taxonID	type	description	language	source
E73B096AFF9EFF87FF50F89410BFF857.taxon	materials_examined	Type material. Holotype male, 4.9 mm, dissected, 4 slides, AM P. 81856, dive site ‘ Winter Wall’, near Horsburgh Island, Cocos (Keeling) Islands, 12 ° 05 ' 15 " S, 96 ° 50 ' 14 " E, 9.5 m, rubble under plate coral, 12 October 2008, coll. K. B. Attwood, MI WA 835. Paratype female, 5 mm, dissected, 4 slides, AM P. 81857, dive site ‘ Winter Wall’, near Horsburgh Island, Cocos (Keeling) Islands, 12 ° 05 ' 15 " S, 96 ° 50 ' 14 " E, 9 m, rubble, 12 October 2008, coll. K. B. Attwood, MI WA 834. Type locality. Near Horsburgh Island, Cocos (Keeling) Islands, Indian Ocean 12 ° 05 ' 15 " S, 96 ° 50 ' 14 " E. Etymology. Derived from the Greek ‘ kallus’, meaning beauty, for the coxa 7 which has a distinctive castelloserrate posterior margin.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF9EFF87FF50F89410BFF857.taxon	description	Description. Of holotype male, 4.9 mm, AM P. 81856. Head. Eyes ovate; lateral cephalic lobe broad, truncated, apically rounded, anteroventral margin with notch / slit, corner rounded. Antenna 1 longer than antenna 2; peduncular article 1 shorter than article 2, without robust setae along posterior margin; peduncular article 2 longer than article 3; flagellum articles as long as broad, with 12 articles; accessory flagellum minute, 3 - articulate. Antenna 2 peduncular article 2 cone gland reaching end of peduncular article 3; peduncular article 4 subequal to article 5; flagellum with 8 articles. Mandible incisor cuspidate; accessory setal row with 4 setae; molar well-developed, triturative; palp well-developed, 3 - articulate; article 1 twice as long as broad, shorter than article 2; article 2 longer than article 3 with many slender setae; article 3 strongly falcate, short, 2.1 as long as broad, longer than article 1, with apical setae and a comb of short robust setae along anterodistal margin. Lower lip inner lobes present, outer lobes with 1 pair of ducts, mandibular lobes apically subacute. Maxilla 1 inner plate subquadrate, with 2 apical plumose setae. Maxilliped palp propodus with small distomedial flap. Pereon. Gnathopod 1 coxa anterior margin straight, anteroventral corner produced, rounded; merus without posterodistal tooth; carpus 1.5 as long as broad, shorter than propodus, length 1.1 propodus, with long slender setae covering surface; propodus medial surface setal comb absent, palm subacute, convex, entire, lined with robust setae, defined by posterodistal corner with 1 long robust seta. Gnathopod 2 sexually dimorphic; subchelate; basis slender, anterodistal corner subquadrate; merus with subquadrate distoventral corner; carpus compressed, lobate, projecting between merus and propodus, length 0.9 breadth; propodus expanded, with	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF9BFF80FF50F90214CEFD8B.taxon	materials_examined	Material examined. Indian Ocean: 1 male, 6 mm, dissected, 4 slides, NTMCr 018337, Ashmore Reef, 12 ° 14 ' S, 12257 ' E, low water line, seagrass flats, green calcareous alga Halimeda sp., 29 July 1986, coll. H. K. Larson; 1 male, NTMCr 018336, Ashmore Reef, 12 ° 14 ' S, 122 ° 59 ' E, low water, sand and reef flat, Thalassia dominant with Halimeda patches, 15 Apr 1987, coll. J. R. Hanley. Type locality. Niui Island, Tuvalu (as Ellice Islands).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF9BFF80FF50F90214CEFD8B.taxon	discussion	Remarks. Elasmopus gracilis has a wide distribution throughout the Indo-Pacific. Material examined here agrees with the original description from Tuvalu (Schellenberg 1938) and subsequent illustrations from Tonga (Myers 1986: figs, 6, 7).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF9BFF80FF50F90214CEFD8B.taxon	distribution	Distribution. Pacific Ocean. Fiji. Viti Levu: Bau (Schellenberg 1938). Micronesia; Kosrae (J. L. Barnard 1965; Myers 1995). Tonga: Tongatabu (Myers 1986). Tuvalu: Niui and Niutao (as Ellice Islands) (Schellenberg 1938). Western Samoa (Myers 1997). Indian Ocean.? Red Sea (Ruffo 1969).? Madagascar (Ledoyer 1967; 1982). Ashmore Reef (current study).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF9AFF8FFF50FA7E116BFEAB.taxon	materials_examined	Type material. Holotype male, 4.5 mm, dissected, 3 slides, AM P. 81496, Swiss Cheese Reef, Norfolk Island, 29 ° 00 ' 23 " S, 167 ° 56 ' 50 " E, 12 m, brown algae Dictyota sp., 19 May 2008, coll. J. K. Lowry, MI NFK 78. Paratypes 16 specimens, AM P. 81483, dive site ‘ The Wall’, Nepean Island, Norfolk Island, 29 ° 04 ' 18 " S, 167 ° 57 ' 41 " E, 10 m, tufted red and brown algae, 15 May 2008, coll. J. K. Lowry, MI NFK 43. Type locality. Slaughter Bay, Norfolk Island, South Pacific, 29 ° 03 ' 30 " S, 167 ° 57 ' E.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF9AFF8FFF50FA7E116BFEAB.taxon	etymology	Etymology. Incomptus from the Latin for unadorned, in reference to the relatively plain male gnathopod 2 propodus palm.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF9AFF8FFF50FA7E116BFEAB.taxon	description	Description. Of holotype male, 4.5 mm, AM P. 81496. Head. Eyes ovate; lateral cephalic lobe broad, truncated, with anteroventral notch / slit, corner rounded. Antenna 1 longer than antenna 2; peduncular article 1 with 1 robust seta on posterior margin, flagellum broken; accessory flagellum minute, with 2 articles. Antenna 2 peduncular article 2 cone gland not reaching to end of peduncular article 3; peduncular article 4 shorter in length than article 5; flagellum with 8 articles. Mandible incisor with cuspidate margin; accessory setal row with 4 setae (illustrated side has row damaged with only 1 seta); molar well-developed; palp well developed, 3 - articulate; article 1 twice as long as broad, shorter than article 2; article 2 subequal to article 3, with several slender setae; article 3 short (twice as long as broad), weakly falcate, with apical setae and a comb of short robust setae along anterodistal margin. Lower lip inner lobes present, outer lobes with single pair of ducts, mandibular lobes apically subacute. Maxilla 1 inner plate subquadrate, with 2 apical plumose setae. Maxilliped palp propodus with small distomedial flap. Pereon. Gnathopod 1 coxa anterior margin straight, anteroventral corner produced, rounded, posteroventral corner without notch; merus without posterodistal tooth; carpus 1.5 × as long as broad, length 0.9 × propodus, anterior margin with a few slender setae; propodus medial surface setal comb absent, palm subacute, convex, entire, defined by posterodistal corner, with 1 robust seta. Gnathopod 2 subchelate; coxa posteroventral corner notch absent; basis slender, anterodistal corner subquadrate; merus with rounded distoventral corner; carpus compressed, lobate, projecting between merus and propodus, length 0.9 × breadth; propodusexpanded, with slender medial setal bunch, palm extremely acute without corner, straight, subtriangular distomedial shelf with group of 5 robust setae on shelf, palm 0.4 × length of propodus, subpalmar surface with long subpalmar seam, with posterodistal robust setae; dactylus apically subacute, closing along andreaching end of palm. Pereopods 5 – 7 basis posterior margin without long slender setae; merus andcarpus not broadened; dactylar ungues simple. Pereopods 5 – 6 basis expanded, posterior margin straight, minutely crenulate, without long slender setae, posteroventral corner broadly rounded; carpus and propodus with few long, slender setae along anterior margin; dactylus unguis anterior margin without accessory spines. Pereopod 7 basis posterior margin convex, weakly crenulate, posteroventral corner broadly rounded; carpus and propodus with long slender setae along posterior margin. Pleon. Epimeron 1 posteroventral corner scalloped. Epimeron 2 posteroventral corner notched. Epimeron 3 ventral margin smooth, posteroventral corner scalloped. Urosomite 1 without carina. Uropod 1 peduncle with 1 basofacial seta; inner ramus shorterthan outer ramus. Uropod 2 inner ramus shorter than outer ramus, subequal in length to peduncle. Uropod 3 inner ramus shorter than outer ramus; outer ramus short, length twice breadth, subequal to peduncle, rami distally truncated, with long and short apical robust setae. Telson as long as broad, deeply cleft (80 %), lobes abutting, truncated distally, each lobe with 2 long apical robust setae, with 2 pair of lateral plumose setae.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF9AFF8FFF50FA7E116BFEAB.taxon	discussion	Remarks. Elasmopus incomptus sp. nov. aligns with E. hawaiiensis Schellenberg, 1938 and E. ecuadorensis, Schellenberg, 1936, based on the male gnathopod 2 propodus palm with dense brush of setae, epimeron 3 posterior margin scalloped, and the shape of the uropod 3 rami. The telson with two long robust setae on each lobe separates E. incomptus sp. nov. from the latter species, where E. hawaiiensis has two short robust setae and E. ecuadorensis has either three long and one short or three long setae on each lobe (following Barnard, 1979). The mandible of the type material has not been figured in full for the latter two species and it could be useful to either separate or show a close affinity to E. incomptus sp. nov. The mandible accessory setal row with only one seta in E. incomptus sp. nov. is unusual for the genus.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF9AFF8FFF50FA7E116BFEAB.taxon	distribution	Distribution. South Pacific Ocean: Tasman Sea: Norfolk Island (current study).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF95FF8FFF50FE8A14E0FC50.taxon	materials_examined	Material examined. Western Australia. 1 male, 5.5 mm, dissected, 4 slides, AM P. 92257, Descartes Island, The Kimberley, 14 ° 11 ' S, 125 ° 40 ' E, 20 July 1988, coll. P. A. Hutchings, Stn 70; 2 specimens, AM P. 92256, Descartes Island, The Kimberley, 14 ° 11 ' S, 125 ° 40 ' E, 20 July 1988, coll. P. A. Hutchings, Stn 70. Type locality. Bora Bora, Society Islands.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF95FF8FFF50FE8A14E0FC50.taxon	discussion	Remarks. The presence of robust setae on the male gnathopod 2 basis anterodistal corner is known for Elasmopus integer, E. hooheno J. L. Barnard, 1970, E. pseudinteger Appadoo & Myers, 2003 and E. arafura Hughes & Lowry, 2011. Of these species, only E. integer and E. pseudinteger have an entire telson. The presence of long slender setae on the margins of the pereopods and uropod 3 rami confirm this material as E. integer and not the closely related E. pseudinteger reported from Mauritius. The telson of specimens examined here varies slightly from the original material with a small suture along the apical margin, not illustrated in the Society Islands material.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF95FF8FFF50FE8A14E0FC50.taxon	distribution	Distribution. Pacific Ocean: Society Islands (Myers 1989). Indian Ocean: Australia Western Australia: The Kimberley (current study).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF93FF89FF50FF7211A8FDE2.taxon	materials_examined	Material examined. 1 male, 4.6 mm, dissected, 4 slides, WAM C 46267, Cassini Island, The Kimberley, Western Australia, 13 ° 55 ' S, 125 ° 37 ' E 1.3 m, 16 October 2010, coll. L. Betteridge, Woodside Kimberley Survey, 32 / K 10, adhoc. Type locality. Oahu, Hawaii.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF93FF89FF50FF7211A8FDE2.taxon	discussion	Remarks. This is the first record of this species in Australia and the most westerly record for this species.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF93FF89FF50FF7211A8FDE2.taxon	distribution	Distribution. Indian Ocean: Australia: Western Australia: Arthur Head, Kalbarri, Ned’s, Camp, Jackson Island, Cassini Island (current study). North Pacific Ocean: Hawaii, Oahu (Barnard 1970).