taxonID	type	description	language	source
E0318797FF9F155BFDF4FB09FDC3D019.taxon	type_taxon	Type species: Caccothryptus compactus Sharp, 1902, by monotypy. urn: lsid: zoobank. org: act: CB 7 B 4207 - FD 54 - 43 A 0 - 9617 - 7 D 15 CAD 62833	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF9F155BFDF4FB09FDC3D019.taxon	type_taxon	Type species: Macrobyrrhinus rouyeri Pic, 1922, by monotypy. urn: lsid: zoobank. org: act: 49 DFB 03 F-B 836 - 4673 - 8 AF 9 - 850 B 9 E 847115	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF9F155BFDF4FB09FDC3D019.taxon	discussion	Remarks This genus is related to five genera in the Mandersia genus group (sensu Hernando & Ribera 2005 a): Resachus Delève, 1968 (revised by Hernando & Ribera 2006), Simplocarina Pic, 1922, Pseudothryptus Hernando & Ribera, 2005, Mandersia Sharp, 1902 and Euthryptus Sharp, 1902. It is also similar to Cyclolimnichus Delève, 1968 (revised by Hernando & Ribera 2000) and Tricholimnichus Hernando & Ribera, 2001 in their large and elongate body.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF9F155BFDF4FB09FDC3D019.taxon	distribution	Distribution SE Asia (Okinawa to India).	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF9C155FFDFBFE81FCA4D509.taxon	description	urn: lsid: zoobank. org: act: 578 B 475 A- 54 DA- 4600 - 9 FFF-F 5 E 7045 F 814 E Figs 1 A, 2 A – E, 4 A – B	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF9C155FFDFBFE81FCA4D509.taxon	diagnosis	Diagnosis Smaller species in the genus; TL = 3.73 – 4.05 mm in male, 3.75 – 4.20 mm in female; parameres weakly pointed in lateral view; apical emargination of parameres deeply U-shaped, with small projections in inner margins.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF9C155FFDFBFE81FCA4D509.taxon	etymology	Etymology The species is named after the type locality.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF9C155FFDFBFE81FCA4D509.taxon	materials_examined	Material examined Holotype TAIWAN: ♂ (TARI), “ Taiwan: Ilan Fushan ---- [Chinese characters] 13. IV. 2011, leg. C. F. Lee ”. Paratypes TAIWAN: 2 ♂♂, 7 ♀♀, 13 ex. (TARI, EUMJ, NMW), same data as for the holotype; 1 ex. (TARI), “ Taiwan: Nantou Nanshanchi 25. VII. 2008, leg. W. - T. Liu ”; 1 ♂, 1 ex. (SEHU), “ TAIWAN: NANTOU Nanshanchi 900 28. III. 1982 Col. T. FUJISAWA ”; 1 ♂ (EUMJ), “ [TAIWAN] Wulu Alt. ca. 700 m Haiduan Township Taitung Country ---- [Chinese characters] 17 – 18. VI. 2011 J. Yamasako leg. ”; 1 ♀ (EUMJ), “ (TAIWAN) Hsinhsien ~ Wulai Taipei Hsien 2. VII, 1970 Y. Hori leg. ”.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF9C155FFDFBFE81FCA4D509.taxon	description	Description Male Body (Fig. 1 A) oblong, convex dorsally and ventrally, shiny, densely covered with short and long silver setae. Coloration of body black; legs and antennae paler. Head densely punctate, slightly convex dorsally. Antennae relatively long, reaching about proximal 1 / 4 of elytra. Pronotum punctate as in head; PW / PL 1.90 – 2.11 (2.03). Scutellar shield equilateral triangular, finely punctate, lateral margins straight. Elytra oblong, widest at middle, lateral margins gently arcuate, coarsely and irregularly punctate; space between punctures same as their diameter; adpressed long silver setae forming obvious zigzag, irregular markings; apex obtuse; humeral parts weakly projecting dorsally; EL / EW 1.17 – 1.37 (1.31), EL / PL 3.00 – 3.50 (3.26), EW / PW 1.17 – 1.37 (1.23), TL / EW 1.53 – 1.78 (1.72). Each claw on foreleg of same size and shape. Sternite VIII (Fig. 2 D) U-shaped, bearing short setae in apical parts. Sternite IX (Fig. 2 E) elongate, with long and stout lateral projections. Aedeagus (Fig. 2 A – C) stout, straightly curved ventrally in apical part, punctate in apical part of median lobe and parameres; median lobe wide in lateral view, rather pointed at apex; apical emargination of parameres deeply U-shaped, with small projections in inner margins; apices of parameres rounded in ventral