taxonID	type	description	language	source
E07B87DEFFCDFF95FF226EB4E2A1FCEF.taxon	description	(Figs 1 – 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFCDFF95FF226EB4E2A1FCEF.taxon	type_taxon	Type species: Guapinannus bierigi Wygodzinsky, 1951	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFCDFF95FF226EB4E2A1FCEF.taxon	diagnosis	Diagnosis: Recognized among Schizopteridae by the relatively small eyes (Fig. 5 C, D), four-segmented labium (Fig. 1 A), flat and wide forewings (Figs 1 – 4), unique forewing venation with long fracture in the costal margin, long rc 1, rc, and tc cells (Figs 1 B, 2, 6 A), C + S, with An 1 / 2 claval process except G. dispar sp. n. (Figs 2, 6 A, D), better developed in males than in females (Fig 1 B), males with wing organ associated with An 1 on corium adjacent to the claval process (Figs 2 – 4, 6 A, D), 2 - 2 - 3 tarsal formula in both sexes, male genitalia simple (Fig. 7) and female with well-developed ovipositor (Figs 1 C, 8). Re-description: Male: Ovoid (greatest length to width ratio 1.57 – 1.85) to elongate ovoid (length to width ratio 3.4), total body length 0.66 – 2.06 mm; length from posterior margin of pronotum to posterior wing margin (0.51 – 1.49 mm). General coloration: uniformly pale or a mix of pale and darker brown or reddish brown. Vestiture: Dorsum with dense setae, setae long or short, setae on posterior margin of forewing long or extremely long. Structure: Head: Eye ranging from small (less than one quarter head height) to relatively large (almost half as high as head) (Fig. 5 C, D); ocellus adjacent to eye (Fig. 5 D), at 11 o’clock position seen in lateral view and about twice the size of one ommatidium; four-segmented labium with segments 1, 2 and 3 of similar length and 4 about 2.5 – 3 times as long as any other segment (Fig. 1 A); muscle scars present on the frons and vertex present (Figs 1 A, 5 C). Thorax: Collar in dorsal view narrow (Fig. 5 C); proepisternal lobe inflated (Fig. 5 D); separation of anterior and posterior lobes (dorsal view) marked by weak depression (Fig. 5 C); anterior pronotal lobe with muscle scars, margin concave or straight; posterior margin slightly concave to almost straight; pits on pronotum ranging from small to large, evenly distributed or grouped (Fig. 1 A); scutellum in dorsal view abruptly or gradually narrowed to tip, tip pointed, rounded, or slightly inflated (Figs 2 – 4). Forewing: With outline elongate oval or broad anteriorly; with (G. dispar, sp. n., Fig. 2) or without (all other species) distinct honey-comb pattern; membrane (cub, dc 1, rc 2 – 3 and areas distal to these cells) contributing about one third or less to forewing length; posterior distal margin rounded or square; clavus twice as long as wide; vein-tracing areoles along margins of a combination of scc, rc 1, rc 1, rc 2, and tc on proximal part of wing absent or present (Fig. 6 A, C); C + Sc narrow (Fig. 2, G. anaticulus, sp. n.), fairly broad (Fig. 2, G. clava, sp. n.), broad (Fig. 2, G. falcis, sp. n.), or very broad (Fig. 3, G. lutuosus, sp. n.), either of uniform width (Fig. 2, G. anaticulus, sp. n.) or wider at widest part of wing (Fig. 2, G. auriculu s, sp. n.); R flattened in distal part (black arrow in Fig. 1 B); distal process of R pronounced (white arrow in Fig. 1 B) or weakly developed; An 1 / 2 claval process well developed in males (except G. tenuis, sp. n.), overlapping base of An 1 wing organ, apex acute and in lateral view pointing dorsad (Figs 2 – 4, 6 A, D); An 1 wing organ on corium strongly inflated in all males, with or without (e. g., Fig. 4, G. tergus, sp. n.) notch articulating with claval process, wing organ rounded, elongate, or acute, with or without median notch; scc about 1.5 times longer than wide (Fig. 6 A) (except ~ 2 times longer than wide in G. castigatus, sp. n.); scc undivided except in G. dispar, sp. n. (Fig. 2); rc 1 about 3 times as long as wide except in G. dispar, sp. n., ranging from parallel-sided to slightly or much wider anteriorly; rc about 4 or 5 times as long as wide; tc 3 times or more than 3 times as long as wide; part of An 1 and Cu forming posterodistal margin of tc of similar width as other veins (e. g., Fig. 2, G. artus, sp. n.) or thicker (e. g., Fig. 2, G. falcis, sp. n.); distal margin of rc 1, rc, and tc cells double s-shaped (Fig. 2, G. anaticulus, sp. n.), s-shaped (Fig. 1, G. bierigi), or angular s-shaped (Fig. 2, G. dispar, sp. n.); rc 2 – 3 almost triangular (e. g., Fig. 2, G. anaticulus, sp. n.), trapezoidal (e. g., Fig. 2, G. castigatus, sp. n.), or rectangular (Fig. 2, G. dispar, sp. n.); cub oval, almost reaching wing margin or distant form wing margin (except elongate in G. dispar; Fig. 2); dc 1 with basal portion fairly narrow and elongate (Fig. 2, G. anaticulus, sp. n.) or relatively wide and short (Fig. 2, G. auriculu s, sp. n.); M beyond Cu short. Legs: males and females with tarsal formula 2 - 2 - 3, with claws long except short in G dispar, sp. n. Genitalia (Fig. 7): Pygophore without lobe on right side (Fig. 7, G. anaticulus, sp. n.), with small (Fig. 7, G. artus, sp. n.) or large lobe (Fig. 7, G. auriculus, sp. n.), or with slightly extended right margin (Fig. 7, G. castigatus, sp. n.). Vesica about one half loop (e. g., Fig. 7, G. castigatus, sp. n.), between half and one loop (e. g., Fig. 7, G. anaticulus, sp. n.), or one loop (e. g., Fig. 7, G. plurilobus, sp. n.), either not reaching right pygophore margin or surpassing margin, at midpoint thick (e. g., Fig. 7, G. graziae, sp. n.), very thick (e. g., Fig. 7, G. robustus, sp. n.), or more slender (e. g., Fig. 7, G. falcis, sp. n.), tapering to blunt or acute tip, tip itself blunt, bottle-opener shaped (e. g., Fig. 7, G. uncus, sp. n.), slightly s-shaped (e. g., Fig. 7, G. falcis, sp. n.), rounded (e. g., Fig. 7, G. artus, sp. n.), or fairly straight (e. g., Fig. 7, G. tatumbia, sp. n.). Anophoric ridge with process present (e. g., Fig. 7, G. artus, sp. n.) or absent (Fig. 7, G. castigatus, sp. n.), process short or long, with single lobe curved or relatively straight, or with multiple lobes, tip pointed, blunt, or combination of pointed and blunt. Right paramere curved or straight, tapering towards apex or broad throughout; basal process of paramere broad, narrow, thumb-shaped, or duck head-shaped. Left paramere straight or curved, tapering towards apex or broad throughout, basal process broad or narrow. Female: As male, but with less well developed An 1 / 2 claval process and without An 1 wing organ on corium (Fig. 1 B). Genitalia: with well-developed ovipositor (Figs 1 C, 8). Reservoir of spermatheca kidney-shaped with short spermathecal duct (Fig. 8).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFCDFF95FF226EB4E2A1FCEF.taxon	distribution	Distribution. Species of Guapinannus are documented from Central and South America, ranging from Veracruz in Mexico to the southern part of Brazil. The greatest species diversity is found in Central America and the northern areas of South America. Specimens have been collected in tropical wet or moist forest, including cloud forest and mixed hardwood forest, from elevation ranging from 5 meters above sea level in Panama to more than 200 meters in Chiapas, Mexico. Collecting method. The majority of specimens for which collection method was recorded were collected using leaf litter sifting and Berlese and Winkler extraction methods. A small number of specimens are derived from Malaise and flight intercept traps.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFCDFF95FF226EB4E2A1FCEF.taxon	discussion	Discussion. The forewing shape, structure with distinct claval suture, and venation set apart Guapinannus from all other genera of Schizopteridae. While Humpatanannus and related genera have not been included in phylogenetic analyses, genitalic features are strikingly different in the two groups, and they are unlikely to be close relatives. The non-genitalic morphology of species of Guapinannus is fairly uniform. Most species are mid-sized and ovoid with some variation, with only few species being distinctly smaller or bigger or markedly elongated ovoid. Some forewing venation features are fairly distinctive and useful as diagnostic characters at the species level, among them the width of the C + Sc and the relative proportion of certain cells. As in many other genera of Schizopteridae, the bulk of diagnostic features is derived from male genitalic structures, in particular the length, curvature, and tip shape of the vesica, shape of the anophoric process, where present, and shape of the pygophore that is expanded into a distinct lobe on the right side in some species. We described as new two species for which only female specimens are known (G. dispar, sp. n. and G. minutus, sp. n.). Both are distinctive based on non-genitalic morphology and G. dispar, sp. n. in addition is geographically isolated from all other known species of Guapinannus. As part of this study we discovered a so far undocumented structure on the forewings that is restricted to males that we interpret as a novel male-specific wing organ. This wing organ is located on the posterior margin of the corium, associated with An 1, consist of a strongly inflated semicircular or elongate area and in most species features a median notch. SEM observation of this wing organ in one species (Fig. 6 D) shows a tuft of hairlike structures that may represent a trichome. Based on our observations, males of all species of Guapinannus possess this wing organ. Among closely related taxa (based on Knyshov et al., 2020), Caucanannus also have a male-specific wing organ, although in a different position on the forewing (Weirauch et al., 2020), while wing organs have so far not been documented in Luachimonannus and Kokeshia.