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF93FF95FF50FA5A1162FF4E.taxon	materials_examined	Type material. Holotype male, 4 mm, dissected, 4 slides, AM P. 81484, Duncombe Bay, Norfolk Island, 29 ° 03 ' 30 " S, 167 ° 57 ' 13 " E, 15 m, algae, sponge and bryozoans, 13 May 2008, coll. J. K. Lowry, MI NFK 15 °. Paratypes: 1 female, 4.8 mm, dissected, 3 slides, AM P. 92244, Cathedral Rock, Norfolk Island, 2900 ' 13 " S, 167 ° 56 ' 57 " E, 10.5 m, overhang wall sponges and ascidians, 13 May 2008, coll. R. T. Springthorpe, MI NFK 6; 13 specimens, AM P. 81490, Cathedral Rock, Norfolk Island, 29 ° 00 ' 13 " S, 167 ° 56 ' 57 " E, 10.5 m, overhang wall sponges and ascidians, 13 May 2008, coll. R. T. Springthorpe, MI NFK 6; 2 specimens, AM P. 81491, Cathedral Rock, Norfolk Island, 29 ° 12 " S, 167 ° 56 ' 57 " E, 10.5 m, overhang wall sponges and ascidians, 13 May 2008, coll. R. T. Springthorpe, MI NFK 6; 1 specimen, AM P. 81487, Gun Club Reef, near Anson Point, Norfolk Island, 29 ° 25 " S, 167 ° 54 ' 44 " E, 25 m, tufted brown and green algae, 17 May 2008, coll. J. K. Lowry, MI NFK 59; 1 specimen, AM P. 81485, Puppies Point, Norfolk Island, 29 ° 1 ' 22 " S, 167 ° 55 ' 8 " E, 13 m, red algae, 17 May 2008, coll. R. T. Springthorpe, MI NFK 53; 1 specimen, AM P. 81493, Fig Valley reef, Norfolk Island, 29 ° 3 ' 20 " S, 167 ° 55 ' 44 " E, 19.6 m, from bommie wall cored in sparse, 19 May 2008, coll. R. T. Springthorpe, MI NFK 66; 1 specimen, AM P. 81486, dive site, ‘ The Wall’, Nepean Island, near Norfolk Island, 29 ° 4 ' 17 " S, 167 ° 57 ' 40 " E, 9.6 m, red coralline algae cf Amphiroa anceps, 15 May 2008, coll. L. E. Hughes, MI NFK 49; 2 specimens, AM P. 81488, dive site ‘ The Wall’, Nepean Island, near Norfolk Island, 29 ° 4 ' 17 " S, 167 ° 57 ' 40 " E, 17 m, green algae Halimeda sp., 15 May 2008, coll. J. K. Lowry, MI NFK 45; 1 specimen, AM P. 81494, dive site ‘ The Wall’, Nepean Island, near Norfolk Island, 29 ° 4 ' 17 " S, 167 ° 57 ' 40 " E, 17 m, green algae Halimeda sp., 15 May 2008, coll. J. K. Lowry MI NFK 45; 1 male, AM P. 81497, Swiss Cheese Reef, Norfolk Island 29 ° 00 ' 23 " S, 167 ° 56 ' 50 " E, 12 m, vertical face encrusted with sponges, under overhang, 15 May 2008, coll. R. T. Springthorpe, MI NFK 37; 1 male, AM P. 81489, Swiss Cheese Reef, Norfolk Island, 29 ° 00 ' 23 " S, 167 ° 56 ' 50 " E, 14 m, overhang, sponges, encrusting algae, 15 May 2008, coll. J. K. Lowry MI NFK 38; 1 male, AM P. 81492, Duncombe Bay, Norfolk Island, 29 ° 03 ' 30 " S, 167 ° 57 ' 13 " E, 15 m, green algae Caulerpa racemosa, 13 May 2008, coll. J. K. Lowry, MI NFK 17. Type locality. Duncombe Bay, Norfolk Island, South Pacific, 29 ° 03 ' 30 " S, 167 ° 57 ' 13 " E.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF93FF95FF50FA5A1162FF4E.taxon	etymology	Etymology. Named for the type locality, Norfolk Island.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF93FF95FF50FA5A1162FF4E.taxon	description	Description. Of holotype male, 4 mm, AM P. 81484. Head. Eyes ovate; lateral cephalic lobe broad, apically rounded, anteroventral margin with notch / slit, corner rounded. Antenna 1 longer than antenna 2; peduncular article 1 subequal in length to article 2, with 2 robust setae on posterodistal margin; peduncular article 2 longer than article 3; flagellum articles as long as broad; with 16 articles; accessory flagellum short, 3 - articulate. Antenna 2 peduncular article 2 cone gland extending beyond peduncular article 3; peduncular article 4 longer than article 5; flagellum with 7 articles. Mandible molar welldeveloped triturative, incisor cuspidate; accessory setal row, with 4 setae; palp 3 - articulate; well-developed, article 3 weakly falcate, long, length 6 width. Lower lip inner lobes present, outer lobes ducts without ducts, mandibular lobes straight apically rounded. Maxilla 1 inner plate subquadrate, with 2 apical plumose setae. Maxilliped palp propodus with small distomedial flap. Pereon. Gnathopod 1 coxa anterior margin concave, anteroventral corner slightly produced, acute; merus without posterodistal tooth; carpus twice as long as broad, longer than propodus, length 1.1 propodus, anterior margin with a few slender setae; propodus anterior margin with a few rows of slender setae, medial surface setal comb absent, palm subacute, straight, entire, defined by posterodistal corner with 4 robust setae. Gnathopod 2 subchelate; coxa posteroventral corner notch absent; basis slender, anterodistal corner subquadrate; merus with rounded distoventral corner; carpus compressed, lobate, projecting between merus and propodus, length 0.9 breadth; propodus expanded, palm acute, smooth, convex, with slender medial setal bunch, palm length 0.8 propodus posterior margin, subquadrate distomedial shelf with group of 10 robust setae, subpalmar surface smooth, palm defining by subpalmar socket with 1 robust seta; dactylus posterior margin with proximal tooth, apically subacute, reaching end of palm and closing into subpalmar socket. Pereopods 5 – 7 basis expanded, posterior margin without long slender setae; merus and carpus not broadened; dactylar ungues simple. Pereopod 5 basis posterior margin straight, weakly serrate, posteroventral corner broadly rounded. Pereopod 6 basis posterior margin serrate, tapering distally, posteroventral corner narrowly rounded to subquadrate. Pereopod 7 basis posterior margin convex, serrate, posteroventral corner narrowly rounded to subquadrate. Pleon. Epimera 1 – 3 posteroventral corner with small acute tooth, ventral margin smooth. Urosomite 1 without carina. Uropod 1 peduncle with 1 basofacial seta; inner ramus longer than outer ramus, 0.8 peduncle length. Uropod 2 inner ramus longer than outer ramus, subequal in length to peduncle. Uropod 3 inner ramus short, length 1.9 breadth, subequal in length to outer ramus, longer than peduncle; rami distally truncated, apical robust setae short. Telson (based on female paratype) deeply cleft (80 %), lobes broader than long, abutting, apically rounded with 1 pair of subapical robust setae. Female (sexually dimorphic characters). Of paratype female, 4.8 mm, AM P. 92244. Gnathopod 2 propodus palm subacute, straight, margin weakly irregular, lined with a few robust setae and defined by 3 loosely clustered robust setae; dactylus reaching end of and closing along palm.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF93FF95FF50FA5A1162FF4E.taxon	discussion	Remarks. The male gnathopod 2 with greatly elongate dactylus more than three quarters the length of the propodus, is similar to Elasmopus aduncus Myers, 1995; E. gracilis Schellenberg, 1938; E. piikoi Barnard, 1970; E. temori Barnard, 1979 and E. spinidactylus Chevreux, 1908. The serrate posterior margin of the pereopod 7 basis separates E. norfolkensis sp. nov. from these species. Other distinctive features of E. norfolkensis sp. nov. include the gnathopod 2 dactylar socket on the proximal mid facial surface of the dactylus and the proximal tooth on dactylus posterior margin.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF93FF95FF50FA5A1162FF4E.taxon	distribution	Distribution. South Pacific Ocean: Tasman Sea: Norfolk Island (current study).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF8FFF94FF50F9E01737FD8B.taxon	materials_examined	Material examined. New Zealand: male, 5.5 mm, dissected, 4 slides, AM P. 98051, Fishing Rock landing, Raoul Island, Kermadec Islands, 29 ° 15 ' 03 " S, 177 ° 54 ' 12 " W, 1 m, invertebrate substrate under boulders, rockpools beside Fishing Rock, some loose rocks and coral cover in pool, algal turf, 18 May 2011, coll. S. J. Keable, M. A. McGrouther and A. Reid, K 2011 - 54 - 1; 15 + specimens, AM P. 90947, Fishing Rock landing, Raoul Island, Kermadec Islands, 29 ° 15 ' 03 " S, 177 ° 54 ' 12 " W, 1 m, under boulders, rockpools beside Fishing Rock, some loose rocks and coral cover in pool, algal turf, 18 May 2011, coll. S. J. Keable, M. A. McGrouther and A. Reid, K 2011 - 54 - 1; 1 specimen, AM P. 90942, Macauley Island, Kermadec Islands, 30 ° 13 ' 40 " S, 178 ° 26 ' 20 " W, 21.2 m, side and top of rock with Porifera, 21 May 2011, coll. S. J. Keable and A. Reid, K 2011 - 70; 2 specimens, AM P. 90943, west side of L'Esperance Rock, Kermadec Islands, 31 ° 21 ' 15 " S, 178 ° 49 ' 36 " W, 12 – 20 m, rockwalls, shelly sediment, sponges and coral scrapings, 26 May 2011, coll. S. J. Keable and A. Reid, K 2011 - 99 - 3; 10 specimens, AM P. 90944, west side of South Meyer Island, Kermadec Islands, 29 ° 14 ' 47 " S, 177 ° 52 ' 53 " W, 12 m, rock-lined gutter about 10 m long, 30 cm deep with rubble, coll. S. J. Keable, M. A. McGrouther and A. Reid, K 201 - 3; 6 specimens, AM P. 90945, Fishing Rock landing, Raoul Island, Kermadec Islands, 29 ° 15 ' 2 " S, 177 ° 54 ' 11 " W, 5 m, scrapings from rock wall, coll. S. J. Keable and A. Reid, K 2011 - 49 - 5; 4 specimens, AM P. 90946, Fishing Rock landing, Raoul Island, Kermadec Islands, 29 ° 15 ' 2 " S, 177 ° 54 ' 11 " W, 5 m, scrapings from rock wall, coll. S. J. Keable and A. Reid, K 2011 - 49 - 7. Type locality. Souillac, Mauritius, Indian Ocean 20 ° 31 ' S, 57 ° 30.7 ' E.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF8FFF94FF50F9E01737FD8B.taxon	discussion	Remarks. Specimens of Elasmopus souillacensis from the Kermadec Island are the first to be recorded since its description from Mauritius and provide a geographic range extension into the South Pacific Ocean.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF8FFF94FF50F9E01737FD8B.taxon	distribution	Distribution. Indian Ocean: Mauritius (Appadoo & Myers 2003). South Pacific Ocean: New Zealand: Kermadec Islands (current study).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF8EFF96FF50F9C81611FF63.taxon	diagnosis	Diagnosis. Head anteroventral corner with notch; eyes ovate. Antenna 1 accessory flagellum minute, with around 3 - article, not extending beyond flagellum article 2. Mandibular molar, well developed triturative; palp article 1 without acutely produced distal corner; article 2 longer than article 3; article 3 strongly falcate, anterodistal margin concave, with setal comb and apical setae. Maxilla 1 inner plate with mainly apical robust setae. Gnathopod 1 coxa anteroventral corner acutely produced, posteroventral corner with notch; propodus palm subacute. Gnathopod 2 (? sexually dimorphic) female propodus symmetrical; palm angle subacute (greater than 90 °); dactylus anterior margin with 1 seta. Pereopod 4 coxa with posterodistal lobe. Pereopods 5 – 7 basis weakly expanded; dactylus simple. Pereonite 7 with dorsal carinae. Pleonites 1 – 3 with dorsal carinae. Urosomites 1 – 3 dorsally smooth. Epimera 2 – 3 ventral margins distally serrate. Uropod 3 rami unequal in length; inner ramus half length of outer ramus; outer ramus 1 articulate, length 1.8 × peduncle, length more than 2.5 × width. Telson moderately cleft (50 %), lobes tapering, with apical robust setae.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF8EFF96FF50F9C81611FF63.taxon	materials_examined	Type species. Leeuwinella mistakensis sp. nov., by present designation and monotypy; gender feminine. Etymology. Named after Leeuwin, the Dutch exploration vessel, meaning lioness, which is most fitting for a genus known only from a female specimen.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF8EFF96FF50F9C81611FF63.taxon	discussion	Remarks. Leeuwinella gen. nov. is most closely aligned to Elasmopus, Hoho Lowry & Fenwick, 1983 and Mallacoota J. L. Barnard, 1972 based on the minute accessory flagellum, presence of dorsal carinae, mandibular palp with short article 1 not distally produced, pereopods 5 – 7 with expanded basis and simple dactylus. The single dorsal carina on the pleonites, the long, apically truncate, uropod 3 outer ramus and reduced inner ramus is similar to Nuuanuidae Lowry & Myers, 2013, althoughthe lack of setae on the urosomite 2 and pereopods 5 – 7 basis weakly expanded, place this genus outside the Nuuanuidae.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF8CFF90FF50FA481021FA47.taxon	materials_examined	Type material. Holotype: female, 5.5 mm, dissected, 4 slide, AM P. 92754, Vancouver Peninsular, near Mistaken Island, King George Sound, Western Australia, 35 ° 4 ' S, 117 ° 56 ' E, 6 m, seagrass, 13 December 1983, coll. R. T. Springthorpe WA 121. Paratypes: 9 females, AM P. 92755, Vancouver Peninsular, near Mistaken Island, King George Sound, 35 ° 4 ' S, 117 ° 56 ' E, 6 m, seagrass, 13 December 1983, coll. R. T. Springthorpe, WA 121. Type locality. Vancouver Peninsular, near Mistaken Island, King George Sound, Western Australia, Indian Ocean, 35 ° 4 ' S, 117 ° 56 ' E.