view, weakly pointed in lateral view. Female Sexual dimorphism indistinct; PW / PL 1.65 – 2.26 (2.02), EL / EW 1.23 – 1.34 (1.30), EL / PL 2.65 – 3.56 (3.25), EW / PW 1.20 – 1.28 (1.23), TL / EW 1.60 – 1.79 (1.71). Ovipositor (Fig. 4 B) well sclerotized; coxite closely punctuate, pointed at apices, about 1.18 times as long as spiculum ventrale (Fig. 4 A); approximate ratio of coxite and baculus (n = 1) 1.0: 3.43. Measurements Males (n = 5): TL 3.73 – 4.05 (3.90) mm, PW 1.80 – 1.90 (1.86) mm, PL 0.88 – 1.00 (0.92) mm, EL 2.85 – 3.15 (2.98) mm, EW 2.10 – 2.50 (2.28) mm. Females (n = 8): TL 3.75 – 4.20 (4.02) mm, PW 1.80 – 2.03 (1.90) mm, PL 0.85 – 1.15 (0.95) mm, EL 2.90 – 3.20 (3.07) mm, EW 2.20 – 2.50 (2.35) mm.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF9C155FFDFBFE81FCA4D509.taxon	distribution	Distribution Only known from the type locality, Taiwan.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF9C155FFDFBFE81FCA4D509.taxon	discussion	Remarks This species belongs to the testudo species group (sensu Hernando & Ribera 2014). It resembles C. sinensis Hernando & Ribera, 2014, known from Fujian, China, in the shape of the parameres, which are broadly and deeply emarginated, and differs from it in the following male genital characteristics: apex of the median lobe rather rounded (rather pointed in C. sinensis); median lobe straightly projecting posteriorly (curved postero-ventrally in C. sinensis).	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF9B155CFD15FD9DFBD6D4D1.taxon	description	urn: lsid: zoobank. org: act: 42 B 7 D 5 F 0 - 7 DC 5 - 4 BD 5 - 8 CCF- 5 A 08 F 9222 A 54 [Japanese name: Okinawa-oo-chibi-doromushi] Figs 1 B, 2 F – J, 4 C – D, 5 A – B	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF9B155CFD15FD9DFBD6D4D1.taxon	diagnosis	Diagnosis Smaller species in the genus; TL = 3.68 – 3.73 mm in male, 3.82 – 4.00 mm in female; parameres rather pointed in lateral view, with furrows in ventral part; apical emargination of parameres deeply U-shaped.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF9B155CFD15FD9DFBD6D4D1.taxon	etymology	Etymology The species is named after “ Orion Beer ” (copyright: Orion Breweries, Ltd), which is a local, but wellknown and favored beer in Okinawa. The type locality of this species is situated near the Nago Factory of Orion Beer.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF9B155CFD15FD9DFBD6D4D1.taxon	materials_examined	Material examined Holotype JAPAN: 1 ♂ (EUMJ), “ Genkagawa, Nago-shi, Okinawa Japan N 26.616704, E 128.062778, 10 - IV- 2015, Masato MORI leg. ”. Paratypes JAPAN: 1 ♂ (EUMJ), same data as for the holotype; 1 ♀, 1 ex. (EUMJ), same data, but collector K. Kitayama; 1 ♀ (EUMJ), “ Zatsun, Kunigami, Okinawa, Ryukyus 21 - VI- 2006 K. Takahashi leg. ”; 2 ♀♀ (EUMJ), “ Genka-ohkawa Okinawa 22 - VII- 1996 C. F. Lee leg. ”.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF9B155CFD15FD9DFBD6D4D1.taxon	description	Description Male Very similar to C. taiwanus sp. nov. in external features; body a little smaller and slender; elytral punctures coarser; PW / PL 1.88 – 1.94 (1.91), EL / EW 1.39 – 1.42 (1.40), EL / PL 3.14 – 3.33 (3.24), EW / PW 1.16 – 1.25 (1.21), TL / EW 1.83 – 1.84 (1.84). Sternite VIII (Fig. 2 I) U-shaped, bearing short setae in apical parts. Sternite IX (Fig. 2 J) rather wide, with long and stout lateral projections. Aedeagus (Fig. 2 F – H) stout, curved ventrally in apical part, punctate in apical part of median lobe and parameres; median lobe wide in lateral view, pointed at apex; apical emargination of parameres deeply U-shaped; apices of parameres rounded in ventral view; pointed in lateral view. Female Sexual dimorphism indistinct; PW / PL 1.85 – 1.94 (1.91), EL / EW 1.36 – 1.45 (1.42), EL / PL 3.15 – 3.44 (3.33), EW / PW 1.18 – 1.29 (1.23), TL / EW 1.77 – 1.91 (1.84). Ovipositor (Fig. 4 D) well sclerotized; coxite closely punctuate, pointed at apices, about 1.11 times as long as spiculum ventrale (Fig. 4 C); approximate ratio of coxite and baculus (n = 1) as 1.0: 3.45. Measurements Males (n = 2): TL 3.68 – 3.73 (3.71) mm, PW 1.60 – 1.75 (1.68) mm, PL 0.85 – 0.90 (0.88) mm, EL 2.83 mm, EW 2.00 – 2.03 (2.02) mm. Females (n = 4): TL 3.82 – 4.00 (3.92) mm, PW 1.70 – 1.75 (1.73) mm, PL 0.90 – 0.92 (0.91) mm, EL 2.90 – 3.10 (3.01) mm, EW 2.00 – 2.20 (2.13) mm.