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFCAFF94FF226BA0E597FA4D.taxon	description	Measurements are based on male specimens, except for G. minutus, sp. n., G. dispar, sp. n. and Guapinannus bierigi Wygodzinsky where only female specimens are known.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFCAFF97FF226ED9E493FD7F.taxon	description	(Figs 2, 7, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFCAFF97FF226ED9E493FD7F.taxon	materials_examined	Holotype. Male. GUATEMALA: San Marcos: Bojonal Road, 14.94786 ° N 91.8837 ° W, 1618 m, 04 Jun 2015, R. Anderson, 1 ♂ (UCR _ ENT 00127123) (UCR).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFCAFF97FF226ED9E493FD7F.taxon	etymology	Etymology: Named for the duck head-shaped basal process of the right paramere after “ anas ” meaning duck in Latin.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFCAFF97FF226ED9E493FD7F.taxon	diagnosis	Diagnosis: Recognized by the mid-sized body length (1.45 mm), oval forewing outline, narrow C + SC (black arrow), rc 1 about 3 times as long as wide and slightly wider anteriorly (white arrow), tc more than 3 times as long as wide (Fig. 2), the short, slender, and straight vesica with blunt, expanded tip, and the broad, duck head-shaped basal process of the right paramere (Fig. 7).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFCAFF97FF226ED9E493FD7F.taxon	description	Description: Male: Total body length 1.45 mm; length from posterior margin of pronotum to posterior wing margin 1.23 mm; greatest width across pronotum 0.63 mm; greatest width across forewings 0.81 mm. General coloration: mix of pale and darker brown or reddish brown. Vestiture: Dorsum with dense setae, setae long; setae on posterior margin of forewing extremely long. Structure: Head: Eye almost half as high as head; muscle scars large. Thorax: Anterior pronotal margin concave, posterior margin slightly concave; pits on pronotum mediumsized, evenly distributed; scutellum in dorsal view abruptly narrowed to tip, tip pointed. Forewing: With outline oval; membrane contributing about one third of forewing length, posterior distal margin rounded, vein-tracing areoles on proximal part of wing absent; Sc + C uniform, relatively narrow; distal process of R pronounced; An 1 wing organ on corium strongly inflated, rounded, with median notch, without notch articulating with claval process; rc 1 about 3 times as long as wide, slightly wider anteriorly; rc about 5 times as long as wide; tc more than 3 times as long as wide; distal margin of rc 1, rc, and tc cells double s-shaped; rc 2 - 3 almost triangular; cub almost reaching wing margin; dc 1 with basal portion fairly narrow and elongate. Legs: Claws long. Genitalia (Fig. 7): Pygophore without lobe on right side; vesica with between half and one loop, surpassing right pygophore margin, thick at midpoint, tapering to blunt, expanded tip, close to apex slightly s-shaped; anophoric ridge with process, process short, with single lobe curved, tip blunt; right paramere curved, tapering, basal process duck head-shaped; left paramere straight, tapering, basal process broad. Female: Unknown. Collecting method and habitat: Collected in cloud forest litter using sifting leaf litter.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC9FF97FF2269CCE0D4F86F.taxon	description	(Figs 2, 7, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC9FF97FF2269CCE0D4F86F.taxon	materials_examined	Holotype. Male. MEXICO: Veracruz: Road to Ruiz Cortinez, 18.5229 ° N 95.15441 ° W, 1114 m, 05 Jun 2016, R. S. Anderson, 1 ♂ (UCR _ ENT 00125959) (UCR). Paratypes: MEXICO: Veracruz: Road to Ruiz Cortinez, 18.5229 ° N 95.15441 ° W, 1114 m, 05 Jun 2016, R. S. Anderson, 1 ♀ (UCR _ ENT 00125960) (UCR). Road to Ruiz Cortinez, 18.52241 ° N 95.14957 ° W, 1170 m, 04 Jun 2016, R. S. Anderson, 1 ♀ (UCR _ ENT 00125961) (UCR). San Andres Tuxtla Co.: 18.52303 ° N 95.15453 ° W, 1109 m, 05 Jun 2016, Manuel Barrios, 2 ♂ (UCR _ ENT 00128215, UCR _ ENT 00128216) (UCR).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC9FF97FF2269CCE0D4F86F.taxon	etymology	Etymology: Named for narrow rc cell of the forewing in dorsal view after “ artus ” Latin for narrow.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC9FF97FF2269CCE0D4F86F.taxon	diagnosis	Diagnosis: Recognized by the mid-sized body length (1.29 mm), elongate oval forewing outline, rc about 5 times as long as wide (black star), tc about 3 times as long as wide (Fig. 2), lateral lobe on right side of pygophore (black star), slender vesica, and short, broad anophoric process with blunt tip (black arrow) (Fig. 7). Similar to G. auriculus, sp. n. that is also known from Veracruz in Mexico but distinguished by the characters listed in the diagnosis.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC9FF97FF2269CCE0D4F86F.taxon	description	Description: Male: Total body length 1.29 mm, length from posterior margin of pronotum to posterior wing margin 1.04 mm; greatest width across pronotum 0.54 mm; greatest width across forewings 0.75 mm. General coloration: mix of pale and darker brown or reddish brown. Vestiture: Dorsum with dense setae, setae long; setae on posterior margin of forewing long. Structure: Head: Eye almost half as high as head; muscle scars large. Thorax: Anterior pronotal margin concave, posterior margin slightly concave; pits on pronotum large, in distinct clusters; scutellum in dorsal view abruptly narrowed to tip, tip rounded. Forewing: with outline broad anteriorly, elongate oval; membrane contributing about one third of forewing length, posterior distal margin squarish, veintracing areoles on proximal part of wing absent; C + Sc wider at widest part of wing; distal process of R pronounced; An 1 wing organ on corium strongly inflated, rounded, with median notch, without notch articulating with claval process; rc 1 about 3 times as long as wide, much wider anteriorly; rc about 5 times as long as wide; tc 3 times as long as wide; part of An 1 and Cu forming posterodistal margin of tc of similar width as other veins; distal margin of rc 1, rc, and tc cells double s-shaped; rc 2 – 3 almost triangular; cub almost reaching wing margin; dc 1 horse headshaped. Legs: Claws long. Genitalia (Fig. 7): Pygophore with lobe on right side; vesica with between half and one loop, not reaching right pygophore margin, relatively slender at midpoint, tapering to acute tip, close to apex rounded; anophoric ridge with process, process short, broad, with single lobe curved, tip blunt; right paramere curved, tapering, basal process narrow; left paramere, tapering, basal process narrow. Female: As in male and generic description. Collecting method and habitat: Collected using sifting leaf litter, Winkler extraction in mixed hardwood forest.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC8FF96FF226BA0E103FA83.taxon	description	(Figs 2, 7, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC8FF96FF226BA0E103FA83.taxon	materials_examined	Holotype. Male. MEXICO: Veracruz: San Andres Tuxtla Co.: 18.52303 ° N 95.15453 ° W, 1109 m, 05 Jun 2016, Manuel Barrios, 4 ♂ (UCR _ ENT 00125965) (UCR). Paratypes: MEXICO: Veracruz: hills above Ruiz Cortinez, 18.5304 ° N 95.14127 ° W, 1239 m, 07 Jun 2016, R. S. Anderson, 1 # f (UCR _ ENT 00126156) (UCR). San Andres Tuxtla Co.: 18.52303 ° N 95.15453 ° W, 1109 m, 05 Jun 2016, Manuel Barrios, 8 ♂ (UCR _ ENT 00125966, UCR _ ENT 00128211, UCR _ ENT 00128214), 13 ♀ (UCR _ ENT 00125967, UCR _ ENT 00126150) (UCR).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC8FF96FF226BA0E103FA83.taxon	etymology	Etymology: Named for the large lobe on the right side of the pygophore after “ auricula ” Latin for lobe.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC8FF96FF226BA0E103FA83.taxon	diagnosis	Diagnosis: Recognized by the mid-sized body length (1.23 mm), rc about 4 times as long as wide (white star), tc less than 3 times as long as wide (Fig. 2), large lateral lobe on right side of pygophore (black star), slender vesica tapering into acute tip, and anophoric process very short, curved, and pointed (black arrow) (Fig. 7). Similar to G. artus, sp. n. that is also known from Veracruz in Mexico but distinguished by the characters listed in the diagnosis. Additionally, G. auriculus, sp. n. and G. castigatus, sp. n. have a similar vesica, but differ in rc 1 shape.