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF8CFF90FF50FA481021FA47.taxon	etymology	Etymology. Named from the type locality.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF8CFF90FF50FA481021FA47.taxon	description	Description. Of holotype female, 5.5 mm, AM P. 92754. Head. Eyes ovate; lateral cephalic lobe broad, truncated, anteroventral margin with notch, corner acute. Antenna 1 longer than antenna 2; peduncular article 1 shorter than article 2, with 1 robust setae on posterodistal margin; peduncular article 2 longer than article 3; flagellum articles longer than broad, with 21 articles; accessory flagellum minute, 3 - articulate. Antenna 2 peduncular article 2 cone gland not reaching end of peduncular article 3; peduncular article 4 subequal to article 5; flagellum with 9 articles. Mandible incisor cuspidate; accessory setal row with 3 setae; molar well-developed, triturative; palp well-developed, 3 - articulate; article 1 twice as long as broad, shorter than article 2; article 2 longer than article 3 with many slender setae; article 3 strongly falcate, anterodistal margin concave, 3 as long as broad, longer than article 1, with apical setae and a comb of short robust setae along anterodistal margin. Lower lip inner lobes present, outer lobes without ducts, mandibular lobes apically acute. Maxilla 1 inner plate subquadrate, with 3 apical plumose setae. Maxilliped palp propodus with small distomedial flap. Pereon. Gnathopod 1 coxa anterior margin concave, anteroventral corner produced, acute, posteroventral margin with notch; merus without posterodistal tooth; carpus 1.9 as long as broad, subequal to propodus, with long slender setae on posterior margin; propodus medial surface setal comb present, palm subacute, convex, entire, lined defined by posterodistal corner with 2 robust setae. Gnathopod 2? sexually dimorphic; subchelate; basis slender, anterodistal corner subquadrate; merus with subquadrate distoventral corner; carpus rectilinear, length 1.9 breadth; propodus subovate to rectilinear, palm acute, near straight, palm weakly defined without corner or robust setae; dactylus apically subacute, closing along and reaching end of palm. Pereopod 4 coxa posterodistal lobe well developed. Pereopods 5 – 7 basis weakly expanded, posterior margin serrate; merus and carpus not broadened; dactylar ungues simple. Pereopod 5 basis posterior margin straight. Pereopod 6 basis posterior margin sinusoidal. Pereopod 7 posterior margin weakly sinusoidal. Pereonite 7 with single dorsal carina. Pleon. Pleonites 1 – 3 with single dorsal carina. Epimera 1 – 3 posterodistal corner with acute tooth. Epimeron 1 posterior and ventral margins smooth. Epimeron 2 posterior margin smooth, ventral margin distally serrate. Epimeron 3 posterior and ventral margins distally serrate. Urosomites 1 – 2 without carina, without dorsal setae. Uropod 1 peduncle with 1 basofacial seta, inner ramus slightly longer than outer ramus, 0.9 peduncle length. Uropod 2 inner ramus subequal to outer ramus. Uropod 3 inner ramus length 2.1 breadth, 0.5 outer ramus length; outer ramus 1.8 peduncle length, distally truncated, apical robust setae short. Telson as long as broad, moderately cleft (80 %), lobes tapering distally (outer margin straight and inner margins angled), each lobe with 2 short apical robust setae.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF8CFF90FF50FA481021FA47.taxon	discussion	Remarks. See remarks for genus. The falcate mandibular palp in Leeuwinella mistakensis gen. et. sp. nov. is also present in Elasmopus, but the presence of dorsal carinae is rare within the genus. Four Elasmopus species, E. japonicas Stephensen, 1932, E. neglectus Chilton, 1915, E. shepherdi Hughes & Lowry, 2011 and E. woodjonesi Hughes & Lowry, 2011 have a unicarinate urosomite 1 only, whereas urosomite 1 is smooth in E. mistakensis sp. nov. Within Maeridae, only two other species have unicarinate pleonites. Hoho carteta J. L. Barnard, 1972 a and Mallacoota unidentata Ren, 1998 have pleonites 1 and 2 only with a single carina. The strongly falcate mandibular palp, single carinae on pereonite 7 and pleonites 1 and 3 along with the serrate distal margins of epimera 2 and 3 make Leeuwinella mistakensis gen. et. sp. nov. distinctive among Australian maerid species.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF8CFF90FF50FA481021FA47.taxon	distribution	Distribution. Indian Ocean: Australia: Western Australia: Vancouver Peninsular (current study).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF8AFF93FF50F96516C4FEF6.taxon	materials_examined	Material examined. Male, 7.5 mm, dissected, 4 slides, NTMCr 18332, Bedout Island, Western Australia, 19 ° 3 ' S, 11 ° 95 ' E, 3 m, reef flat, 4 June 1985, coll. B. C. Russell, BCR 85 - 9; female, 6.5 mm, dissected, 1 slides, NTMCr 18333, Bedout Island, Western Australia, 19 ° 35 ' S, 11 ° 95 ' E, 3 m, reef flat, 4 June 1985, coll. B. C. Russell, BCR 85 - 9; 7 female specimens (oostegites narrow, not inflated), NTMCr 18331, Bedout Island, Western Australia, 19 ° 35 ' S, 11 ° 95 ' E, 3 m, reef flat, 4 June 1985, coll. B. C. Russell, BCR 85 - 9. Type locality. Between Bird Islet and South Island, Lizard Island, Queensland, Australia, 14 ° 42 ' S, 145 ° 28 ' E. Remarks. Material examined here extends the distribution of M. griffini from Queensland to Western Australia.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF8AFF93FF50F96516C4FEF6.taxon	distribution	Distribution. Indian Ocean: Australia: Western Australia: Bedout Island (current study). South Pacific Ocean: Australia: Queensland: Lizard Island (Berents 1983; Krapp-Schickel 2009).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF86FF9FFF50FA76163CFED3.taxon	materials_examined	Material examined. South Australia: Several specimens, SAMA C 6526, 18.6 miles west north west of Rapid Head, Gulf St Vincent, 35 ° 31.14 ' S, 138 ° 10.02 ' E, prawn trawl, 23 June 1987, coll. K. L. Gowlett-Homes and S. Corigliano, FV Rivoli Queen. Tasmania: 2 specimens, AM P. 83528, off Hannants Bight, Cape Sorell, 42 ° 11 ' 30 " S, 145 ° 11 ' 18 " E, 16 m, red algae with epiphytic algae, bryozoa and sponges, 27 April 1991, coll. R. T. Springthorpe and P. M. Berents, TAS 289; 4 specimens, AM P. 83530, Cemetery Bluff, Adventure Bay, Bruny Island, 43 ° 19 ' 48 " S, 147 ° 19 ' 12 " E, 15 m, sponges, 23 April 1991, coll. R. T. Springthorpe and P. M. Berents, TAS 249; 2 specimens, AM P. 83529, Cemetery Bluff, Adventure Bay, Bruny Island, 43 ° 19 ' 48 " S, 147 ° 19 ' 12 " E, 15 m, lace bryozoan, 23 April 1991, coll. R. T. Springthorpe and P. M. Berents, TAS 247. Tasman Sea. Many specimens, AM P. 73317, North Norfolk Ridge, 28 ° 54 ' 23 " S, 167 ° 41 ' 3 " E, 111 m, beam trawl, coll. P. B. Berents, 15 May 2003. Type locality. Between Troubridge Light and Cape Jervis, South Australia, 3520 ' S, 13740 ' E.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF86FF9FFF50FA76163CFED3.taxon	discussion	Remarks. The current record extends M. thetis further south from the southern coast of Australia to Tasmania and west to the North Norfolk Ridge in the Tasman Sea between Australia and New Zealand.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF86FF9FFF50FA76163CFED3.taxon	distribution	Distribution. Australia: Tasmania: Cape Sorrell, Bruny Island (current study); New South Wales: Coogee, Wollongong (Stebbing 1910); South Australia: Cape Jervis, Gulf St Vincent (Lowry & Springthorpe 2005; current study). Tasman Sea: North Norfolk Ridge (current study).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF85FF9FFF50FE5217D5FA68.taxon	diagnosis	Diagnosis. Head anteroventral corner with weak notch; eyes poorlydeveloped, ommatidia scattered, ovate. Antenna 1 accessory flagellum minute, 2 - article, not extending beyond flagellum article 2. Mandibular molar reduced, columnar, triturative; palp article 1 without acutely produced distal corner; article 2 longer than article 3; article 3 strongly falcate, anterodistal margin concave, with setal comb and apical setae. Maxilla 1 inner plate subtriangular with setae along entire margin. Coxa 1 – 4 reduced, broader than deep, less than the pereonites. Gnathopod 1 coxa anteroventral corner not produced, posteroventral corner without notch; propodus palm subacute. Gnathopod 2 (sexually dimorphic) propodus symmetrical; palm angle obtuse with pollex or subacute (greater than 90 °); dactylus anterior margin with 1 seta. Pereopod 4 coxa with posterodistal lobe. Pereopods 5 – 7 basis and merus rectilinear, not expanded or expanded; dactylus simple. Pereonites, Pleonites and Urosomites dorsally smooth. Epimera 1 – 3 margins smooth. Uropod 3 rami subequal, length twice × width, longer or shorter than peduncle; apically rounded; without robust setae, with apical slender setae only. Telson deeply cleft (> 66 %), lobes abutting, tapering distally, with apical slender setae.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF85FF9FFF50FE5217D5FA68.taxon	materials_examined	Type species. Pseudelasmopus cheliferus Ledoyer, 1978, by original designation and monotypy. Included species. Pseudelasmopus cheliferus Ledoyer, 1978; and P. walkerae sp. nov.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF85FF9FFF50FE5217D5FA68.taxon	discussion	Remarks. The main characters used to diagnose Pseudelasmopus, uropod 3 and telson reduced, are no longer sufficient to define the genus from other Maeridae genera (Ledoyer 1978). Therefore, the diagnosis is updated and expanded here to confirm Pseudelasmopus as a valid and distinct genus. Characters including the reduced eyes, reduced coxae, setose margin of the maxilla 1 inner plate, uropod 3 with slender setae onlyand telson without apical robust setae and only slender setae, separate this genus from the closely aligned and more speciose Elasmopus. As much as the latter characters group P. cheliferus and P. walkerae together in comparison to other Maeridae, several characters usually of generic level vary between the two known species — the gnathopod 2 chelation, expansion of the basis and merus in pereopods 5 – 7, uropod 3 peduncle lengthand telson size. As P. cheliferus was described from a male and P. walkerae is known only from females, there is a possibility that some of these characters relate to sexual dimorphism. Until more is known about these species, it seems more appropriate to group them together in the same genus within the Maeridae, than to establish another monotypic genus for a species only known from limited material.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF85FF98FF50F9D4116BFA76.taxon	materials_examined	Type material. Holotype male, 3 mm, dissected, 4 slides, AM P. 92517, Emily Bay, Norfolk Island, 29 ° 3 ' 36 " S, 167 ° 57 ' 12 " E, 3.7 m, airlift, rubble on sand, 20 May 2008, coll. R. T. Springthorpe, MI NFK 82. Paratypes: 1 specimen, AM P. 92518, Emily Bay, Norfolk Island, 29 ° 3 ' 36 " S, 167 ° 57 ' 12 " E, 3.7 m, airlift, rubble on sand, 20 May 2008, coll. R. T. Springthorpe MI NFK 82; 8 specimens, AM P. 92522, Emily Bay, Norfolk Island, 29 ° 3 ' 36 " S, 167 ° 57 ' 12 " E, 3.7 m, airlift, rubble on sand, 20 May 2008, coll. R. T. Springthorpe, MI NFK 82; 1 specimen, AM P. 92519, Emily Bay, Norfolk Island, 29 ° 3 ' 36 " S, 167 ° 57 ' 12 " E, 3.7 m, airlift, rubble and algae, 20 May 2008, coll. R. T. Springthorpe, MI NFK 84; 1 specimen, AM P. 92521, Swiss cheese reef, Norfolk Island, 29 ° 00 ' 23 " S, 167 ° 56 ' 50 " E, 12 m, airlift, under overhang, vertical face encrusted with sponges, coll. 15 May 2008, coll. R. T. Springthorpe, MI NFK 37; 5 specimens, AM P. 92520, east of Sail Rock, Phillip Island, near Norfolk Island, 29 ° 6 ' 54 " S, 167 ° 57 ' 31 " E, 14.8 m, airlift, under stones, rubble, 14 May 2008, coll. R. T. Springthorpe MI NFK 20; 16 specimens, AM P. 92523, east of Sail Rock, Phillip Island, near Norfolk Island, 29 ° 6 ' 54 " S, 167 ° 57 ' 31 " E, 13 m, airlift, under stones, 14 May 2008, coll. R. T. Springthorpe, MI NFK 22. Type locality. Emily Bay, Norfolk Island, South Pacific, 29 ° 3 ' 36 " S, 167 ° 57 ' 12 " E.