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF9B155CFD15FD9DFBD6D4D1.taxon	distribution	Distribution Only known from the type locality, Okinawa-jima, the Ryukyus, Japan.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF9B155CFD15FD9DFBD6D4D1.taxon	discussion	Remarks This species belongs to the testudo species group (sensu Hernando & Ribera 2014). It is similar to C. taiwanus sp. nov. in the shape of the male genitalia, but differs from it in the following characteristics: parameres rather pointed in lateral view, with furrows in the ventral part; apex of the median lobe rather pointed in lateral view.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF9B155CFD15FD9DFBD6D4D1.taxon	biology_ecology	Biological notes The type locality (Fig. 5 A – B) is a small river and the specimens were collected from under the surface of water-logged wood (personal communication from Mr. Masato Mori).	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF981553FDFAFBB5FD07D6CE.taxon	description	urn: lsid: zoobank. org: act: 3 CE 4012 A- 96 D 7 - 4568 - 9 DB 7 - F 2175 F 0 B 1278 Figs 1 C, 3 A – D, 4 E – F	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF981553FDFAFBB5FD07D6CE.taxon	diagnosis	Diagnosis Medium size in the genus; TL = 4.65 mm in male, 4.40 – 4.70 mm in female; apical emargination of parameres deeply U-shaped.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF981553FDFAFBB5FD07D6CE.taxon	etymology	Etymology The species is named after the type locality.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF981553FDFAFBB5FD07D6CE.taxon	materials_examined	Material examined Holotype TIBET: ♂ (EUMJ), “ Cha Yu, 1824 m S. E. Tibet 9 - IX- 1996 C. I. Li leg. ”. Paratypes TIBET: 3 ♀♀ (EUMJ, NMW), same data as for the holotype.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF981553FDFAFBB5FD07D6CE.taxon	description	Description Male Body oblong, convex strongly in dorsal and slightly in ventral parts, shiny, densely covered with short silver setae. Coloration of body dark brown, but femora and tarsi paler. Head slightly convex dorsally, densely covered with fine punctures. Antennae relatively long, reaching about proximal 1 / 5 of elytra. Pronotum punctate as in head; PW / PL 2.10. Scutellar shield equilateral triangular, finely punctate, lateral margins straight. Elytra oblong, subparallel-sided near base to apical 1 / 4, gently tapering in apical 1 / 4, densely and regularly punctate; space between punctures smaller than their diameter; adpressed silver setae forming zigzag markings; apex relatively pointed; humeral parts distinctly projecting dorsally; EL / EW 1.40, EL / PL 3.65, EW / PW 1.24, TL / EW 1.79. Each claw on forelegs of same size and shape. Sternite IX (Fig. 3 D) elongate, with long and slender lateral projections. Aedeagus (Fig. 3 A – C; basal part somewhat damaged) stout, curved ventrally in apical part, finely punctate in apical part of median lobe and parameres; median lobe slender in lateral view, pointed at apex; apical emargination of parameres deeply U-shaped; apices of parameres pointed in ventral and lateral views. Female Sexual dimorphism indistinct; PW / PL 1.98 – 2.08 (2.02), EL / EW 1.32 – 1.42 (1.37), EL / PL 3.40 – 3.76 (3.62), EW / PW 1.24 – 1.35 (1.31), TL / EW 1.68 – 1.80 (1.76). Ovipositor (Fig. 4 F) well sclerotized; coxite closely punctuate, pointed at apices, about 1.03 times as long as spiculum ventrally (Fig. 4 E); approximate ratio of coxite and baculus (n = 1) 1.0: 3.41. Measurements Male (n = 1): TL 4.65 mm, PW 2.10 mm, PL 1.00 mm, EL 3.65 mm, EW 2.60 mm. Females (n = 3): TL 4.40 – 4.70 (4.59) mm, PW 1.95 – 2.08 (2.00) mm, PL 0.98 – 1.00 (0.99) mm, EL 3.40 – 3.70 (3.59) mm, EW 2.45 – 2.80 (2.62) mm.