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC8FF96FF226BA0E103FA83.taxon	description	Description: Male: Total body length 1.23 mm; length from posterior margin of pronotum to posterior wing margin 0.9 mm; greatest width across pronotum 0.55 mm; greatest width across forewings 0.75 mm. General coloration: mix of pale and darker brown or reddish brown. Vestiture: Dorsum with dense setae, setae long; setae on posterior margin of forewing extremely long. Structure: Head: Eye almost half as high as head; muscle scars large. Thorax: Anterior pronotal margin concave, posterior margin slightly concave; pits on pronotum large, evenly distributed; scutellum in dorsal view abruptly narrowed to tip, tip rounded. Forewing: With outline broad anteriorly; membrane contributing about one third of forewing length, posterior distal margin rounded, vein-tracing areoles on proximal part of wing absent; C + Sc broad, wider at widest part of wing; distal process of R pronounced; An 1 wing organ on corium strongly inflated, rounded, with median notch, without notch articulating with claval process; rc 1 about 3 times as long as wide, much wider anteriorly; rc about 4 times as long as wide; tc 3 less than 3 times as wide; part of An 1 and Cu forming posterodistal margin of tc of similar width as other veins; distal margin of rc 1, rc, and tc cells double s-shaped; rc 2 - 3 almost triangular; cub almost reaching wing margin; dc 1 with basal portion relatively wide and short. Legs: Claws long. Genitalia (Fig. 7): Pygophore with large lobe on right side; vesica with between half and one loop, not reaching right pygophore margin, thick at midpoint, tapering to acute tip, close to apex rounded; anophoric ridge with process, process short, with single lobe curved, tip pointed; right paramere curved, tapering, basal process narrow; left paramere straight, tapering, basal process narrow. Female: As in male and generic description. Collecting method and habitat: Collected using sifting leaf litter, Winkler extraction in broadleaf forest and tree ferns.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC8FF99FF226E90E3ACFEB3.taxon	description	(Figs 1, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC8FF99FF226E90E3ACFEB3.taxon	diagnosis	Diagnosis: Recognized by the mid-sized body length (1.45 mm), part of An 1 and Cu forming posterodistal margin of tc more pronounced than other veins, and the great distance of cub from the margin of the wing. Most similar to G. falcis, sp. n., that is also known from Costa Rica, but distinguished by the much greater distance of cub from the wing margin compared to G. falcis, sp. n. Re-description: Female: Total body length 1.45 mm, greatest width across pronotum 0.67 mm. General coloration: mix of pale and darker brown or reddish brown. Vestiture: Dorsum with sparse setae, setae short; setae on posterior margin of forewing long. Structure: Head: Eye almost half as high as head; muscle scars large. Thorax: Anterior pronotal margin concave, posterior margin straight; pits on pronotum medium-sized, evenly distributed; scutellum in dorsal view abruptly narrowed to tip, tip pointed. Forewing: With outline broad anteriorly; membrane contributing about one third of forewing length, posterior distal margin squarish, vein-tracing areoles on proximal part of wing absent; C + Sc broad, wider at widest part of wing; distal process of R pronounced; An 1 wing organ on corium absent (as in all females); rc 1 about 3 times as long as wide, slightly wider anteriorly; rc about 4 times as long as wide; tc about three times as long as wide; part of An 1 and Cu forming posterodistal margin of tc more pronounced than other veins; distal margin of rc 1, rc, and tc cells s-shaped; rc 2 - 3 almost triangular; cub removed from wing margin; dc 1 with basal portion fairly narrow and elongate. Legs: Claws long. Genitalia: As in generic description (Fig. 1). Male: Unknown. Specimens examined: Holotype: COSTA RICA: Limon: Guapiles, 10.2167 ° N 83.7833 ° W, 268 m, 05 Feb 1946 – 13 Feb 1946, A. Bierig, 1 ♀ (AMNH _ IZC 00150354) (AMNH).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC7FF99FF226A80E463FA33.taxon	description	(Figs 2, 7, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC7FF99FF226A80E463FA33.taxon	materials_examined	Holotype. Male. MEXICO: Chiapas: Palenque, 15.63333 ° N 92.65 ° W, 2500 m, 02 Jul 1983 – 30 Jul 1983, S. B. Peck, J. Kukalova-Peck, R. S. Anderson, 1 ♂ (UCR _ ENT 00101251) (FMNH).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC7FF99FF226A80E463FA33.taxon	etymology	Etymology: Named for the stout vesica after “ castigatus ” Latin for stout.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC7FF99FF226A80E463FA33.taxon	diagnosis	Diagnosis: Recognized by the mid-sized body length (1.48 mm), elongate oval forewing outline, wide C + Sc (black arrow), small vein-tracing areoles along scc, rc, rc 1 (white arrow) (Fig. 2), slightly extended right pygophore margin (black star), vesica relatively short and thick, tapering into acute tip, and without anophoric process (Fig. 7). Similar to G. auriculus, sp. n. in length and shape of vesica but differentiated by differentiated from that species by the slender and long rc 1.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC7FF99FF226A80E463FA33.taxon	description	Description: Male: Total body length 1.48 mm; length from posterior margin of pronotum to posterior wing margin 1.27 mm; greatest width across pronotum 0.66 mm; greatest width across forewings 0.88 mm. General coloration: mix of pale and darker brown or reddish brown. Vestiture: Dorsum with dense setae, setae long; setae on posterior margin of forewing long. Structure: Head: Eye almost half as high as head; muscle scars large. Thorax: Anterior pronotal margin concave, posterior margin slightly concave; pits on pronotum medium-sized, evenly distributed; scutellum in dorsal view gradually narrowed to tip, tip pointed. Forewing: With outline elongate oval; membrane contributing about one third of forewing length, posterior distal margin squarish, vein-tracing areoles on proximal part of wing present; areoles small, distributed along scc, rc, rc 1, and tc; C + Sc broad, wider at widest part of wing; distal process of R pronounced; An 1 wing organ on corium strongly inflated, elongate, with median notch, without notch articulating with claval process; scc about 2 times as wide as long; rc 1 about 3 times as long as wide, slightly wider anteriorly; rc about 4 times as long as wide; tc about 3 times as long as wide; part of An 1 and Cu forming posterodistal margin of tc of similar width as other veins; distal margin of rc 1, rc, and tc cells double s-shaped; rc 2 - 3 trapezoidal; cub almost reaching wing margin; dc 1 with basal portion relatively wide and short. Legs: Claws long. Genitalia (Fig. 7): Pygophore with slightly extended right margin; vesica with about half loop, not reaching right pygophore margin, thick at midpoint, tapering to acute tip, close to apex rounded; anophoric ridge without process; right paramere curved, tapering, basal process narrow; left paramere straight, tapering, basal process narrow. Female: Unknown. Collecting method and habitat: Collected using flight intercept trap in rainforest.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC7FF98FF226D00E166FBC5.taxon	description	(Figs 2, 5 – 7, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC7FF98FF226D00E166FBC5.taxon	materials_examined	Holotype. Male. PANAMA: Bocas del Toro: Almirante, at dam on Nigua Creek, 9.25 ° N 82.65 ° W, 810 m, 23 Mar 1959, H. S. Dybas, 1 ♂ (UCR _ ENT 00097823) (FMNH). Paratypes: COSTA RICA: Puntarenas: 5 km SW Finca Las Cruces, La Fila, 8.78937 ° N 82.98133 ° W, 1433 m, 21 Mar 1973, J. A. Wagner, J. B. Kethley, 1 ♂ (UCR _ ENT 00090603) (FMNH). PANAMA: Bocas del Toro: Almirante, at dam on Nigua Creek, 9.25 ° N 82.65 ° W, 1080 m, 23 Mar 1959, H. S. Dybas, 2 ♂ (UCR _ ENT 00090784, UCR _ ENT 00097822), 7 ♀ (UCR _ ENT 0009782, UCR _ ENT 00097824 - UCR _ ENT 00097829) (FMNH); 25 Mar 1959, H. S. Dybas, 1 ♂ (UCR _ ENT 00090740) (FMNH). Almirante, trail to dam on Nigua Creek, 9.3003 ° N 82.40214 ° W, 5 m, 23 Mar 1959, H. S. Dybas, 3 ♀ (UCR _ ENT 00090748, UCR _ ENT 00090778, UCR _ ENT 00090779), 4; u (UCR _ ENT 00090757, UCR _ ENT 00090758, UCR _ ENT 00090760, UCR _ ENT 00090761), 2 ♂ (UCR _ ENT 00090756, UCR _ ENT 00090777) (FMNH). Bocas del Toro: Almirante, trail to dam on Nigua Creek, 9.29792 ° N 82.4211 ° W, 15 m, 31 Mar 1959, H. S. Dybas, 2 ♂ (UCR _ ENT 00090715, UCR _ ENT 00090717) (FMNH).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC7FF98FF226D00E166FBC5.taxon	etymology	Etymology: Named for the club-shaped vesica after “ clava ” Latin for club (noun in apposition).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC7FF98FF226D00E166FBC5.taxon	diagnosis	Diagnosis: Recognized by the mid-sized body length (1.54 mm), oval forewing outline (Fig. 2), fairly broad C + Sc (black arrow), rc (black star) and rc 1 relatively short and wide, large vein-tracing areoles associated with scc, rc 1, and rc (white arrow) (Fig. 6 A), vesica short and thick with thick, rounded, club-shaped tip, and long, thin, and curved anophoric process (Figs 6 F, 7).