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF85FF98FF50F9D4116BFA76.taxon	etymology	Etymology. Named in honour of Wendy Walker, the zoological technician who performed the preliminary microscope processing of the Norfolk Island samples and has been a regular volunteer in the Australian Museum Marine Invertebrates section for over 20 years.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF85FF98FF50F9D4116BFA76.taxon	description	Description. Of holotype male, 3 mm, AM P. 92517. Head. Eyes round; lateral cephalic lobe broad, apically rounded, anteroventral margin with notch / slit, anteroventral corner rounded. Antenna 1 longer than antenna 2; peduncular article 1 length 1.1 × article 2, without robust setae on posterior margin; peduncular article 2 longer than article 3; flagellum articles longer than broad, with 17 articles; accessory flagellum short, less than half length of primary flagellum, with 2 articles. Antenna 2 peduncular article 4 longer than article 5; flagellum with 9 articles. Mandible accessory setal row, well developed with 5 setae; palp well developed, article 1 length 1.5 × width, shorter than article 2, inner margin not distally produced; article 2 length subequal to article 3, with long slender setae; article 3 falcate, twice as long as broad, with well-developed setal comb. Lower lip without apical ducts, mandibular lobes subacute. Maxilla 1 inner plate with medial setal row; outer plate palp, 2 articulate slender, article 2 with 4 apical slender setae. Pereon. Gnathopod 1 coxa anterior margin straight, anteroventral corner not produced, broadly rounded, ventral margin without robust setae, posteroventral corner without notch; basis anterodistal corner subquadrate; ischium anterodistal corner without lobes; merus with posteroventral corner subacute; carpus length 1.8 × width, subequal to propodus length, anterior margin with a few long slender setae, posterior margin lined with long slender setae, without dense rows of setae covering medial surface; propodus subovate, medial surface setal comb absent, palm subacute, straight, entire, lined with short robust setae, palm with weakly defined corner and no robust setae; dactylus posterior closing along palm, unguis present. Gnathopod 2 symmetrical, subchelate; coxa posteroventral corner without notch, ventral margin without robust setae; basis slender, anterodistal corner subquadrate; ischium anterodistal corner without lobes; merus posteroventral corner acute; carpus compressed, length 0.7 × width, anterior margin with 1 slender seta, posterior margin with rows of slender setae; propodus subovate, length 1.4 × width, posterior margin with clusters of long slender setae, palm length half of propodus posterior margin, angle subacute, distal shelf absent, palm smooth, entire, defined by posteroventral corner with small subacute tooth androbust seta; dactylus reaching end of and closing along palm, posterior margin smooth, unguis absent. Pereopods 3 – 4 coxa subquadrate; basis long, length 3.5 × width; merus, carpus and propodus not broadened; dactylus unguissimple. Pereopods 5 – 7 basis slender posterior margin straight; merus, carpus and propodus slender, margins lined with long slender setae; dactylus unguis simple. Pleon. Pleonites 1 – 3 dorsally smooth. Epimera 1 – 3 posterior and ventral margins smooth, posteroventral corner with subacute to acute tooth. Urosomite 1 dorsally smooth. Uropod 1 peduncle with 1 basofacial seta, without interramal spur, peduncle length 0.8 × outer ramus; inner ramus subequal to outer ramus length. Uropod 2 peduncle length 0.9 × outer ramus; inner ramus slightly shorter than outer ramus. Uropod 3 peduncle 0.8 × inner ramus; inner ramus subequal to outer ramus, length twice width, apically rounded, without lateral setae, with apical long slender setae. Telson broader than long, deeply cleft (80 %), lobes abutting, tapering distally, lobes apically convex, each lobe with 2 short slender apical setae.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF85FF98FF50F9D4116BFA76.taxon	discussion	Remarks. Pseudelasmopus walkerae sp. nov. has a subchelate gnathopod 2 distinguishing it from P. cheliferus Ledoyer, 1978 of Mauritius which has a chelate gnathopod. Other characters that separate these species include the smooth posterior margin of pereopods 5 – 7 and the much longer than broad uropod 3 rami in P. walkerae sp. nov. The telson is well developed in P. walkerae sp. nov. unlike that of P. cheliferus in which the telson is vestigial and reduced as two small lobes. This is only the second species described from this fascinating genus. See also generic remarks for Pseudelasmopus.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF85FF98FF50F9D4116BFA76.taxon	distribution	Distribution. South Pacific Ocean: Tasman Sea: Norfolk Island (current study).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF82FF9AFF50F9AA165BF909.taxon	materials_examined	Type material. Holotype male, 4.5 mm, dissected, 3 slides, AM P. 92677, Cape Leveque, Cygnet Bay, Western Australia, 16 ° 28 ' 40 " S, 123 ° 02 ' 06 " E, hand collected on snorkel, 10 April 2008, coll. J. Jelbart, MI WA 1068. Paratypes: 1 female, 4.5 mm, dissected, 3 slides, AM P. 92790 Cape Leveque, Cygnet Bay, Western Australia, 16 ° 28 ' 40 " S, 123 ° 02 ' 06 " E, hand collected on snorkel, 10 April 2008, coll. J. Jelbart, MI WA 1068; 12 specimens, AM P. 80976, Cape Leveque, Cygnet Bay, Western Australia, 16 ° 28 ' 40 " S, 123 ° 02 ' 06 " E, hand collected on snorkel, 10 April 2008, coll. J. Jelbart, MI WA 1068. Type locality. Cape Leveque, Cygnet Bay, Western Australia 1628 ' 40 " S, 12302 ' 06 " E.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF82FF9AFF50F9AA165BF909.taxon	etymology	Etymology. Named in honor of the Brown family, owners of the Cygnet Bay Pearl Farm, whose gracious support made collecting in Cape Leveque possible.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF82FF9AFF50F9AA165BF909.taxon	description	Description. Of holotype male, 4.5 mm AM P. 92677. Head. Eyes round; lateral cephalic lobe broad, apically truncate, anteroventral margin with excavate, anteroventral corner acute. Antenna 1 longer than antenna 2; peduncular article 1 shorter than article 2, with 1 long distal slender robust setae on posterior margin; peduncular article 2 longer than article 3; flagellum articles as long as broad, or broader than long, with 13 articles; accessory flagellum long, more than half length of primary flagellum, with 6 articles. Antenna 2 peduncular article 4 longer than article 5; flagellum with 8 articles. Mandible accessory setal row, well developed with 4 setae; palp well developed, article 1 length 1.9 × width, shorter than article 2, inner margin weakly produced rounded; article 2 length 0.9 × article 3, lined with long slender setae; article 3 rectilinear, long, 2.5 × as long as broad, longer than article 1, with 3 lateral and 3 apical slender setae. Pereon. Gnathopod 1 coxa anterior margin straight, anteroventral corner subquadrate, not produced, ventral margin without robust setae, posteroventral corner without notch; basis anterodistal corner without lobes; ischium anterodistal corner without lobes; merus with posteroventral corner subquadrate; carpus length twice width, 1.5 × propodus length, posterior margin lined with long setae, with rows of setae covering medial surface; propodus subovate, medial surface setal comb absent, palm subacute, weakly convex, entire, lined with slender setae, defined by posterodistal corner with 1 robust seta; dactylus closing along palm, unguis present. Gnathopod 2 subchelate; coxa posteroventral corner without notch, ventral margin without robust setae; basis broad, anterodistal corner lateral margin with well-developed lobe; ischium anterodistal corner without lobe; merus with subquadrate posteroventral corner; carpus compressed, length 0.6 × width, anterior margin with 1 robust seta, posterior margin with a few slender setae; propodus quadrate, length 1.1 × width, posterior margin with a few slender setae, palm length one third of propodus posterior margin, angle transverse, distal shelf subquadrate with 4 pairs of robust setae, palm with rounded medial excavation and subacute projection margin lined with robust setae, palm defined by posteroventral corner (90 ° angle, large subacute tooth and 1 robust seta; dactylus broad, falcate, posterior margin with medial sinus, reach end of and closing along palm. Pereopods 3 – 4 coxa subquadrate; basis long, length 3 × width; merus, carpus and propodus not broadened; dactylus unguis bicuspidate. Pereopods 5 – 7 basis expanded, posterior margin weakly serrate, posteroventral corner rounded; merus and carpus expanded; dactylus unguis bicuspidate. Pereopod 7 propodus expanded. Pleon. Pleonites 1 – 3 dorsally smooth. Epimera 1 – 3 posterior and ventral margins smooth, posteroventral corner with acute tooth. Uropod 1 peduncle with 1 basofacial seta, length 1.4 × outer ramus; inner ramus slightly longer than outer ramus. Uropod 2 peduncle shorter than outer ramus; inner ramus slightly shorter than outer ramus. Uropod 3 peduncle 0.9 × inner ramus; inner ramus shorter than outer ramus, length 1.6 × width, apically truncate with long robust setae. Telson as long as broad, deeply cleft (80 %), lobes weakly divergent, scarely tapering distally, lobes apically truncate, each lobe with 5 – 7 apical robust setae.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF82FF9AFF50F9AA165BF909.taxon	discussion	Remarks. The convex palm of gnathopod 2 and dactylus posterior margin with sinus align Q. brownorum sp. nov. with Q. gregoryi sp. nov also from Western Australia, as well as Q. inaequipes (A. Costa, 1851) from the Caribbean and Q. massavensis (Kossman, 1880) from north east Africa and female specimens of Q. santiniae Krapp-Schickel & Ruffo, 2006 from the isolated Amsterdam Island in the Indian Ocean. The more distally broad dactylus of gnathopod 2 in males and females separates Q. brownorum sp. nov. from the latter species. The telson apical margins are also more densely setose in Q. brownorum sp. nov with 5 long robust setae while Q. gregoryi and Q. massavensis have only 3 robust setae, Q. inaequipes 2 or 3 robust setae and only 2 robust setae in Q. santiniae.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF82FF9AFF50F9AA165BF909.taxon	distribution	Distribution. Western Australia: Cygnet Bay (current study).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF80FFA6FF50F8E210B8F85D.taxon	materials_examined	Type material. Holotype male, 5 mm, dissected, 3 slides, AM P. 92788, off end of South Mole, Arthur Head, Fremantle, Western Australia, 32 ° 03 ' S, 115 ° 44 ' E, 6 m, sponges, 25 December 1983, coll. R. T. Springthorpe, WA 276. Paratypes: B female, 7.2 mm, dissected, 1 slide, AM P. 92789, off end of South Mole, Arthur Head, Fremantle, Western Australia 32 ° 03 ' S, 115 ° 44 ' E, 6 m, sponges, 25 December 1983, coll. R. T. Springthorpe WA 276; 19 specimens 10 males, 4 gravid females and 5 juveniles, AM P. 79370, off end of South Mole, Arthur Head, Fremantle, Western Australia, 32 ° 03 ' S, 115 ° 44 ' E, 6 m, sponges, 25 December 1983, coll. R. T. Springthorpe, WA 276; 25 + specimens, AM P. 79308, off end of South Mole, Arthur Head, Fremantle, Western Australia 32 ° 03 ' S, 115 ° 44 ' E, 6 m, spirorbid tube worms, 25 December 1983, coll. J. K. Lowry, WA 283. Additional material examined. 31 specimens (1 male, 5 gravid females and 25 juveniles, AM P. 79413, 1 km west of Red Bluff, Kalbarri, Western Australia, 27 ° 42 ' S, 114 ° 09 ' E, 18 m, macroalga Ecklonia sp. bed, rocky bottom, Ecklonia sp. holdfasts, 9 January 1984, coll. R. T. Springthorpe, WA 453; 16 specimens, (1 male, 3 gravid females and 12 juveniles), AM P. 79409, inshore limestone reef, Neds Camp, Cape Range National Park, Western Australia, 21 ° 59 ' S, 113 ° 55 ' E, 1 m, green / brown algae, 2 January 1984, coll. R. T. Springthorpe, WA 373. Type locality. South Mole, Arthur Head, Fremantle, Western Australia, 3203 ' S, 11544 ' E.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF80FFA6FF50F8E210B8F85D.taxon	etymology	Etymology. Name for the explorers Sir Augustus Charles Gregory and his brother Francis Thomas Gregory, who conducted survey expeditions along the Western Australian coast.