown setae; surface almost flat, with one longitudinal series of long bristles on each side, converging towards base.</p><p>Antennae with 10 antennomeres, club about 1.5 times the length of antennomeres 2–7 together.</p><p>Pronotum 1.5 times as wide as long, widest near base, sides more strongly convergent to apex on anterior half; anterior border widely and arcuately projected in middle; basal angles widely rounded; basal border without marginal bead, almost uniformly arcuate, slightly more projected in central third; surface smooth (40X); punctures annulate, moderately fine and dense, the distance between most punctures on disc as long as or longer than diameter of punctures, punctures gradually larger towards sides, the diameter of a puncture about 1.5 times the diameter of those on center of disc, more dense on basal angles, where the distance between most punctures is as large as or smaller than diameter of punctures; all surface also covered with numerous minute punctures (50X or more). Scutellum triangular, sides feebly arcuate, apex briefly rounded; punctures much finer than those of pronotal disc.</p><p>Elytra 1.2 times as long as wide, arcuately widened from first quarter to middle, then rounded to apex, sutural angle well defined, but rounded; humeral and apical umbones slight; a feeble, short, elongate tumescence behind humeral umbones on external border; surface smooth, with large, shallow, annulate punctures, much smaller on sides and apex, double rows not evident, also with numerous minute punctures (50X or more) scattered over all the surface; sutural interstria flat, weakly elevated on apical half. Epipleura narrow, more so apically.</p><p>Pygidium 1.9 times as wide as long; in posterior view the inferior borders almost straight, converging to widely and feebly convex-truncate apex at an angle of about 90°, in lateral view (Fig. 67) uniformly convex, more strongly on apical half; surface smooth, shiny, with small and dense annulate punctures, distances between most punctures as large as or smaller than diameter of punctures on basal half; punctures on apical region much smaller, simple, the distances between most punctures much longer than the diameter of punctures.</p><p>Protibiae (Fig. 31) tridentate, short basal tooth separated from intermediate tooth by a distance 1.2 times as long as that between intermediate and anterior teeth; intermediate and anterior teeth well developed, basal tooth with its anterior border more or less perpendicular to long axis of tibiae; apical spur almost straight, surpassing apex of first tarsomere. Protarsi (Fig. 40) enlarged; tarsomere 5 shorter than 1–4 together, without ventro-internal carina, ventral side with very short and very fine longitudinal striae on inner side of apex; tarsomere 4 longitudinally striate on apical third, tarsomere 3 on apical fifth; largest claw (Fig. 48) wide, flattened, moderately narrowly cleft at apex, external border of small branch continuous with internal curvature of claw. Metafemur (Fig. 55) 2.3 times as long as its maximum width. Metatibiae (Fig. 55) robust, widened at apex, bristles on diagonal carina and apex very strong. Metatarsi subequal in length to tibiae, with first tarsomere scarcely widened towards apex, with apical bristles similar to those of apex of tibiae.</p><p>Prosternal projection high, apex alutaceous, anterior part transversely oval, not externally pointed; posterior part not wider than anterior one, posterior border fimbriate with long bristles.</p><p>Pronotal hypomera alutaceous, with sparse erect setae. Mesopleura, metapleura, and sides of metasternum alutaceous, with large and very dense confluent, annulate punctures; disc of metasternum smooth, with small, disperse punctures; metacoxae with punctures similar to pleurae but smaller and more disperse. Ventrites with fine, annulate, moderately dense punctures.</p><p>Aedeagus: Figs. 76–77.</p><p>Allotype female. differs from the male in the following characters: punctation generally stronger; protibiae as in Fig. 32; protarsi slender; metafemora (Fig. 56) somewhat more robust; epipleura (Fig. 61) relatively wide in relation to other species in the group (see Figs. 58–60), gradually narrowed to beginning of second ventrite; pygidium in lateral view as in Fig. 68.</p><p>Etymology. This species is named in honor of my friend Anibal Chacón.</p><p>Distribution. Venezuela.</p><p>Diagnosis. Cyclocephala anibali can be differentiated from all other species in the Cyclocephala bicolor species group by the following combination of character states: Surface of clypeus finely alutaceous with discrete punctures; frontoclypeal suture posteriorly projecting angularly; antennal club about 1.5 times as long as antennomeres 2–7 together; punctures on scutellum very fine, distinctly finer than those on middle of base of pronotum; punctures on basal half of pygidium small and dense, distinctly finer on apical half; prosternal projection with anterior part not externally pointed; basal tooth of protibia not forwardly directed, its anterior border more or less perpendicular to longitudinal axis of tibia; pygidium about 1.9 times as wide as long. The aedeagus is relatively wide with apical external angles rounded</p><p>Cyclocephala sarahae is very similar to C. anibali, but it can be differentiated because most punctures on clypeus are confluent, the frontoclypeal suture is sinuate, not posteriorly projecting angularly, the antennal club about 2.0 times as long as antennomeres 2–7 together, punctures on scutellum similar to those on pronotal disc, punctures on basal half of pygidium large, smaller on apical half, the prosternal projection with anterior part externally pointed, the pygidium about 2.2 times as wide as long and the aedeagus is pointed on external apical angles.</p></div>	https://treatment.plazi.org/id/E2083F003B02FFBDFF74F8BEFE5D650D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Joly, Luis J.	Joly, Luis J. (2009): Review of the species in the Cyclocephala bicolor Laporte species group (Coleoptera: Scarabaeidae: Dynastinae). Zootaxa 2048: 47-64, DOI: 10.5281/zenodo.186485