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC7FF98FF226D00E166FBC5.taxon	description	Description: Male: Total body length 1.54 mm; length from posterior margin of pronotum to posterior wing margin 1.16 mm; greatest width across pronotum 0.66 mm; greatest width across forewings 0.87 mm. General coloration: mix of pale and darker brown or reddish brown. Vestiture: Dorsum with sparse setae, setae short; setae on posterior margin of forewing long. Structure: Head: Eye almost half as high as head; muscle scars large. Thorax: Anterior pronotal margin concave, posterior margin slightly concave; pits on pronotum medium-sized, evenly distributed; scutellum in dorsal view abruptly narrowed to tip, tip pointed. Forewing: With outline broad anteriorly, oval; membrane contributing about one third of forewing length, posterior distal margin squarish, veintracing areoles large, distributed along scc, rc 1, and rc; C + Sc fairly broad, wider at widest part of wing; distal process of R pronounced; An 1 wing organ on corium weakly inflated, rounded, with median notch, without notch articulating with claval process; rc 1 about 3 times as long as wide, much wider anteriorly; rc about 4 times as long as wide; tc about 3 times as long as wide; part of An 1 and Cu forming posterodistal margin of tc slightly wider than other veins distal margin of rc 1, rc, and tc cells double s-shaped; rc 2 – 3 trapezoidal; cub almost reaching wing margin; dc 1 horse head-shaped. Legs: Caws long. Genitalia (Fig. 7): Pygophore without lobe on right side; vesica with about half loop, not reaching right pygophore margin, thick at midpoint, with blunt apex, close to apex rounded; anophoric ridge with process, process long, with single lobe curved, tip pointed; right paramere curved, tapering, basal process broad; left paramere straight, tapering, basal process narrow. Female: As in male and generic description. Collecting method and habitat: Collected using Berlese extraction, Winkler extraction in fibrous center of decaying palm log.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC6FF9AFF226F54E15EFEB2.taxon	description	(Figs 2, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC6FF9AFF226F54E15EFEB2.taxon	materials_examined	Holotype. Female. BRAZIL: Mato Grosso: Jacare Alto Xingu, 10.54341 ° S 53.75637 ° W, 330 m, 01 Nov 1965, Alvarenga & Werner, 1 ♀ (UCR _ ENT 00090847) (FMNH).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC6FF9AFF226F54E15EFEB2.taxon	etymology	Etymology: Named for the characteristic wing venation after “ dispar ” Latin for different.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC6FF9AFF226F54E15EFEB2.taxon	diagnosis	Diagnosis: Recognized by the very small size (total body length 0.66 mm), forewing with distinct honey-comb pattern (black arrow), absence of An 1 / 2 claval process, expanded scc divided by sinuous, transverse vein (white arrow), length and shape of rc 1 (black star), distinctly surpassing apex of rc and tc, rectangular rc 2 – 3 (white star), and slender and elongate cub (grey star) and tc (Fig. 2).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC6FF9AFF226F54E15EFEB2.taxon	description	Description: Female: Total body length 0.66 mm; length from posterior margin of pronotum to posterior wing margin 0.51 mm; greatest width across pronotum 0.28 mm; greatest width across forewings 0.42 mm. General coloration: mix of pale and darker brown or reddish brown. Vestiture: Dorsum with sparse, short setae; setae on posterior margin of forewing long. Structure: Head: Eye almost half as high as head; muscle scars small. Thorax: Anterior pronotal margin concave, posterior margin almost straight; pits on pronotum large, evenly distributed; scutellum in dorsal view gradually narrowed to tip, tip pointed. Forewing: with outline broad anteriorly, elongate oval; with distinct honey-comb pattern; membrane contributing less than one third of forewing length, posterior distal margin rounded, vein-tracing areoles on proximal part of wing absent; C + Sc narrow, wider at widest part of wing; distal process of R weakly developed; An 1 / 2 process on clavus absent; An 1 wing organ absent (as in all females); rc 1 about 4 times as long as wide, much wider anteriorly; rc about 4 times as long as wide; tc about 3 times as long as wide; part of An 1 and Cu forming posterodistal margin of tc of similar width as other veins; distal margin of rc 1, rc, and tc cells angular s-shaped; rc 2 - 3 rectangular; cub removed from wing margin; dc 1 with basal portion relatively wide and short. Legs: Claws short. Genitalia: As in generic description. Male: Unknown. Collecting method and habitat: Unknown.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC6FF9AFF226F54E15EFEB2.taxon	discussion	Discussion: This species is set apart from all other species of Guapinannus both in terms of morphology (size, forewing sculpture, wing venation) and distribution (the only documented species of Guapinannus from Brazil). We decided to include this species into Guapinannus and to somewhat expand the concept of this genus, because G. dispar, sp. n., is more similar to species of “ core ” Guapinannus than to any other species of Schizopteridae we have seen from the Neotropical region. Once male specimens become available for this species, our hypothesis should be further tested.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC4FF9AFF226A81E460F92D.taxon	description	(Figs 2, 7, 8, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC4FF9AFF226A81E460F92D.taxon	materials_examined	Holotype. Male. COSTA RICA: Heredia: 6 km ENE Vara Blanca, 10.18333 ° N 84.11667 ° W, 2000 m, 16 Mar 2002, INBio-OET-ALAS, 1 ♂ (UCR _ ENT 00014616) (INBIO). Paratypes: COSTA RICA: Alajuela: Upala Co.: Bijagua, San Miguel, Finca Inti-Aura, 10.75778 ° N 85.00939 ° W, 300 m, 16 Apr 2011 – 20 May 2011, I. Chacon, 1 ♂ (UCR _ ENT 00014608) (INBIO). Heredia: Sarapiqui Co.: P. N. B. Carrillo 5 km. E Vara Blanca, 10.17226 ° N 84.11213 ° W, 2100 m, 20 Apr 2002, INBio-OET-ALAS, 1 ♀ (UCR _ ENT 00014611) (INBIO). P. N. B. Carrillo 6 km. ENE Vera Blanca, 10.15 ° N 83.91667 ° W, 2000 m, 17 Feb 2002 – 21 Feb 2002, INBio-OET-ALAS, 1 ♀ (UCR _ ENT 00014612) (INBIO); 10 Mar 2002, INBio-OET-ALAS, 1 ♀ (UCR _ ENT 00014610), 1 ♂ (UCR _ ENT 00014613) (INBIO). 6 km ENE Vara Blanca, 10.18333 ° N 84.11667 ° W, 2000 m, 16 Mar 2002, INBio-OET-ALAS, 1 ♀ (UCR _ ENT 00014614) (INBIO); 22 Mar 2002, INBio-OET-ALAS, 4 ♀ (UCR _ ENT 00014618 - UCR _ ENT 00014621) (INBIO).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC4FF9AFF226A81E460F92D.taxon	etymology	Etymology: Named for sickle shaped vesica after “ flax ” Latin for sickle.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC4FF9AFF226A81E460F92D.taxon	diagnosis	Diagnosis: Recognized by the mid-sized body length (1.56 mm), dense vestiture, part of An 1 and Cu forming posterodistal margin of tc more pronounced than other veins (black arrow) (Fig. 2), long, relatively slender vesica, with s-shaped bent proximal of acute tip, and short, curved anophoric process with pointed tip (black arrow) (Fig. 7). Most similar to G. bierigi, sp. n., that is also known from Costa Rica, but distinguished by the much smaller distance of cub from the wing margin compared to G. bierigi, sp. n.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC4FF9AFF226A81E460F92D.taxon	description	Description: Male: Total body length 1.56 mm; length from posterior margin of pronotum to posterior wing margin 1.27 mm; greatest width across pronotum 0.65 mm; greatest width across forewings 0.87 mm. General coloration: mix of pale and darker brown or reddish brown. Vestiture: Dorsum with dense, long setae; setae on posterior margin of forewing long. Structure: Head: Eye about one third as high as head; muscle scars not observed. Thorax: Anterior pronotal margin concave, posterior margin slightly concave; pits on pronotum large, evenly distributed; scutellum in dorsal view abruptly narrowed to tip, tip pointed. Forewing: With outline broad anteriorly, elongate oval; membrane contributing about one third of forewing length, posterior distal margin squarish, vein-tracing areoles on proximal part of wing absent; C + Sc uniformly broad; distal process of R pronounced; An 1 wing organ on corium inflated, elongate, with median notch; without notch articulating with claval process; rc 1 about 3 times as long as wide, slightly wider anteriorly; rc about 5 times as long as wide; tc more than 3 times as long as wide; part of An 1 and Cu forming posterodistal margin of tc more pronounced than other veins; distal margin of rc 1, rc, and tc cells double s-shaped; rc 2 - 3 almost triangular; cub almost reaching wing margin; dc 1 with basal portion relatively wide and short. Legs: Claws long. Genitalia (Fig. 7): Pygophore without lobe on right side; vesica with between half and one loop, surpassing right pygophore margin, relatively slender at midpoint, tapering to acute tip, close to apex slightly s-shaped; anophoric ridge with process, process short, with single lobe curved, tip pointed; right paramere curved, broad throughout, basal process broad; left paramere straight, tapering, basal process broad. Female: As in male and generic description. Collecting method and habitat: Collected using Malaise trap in unknown habitat.