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFF80FFA6FF50F8E210B8F85D.taxon	description	Description. Of holotype male, 5 mm, AM P. 92788. Head. Eyes ovate; lateral cephalic lobe broad, apically truncate, anteroventral margin with excavate, anteroventral corner acute. Antenna 1 longer than antenna 2; peduncular article 1 shorter than article 2, with 3 robust setae on posterior margin; peduncular article 2 longer than article 3; flagellum articles as long as broad, or longer than broad, with 9 articles; accessory flagellum long, more than half length of primary flagellum, with 8 articles. Antenna 2 peduncular article 4 longer in length than article 5; flagellum with 8 articles. Mandible accessory setal row, well developed with 5 setae; palp well developed, article 1 length 1.2 × width, shorter than article 2, inner margin distally produced rounded; article 2 length 0.9 × article 3, lined with long slender setae; article 3 rectilinear, long, 2.8 × as long as broad, longer than article 1, with 5 lateral and 2 apical slender setae. Lower lip inner lobes present; outer lobes without apical ducts, mandibular lobes apically rounded to subacute (not illustrated). Maxilla 1 inner plate with 3 slender setae. Pereon. Gnathopod 1 coxa anterior margin concave, anteroventral corner produced, acute, ventral margin without robust setae, posteroventral corner without notch; basis anterodistal corner without lobes; ischium anterodistal corner without lobes; merus with posteroventral corner subquadrate; carpus length twice width, 1.2 × propodus length, posterior margin lined with long setae, with rows of setae covering medial surface; propodus subovate, medial surface setal comb absent, palm subacute, weakly convex, entire, lined with slender setae, defined by posterodistal corner with 3 robust setae; dactylus posterior margin lined with setules, closing along palm, unguis present. Gnathopod 2 subchelate; coxa posteroventral corner without notch, ventral margin without robust setae; basis broad, anterodistal corner with pair of well-developed rounded lobes; ischium anterodistal corner with pair of well-developed rounded lobes; merus with acute posteroventral corner; carpus compressed, length 0.7 × width, anterior margin with 2 robust setae, posterior margin a few slender setae; propodus quadrate, length 1.1 × width, anterior and posterior margin with clusters of long slender setae, palm length less than one third of propodus posterior margin, angle transverse, distal shelf subquadrate with 3 pairs of robust setae, palm	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFBBFFA1FF50FBF914E9F804.taxon	materials_examined	Material examined. Male, 4.6 mm, dissected 4 slides, AM P. 92526, south end of McCluer Island, Northern Territory, 11 ° 6 ' S, 133 ° 00 ' E, 0 m, red algae, 18 October 1982, coll. J. K. Lowry, NT 63; 1 male, 4 mm, dissected, 3 slides, AM P. 92527, south end of McCluer Island, Northern Territory, 11 ° 6 ' S, 133 ° E, 0 m, red algae, 18 October 1982, coll. J. K. Lowry NT 63; 1 c female, 4.2 mm, dissected, 1 slide, AM P. 92528, south end of McCluer Island, Northern Territory, 11 ° 6 ' S, 133 ° 00 ' E, 0 m, red algae, 18 October 1982, coll. J. K. Lowry, NT 63; 10 specimens, AM P. 78319, south end of McCluer Island, Northern Territory, 11 ° 6 ' S, 133 ° E, 0 m, red algae, 18 October 1982, coll. J. K. Lowry, NT 63; 2 specimens, AM P. 77903, reef at west end of Oxley Island, 1100 ' S, 13249 ' E, 5 m, red sponge on dead hard coral Acropora sp., 20 October 1982, coll. J. K. Lowry, NT 79; 1 juvenile specimen, AM P. 78318, reef on north side of New Year Island, 10 ° 54 ' S, 133 ° 02 ' E, 2 m, red algae, 13 October 1982, coll. G. C. B. Poore, NT 1. Type locality. East of Metinaro, Secret Garden Reef, Timor-Leste, 8 ° 29 ' 15 " S, 125 ° 49 ' 53 " E.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFBBFFA1FF50FBF914E9F804.taxon	description	Variation. Adult male of 4.6 mm (AM P. 92526) with well developed lobes on the gnathopod 2 basis, which are not present on smaller male and female specimens of 4 mm and 4.2 mm body length (AM P. 92527 and AM P. 92528).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFBBFFA1FF50FBF914E9F804.taxon	discussion	Remarks. Specimens from the Northern Territory have only three robust setae on each telsonic lobe, for both juveniles and adults, in contrast to the type material from Timor-Leste, which has four robust setae. With only one seta difference the, material is attributed to Q. metinaro. Quadrimaera mirandelle Appadoo & Myers, 2002 from Mauritius has a similar gnathopod 2 propodus palm to Q. metinaro, and the telson has 5 robust setae. With Q. metinaro from Northern Territory presenting an intermediate telson setal number the possibility that these latter species is a junior synonym was considered, but the gnathopod 2 dactylus is more medially broad in Q. mirandelle.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFBBFFA1FF50FBF914E9F804.taxon	distribution	Distribution. Timor-Leste (Hughes 2015). Australia: Northern Territory: McCluer Island, Oxley Island, New Year Island (current study).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFBAFFADFF50FAF31696FEAA.taxon	materials_examined	Material examined. Cocos (Keeling) Islands: 1 male, 5.6 mm, dissected, 4 slides, AM P. 98053, dive site ‘ Winter Wall’, near Horsburgh Island, 12 ° 05 ' 15 " S, 96 ° 50 ' 14 " E, 8 m, rubble and green calcareous alga Halimeda sp., 12 October 2008, coll. J. K. Lowry, MI WA 836; 1 specimen, AM P. 98054, dive site ‘ Winter Wall’, near Horsburgh Island, 12 ° 05 ' 15 " S, 96 ° 50 ' 14 " E, 8 m, rubble and green calcareous alga Halimeda sp., 12 October 2008, coll. J. K. Lowry, WA 836; 1 female, dissected, 4 slides, AM P. 98055, dive site " Surf Shack Drift ", western side of West Island, 12 ° 9 ' S, 96 ° 48 ' E, 10 m, rubble with epiphytes, 15 October 2008, coll. J. K. Lowry and L. E. Hughes, MI WA 859; 2 specimens, AM P. 82194, dive site ‘ Winter Wall’, near Horsburgh Island, 12 ° 5 ' 15 " S, 96 ° 50 ' 13 " E, coral rubble, 12 October 2008, coll. K. B. Attwood, MI WA 835; 1 specimen, AM P. 82195, northern side of Direction Island, 12 ° 5 ' 4 " S, 96 ° 52 ' 54 " E, fine sediment, 8 October 2008, coll. J. K. Lowry and K. B. Attwood, MI WA 784; 1 specimen, AM P. 82197, centre of lagoon, 128 ° ' 49 " S, 96 ° 51 ' 13 " E, Acropora sp. coral dead with red epiphytic algae, 14 October 2008, coll. K. B. Attwood and L. E. Hughes, MI WA 845; 1 specimen, AM P. 82196, north east corner of the lagoon, 12 ° 6 ' 17 " S, 96 ° 51 ' 11 " E, seagrass base and stems based covered in epiphytes, seagrass beds Thalassia hemprichii, 10 October 2008, coll. J. K. Lowry MI WA 810; 2 specimens, AM P. 82193, dive site " Two Trees ", western side of West Island, 12 ° 9 ' 34 " S, 96 ° 48 ' 59 " E, 15 October 2008, coral Acropora sp. plate rubble, coll. L. E. Hughes and K. B. Attwood, MI WA 854. Western Australia: 1 male, 6.5 mm, dissected, 3 slides, AM P. 91142, Shenton Bluff, Cape Leveque, Cygnet Bay, 16 ° 28 ' 38 " S, 123 ° 02 ' 38 E, 5.1 m, green calcareous alga Halimeda sp., 22 May 2010, coll. K. B. Attwood, MI WA 1076; 1 female specimen (damaged gnathopods), AM P. 91141, Shenton Bluff, Cape Leveque, Cygnet Bay, 16 ° 28 ' 38 " S, 123 ° 02 ' 38 E, 5.1 m, green calcareous alga Halimeda sp., 22 May 2010, coll. K. B. Attwood, MI WA 1076. Type locality. La Cuvette, Mauritius, 20 ° 00 ' S, 57 ° 34.2 ' E.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFBAFFADFF50FAF31696FEAA.taxon	discussion	Remarks. These records extend the distribution of Q. micheli further east into the Indian Ocean, from Mauritius to Cocos (Keeling) Islands and Western Australia.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFBAFFADFF50FAF31696FEAA.taxon	distribution	Distribution. Indian Ocean: Cocos (Keeling) Islands (current study). Western Australia: Cygnet Bay (current study). Mauritius (Appadoo et al. 2002).? Madagascar (Ledoyer 1982).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFB7FFACFF50FACA1729F842.taxon	materials_examined	Material examined. Queensland: many specimens, AM P. 75779, 200 m off the beach on the southern side of Thursday Island, Torres Strait, 10 ° 35 ' 17 " S, 142 ° 12 ' 58 " E, 2 m, low turfing brown algae on mooring rope, 30 September 2006, coll. J. K. Lowry and M. Capa, QLD 1894; 17 specimens, AM P. 75780, Sunken reef, north of Goodes Island, Torres Strait, 10 ° 31 ' 29 " S, 14 ° 23 ' 18 " E, 12 m, short complex red algae, 29 September 2006, coll. M. Capa and J. K. Lowry, QLD 1860; 7 specimens, AM P. 75791, Sunken reef, north of Goodes Island, Torres Strait, 10 ° 31 ' 29 " S, 14 ° 23 ' 18 " E, 10 m, plates of coral rubble, 29 September 2006, coll. M. Capa and J. K. Lowry, QLD 1868; 1 specimen, AM P. 75793, Sunken reef, north of Goodes Island, Torres Strait, 10 ° 31 ' 29 " S, 14 ° 23 ' 18 " E, 12 m, rocks with 15 cm tall brown alga Sargassum sp., 29 September 2006, coll. M. Capa and J. K. Lowry, QLD 1866; 3 specimens, AM P. 75794, Sunken reef, north of Goodes Island, Torres Strait, 10 ° 31 ' 29 " S, 14 ° 23 ' 18 " E, 12 m, large sticks of coral rubble, 29 September 2006, coll. M. Capa and J. K. Lowry, QLD 1863; 1 specimen, AM P. 75786, Number One reef, Katai Nab, North of Horn Island, Torres Strait, 10 ° 32 ' 30 " S, 142 ° 10 ' 15 " E, 6 m, submerged rope fibre caught on reef, encrusted with red epiphytic algae on tips, 28 September 2006, coll. J. K. Lowry, QLD 1846; 5 specimens, AM P. 75785, Number One reef, Katai Nab, North of Horn Island, Torres Strait, 10 ° 32 ' 30 " S, 142 ° 10 ' 15 " E, 8 m, large brick covered in fine sediment, epiphytes, brown alga Padina sp., low turfing algae, 28 September 2006, coll. J. K. Lowry and L. E. Hughes, QLD 1851; 2 specimens, AM P. 75781, Number One reef, Katai Nab, North of Horn Island, Torres Strait, 10 ° 32 ' 30 " S, 142 ° 10 ' 15 " E, 5.6 m, coarse coral rubble, 28 September 2006, coll. L. E. Hughes, QLD 1849; 2 specimens, AM P. 75790, Number One reef, Katai Nab, north of Horn Island, Torres Strait, 10 ° 32 ' 30 " S, 142 ° 10 ' 15 " E, 7.8 m, brown alga Sargassum sp. 10 – 15 cm tall, 28 September 2006, coll. L. E. Hughes, QLD 1848; 5 specimens, AM P. 75789, drop off at the north western corner of Hammond Island, Torres Strait, 10 ° 32 ' 24 " S, 142 ° 11 ' 19 " E, 15 m, red macrophyte, 28 September 2006, coll. J. K. Lowry and L. E. Hughes, QLD 1882; 1 specimen, AM P. 75782, western side of Goodes Island, Torres Strait, 10 ° 34 ' 9 " S, 14 ° 28 ' 51 " E, 9 m, fine sediment, 2 October 2006, coll. J. K. Lowry and L. E. Hughes QLD 1924; 7 specimens, AM P. 75783, north-west corner of Goodes Island, Torres Strait, 10 ° 33 ' 34 " S, 14 ° 29 ' 8 " E, 5 m, live rock, 29 September 2006, coll. M. Capa and L. E. Hughes QLD 1877; 2 specimens, AM P. 75787, Northwest Islet, Torres Strait, 10 ° 18 ' 54 " S, 14 ° 25 ' 26 " E, 5 m, dead red algae, 1 October 2006, coll. L. E. Hughes and M. Capa, QLD 1911; 1 specimen, AM P. 75792, Northwest Islet, Torres Strait, 10 ° 18 ' 54 " S, 14 ° 25 ' 26 " E, 3.8 m, plastic fabric attached to dead coral covered in epiphytes, 1 October 2006, coll. M. Capa and L. E. Hughes, QLD 1918; 2 specimens, AM P. 75856, near shore, western side of Hawkesbury Island, Torres Strait, 10 ° 21 ' 44 " S, 14 ° 27 ' 4 " E, 1.7 m, coral rubble, 1 October 2006, coll. J. K. Lowry and M. Capa, QLD 1909; 2 specimens, AM P. 75788, near shore, western side of Hawkesbury Island, Torres Strait, 10 ° 21 ' 44 " S, 14 ° 27 ' 4 " E, 8 m, soft sediment, 1 October 2006, coll. J. K. Lowry and M. Capa, QLD 1900; 4 specimens, AM P. 75858, largest bommie in lagoon at the ' Entrance', One Tree Island, 23 ° 29 ' 16 " S, 15 ° 24 ' 46 " E, 4.2 m, very fine silty sediment, 24 October 2006, coll. L. E. Hughes andI. Takeuchi, QLD 1936; 2 specimens, AM P. 75768, largest bommie in lagoon at the ' Entrance', One Tree Island, 23 ° 29 ' 16 " S, 15 ° 24 ' 46 " E, 4.5 m, large sticks of coral rubble, 25 October 2006, coll. J. K. Lowry and L. E. Hughes, QLD 1949; 1 specimen, AM P. 75766, north of No Tree Island, One Tree Island, 23 ° 29 ' 47 " S, 15 ° 23 ' 25 " E, 2.1 m, soft sediment, 25 October 2006, coll. I. Takeuchi and L. E. Hughes, QLD 1955; 1 specimen, AM P. 75857, outside barrier reef near the ' Entrance', One Tree Island, 23 ° 29 ' 2 " S, 15 ° 25 ' 22 " E, 10 m, large brown hydroid, 26 October 2006, coll. I. Takeuchi and L. E. Hughes, QLD 1962; 1 specimen, AM P. 75773, outside barrier reef near the ' Entrance', One Tree Island, 23 ° 29 ' 2 " S, 15 ° 25 ' 22 " E, 10 m, mixed red algae, 26 October 2006, coll. I. Takeuchi andL. E. Hughes, QLD 1960; 1 specimen, AM P. 75769, largest bommie in lagoon at the ' Entrance', One Tree Island, 23 ° 29 ' 16 " S, 15 ° 24 ' 46 " E, 4 m, red algae, 25 October 2006, coll. J. K. Lowry and L. E. Hughes, QLD 1950; 11 specimens, AM P. 75775, largest bommie in lagoon at the ' Entrance', One Tree Island, 23 ° 29 ' 16 " S, 15 ° 24 ' 46 " E, 4 m, coral rubble with coarse sand, coll. J. K. Lowry and L. E. Hughes, 25 October 2006, QLD 1950; many specimens, AM P. 75767, centre bommies, First lagoon, One Tree Island, 23 ° 30 ' 12 " S, 15 ° 25 ' 19 " E, 1.7 m, dead coral with epiphytes, 29 October 2006, coll. L. E. Hughes and J. K. Lowry, QLD 2006; 10 specimens, AM P. 75774, channel on western side, One Tree Island, 23 ° 30 ' 12 " S, 15 ° 25 ' 19 " E, 2 m, coral rubble, 28 October 2006, coll. I. Takeuchi and L. E. Hughes, QLD 1988; 1 specimen, AM P. 75777, outer reef slope, south of One Tree Island, 23 ° 30 ' 47 " S, 15 ° 25 ' 10 " E, 13.8 m, large sticks of coral rubble, 28 October 2006, coll. I. Takeuchi and L. E. Hughes, QLD 1994; many specimens, AM P. 75776, outer reef, Steves bommie, near Two Trees Islet, One Tree Island, 23 ° 29 ' 3 " S, 15 ° 25 ' 27 " E, 13 m, coral rubble with epiphytes and solitary ascidians, 27 October 2006, coll. L. E. Hughes and J. K. Lowry, QLD 1978; 2 specimens, AM P. 75772, outer reef, Steves bommie, near Two Trees Islet, One Tree Island, 23 ° 29 ' 3 " S, 15 ° 25 ' 27 " E, 13.8 m, low turfing algae and fine sediment, 27 October 2006, coll. L. E. Hughes and J. K. Lowry, QLD 1975; 8 specimens, AM P. 75778, outer reef, Steves bommie, near Two Trees Islet, One Tree Island, 23 ° 29 ' 3 " S, 15 ° 25 ' 27 " E, 13.2 m, coral rubble with algae, 27 October 2006, coll. L. E. Hughes and J. K. Lowry, QLD 1974; 1 specimen, AM P. 75770, outer reef, north of Third Lagoon, One Tree Island, 23 ° 29 ' 4 " S, 15 ° 24 ' 7 " E, 18 m, coral rubble, 27 October 2006, coll. I. Takeuchi and J. K. Lowry, QLD 1982; 1 specimen, AM P. 75771, outer reef, north of Third Lagoon, One Tree Island, 23 ° 29 ' 4 " S, 15 ° 24 ' 7 " E, 18 m, fine sediment, 27 October 2006, coll. I. Takeuchi and J. K. Lowry, QLD 1979. New Zealand: 10 specimens, AM P. 90951, west side of North Chanter Island, Herald Islands, Kermadec Islands, 29 ° 15 ' 6 " S, 177 ° 51 ' 21 " W, 10 m, 16 May 2011, coll. A. Ballance and S. J. Keable, K 2011 - 42. Type locality. Makin, Kiribati, ~ 140 ' S, 17922 ' W.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFB7FFACFF50FACA1729F842.taxon	distribution	Distribution. South Pacific Ocean. Australia: Queensland: Torres Strait, One Tree Island (current study); New Zealand: Kermadec Islands (current study); Fiji (Schellenberg 1938; Myers 1985). North Pacific Ocean: Hawaii (Schellenberg 1938; J. L. Barnard 1970); Kiribati (Schellenberg 1938). Indian Ocean: Madagascar (Ledoyer 1972); Mauritius (Appadoo et al. 2002).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFB5FFAEFF50FF721160FE66.taxon	materials_examined	Material examined. Cocos (Keeling) Islands: 1 specimen, AM P. 82199, Turks Reef, 12 ° 06 ' 44 " S, 96 ° 49 ' 43 " E, 3.8 m, sticks of rubble and algae, coll. L. E. Hughes, 10 October 2008, MI WA 818; 2 specimens, AM P. 82198, Turks Reef, 12 ° 06 ' 44 " S, 96 ° 49 ' 43 " E, 3.8 m, sticks of rubble and algae, coll. L. E. Hughes, 10 October 2008, MI WA 818. Western Australia: 6 specimens, WAM C 45875, Cassini Island, The Kimberley, 13 ° 57 ' S 12 ° 537 ' E, 2 m, 18 October 2010, coll. Andrew Hosie, Woodside Kimberley Survey 2010, 37 / K 10 - Q 1; 4 specimens, WAM C 46382, Cassini Island, The Kimberley, 13 ° 57 ' S 125 ° 37 ' E, 2 m, 18 October 2010, coll. Lee Paterson, Woodside Kimberley Survey 2010, 37 / K 10 - Adhoc; 3 specimens, WAM C 47507, Cassini Island, The Kimberley, 13 ° 57 ' S 125 ° 37 ' E, 2 m, coarse coral rubble, 18 October 2010, coll. Andrew Hosie, Woodside Kimberley Survey 2010, 37 / K 10 - Q 1; 12 specimens, WAM C 45893, Long Reef, The Kimberley, 13 ° 48 ' S 125 ° 49 ' E, 2 m, 13 October 2010, coll. Lexie Walker, Woodside Kimberley Survey 2010, 52 / K 10 - Q 3; 4 specimens, AM P. 98056, reef front, south of Tantabiddy, Ningaloo Reef, 21 ° 54 ' 37 " S 113 ° 55 ' 42 " E, 9 m, coarse coral rubble, 12 June 2008, coll. Neil. L. Bruce and M. Blazewicz-Paszkowycz, MI WA 973, NIN 10 c; 5 specimens, AM P. 98057, reef front, south of Tantabiddy, Ningaloo Reef, 21 ° 55 ' 42 " E 113 ° 55 ' 11 " E, 13 m, coarse coral rubble, 17 June 2008, coll. Neil. L. Bruce and M. Blazewicz-Paszkowycz, MI WA 979, NIN 17 a; 4 specimens, AM P. 91143, Ningaloo Reef, 22 ° 39 ' 30 " S, 113 ° 37 ' 05 " E, 6.8 m, 25 May 2009, coll. L. E. Hughes, CReefs Ningaloo 2009, NR 09 - 60; 4 specimens, AM P. 91143, Ningaloo Reef, 22 ° 39 ' 30 " S, 113 ° 37 ' 5 " E, 6.8 m, dead coral head, 25 May 2009, coll. L. E. Hughes, MI WA 1054; 4 specimens, AM P. 79349, inside of outer Ningaloo Reef, off Neds Camp, Cape Range National Park, 21 ° 59 ' 30 " S, 113 ° 54 ' 30 " E, 2 m, small brown dictyotalean, 1 January 1984, coll. R. T. Springthorpe and J. K. Lowry WA 342; many specimens, AM P. 83829, Riddell Point, Cygnet Bay, Cape Leveque, 16 ° 27 ' 52 " S, 12 ° 31 ' 10 " E, 6.8 m, coral rubble, filamentous red turfing alga, green alga Halimeda sp., 24 May 2010, coll. K. B. Attwood, MI WA 1102; 1 specimen, AM P. 83830, Riddell Point, Cygnet Bay, Cape Leveque, 16 ° 27 ' 52 " S, 12 ° 31 ' 10 " E, 6.8 m, rubble, 24 May 2010, coll. J. K. Lowry, MI WA 1105; many specimens, AM P. 83832, Riddell Point, Cygnet Bay, Cape Leveque, 16 ° 27 ' 52 " S, 12 ° 31 ' 10 " E, 9.3 m, dead coral Seriatopora hystrix, 25 May 2010, coll. K. B. Attwood, MI WA 1125; 4 specimens, AM P. 83833, Riddell Point, Cygnet Bay, Cape Leveque, 1627 ' 52 " S, 1231 ' 10 " E, 9.9 m, rubble with red coralline algae and sponge, 25 May 2010, coll. K. B. Attwood, MI WA 1126; many specimens, AM P. 83831, Riddell Point, Cygnet Bay, Cape Leveque, 16 ° 27 ' 52 " S, 12 ° 31 ' 10 " E, 9.8 m, tuft of red algae, 25 May 2010, coll. L. E. Hughes, MI WA 1121; many specimens, AM P. 83834, Bird Rock, Cygnet Bay, Cape Leveque, 16 ° 29 ' 4 " S, 123 ° 35 " E, 12 m, dead shell bottom, 26 May 2010, coll. J. K. Lowry, MI WA 1130; 2 specimens, AM P. 83835, Bird Rock, Cygnet Bay, Cape Leveque, 16 ° 29 ' 4 " S, 123 ° 35 " E, 10 m, coarse sediment, 26 May 2010, coll. K. B. Attwood, MI WA 1135; 7 specimens, AM P. 83828, Mission Beach at bar, Cygnet Bay, Cape Leveque, 16 ° 27 ' 38 " S, 123 ° 00 ' 40 " E, 7.8 m, live rock, 24 May 2010, coll. L. E. Hughes, MI WA 1097. Queensland: many specimens, AM P. 75797 near shore, western side of Hawkesbury Island, Torres Strait, 10 ° 21 ' 44 " S, 14 ° 27 ' 4 " E, 8 m, coral rubble with sponges, 1 October 2006, coll. J. K. Lowry and M. Capa, QLD 1905; 3 specimens, AM P. 75795, outer reef slope, south of One Tree Island, 23 ° 30 ' 47 " S, 15 ° 25 ' 10 " E, 16 m, red algal tufts, 28 October 2006, coll. I. Takeuchi and L. E. Hughes, QLD 1992; many specimens, AM P. 75796, outer reef, Steves bommie, near Two Trees Islet, One Tree Island, 23 ° 29 ' 3 " S, 15 ° 25 ' 27 " E, 13 m, red coralline alge with epiphytes, Dictyota sp. and Halimeda sp., 27 October 2006, coll. L. E. Hughes and J. K. Lowry, QLD 1977; 1 specimen, AM P. 52786, Middle Reef, North Stradbroke Island, 27 ° 24 ' 24 " S, 153 ° 32 ' 00 " E, 18 m, red algae, brown alga Zonaria sp., crinoid, sponge hand collected, on scuba, 4 June 1993, coll. A. R. Parker, QLD 867. New South Wales: many specimens, AM P. 92461, Korffs Islet, Solitary Islands, 30 ° 19 ' 8 " S, 15 ° 39 ' 12 " E, depth unknown, 2002, coll. L. E. Hughes; many specimens, AM P. 92460, Muttonbird Island, Coffs Harbour, 30 ° 17 ' S, 153 ° 10 ' E, depth unknown, 2002, coll. L. E. Hughes. Type locality. Fiji, ~ 1740 ' S, 17831 ' E.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFB5FFAEFF50FF721160FE66.taxon	distribution	Distribution. Australia. Queensland: Lizard Island, Torres Strait, North Stradbroke Island (Berents 1983; Krapp-Schickel 2009; current study). New South Wales: Korffs Islet, Muttonbird Island (current study). Indian Ocean. Western Australia: Cassini Island, Long Reef, The Kimberley; Ningaloo Reef (current study). Cocos (Keeling) Islands (current study). Madagascar (Ledoyer 1972, 1983). India (Sivaprakasam 1968, 1970). Pacific Ocean. Marshall Islands and Kiribati (Schellenberg 1938; J. L. Barnard 1965). Fiji (Dana 1853; Schellenberg 1938, Myers 1985). North Pacific Ocean. Hawaii (Schellenberg 1938; J. L. Barnard 1970).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFB4FFA9FF50FB0E1482FF4E.taxon	materials_examined	Material examined. Western Australia. 3 specimens, AM P. 91144, Ningaloo Reef, 22 ° 39 ' 30 " S, 113 ° 37 ' 05 " E, 6.8 m, 25 May 2009, coll. L. E. Hughes, CReefs Ningaloo 2009, NR 09 - 60; many specimens, AM P. 83916, Cygnet Bay Pearl Farm outside lines, Cygnet Bay, Cape Leveque, 16 ° 28 ' 49 " S, 12 ° 32 ' 15 " E, epiphytic growth from pearl shell panels, 23 May 2010, coll. J. K. Lowry and K. B. Attwood, MI WA 1094; many specimens, AM P. 83917, Cygnet Bay Pearl Farm inside lines, Cygnet Bay, Cape Leveque, 16 ° 29 ' 43 " S, 12 ° 31 ' 46 " E, 2 m, epiphytic growth from pearl shell panels, 23 May 2010, coll. J. K. Lowry and K. B. Attwood, MI WA 1095. New South Wales. 1 specimen, AM P. 78432, Park Beach Bommie, east of Coffs Harbour, 30 ° 17 ' 42 " S, 153 ° 12 ' E, 14 m, bryozoan Biflustra perfragilis, hand collected on SCUBA, 3 May 2005, coll. K. B. Attwood, NSW 2827. Queensland: 1 specimen, AM P. 75798, centre bommie, First Lagoon, One Tree Island, 23 ° 30 ' 12 " S, 15 ° 25 ' 19 " E, 1.7 m, 29 October 2006, dead coral with epiphytes, coll. L. E. Hughes and J. K. Lowry, QLD 2006. Type locality. Espiritu Santo Island, Gulf of California.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFB4FFA9FF50FB0E1482FF4E.taxon	distribution	Distribution. South Pacific Ocean: Australia: New South Wales: Coffs Harbour (current study); Queensland: Lizard Island, One Tree Island (Berents 1983; current study). Indian Ocean: Western Australia: Ningaloo Reef, Cygnet Bay (current study). Eastern Pacific Ocean: United States of America: Gulf of California; Galapagos Islands (J. L. Barnard 1979); Ecuador (J. L. Barnard 1979); Mexico: Gulf of Tehuantepec (J. L. Barnard 1979; Garcia-Madrigal 2010).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFB3FFABFF50FBC010A7FA2A.taxon	materials_examined	Material examined. Cocos (Keeling) Islands. 1 specimen, AM P. 82202, dive site " Rose Wall ", near Horsburgh Island, 12 ° 05 ' 45 " S, 96 ° 50 ' 32 " E, 13.8 m, green calcareous alga Halimeda sp., coll. K. B. Attwood, 12 October 2008, MI WA 827; 7 specimens, AM P. 82203, dive site " Rose Wall ", near Horsburgh Island, 12 ° 05 ' 45 " S, 96 ° 50 ' 32 " E, 11 m, rubble some green calcareous alga Halimeda sp., coll. L. E. Hughes, 12 October 2008, MI WA 830; 1 specimen, AM P. 82200, off West Island Jetty, 12 ° 05 ' 15 " S, 96 ° 50 ' 14 " E, 1.5 m, rubble, coll. L. E. Hughes, 14 October 2008, MI WA 843; 1 specimen, AM P. 82201, off West Island Jetty, 12 ° 05 ' 15 " S, 96 ° 50 ' 14 " E, 0 m, brown alga Turbinaria sp. as floating wrack, coll. L. E. Hughes, 14 October 2008, MI WA 844; 2 specimens, AM P. 82204, off West Island Jetty, 12 ° 05 ' 15 " S, 96 ° 50 ' 14 " E, 2 m, brown alga Padina sp., coll. L. E. Hughes and J. K. Lowry, 14 October 2008, MI WA 841. Western Australia. 