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF981553FDFAFBB5FD07D6CE.taxon	distribution	Distribution Only known from the type locality, Tibet.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF981553FDFAFBB5FD07D6CE.taxon	discussion	Remarks This species belongs to the testudo species group (sensu Hernando & Ribera 2014). It is related to C. sinensis, but differs from it in having the median lobe straightly projecting posteriorly, and the emargination of parameres somewhat wider.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF971550FDF7FE57FD3DD508.taxon	description	urn: lsid: zoobank. org: act: 43136 CDF- 3 F 5 C- 4 AD 4 - A 7 DC- 3 F 57299013 EC Figs 1 D, 3 E – I	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF971550FDF7FE57FD3DD508.taxon	description	Figs 1 D, 3 E – I	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF971550FDF7FE57FD3DD508.taxon	diagnosis	Diagnosis Larger species in the genus; TL = 5.90 mm in male; median lobe minutely serrate in ventral part; apical emargination of parameres V-shaped.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF971550FDF7FE57FD3DD508.taxon	etymology	Etymology The species is named after the type locality.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF971550FDF7FE57FD3DD508.taxon	materials_examined	Material examined Holotype TIBET: ♂ (EUMJ), “ Cha Yu, 1824 m S. E. Tibet 9 - IX- 1996 C. I. Li leg. ”.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF971550FDF7FE57FD3DD508.taxon	description	Description Male Body oblong, convex strongly in dorsal and slightly in ventral parts, shiny, densely covered with short silver setae. Coloration of body dark brown, but femora and tarsi paler. Head convex dorsally, densely covered with fine punctures. Antennae relatively long, reaching about proximal 1 / 5 of elytra. Pronotum punctate as in head; PW / PL 2.04. Scutellar shield equilateral triangular, finely punctate, lateral margins straight. Elytra oblong, subparallel-sided near base to apical 1 / 4, gently tapering in apical 1 / 4, relatively pointed at apex, sparsely and irregularly punctate; space between punctures larger than their diameter; adpressed silver setae forming zigzag markings; humeral parts distinctly convex dorsally; EL / EW 1.50, EL / PL 4.17, EW / PW 1.36, TL / EW 1.86. Each claw on foreleg of same size and shape. Sternite VIII (Fig. 3 H) U-shaped, bearing short setae in apical parts. Sternite IX (Fig. 3 I) elongate, with long and slender lateral projections. Aedeagus (Fig. 3 E – G) slender, slightly curved ventrally, finely punctate in apical part of median lobe and parameres; median lobe wide in lateral view, rather pointed at apex, minutely serrate in ventral part; apical emargination of parameres V-shaped; apices of parameres pointed in ventral view, forming thumb-like projection in lateral view. Female Unknown. Measurements Male (n = 1): TL 5.9 mm, PW 2.35 mm, PL 1.15 mm, EL 4.80 mm, EW 3.20 mm.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF971550FDF7FE57FD3DD508.taxon	distribution	Distribution Only known from the type locality, Tibet.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
E0318797FF971550FDF7FE57FD3DD508.taxon	discussion	Remarks This species belongs to the testudo species group (sensu Hernando & Ribera 2014). It is related to C. fujianensis Hernando & Ribera, 2014 known from China (Fujian), C. malickyi Hernando & Ribera, 2014 from Vietnam and C. jendeki Hernando & Ribera, 2014 from India, but differs from them in the following characteristics: median lobe with minute serrae in the ventral part, pointed at apex; emargination of parameres rather shallow.	en	Yoshitomi, Hiroyuki (2015): Four new species of the genus Caccothryptus (Coleoptera, Limnichidae). European Journal of Taxonomy 147: 1-17, DOI: 10.5852/ejt.2015.147