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC4FF9DFF226C7BE0D4FBAB.taxon	description	(Figs 3, 7, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC4FF9DFF226C7BE0D4FBAB.taxon	materials_examined	Holotype. Male. NICARAGUA: Region Autonoma del Atlantico Sur: 13 km WNW Rama, 12.19472 ° N 84.33667 ° W, 190 m, 18 Apr 2011, J. T. Longino, 1 ♂ (UCR _ ENT 00102045) (FMNH). Paratypes: NICARAGUA: Region Autonoma del Atlantico Sur: 13 km WNW Rama, 12.19472 ° N 84.33667 ° W, 190 m, 18 Apr 2011, J. T. Longino, 3 ♀ (UCR _ ENT 00102046 - UCR _ ENT 00102048), 1 ♂ (UCR _ ENT 00101103) associated with Carduus occidentalis Nutt. (Asteraceae), 1 ♂ (UCR _ ENT 00102049) (FMNH). Rivas: Matagalpa: RN Cerro Musún, 12.95944 ° N 85.225 ° W, 800 m, 01 May 2011, collector unknown, 1 ♂ (UCR _ ENT 00101361) (FMNH).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC4FF9DFF226C7BE0D4FBAB.taxon	etymology	Etymology: Named after our dear mentor, colleague, and friend, the grande dame of South American heteropteran systematics, Jocelia Grazia.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC4FF9DFF226C7BE0D4FBAB.taxon	diagnosis	Diagnosis: Recognized by the small body length (1.16 mm), very broad C + Sc margin (white arrow) (Fig. 3), slightly s-shaped vesica, and short, curved anophoric process (black arrow) (Fig. 7).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC4FF9DFF226C7BE0D4FBAB.taxon	description	Description: Male: Total body length 1.16 mm; length from posterior margin of pronotum to posterior wing margin 0.89 mm; greatest width across pronotum 0.56 mm; greatest width across forewings 0.69 mm. General coloration: mix of pale and darker brown or reddish brown. Vestiture: Dorsum with sparse setae, setae short; setae on posterior margin of forewing long. Structure: Head: Eye almost half as high as head; muscle scars large. Thorax: Anterior pronotal margin concave, posterior margin slightly concave; pits on pronotum medium-sized, evenly distributed; scutellum in dorsal view abruptly narrowed to tip, tip pointed. Forewing: With outline broad anteriorly; membrane contributing about one third of forewing length, posterior distal margin squarish, veintracing areoles on proximal part of wing absent; C + Sc very broad, wider at widest part of wing; distal process of R pronounced; An 1 wing organ on corium strongly inflated, elongate, with median notch, without notch articulating with claval process; rc 1 about 3 times as long as wide, slightly wider anteriorly; rc about 4 times as long as wide; tc about 3 times as long as wide; part of An 1 and Cu forming posterodistal margin of tc of similar width as other veins; distal margin of rc 1, rc, and tc cells s-shaped; rc 2 – 3 trapezoidal; cub almost reaching wing margin; dc 1 with basal portion relatively wide and short. Legs: Claws long. Genitalia (Fig. 7): Pygophore without lobe on right side; vesica with between half and one loop, not reaching right pygophore margin, thick at midpoint, tapering to acute tip, close to apex slightly s-shaped; anophoric ridge with process, process short, with single lobe curved, tip pointed; right paramere curved, tapering, basal process narrow; left paramere straight, tapering, basal process narrow. Female: As in male and generic description. Collecting method and habitat: Collected using Winkler extraction in second growth forest and tropical wet forest.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC3FF9CFF226FF8E2B2FF66.taxon	description	(Figs 3, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC3FF9CFF226FF8E2B2FF66.taxon	materials_examined	Holotype. Female. COSTA RICA: Heredia: Sarapiqui Co.: P. N. Braulio Carrillo, 16 Km SSE La Virgen, 10.26784 ° N 84.084 ° W, 1050 m, 20 Feb 2001 – 23 Feb 2001, INBio-OET-ALAS, 1 ♀ (UCR _ ENT 00014609) (INBIO).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC3FF9CFF226FF8E2B2FF66.taxon	etymology	Etymology: Named for the small body size after “ minutus ” Latin for small.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC3FF9CFF226FF8E2B2FF66.taxon	diagnosis	Diagnosis: Recognized by the small size (1.14 mm), inflated tip of scutellum (white arrow), well-defined proximal part of Cu (black arrow), and dark coloration (female-based) (Fig. 3).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC3FF9CFF226FF8E2B2FF66.taxon	description	Description: Female: Total body length 1.14 mm; length from posterior margin of pronotum to posterior wing margin 0.90 mm; greatest width across pronotum 0.52 mm; greatest width across forewings 0.71 mm. General coloration: mix of pale and darker brown or reddish brown. Vestiture: Dorsum with dense setae, setae short; setae on posterior margin of forewing long. Structure: Head: Eye almost half as high as head; muscle scars large. Thorax: Anterior pronotal margin concave, posterior margin slightly concave; pits on pronotum medium-sized, evenly distributed; scutellum in dorsal view abruptly narrowed to tip, tip inflated. Forewing: With outline elongate oval; membrane contributing about one third of forewing length, posterior distal margin rounded, vein-tracing areoles on proximal part of wing absent; C + Sc uniformly fairly broad; distal process of R pronounced; An 1 wing organ on corium absent (as in all females); rc 1 about 3 times as long as wide, slightly wider anteriorly; rc about 4 times as long as wide; tc about 3 times as long as wide; part of An 1 and Cu forming posterodistal margin of tc of similar width as other veins; distal margin of rc 1, rc, and tc cells double s-shaped; rc 2 - 3 almost triangular; cub almost reaching wing margin; dc 1 with basal portion fairly narrow and elongate. Legs: Claws long. Genitalia: As in generic description. Male: Unknown. Collecting method and habitat: Unknown.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC2FF9CFF226A35E431FAAE.taxon	description	(Figs 3, 7, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC2FF9CFF226A35E431FAAE.taxon	materials_examined	Holotype. Male. PANAMA: Bocas del Toro: Bocas del Toro: Almirante, trail to dam on Nigua Creek, 9.29792 ° N 82.4211 ° W, 32 m, 31 Mar 1959, H. S. Dybas, 1 ♂ (UCR _ ENT 00090718) (FMNH). Paratypes: PANAMA: Bocas del Toro: Bocas del Toro: Almirante, trail to dam on Nigua Creek, 9.29792 ° N 82.4211 ° W, 12 m, 31 Mar 1959, H. S. Dybas, 1 ♂ (UCR _ ENT 00097813), 4 ♀ (UCR _ ENT 00097814 - UCR _ ENT 00097817), 2 nymphs (UCR _ ENT 00097818, UCR _ ENT 00097819) (FMNH). Chiriqui: Near Nueva California W. of Finca Palo Santo, 8.5 ° N 82.41667 ° W, 1524 m, 10 Mar 1959, H. S. Dybas, 1 ♂ (UCR _ ENT 00090796) (FMNH).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC2FF9CFF226A35E431FAAE.taxon	diagnosis	Diagnosis: Recognized by the small mid-sized body length (1.26 mm), elongate body shape (Fig. 3), one-loop, relatively thick vesica with pointed tip, and short, slender, pointed anophoric process (black arrow) (Fig. 7).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC2FF9CFF226A35E431FAAE.taxon	description	Description: Male: Total body length 1.26 mm; length from posterior margin of pronotum to posterior wing margin 0.9 mm; greatest width across pronotum 0.53 mm; greatest width across forewings 0.68 mm. General coloration: uniformly pale. Vestiture: Dorsum with sparse setae, setae short; setae on posterior margin of forewing long. Structure: Head: Eye about one third as high as head; muscle scars large. Thorax: Anterior pronotal margin concave, posterior margin slightly concave; pits on pronotum medium-sized, evenly distributed; scutellum in dorsal view abruptly narrowed to tip, tip rounded. Forewing: With outline elongate oval; membrane contributing about one third of forewing length, posterior distal margin squarish, vein-tracing areoles on proximal part of wing absent; C + Sc uniformly fairly broad; distal process of R pronounced; An 1 wing organ on corium inflated, elongate, with median notch; without notch articulating with claval process; rc 1 about 3 times as long as wide, slightly wider anteriorly; rc about 4 times as long as wide; tc about 3 times as long as wide; part of An 1 and Cu forming posterodistal margin of tc slightly wider than other veins; distal margin of rc 1, rc, and tc cells double s-shaped; rc 2 - 3 trapezoidal; cub almost reaching wing margin; dc 1 horse head-shaped. Legs: Claws long. Genitalia (Fig. 7): Pygophore without lobe on right side; vesica with between half and one loop, not reaching right pygophore margin, thick at midpoint, tapering to acute tip, close to apex rounded; anophoric ridge with process, process short, slender, with single lobe curved, tip pointed; right paramere curved, tapering, basal process broad; left paramere straight, tapering, basal process narrow. Female: As in male and generic description. Collecting method and microhabitat: Collected using Berlese and Winkler extraction from the cortex of a hollowed palm stump and from floor litter at base of a log and cut stump.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC2FF9CFF226A35E431FAAE.taxon	etymology	Etymology: Named for the rounded vesica after “ orbiculatus ” Latin for round.