2 specimens, WAM C 47855, Long Reef, The Kimberley, 13 ° 53 ' S, 125 ° 44 ' E, 2 m, 20 October 2010, coll. Andrew Hosie, Woodside Kimberley Survey 2010, 44 / K 10 - T 1; 1 specimen, WAM C 47856, Cassini Island, The Kimberley, 13 ° 55 ' S, 125 ° 37 ' E, 2 m, 16 October 2010, coll. Andrew Hosie, Woodside Kimberley Survey, 32 / K 10 - T 1; 1 specimen, AM P. 98058, inner lagoon, Ningaloo Reef, 21 ° 53 ' 9 " S, 113 ° 59 ' 26 " E, 1.5 m, brown alga Lobophora sp., 18 June 2008, coll. L. E. Hughes MI WA 987, NR 18; 1 specimen, AM P. 83824, Shenton Bluff, Cygnet Bay, Cape Leveque, 16 ° 28 ' 37 " S, 12 ° 32 ' 38 " E, 5.1 m, brown algae Sargassum sp. on rubble, 22 May 2010, coll. K. B. Attwood, MI WA 1080; 2 specimens, AM P. 83825, emerging lagoon at low tide near Shenton Bluff, Cygnet Bay, Cape Leveque, 16 ° 28 ' 58 " S, 12 ° 32 ' 40 " E, 6.9 m, brown algae Sargassum sp., 23 May 2010, coll. K. B. Attwood, MI WA 1090; 3 specimens, AM P. 83826, Riddell Point, Cygnet Bay, Cape Leveque, 16 ° 27 ' 59 " S, 12 ° 31 ' 41 " E, 9.9 m, rubble, 25 May 2010, coll. K. B. Attwood, MI WA 1127; 2 specimens, AM P. 83827, Jackson Island site A, Cape Leveque, 16 ° 25 ' 19 " S, 12 ° 35 ' 19 " E, 5.8 m, rubble, 27 May 2010, coll. L. E. Hughes, MI WA 1148. Northern Territory: 2 specimens, AM P. 78314, patch reef on north side of New Year Island, 10 ° 54 ' S, 13 ° 32 ' E, 10 m, hydroids on coral, 14 October 1982, coll. G. C. B. Poore, NT 17; 1 specimen, AM P. 78316, south end of McCluer Island, 11 ° 06 ' S, 133 ° E, 8 m, had coral Acropora sp. base, 17 October 1982, coll. P. Horner, NT 59; 13 specimens, AM P. 78315, bommies at north-west end of McCluer Island, 11 ° 2 ' S, 132 ° 58 ' E, 8 m, Stylophora bases, 16 October 1982, coll. P. Horner, NT 36; 11 specimens, AM P. 78317, south end of McCluer Island, 11 ° 6 ' S, 133 ° 00 ' E, 8 m, coral Seriatopora hystrix, 17 October 1982, coll. J. K. Lowry, NT 61. Queensland: 1 specimen, AM P. 75800, largest bommie in lagoon at the ' Entrance', One Tree Island, 23 ° 29 ' 16 " S, 15 ° 24 ' 46 " E, 4.1 m, 25 October 2006, coral rubble, coll. J. K. Lowry and L. E. Hughes, QLD 1946; many specimens, AM P. 75799, centre bommie, First Lagoon, One Tree Island, 23 ° 30 ' 12 " S, 15 ° 25 ' 19 " E, 1.7 m, dead coral with epiphytes, 29 October 2006, coll. L. E. Hughes and J. K. Lowry, QLD 2006; many specimens, AM P. 80624, Cockle Bay, Magnetic Island, 19 ° 11 ' S, 146 ° 49 ' E, depth unknown, seagrasses Cymodocea serrulata, Halodule uninervis and Halophila ovalis, 17 February 1999, coll. D. W. Klumpp and S. N. Kwak, Original Vial Label Data: 290499; many specimens, AM P. 80625, Cockle Bay, Magnetic Island, 19 ° 11 ' S, 146 ° 49 ' E, depth unknown, seagrasses Cymodocea serrulata, Halodule uninervis and Halophila ovalis, 17 February 1999, coll. D. W. Klumpp and S. N. Kwak, Original Vial Label Data: 170299. Norfolk Island: 1 male, 9 mm, dissected, 3 slides, AM P. 92512, Fig Valley reef, 29 ° 3 ' 20 " S, 167 ° 55 ' 44 " E, 18.7 m, red coralline algae Amphiroa anceps and other mixed red algae, 19 May 2008, coll. L. E. Hughes, MI NFK 71; 1 b female, 6.4 mm, dissected, 1 slide, AM P. 92513, Fig Valley reef, 29 ° 3 ' 20 " S, 167 ° 55 ' 44 " E, 18.7 m, red coralline algae Amphiora anceps and other mixed red algae, 19 May 2008, coll. L. E. Hughes, MI NFK 71; 1 c male, 6 mm, AM P. 92514, Fig Valley reef, 29 ° 3 ' 20 " S, 167 ° 55 ' 44 " E, 18.7 m, red coralline algae Amphiora anceps and other mixed red algae, 19 May 2008, coll. L. E. Hughes, MI NFK 71; 3 specimens, AM P. 81503, Fig Valley reef, 29 ° 3 ' 20 " S, 167 ° 55 ' 44 " E, 18.7 m, red coralline algae Amphiroa anceps and other mixed red algae, 19 May 2008, coll. L. E. Hughes, MI NFK 71; 1 specimen, AM P. 81505, Fig Valley reef, 29 ° 3 ' 20 " S, 167 ° 55 ' 44 " E, 20 m, red sponge like algae, 19 May 2008, coll. L. E. Hughes, MI NFK 69; 10 specimens, AM P. 81502, dive site ‘ The Wall’, Nepean Island, 29 ° 4 ' 17 " S, 167 ° 57 ' 40 " E, 17 m, green algae Halimeda sp., 15 May 2008, coll. J. K. Lowry, MI NFK 45. Lord Howe Island. Many specimens, AM P. 98059, north end of Old Settlement Beach, 31 ° 31 ' 12 " S, 15 ° 93 ' 36 " E, under intertidal rocks, 11 May 1977, coll. G. D. Fenwick and J. K. Lowry, LHA 13. Type locality. Kiribati, 140 ' S, 17922 ' W.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFB3FFABFF50FBC010A7FA2A.taxon	distribution	Distribution. South Pacific Ocean: Australia: Queensland: Lizard Island, One Tree Island, Magnetic Island, (Berents 1983; Krapp-Schickel 2009; current study); Northern Territory: New Year Island, McCluer Island (current study); Tasman Sea: Norfolk Island; Lord Howe Island (current study). Marshall Islands and Kiribati (Schellenberg 1938; J. L. Barnard 1965); Fiji (Schellenberg 1938; Myers 1985). Indian Ocean: Australia: Western Australia: Long Reef, Cassini Island, The Kimberley; Cape Leveque (current study); Cocos (Keeling) Islands (current study); Mauritius (Appadoo et al. 2002); Madagascar (Ledoyer 1967, 1983); South Africa (Griffiths 1973); Red Sea (Ruffo 1969); India (Sivaprakasam 1968). North Pacific Ocean: Hawaii (Schellenberg 1938; J. L. Barnard 1970).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFB1FFB5FF50FA0215C7F873.taxon	materials_examined	Type material. Holotype male, 4.0 mm, dissected 3 slides, AM P. 92505, dive site ‘ The Wall’, Nepean Island, near Norfolk Island, 29 ° 04 ' 17 " S, 167 ° 57 ' 40 " E, 10 m, tufted red and brown algae, 15 May 2008, coll. J. K. Lowry, MI NFK 43. Paratypes: male, 4.4 mm, dissected, 1 slide, AM P. 92506; male, 4.0 mm, dissected, 1 slide, AM P. 98702; 11 specimens, AM P. 81504, same location as holotype. Additional material examined. New Zealand. 1 specimen, AM P. 90948, west side of South Meyer Island, Kermadec Islands, 29 ° 14 ' 47 " S, 177 ° 52 ' 53 " W, 10 – 12 m, rubble, 12 May 2011, coll. S. J. Keable, M. A. McGrouther and A. Reid, K 2011 - 3 - 2; 3 specimens, AM P. 90949, Milne Islets, Raoul Island, Kermadec Islands, 29 ° 16 ' 57 " S, 177 ° 54 ' 10 " W, 22 – 27 m, tufting algae, 15 May 2011, coll. S. J. Keable, M. A. McGrouther and A. Reid, K 2011 - 23 - 4; 4 specimens, AM P. 90950, Fishing Rock landing, Raoul Island, Kermadec Islands, 29 ° 15 ' 03 " S, 17 ° 754 ' 12 " W, 5 m, scrapings from rock wall, 17 May 2011, coll. S. J. Keableand A. Reid, K 2011 - 49 - 7. Type locality. Dive site ‘ The Wall’, Nepean Island, near Norfolk Island, Tasman Sea, 29 ° 4 ' 17 " S, 167 ° 57 ' 40 " E.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFB1FFB5FF50FA0215C7F873.taxon	etymology	Etymology. Latin for wall, in reference to the type locality.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFB1FFB5FF50FA0215C7F873.taxon	description	Description. Of holotype male, 4.0 mm AM P. 92505. Head. Eyes ovate; lateral cephalic lobe broad, apically truncate, anteroventral margin with excavate, anteroventral corner acute. Antenna 1 longer than antenna 2; peduncular article 1 shorter than article 2, with 2 proximal robust setae on posterior margin; peduncular article 2 longer than article 3; flagellum articles as long as broad, or broader than long, with 13 articles; accessory flagellum long, more than half length of primary flagellum, with 7 articles. Antenna 2 peduncular article 4 longer than article 5; flagellum with 8 articles. Mandible accessory setal row, well developed with 4 setae; palp well developed, article 1 length twice width, shorter than article 2, inner margin weakly produced rounded; article 2 length 0.9 × article 3, lined with long slender setae; article 3 rectilinear, long, 2.8 × as long as broad, longer than article 1, with 5 lateral and 4 apical slender setae. Pereon. Gnathopod 1 coxa anterior margin straight, anteroventral corner subquadrate, not produced, ventral margin without robust setae, posteroventral corner without notch; basis anterodistal corner without lobes; ischium anterodistal corner without lobes; merus with posteroventral corner subquadrate; carpus length 2.1 × width, 1.3 × propodus length, posterior margin lined with long setae, with rows of setae covering medial surface; propodus subovate, medial surface setal comb absent, palm subacute, weakly convex, entire, lined with slender setae, defined by posterodistal corner with 1 robust seta; dactylus closing along palm, unguis present. Gnathopod 2 subchelate; coxa posteroventral corner without notch, ventral margin without robust setae; basis broad, anterodistal corner with pair of well-developed lobes; ischium anterodistal corner with pair of well-developed lobes; merus with subacute posteroventral corner; carpus compressed, length 0.7 × width, anterior margin with 1 robust seta, posterior margin with a few slender setae; propodus quadrate, length 1.1 × width, posterior margin with clusters of long slender setae, palm length less than one third of propodus posterior margin, angle transverse, distal shelf subquadrate with 3 pairs of robust setae, palm with shallow rounded excavation, distal palm margin lined with robust setae, palm defined by posteroventral corner (90 ° angle, large subacute tooth and 1 robust seta; dactylus reach end of and closing along palm. Pereopods 3 – 4 coxa subquadrate; basis long, length 3 × width; merus, carpus and propodus not broadened; dactylus unguis bicuspidate. Pereopods 5 – 7 slender distally; basis rectilinear; merus, carpus and propodus slender, margins lined with a few long slender setae; dactylus unguis bicuspidate. Pleon. Pleonites 1 – 3 dorsally smooth. Epimera 1 – 3 posterior and ventral margins smooth, posteroventral corner with acute tooth. Uropod 1 peduncle with 1 basofacial seta, length 1.4 × outer ramus; inner ramus slightly longer than outer ramus. Uropod 2 peduncle length subequal to outer ramus; inner ramus slightly longer than outer ramus. Uropod 3 peduncle 0.5 × inner ramus; inner ramus subequal to outer ramus, length 4 × width, apically truncate with long robust setae. Telson as long as broad, deeply cleft (80 %), lobes divergent, not tapering distally, lobes apically truncate, each lobe with 5 apical robust setae.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFB1FFB5FF50FA0215C7F873.taxon	discussion	Remarks. Quadrimaera vallaris sp. nov. has a plain gnathopod 2 palm without teeth or deep excavations and, 5 apical setae on telson similar to Q. schellenbergi (Ruffo, 1938). The exceptionally long rami of uropod 3, length 4 times the breadth, distinguished this species from Q. schellenbergi and other Quadrimaera species. Although uropod 3 length is known to variable with growth stage (Krapp-Schickel 2009), the length of rami in Q. vallaris sp. nov. is exceptionally elongate distinguishing it from currently known species. The male figured here is 4.0 mm, so it is possible that the palm development is not that of a terminal male given the relatively propodus palm sculpturing and lack of sinus or tooth on the dactylus posterior margin.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFB1FFB5FF50FA0215C7F873.taxon	distribution	Distribution. South Pacific Ocean: Tasman Sea: Norfolk Island; New Zealand: Kermadec Islands (current study).