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC2FF9FFF226EFDE34AFD27.taxon	description	(Figs 3, 7, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC2FF9FFF226EFDE34AFD27.taxon	materials_examined	Holotype. Male. PERU: Amazonas: Pacaya – Samiria National Reserve, 5.2012 ° S 74.59319 ° W, 280 m, 07 Nov 1980, CI. Vaucher, 1 ♂ (UCR _ ENT 00089590) (MHNG). Paratypes: PERU: Amazonas: Pacaya – Samiria National Reserve, 5.2012 ° S 74.59319 ° W, 280 m, 07 Nov 1980, CI. Vaucher, 4 ♂ (UCR _ ENT 00089589, UCR _ ENT 00089591 - UCR _ ENT 00089593), 5 ♀ (UCR _ ENT 00089594 - UCR _ ENT 00089598) (MHNG). Unknown locality in Peru, no date provided, collector unknown, 1 ♀ (UCR _ ENT 00089582) (MHNG).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC2FF9FFF226EFDE34AFD27.taxon	etymology	Etymology: Named for the multiple lobes on the anophoric process after “ pluri ” and “ lobus ” Latin for multiple lobes.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC2FF9FFF226EFDE34AFD27.taxon	diagnosis	Diagnosis: Recognized by the small body length (1.14 mm), relatively short rc 1, rc, and tc cells (black star) (Fig. 3), slender vesica with one loop, and multilobed anophoric process, with combination of blunt and acute tips (black arrow) (Fig. 7).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC2FF9FFF226EFDE34AFD27.taxon	description	Description: Male: Total body length 1.14 mm; length from posterior margin of pronotum to posterior wing margin 0.8 mm; greatest width across pronotum 0.44 mm; greatest width across forewings 0.67 mm. General coloration: uniformly pale. Vestiture: Dorsum with sparse setae, setae short; setae on posterior margin of forewing long. Structure: Head: Eye almost half as high as head; muscle scars large. Thorax: Anterior pronotal margin concave, posterior margin slightly concave; pits on pronotum medium-sized, evenly distributed; scutellum in dorsal view abruptly narrowed to tip, tip rounded. Forewing: With outline broad anteriorly; membrane contributing about one third of forewing length, posterior distal margin squarish, vein-tracing areoles on proximal part of wing absent; C + Sc broad, wider at widest part of wing; distal process of R pronounced; An 1 wing organ on corium weakly inflated, elongate, with median notch; without notch articulating with claval process; rc 1 about 3 times as long as wide, slightly wider anteriorly; rc about 4 times as long as wide; tc about 3 times as long as wide; part of An 1 and Cu forming posterodistal margin of tc slightly wider than other veins; distal margin of rc 1, rc, and tc cells double s-shaped; rc 2 - 3 almost triangular; cub almost reaching wing margin; dc 1 with basal portion fairly narrow and elongate. Legs: Claws long. Genitalia (Fig. 7): Pygophore without lobe on right side; vesica with one loop, not reaching right pygophore margin, relatively slender at midpoint, tapering to blunt tip, close to apex rounded; anophoric ridge with process, process long, with multiple lobes, tip blunt; right paramere curved, broad throughout, basal process broad; left paramere straight, tapering, basal process narrow. Female: As in male and generic description. Collecting method: Collected using Winkler extraction.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC1FF9FFF226874E31DF89B.taxon	description	(Figs 3, 7, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC1FF9FFF226874E31DF89B.taxon	materials_examined	Holotype. Male. COLOMBIA: Cauca: PNN Gorgona, Antigua Laguna, 2.96667 ° N 78.18333 ° W, 70 m, 18 Dec 2000 – 03 Jan 2001, H. Torres, 1 ♂ (UCR _ ENT 00109003) (UCR). Paratype: COLOMBIA: Cauca: PNN Gorgona, Alto el Mirador, 2.96666 ° N 78.18333 ° W, 180 m, 06 Oct 2000 – 23 Oct 2000, R. Duque, 1 ♀ (UCR _ ENT 00115594) (UCR).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC1FF9FFF226874E31DF89B.taxon	etymology	Etymology: Named for the thumb-shaped basal process of the right paramere after “ policis ” Latin for thumb.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC1FF9FFF226874E31DF89B.taxon	diagnosis	Diagnosis: Recognized by the small body length (1.03 mm), elongate oval forewing outline, short and wide rc, rc 1, and tc (black star) (Fig. 3), slender vesica with acute tip, and thumb-shaped basal process of the right paramere (black arrow) (Fig. 7).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFC1FF9FFF226874E31DF89B.taxon	description	Description: Male: Total body length 1.03 mm; length from posterior margin of pronotum to posterior wing margin ambiguous 0.7 mm; greatest width across pronotum 0.47 mm; greatest width across forewings 0.63 mm. General coloration: mix of pale and darker brown or reddish brown. Vestiture: Dorsum with sparse setae, setae short; setae on posterior margin of forewing long. Structure: Head: Eye almost half as high as head; muscle scars small. Thorax: Anterior pronotal margin concave, posterior margin slightly concave; pits on pronotum large, evenly distributed; scutellum in dorsal view gradually narrowed to tip, tip pointed. Forewing: With outline broad anteriorly, elongate oval; membrane contributing about one third of forewing length, posterior distal margin squarish, veintracing areoles on proximal part of wing absent; C + Sc broad, wider at widest part of wing; distal process of R pronounced; An 1 wing organ on corium inflated, elongate, with median notch; without notch articulating with claval process; rc 1 about 3 times as long as wide, much wider anteriorly; rc about 3 times as long as wide; tc about 3 times as long as wide; part of An 1 and Cu forming posterodistal margin of tc of similar width as other veins; distal margin of rc 1, rc, and tc cells double s-shaped; rc 2 – 3 trapezoidal; cub almost reaching wing margin; dc 1 horse headshaped. Legs: Claws long. Genitalia (Fig. 7): Pygophore without lobe on right side; vesica with between half and one loop, not reaching right pygophore margin, relatively slender at midpoint, tapering to acute tip, close to apex rounded; anophoric ridge with process, process short, with single lobe curved, tip pointed; right paramere curved, broad throughout, basal process thumb-shaped; left paramere straight, tapering, basal process narrow. Female: As in male and generic description. Collecting method: Collected using a Malaise Trap.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDFFF81FF226BA0E350FB8C.taxon	description	(Figs 3, 7, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDFFF81FF226BA0E350FB8C.taxon	materials_examined	Holotype. Male. MEXICO: Chiapas: RSV A El Triunfo, Est. El Triunfo, 16.20286 ° N 91.86044 ° W, 1950 m, 08 Aug 1996 – 11 Aug 1996, A. R. Gillogly, 1 ♂ (UCR _ ENT 00094282) (UCR).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDFFF81FF226BA0E350FB8C.taxon	etymology	Etymology: Named for the extremely thick vesica after “ robustus ” Latin for robust.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDFFF81FF226BA0E350FB8C.taxon	diagnosis	Diagnosis: Recognized by the mid-sized body length (1.49 mm), small vein-tracing areoles distributed along scc, rc and rc 1 (black arrow) (Fig. 3), slight extensions of the right pygophore margin (black star), and very thick vesica with blunt tip (Fig. 7).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDFFF81FF226BA0E350FB8C.taxon	description	Description: Male: Total body length 1.49 mm; length from posterior margin of pronotum to posterior wing margin 1.1 mm; greatest width across pronotum 0.28 mm. General coloration: mix of pale and darker brown or reddish brown. Vestiture: Dorsum with dense setae, setae long; setae on posterior margin of forewing extremely long. Structure: Head: Eye almost half as high as head; muscle scars large. Thorax: Anterior pronotal margin concave, posterior margin slightly concave; pits on pronotum medium-sized, evenly distributed; scutellum in dorsal view gradually narrowed to tip, tip rounded. Forewing: With outline elongate oval; membrane contributing about one third of forewing length, posterior distal margin rounded, vein-tracing areoles small, distributed along scc, rc and rc 1; C + Sc broad, wider at widest part of wing; distal process of R pronounced; An 1 wing organ on corium strongly inflated, triangular, with median notch, without notch articulating with claval process; rc 1 about 3 times as long as wide, slightly wider anteriorly; rc about 5 times as long as wide; tc about 3 times as long as wide; part of An 1 and Cu forming posterodistal margin of tc of similar width as other veins; distal margin of rc 1, rc, and tc cells double s-shaped; rc 2 - 3 trapezoidal; cub almost reaching wing margin; dc 1 with basal portion fairly narrow and elongate. Legs: Claws long. Genitalia (Fig. 7): Pygophore with slightly extended right margin; vesica with about half loop, not reaching right pygophore margin, very thick at midpoint, tapering to blunt tip, close to apex slightly s-shaped; anophoric ridge with process, process short, with single lobe relatively straight, tip pointed; right paramere curved, broad throughout, basal process broad; left paramere straight, tapering, basal process narrow. Female: Unknown. Collecting method: Collected using a flight intercept trap.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDFFF80FF226F9CE3D8FE5F.taxon	description	(Figs 3, 7, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDFFF80FF226F9CE3D8FE5F.taxon	materials_examined	Holotype. Male. MEXICO: Chiapas: Palenque, 15.63333 ° N 92.65 ° W, 1640 m, 02 Jul 1983 – 30 Jul 1983, S. B. Peck, J. Kukalova-Peck, R. S. Anderson, 1 ♂ (UCR _ ENT 00102050) (FMNH). Paratypes: MEXICO: Chiapas: El Bosque (6.6 mi SW), 17.01667 ° N 92.78333 ° W, 1463 m, 29 Aug 1973, A. F. Newton, 7 ♀ (UCR _ ENT 00101241, UCR _ ENT 00102051 - UCR _ ENT 00102053) (FMNH). Playón de la Gloria, 16.16028 ° N 90.90139 ° W, 160 m, 26 Jun 2008, collector unknown, 1 ♂ (UCR _ ENT 00120069) (MTEC).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDFFF80FF226F9CE3D8FE5F.taxon	etymology	Etymology: Named for the s-shaped vesica after “ sinuosus ” Latin for bending.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDFFF80FF226F9CE3D8FE5F.taxon	diagnosis	Diagnosis: Recognized by the mid-sized body length (1.49 mm), C + Sc margin broad anteriorly and more slender posteriorly, distinctly double s-shaped margin of rc 1, rc, tc cells (black arrow) (Fig. 3), and relatively tick and s-shaped vesica (Fig. 7).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDFFF80FF226F9CE3D8FE5F.taxon	description	Description: Male: Total body length 1.49 mm; length from posterior margin of pronotum to posterior wing margin 0.9 mm; greatest width across pronotum 0.54 mm. General coloration: uniformly pale. Vestiture: Dorsum with sparse setae, setae short; setae on posterior margin of forewing long. Structure: Head: Eye almost half as high as head; muscle scars large. Thorax: Anterior pronotal margin concave, posterior margin slightly concave; pits on pronotum small, evenly distributed; scutellum in dorsal view gradually narrowed to tip, tip pointed. Forewing: With outline elongate oval; membrane contributing about one third of forewing length, posterior distal margin rounded, vein-tracing areoles on proximal part of wing absent; C + Sc uniformly broad; distal process of R weakly developed; An 1 wing organ on corium strongly inflated, without notch articulating with claval process, rounded, without median notch; rc 1 about 3 times as long as wide, slightly wider anteriorly; rc about 5 times as long as wide; tc about 3 times as long as wide; distal margin of rc 1, rc, and tc cells double s-shaped; rc 2 – 3 almost triangular; cub almost reaching wing margin; dc 1 with basal portion relatively wide and short. Legs: Claws long. Genitalia (Fig. 7): Pygophore without lobe on right side; vesica with between half and one loop, not reaching right pygophore margin, thick at midpoint, tapering to acute tip, close to tip slightly s-shaped; anophoric ridge with process, process short, with single lobe curved, tip pointed; right paramere curved, tapering, basal process broad; left paramere straight, tapering, basal process narrow. Female: As in male and generic description. Collecting method and habitat: Collected using Berlese extraction, flight intercept trap, and sifting leaf litter in closed forest, cloud forest with pines, mature wet forest, and rainforest.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDEFF80FF226AECE2B2F9FB.taxon	description	(Figs 4, 7, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDEFF80FF226AECE2B2F9FB.taxon	materials_examined	Holotype. Male. HONDURAS: Francisco Morazan: Tatumbia Montaña del Aguacate, 14 ° N 87.08333 ° W, 1425 m, 14 Mar 1996, R. Cave, 1 ♂ (UCR _ ENT 00078556) (EAPZ). Paratypes: HONDURAS: Francisco Morazan: Tatumbia Montaña del Aguacate, 14 ° N 87.08333 ° W, 1425 m, 14 Mar 1996, R. Cave, 5 ♂ (UCR _ ENT 00078555, UCR _ ENT 00078557 - UCR _ ENT 00078560), 5 ♀ (UCR _ ENT 00078561 - UCR _ ENT 00078565) (EAPZ). Olancho: La Union Parq Nac La Maralla, 15.07949 ° N 86.76175 ° W, 1060 m, 14 Sep 1994, R. Cordero, 1 ♀ (UCR _ ENT 00078566) (EAPZ).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDEFF80FF226AECE2B2F9FB.taxon	etymology	Etymology: Named for the type locality “ Tatumbia Montana ” in Honduras (noun in apposition).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDEFF80FF226AECE2B2F9FB.taxon	diagnosis	Diagnosis: Recognized by the mid-sized body length (1.51 mm), elongate oval outline of forewings, inflated tip of scutellum (black arrow) (Fig. 4), vesica relatively thick and straight with acute tip, not reaching pygophore margin, and without anophoric process (Fig. 7).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDEFF80FF226AECE2B2F9FB.taxon	description	Description: Male: Total body length 1.51 mm; length from posterior margin of pronotum to posterior wing margin 1.27 mm; greatest width across pronotum 0.62 mm; greatest width across forewings 0.88 mm. General coloration: mix of pale and darker brown or reddish brown. Vestiture: Dorsum with dense setae, setae long; setae on posterior margin of forewing extremely long. Structure: Head: Eye about one third as high as head; muscle scars large. Thorax: Anterior pronotal margin concave, posterior margin slightly concave; pits on pronotum small, evenly distributed; scutellum in dorsal view abruptly narrowed to tip, tip rounded. Forewing: With outline broad anteriorly; membrane contributing about one third of forewing length, posterior distal margin squarish, vein-tracing areoles on proximal part of wing absent; C + Sc broad, wider at widest part of wing; distal process of R pronounced; An 1 wing organ on corium inflated, elongate, with median notch, without notch articulating with claval process; rc 1 about 3 times as long as wide, much wider anteriorly; rc about 4 times as long as wide; tc about 3 times as long as wide; part of An 1 and Cu forming posterodistal margin of tc of similar width as other veins; distal margin of rc 1, rc, and tc cells double s-shaped; rc 2 – 3 trapezoidal; cub almost reaching wing margin; dc 1 with basal portion relatively wide and short. Legs: Claws long. Genitalia (Fig. 7): Pygophore without lobe on right side; vesica with between half and one loop, not reaching right pygophore margin, thick at midpoint, tapering to acute tip, close to apex fairly straight; anophoric ridge without process; right paramere curved, broad throughout, basal process broad; left paramere straight, tapering, basal process narrow. Female: As in male and generic description. Collecting method and habitat: Unknown.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDEFF82FF226D48E2B2FE07.taxon	description	(Figs 4, 7, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDEFF82FF226D48E2B2FE07.taxon	materials_examined	Holotype. Male. COSTA RICA: Unknown locality, no date provided, collector unknown, 1 ♂ (UCR _ ENT 00014617) (INBIO).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDEFF82FF226D48E2B2FE07.taxon	etymology	Etymology: Named for the slender vesica after “ tenuis ” Latin for slender.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDEFF82FF226D48E2B2FE07.taxon	diagnosis	Diagnosis: Recognized by the large body length (2.06 mm), narrow C + Sc (black arrow), distal margin of rc 1, rc, and tc cells distinctly double s-shaped (white arrow) (Fig. 4), relatively long s-shaped vesica, and absence of anophoric process (Fig. 7).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDEFF82FF226D48E2B2FE07.taxon	description	Description: Male: Total body length 2.06 mm; length from posterior margin of pronotum to posterior wing margin 1.49 mm; greatest width across pronotum 0.69 mm; wing missing. General coloration: uniformly pale. Vestiture: Dorsum with sparse setae, setae short; setae on posterior margin of forewing long. Structure: Head: Eye about one third as high as head; muscle scars large. Thorax: Anterior pronotal margin straight, posterior margin slightly concave; pits on pronotum large, in distinct clusters; scutellum in dorsal view abruptly narrowed to tip, tip rounded. Forewing: With outline elongate oval; membrane contributing about one third of forewing length, posterior distal margin rounded, vein-tracing areoles on proximal part of wing absent; C + Sc uniformly narrow; distal process of R weakly developed; An 1 wing organ on corium weakly developed; rc 1 about 3 times as long as wide, much wider anteriorly; rc about 5 times as long as wide; tc about 3 times as long as wide; part of An 1 and Cu forming posterodistal margin of tc of similar width as other veins; distal margin of rc 1, rc, and tc cells s-shaped; rc 2 - 3 almost triangular; cub removed from wing margin; dc 1 with basal portion relatively wide and short. Legs: Claws long. Genitalia (Fig. 7): Pygophore without lobe on right side; vesica with between half and one loop, not reaching right pygophore margin, thick at midpoint, tapering to acute tip, close to apex slightly s-shaped; anophoric ridge with process, process short, with single lobe curved, tip pointed; right paramere straight, tapering, basal process broad; left paramere straight, tapering, basal process broad. Female: Unknown. Collecting method and habitat: Unknown.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDCFF82FF226914E4A5F987.taxon	description	(Figs 4, 7, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDCFF82FF226914E4A5F987.taxon	materials_examined	Holotype. Male. ECUADOR: Pastaza: 25 km N Puyo, 1.2855 ° S 78.02291 ° W, 1000 m, 13 Jul 1976, S. B. Peck, 1 ♂ (UCR _ ENT 00090642) (FMNH). Paratypes: ECUADOR: Pastaza: Pastaza, 1.43583 ° S 78.00194 ° W, 972 m, 13 Jul 1976, S. B. Peck, 4 ♂ (UCR _ ENT 00090643 - UCR _ ENT 00090646) (FMNH).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDCFF82FF226914E4A5F987.taxon	etymology	Etymology: Named for the long ridge-like process on the anophoric after “ tergus ” Latin for ridge (noun in apposition).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDCFF82FF226914E4A5F987.taxon	diagnosis	Diagnosis: Recognized by the very small body length (0.67 mm), small vein-tracing areoles along rc and rc 1 (black arrow) (Fig. 4), almost full-loop, circular vesica, and long anophoric process with curved single tip (black arrow) (Fig. 7).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDCFF82FF226914E4A5F987.taxon	description	Description: Male: Total body length 0.67 mm; length from posterior margin of pronotum to posterior wing margin 0.52 mm; greatest width across pronotum 0.28 mm; greatest width across forewings 0.42 mm. General coloration: mix of pale and darker brown or reddish brown. Vestiture: Dorsum with sparse setae, setae short; setae on posterior margin of forewing long. Structure: Head: Eye almost half as high as head; muscle scars large. Thorax: Anterior pronotal margin concave, posterior margin slightly concave; pits on pronotum medium-sized, evenly distributed; scutellum in dorsal view abruptly narrowed to tip, tip pointed. Forewing: With outline broad anteriorly; membrane contributing about one third of forewing length, posterior distal margin squarish, vein-tracing areoles small, distributed along rc and rc 1; C + Sc fairly narrow, wider at widest part of wing; distal process of R pronounced; An 1 wing organ on corium strongly inflated, rounded, with median notch, without notch articulating with claval process; rc 1 about 3 times as long as wide, much wider anteriorly; rc about 4 times as long as wide; tc about 3 times as long as wide; part of An 1 and Cu forming posterodistal margin of tc of similar width as other veins; distal margin of rc 1, rc, and tc cells double s-shaped; rc 2 – 3 trapezoidal; cub almost reaching wing margin; dc 1 horse head-shaped. Legs: Claws long. Genitalia (Fig. 7): Pygophore without lobe on right side; vesica with between half and one loop, not reaching right pygophore margin, thick at midpoint, tapering to acute tip, close to apex rounded; anophoric ridge with process, process long, with single lobe curved, tip pointed; right paramere curved, broad throughout, basal process narrow; left paramere straight, broad throughout, basal process narrow. Female: Unknown. Collecting method and habitat: Collected using Berlese extraction in cloud forest litter.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDCFF87FF226D94E0D4FE07.taxon	description	(Figs 4, 7, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDCFF87FF226D94E0D4FE07.taxon	materials_examined	Holotype. Male. MEXICO: Veracruz: San Andres Tuxtla Co.: 18.52303 ° N 95.15453 ° W, 1109 m, 05 Jun 2016, Manuel Barrios, 1 ♂ (UCR _ ENT 00128210) (UCR). Paratypes: MEXICO: Veracruz: Road to Ruiz Cortinez, 18.5229 ° N 95.15441 ° W, 1114 m, 05 Jun 2016, R. S. Anderson, 1 ♂ (UCR _ ENT 00125950), 2 ♀ (UCR _ ENT 00125951) (UCR).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDCFF87FF226D94E0D4FE07.taxon	etymology	Etymology: Named for the three lobes on the anophoric process after “ trilobus ” Latin for three lobes.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDCFF87FF226D94E0D4FE07.taxon	diagnosis	Diagnosis: Recognized by the mid-sized body length (1.40 mm), broad C + Sc (black arrow) (Fig. 4), vesica with almost one circular loop with acute tip, and anophoric process with multiple pointed tips (black arrows) (Fig. 7).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFDCFF87FF226D94E0D4FE07.taxon	description	Description: Male: total body length 1.40 mm; length from posterior margin of pronotum to posterior wing margin 1.03 mm; greatest width across pronotum 0.61 mm; greatest width across forewings 0.85 mm. General coloration: mix of pale and darker brown or reddish brown. Vestiture: Dorsum with sparse setae, setae long; setae on posterior margin of forewing extremely long. Structure: Head: Eye almost half as high as head; muscle scars large. Thorax: Anterior pronotal margin concave, posterior margin slightly concave; pits on pronotum large, evenly distributed; scutellum in dorsal view abruptly narrowed to tip, tip pointed. Forewing: With outline broad anteriorly; membrane contributing about one third of forewing length, posterior distal margin squarish, vein-tracing areoles on proximal part of wing absent; C + Sc broad, wider at widest part of wing; distal process of R pronounced; An 1 wing organ on corium inflated, elongate, with median notch, without notch articulating with claval process; rc 1 about 3 times as long as wide, much wider anteriorly; rc about 4 times as long as wide; tc about 3 times as long as wide; part of An 1 and Cu forming posterodistal margin of tc slightly wider than other veins; distal margin of rc 1, rc, and tc cells double s-shaped; rc 2 – 3 almost triangular; cub almost reaching wing margin; dc 1 with basal portion fairly narrow and elongate. Legs: Claws long. Genitalia (Fig. 7): Pygophore without lobe on right side; vesica with almost one loop, not reaching right pygophore margin, relatively slender at midpoint, tapering to acute tip, close to apex rounded; anophoric ridge with process, process long, with multiple pointed tips; right paramere curved, broad throughout, basal process broad; left paramere curved and tapering, basal process broad. Female: As in male and generic description. Collecting method and habitat: Collected using leaf litter sifting and Winkler extraction from mixed hardwood forest.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFD9FF87FF226914E3D2F9DF.taxon	description	(Figs 4, 7, 9)	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFD9FF87FF226914E3D2F9DF.taxon	materials_examined	Holotype. Male. BELIZE: Belize, 17.18988 ° N 88.49765 ° W, 55 m, no date provided, collector unknown, 1 ♂ (UCR _ ENT 00090858) (FMNH). Paratypes: BELIZE: Orange Walk: Rio Bravo Cons. Area 3 Well Tran (Near Res. Station), 18.06667 ° N 88.55 ° W, 10 m, 10 Sep 1995, P. W. Kovarik, 1 ♂ (UCR _ ENT 00094283) (UCR).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFD9FF87FF226914E3D2F9DF.taxon	etymology	Etymology: Named for the hook-shaped vesica after “ uncus ” Latin for hook (noun in apposition).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFD9FF87FF226914E3D2F9DF.taxon	diagnosis	Diagnosis: Recognized by the small mid-sized body length (0.67 mm), elongate oval body shape (length to width ratio 3.4) (Fig. 4), and apex of vesica hook-shaped (Fig. 7).	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
E07B87DEFFD9FF87FF226914E3D2F9DF.taxon	description	Description: Male: Total body length 1.19 mm; length from posterior margin of pronotum to posterior wing margin 0.97 mm; greatest width across pronotum 0.27 mm; greatest width across forewings 0.35 mm. General coloration: uniformly pale. Vestiture: Dorsum with sparse setae, setae short; setae on posterior margin of forewing long. Structure: Head: Eye almost half as high as head; muscle scars indistinct. Thorax: Anterior pronotal margin concave, posterior margin slightly concave; pits on pronotum small, evenly distributed; scutellum in dorsal view gradually narrowed to tip, tip pointed. Forewing: With outline elongate oval; membrane contributing about one third of forewing length, posterior distal margin rounded, vein-tracing areoles on proximal part of wing absent; C + Sc broad, wider at widest part of wing; distal process of R pronounced; An 1 wing organ on corium inflated, elongate, with median notch, without notch articulating with claval process; rc 1 about 3 times as long as wide, slightly wider anteriorly; rc about 4 times as long as wide; tc about 3 times as long as wide; part of An 1 and Cu forming posterodistal margin of tc slightly wider than other veins; distal margin of rc 1, rc, and tc cells double s-shaped; rc 2 – 3 trapezoidal; cub almost reaching wing margin; dc 1 with basal portion fairly narrow and elongate. Legs: Claws long. Genitalia (Fig. 7): Pygophore without lobe on right side; vesica with between half and one loop, surpassing right pygophore margin, thick at midpoint, with bottle opener-shaped apex, close to apex rounded; anophoric ridge with process, process short, with single lobe curved, tip pointed; right paramere curved, broad throughout, basal process broad; left paramere straight, tapering, basal process broad. Female: Unknown. Collecting method and habitat: Collected using flight intercept trap.	en	Frankenberg, Sarah, Knyshov, Alexander, Hoey-Chamberlain, Rochelle, Weirauch, Christiane (2021): Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species. Zootaxa 4958 (1): 261-286, DOI: 10.11646/zootaxa.4958.1.14