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFAEFFB1FF50FA05171CF91D.taxon	materials_examined	Material examined. Western Australia: 6 specimens, AM P. 79346, outer edge of Ningaloo Reef off Neds Camp, Cape Range National Park, 21 ° 59 ' 00 " S, 113 ° 54 ' 30 " E, 12 m, encrusting invertebrates on plate coral, 2 January 1984, coll. J. K. Lowry, WA 359; 8 specimens, AM P. 79324, inshore limestone reef off Neds Camp, Cape Range National Park, 21 ° 59 ' S, 113 ° 55 ' E, 1.5 m, " crinkly " brown alga, 2 January 1984, coll. R. T. Springthorpe, WA 379; 15 specimens, AM P. 79317, off Possession Point, King George Sound, 35 ° 03 ' S, 11 ° 758 ' E, 7 m, finger sponges and algae, 14 December 1983, coll. R. T. Springthorpe, WA 136; many specimens, AM P. 79365, off Possession Point, King George Sound, 35 ° 03 ' S, 117 ° 58 ' E, 7 m, mixed sponges and algae, 14 December 1983, coll. J. K. Lowry, WA 135; many specimens, AM P. 79314, near Mistaken Island, King George Sound, Vancouver Peninsula, 35 ° 04 ' S, 117 ° 56 ' E, 2 m, soft corals, 13 December 1983, coll. J. K. Lowry, WA 105; 16 specimens, AM P. 79420, near Mistaken Island, Vancouver Peninsula, King George Sound, 35 ° 04 ' S, 117 ° 56 ' E, 6 m, seagrass, 13 December 1983, coll. R. T. Springthorpe, WA 121; 16 specimens, AM P. 79339, near Mistaken Island, Vancouver Peninsula, King George Sound, 35 ° 04 ' S, 117 ° 56 ' E, 2 m, grey sponge with crinoids, 13 December 1983, coll. R. T. Springthorpe, WA 114; many specimens, AM P. 79360, rocks near Migo Island, Torbay Bay, Port Harding, 35 ° 04 ' S, 117 ° 39 ' E, 6 – 7 m, branched alga with compound tunicate on underside of branches, 15 December 1983, coll. R. T. Springthorpe and J. K. Lowry, WA 152; 8 specimens, AM P. 79376, rocky shore, Red Bluff, Kalbarri, 27 ° 42 ' S, 11 ° 49 ' E, 3 – 4 m, mixed algae, sediment, 10 January 1984, coll. R. T. Springthorpe, WA 474; 26 specimens, AM P. 79335, rocky shore, Red Bluff, Kalbarri, 27 ° 42 ' S, 114 ° 09 ' E, 1 m, stalked ascidians on rock ledge, 9 January 1984, coll. H. E. Stoddart, WA 445; 14 specimens, AM P. 79328, rocks near Migo Island, Torbay Bay, Port Harding, 35 ° 04 ' S, 117 ° 39 ' E, 6 – 7 m, mixed sponges, 15 December 1983, coll. J. K. Lowry, WA 141; 4 specimens, AM P. 79384, rocks near Migo Island, Port Harding, Torbay Bay, 35 ° 04 ' S, 117 ° 39 ' E, 6 – 7 m, red sponges with zoanthid, black-spotted white ascidian, 15 December 1983, coll. J. K. Lowry, WA 151; 14 specimens, AM P. 79333, reef west of groyne, 2 km south of Cape Peron, 32 ° 16 ' S, 115 ° 41 ' E, 3 m, tough red alga, 26 December 1983, coll. R. T. Springthorpe, WA 299; 1 specimen, NMV J 56565, point at north end of Little Beach, Two Peoples Bay, 34 ° 58 ' 12 " S, 118 ° 10 ' 48 " E, 5 m, tufted red algae and soft erect bryozoan, 20 April 1985, coll. G. C. B. Poore, SWA 72; 15 specimens, NMV J 57014, northeastern end, Vancouver Peninsula, 35 ° 03 ' 24 " S, 117 ° 56 ' 12 " E, 3 m, tufted red algae, soft coral and sponges, 8 April 1985, coll. G. C. B. Poore and H. M. Lew Ton, SWA 22; 7 specimens, NMV J 56990, north side, Cape Riche, 34 ° 37 ' 00 " S, 118 ° 47 ' 00 " E, 7 m, date, SWA 49; 6 specimens, NMV J 57002, north of False Island, King George Sound, 35 ° 00 ' 42 " S, 118 ° 10 ' 06 " E, 27 m, red algae and bryozoan, 15 April 1985, coll. G. C. B. Poore and H. M. Lew Ton, SWA 57; 10 specimens, NMV J 56982, north side, Cape Riche, 34 ° 37 ' 00 " S, 118 ° 47 ' 00 " E, 6 m, sponges on vertical rock face, 12 April 1985, coll. G. C. B. Poore and H. M. Lew Ton, SWA 47; 2 specimens, NMV J 56995, northeastern end, Vancouver Peninsula, 35 ° 03 ' 24 " S, 117 ° 56 ' 12 " E, 10 m, red algae, 8 April 1985, coll. G. C. B. Poore and H. M. Lew Ton, SWA 19; 7 specimens, NMV J 56904, northeastern end, Vancouver Peninsula, 35 ° 03 ' 24 " S, 117 ° 56 ' 12 " E, 10 m, red algae, 8 April 1985, coll. G. C. B. Poore and H. M. Lew Ton, SWA 24; 10 specimens, NMV J 56591, northern side, West Island, 35 ° 37 ' 00 " S, 138 ° 36 ' 00 " E, 4 m, macroalga Ecklonia sp. holdfast, 21 March 1985, coll. G. C. B. Poore and H. M. Lew Ton, SA 45; 3 specimens, NMV J 56589, 1 km offshore, Seven Mile Beach, North of Dongara, 29 ° 12 ' 00 " S, 114 ° 53 ' 00 " E, 9 m, 27 April 1986, SWA 98; 7 specimens, NMV J 56590, 1 km offshore, Seven Mile Beach, north of Dongara, 29 ° 12 ' 00 " S, 114 ° 53 ' 00 " E, 9 m, 27 April 1986, SWA 99; 1 specimens, NMV J 56588, Seven Mile Beach, north of Dongara, 29 ° 12 ' 00 " S, 114 ° 53 ' 00 " E, 1 m, 22 April 1986, SWA 83; 2 specimens, NMV J 56592, Seven Mile Beach, north of Dongara, 29 ° 12 ' 00 " S, 114 ° 53 ' 00 " E, 1.0 m, 24 April 1986, SWA 91; 2 specimens, NMV J 56566, point at north end of Little Beach, Two Peoples Bay, 34 ° 58 ' 12 " S, 118 ° 10 ' 48 " E, 5.0 m, 18 April 1986, SWA 73; 8 specimens, NMV J 56567, point at north end of Little Beach, Two Peoples Bay, 34 ° 58 ' 12 " S, 118 ° 10 ' 48 E, 5.0 m, 18 April 1986, SWA 78. South Australia: several specimens, SAMA C 6530, 1300 m offshore, Cape Northumberland, 38 ° 3 ' 29 " S, 140 ° 39 ' 43 " E, red alga community, 1976 – 1977, coll. S. A. Shepherd; several specimens, SAMA C 6735, Fowlers Bay Jetty, west of Ceduna, 31 ° 59 ' 24 " S, 132 ° 26 ' 20 " E, 2 March 1993, coll. W. Zeidler, K. L. Gowlett-Holmes and B. McHenry; several specimens, SAMA C 6529, Greenly Island, 34 ° 38 ' 46 " S, 134 ° 46 ' 59 " E, in weed, 28 November 1976, coll. W. Zeidler; 2 specimens, NMV J 56564, " The Hotspot " reef, 4 – 5 nautical miles west of northern end of Flinders Island, 33 ° 40 ' 30 " S, 134 ° 22 ' 00 " E, 17 m, macroalga Cystophora sp. holdfast on exposed rocky bottom, 19 April 1985, coll. G. C. B. Poore, SA 61; 2 specimens, NMV J 56563, northeastern side of Topgallant Island, Investigator Group, 33 ° 43 ' 00 " S, 134 ° 36 ' 36 " E, 7.0 m, brown alga Acrocarpia aniculata and red algae, 22 April 1985, coll. G. C. B. Poore, SA 83; 1 specimen, NMV J 56562, " The Hotspot " reef, 5 nautical miles west of Flinders Island, 33 ° 40 ' 48 " S, 134 ° 22 ' 30 " E, 21 m, tufted red algae and soft red bryozoan, 20 April 1985, coll. G. C. B. Poore, SA 72; 1 specimen, NMV J 56561, northeastern side of Topgallant Island, Investigator Group, 33 ° 43 ' 00 " S, 134 ° 36 ' 36 " E, 16 m, alga Cystophora sp., 22 April 1985, coll. G. C. B. Poore, SA 84; several specimens, SAMA C 6531, 18.6 miles west north west of Rapid Head, Gulf of St Vincent, 35 ° 31 ' 08 " S, 138 ° 10 ' 01 " E, in sponges and bryozoans, in prawn trawl, 5 June 1987, coll. K. L. Gowlett-Homes and S. Corigliano, FV Rivoli Queen; several specimens, SAMA C 6533, East Bay, St Francis Island, Nuyts Archipelago, 33 ° 30 ' 6 " S, 133 ° 17 ' 24 ” E, 1 February 2002, coll. A. Hirst, SARDI Encounter 2002 Expedition; several specimens, SAMA C 6532, Western side of Smooth Island, Nuyts Archipelago, 33 ° 30 ' 6 " S, 133 ° 17 ' 24 ” E, kelp dominated, February 2002, coll. A. Hirst, SARDI Encounter 2002 Expedition; several specimens, SAMA C 6705, Moonta Bay Jetty, YorkePeninsula, 34 ° 2 ' 30 " S, 137 ° 33 ' 0 " E, Jetty pylons, rubble, sand and seagrass Posidonia sp., 15 December 1994, coll. W. Zeidler and K. L. Gowlett-Holmes; several specimens, SAMA C 6543, approximately 1.5 km W of Cape Jaffa, 36 ° 56 ' 43 " S, 139 ° 39 ' 57 " E, undulating reef algae, 15 February 1989, coll. W Zeidler; several specimens, SAMA C 6537, Lagoon East of Margaret Brock Reef Lighthouse, off Cape Jaffa, 36 ° 57 ' 09 " S, 139 ° 39 ' 52 " E, stringy ascidian, 18 February 1989, coll. W. Zeidler and K. Gowlett-Holmes; several specimens, SAMA C 6534, approximately 3 km west north west of Margaret Brock Reef Lighthouse, South East, Cape Jaffa, 36 ° 57 ' 9 " S, 139 ° 39 ' 52 " E, reef, ridges, algae, sand, 17 February 1989 coll. W. Zeidler and K. L. Gowlett-Holmes; several specimens, SAMA C 6542, South East, between Godfrey Islands, Cape Thomas, 37 ° 5 ' 19 " S, 139 ° 43 ' 20 " E, soft rock, reef, algae, sand, 16 February 1989, coll. W. Zeidler and K. L. Gowlett-Holmes; several specimens, SAMA C 6576, Fanny Point, Boston Island, Eyre Penninsula, 34 ° 44 ' 02 " S, 135 ° 55 ' 45 ' E, on Sargassum sp. algae, 17 February 1988, coll. W. Zeidler and K. L. Gowlett-Holmes; several specimens, SAMA C 6536, Penneshaw Jetty, Kangaroo Island, 35 ° 43 ' 00 " S, 137 ° 56 ' 30 " E, Pilons, sand, weed, 31 January 1989, coll. K. L. Gowlett-Holmes; several specimens, SAMA C 6538, northern side of Point Ellen Reef, Vivonne Bay, Kangaroo Island, 35 ° 59 ' 13 " S, 137 ° 12 ' 25 " E, boulders, sand, 26 January 1989, coll. W. Zeidler and K. L. Gowlett-Holmes; several specimens, SAMA C 6539, NorthPoint St. Francis Island, Nuyts Archipelago, 33 ° 30 ' 30 " S, 133 ° 14 ' 48 " E, kelp-dominated, February 2002, coll. A. Hirst, SARDI Encounter 2002 Expedition; several specimens, SAMA C 6540, North west of Freeling Island, Nuyts Archipelago, 33 ° 30 ' 30 " S, 133 ° 14 ' 48 " E, mixed fucoid, February 2002, coll. A. Hirst, SARDI Encounter 2002 Expedition; several specimens, SAMA C 6535, Edithburg Jetty, Yorke Penninsula, 35 ° 6 ' 0 " S, 137 ° 43 ' 60 " E, in bryozoan-sponge mass, 26 November 1985, coll. W. Zeidler and K. L. Gowlett-Holmes; several specimens, SAMA C 6541, Edithburg Jetty, Yorke Penninsula, 35 ° 6 ' 0 " S, 137 ° 43 ' 60 " E, in sponge bryozoan mass, 26 November 1985, coll. W. Zeidler, K. L. Gowlett- Homes and T. Stranks; 3 specimens, NMV J 56568, " The " Hotspot " reef, 5 nautical miles west of northern end of Flinders Island, 33 ° 40 ' 30 " S, 134 ° 22 ' 00 " E, 17 m, 19 April 1985, coll. G. C. B. Poore, SA 65. New South Wales: many specimens, AM P. 72913, East Wall, north of Burrewarra Point, 35 ° 50 ' 1 " S, 150 ° 14 ' 9 " E, 25 m, macroalga Corallina berteri, 25 October 2002, coll. G. D. F. Wilson, A. J. Millar andN. Yee, NSW 1987. Tasmania: 2 specimens, AM P. 83563, Joes Bight, Freycinet Peninsula, 42 ° 16 ' 42 " S, 148 ° 18 ' 42 " E, 17 m, flat red algae with under surface heavily encrusted with epiphytes, 1 May 1991, coll. R. T. Springthorpe and S. J. Keable, TAS 346; 4 specimens, AM P. 83564, Joes Bight, Freycinet Peninsula, 42 ° 16 ' 42 " S, 148 ° 18 ' 42 " E, 17 m, Ecklonia sp. holdfasts, 1 May 1991, coll. R. T. Springthorpe and S. J. Keable, TAS 329; many specimens, AM P. 83567, Rocky Cape, 40 ° 51 ' S, 145 ° 31 ' E, 6 m, algae, 8 December 1977, coll. C. J. Short, TAS- 5; 1 specimen, AM P. 83560, east of Hannants Bight, Cape Sorell, 42 ° 11 ' 30 " S, 145 ° 11 ' 30 " E, 15 m, mixed sponges, 28 April 1991, coll. R. T. Springthorpe and P. M. Berents, TAS 308; 4 specimens, AM P. 83557, north side of Esperance Point, D'Entrecasteaux Channel, 43 ° 19 ' 30 " S, 14 ° 75 ' 30 " E, 14 m, mixed red algae, sponges and Caulerpa sp., 18 April 1991, coll. S. J. Keable, J. K. Lowry and R. T. Springthorpe, TAS 179; many specimens, AM P. 83562, north side of Esperance Point, D'Entrecasteaux Channel, 43 ° 19 ' 30 " S, 14 ° 75 ' 30 " E, 13 m, sponges, lace bryozoan, red algae and? Vittaticella, 18 April 1991, coll. S. J. Keable, J. K. Lowry and R. T. Springthorpe, TAS 187; many specimens, AM P. 83559, Cemetery Bluff, Adventure Bay, Bruny Island, 43 ° 19 ' 48 " S, 147 ° 19 ' 12 " E 15 m, macroalga Ecklonia sp. holdfasts, 23 April 1991, coll. R. T. Springthorpe and P. M. Berents, TAS- 252; 1 specimen, AM P. 83561, Cemetery Bluff, Adventure Bay, Bruny Island, 43 ° 19 ' 48 " S, 147 ° 19 ' 12 " E, 15 m, red algae, 23 April 1991, coll. R. T. Springthorpe and P. M. Berents, TAS- 253; 1 specimen, AM P. 83558, Cemetery Bluff, Adventure Bay, Bruny Island, 43 ° 19 ' 48 " S, 147 ° 19 ' 12 " E, 15 m, sponges, 23 April 1991, coll. R. T. Springthorpe and P. M. Berents, TAS- 249; 4 specimens, AM P. 83566, Quiet Corner, Adventure Bay, South Bruny Island, 43 ° 21 ' 30 " S, 147 ° 19 ' 30 " E, 3 m, orange " frilly " bryozoan, 22 April 1991, coll. P. B. Berents and P. M. Berents, TAS 235; 6 specimens, AM P. 83565, Quiet Corner, Adventure Bay, South Bruny Island, 43 ° 21 ' 30 " S, 147 ° 19 ' 30 " E, 1 – 2 m, kelp holdfasts, 22 April 1991, coll. P. B. Berents and P. M. Berents, TAS 233. Norfolk Island: 1 specimen, AM P. 81498, east of Sail Rock, Phillip Island, 29 ° 06 ' 54 " S, 167 ° 57 ' 31 " E, 10.6 m, red algae with encrusting fauna, 14 May 2008, coll. L. E. Hughes, MI NFK 23; 1 specimen, AM P. 81499, Cow Bay, Phillip Island, 29 ° 09 " S, 167 ° 57 ' 03 " E, 12.8 m, red sponge like algae, 14 May 2008, coll. L. E. Hughes, MI NFK 33; 1 male, dissected, 4 slides, AM P. 81500, dive site ‘ The Wall’, Nepean Island, 29 ° 04 ' 17 " S, 167 ° 57 ' 40 " E, 10 m, tufted red and brown algae, 15 May 2008, coll. J. K. Lowry, MI NFK 43. Type locality. Clark Island, Port Jackson, New South Wales, Australia.	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
E73B096AFFAEFFB1FF50FA05171CF91D.taxon	distribution	Distribution. Australia: Western Australia: Cottesloe; Albany; King George Sound; Cape Range National Park; Port Harding; Kalbarri; Cape Peron; Vancouver Peninsula; Cape Riche; Dongara; Two Peoples Bay (J. L. Barnard 1972 a; current study); South Australia: Flinders Island; Investigator Group; Cape Northumberland; Ceduna; Greenly Island; Gulf of St Vincent; Nuyts Archipelago; Yorke Peninsula; Kangaroo Island; Point Noarlunga; Eyre Penninsula; Cape Jaffa; Cape Thomas (J. L. Barnard 1972 a; current study); Victoria: Port Phillip (J. L. Barnard 1972 a); New South Wales: Port Jackson; Burrewarra Point (Haswell 1879; Chiton 1916; current study); Tasmania: eastern slope of Bass Strait; Freycinet Peninsula; Rocky Cape; Cape Sorell; D'Entrecasteaux Channel; Bruny Island (Chilton 1921; current study); Lord Howe Island (Chilton 1916); Norfolk Island: Phillip Island; Nepean Island (current study).	en	Hughes, Lauren E. (2015): Maeridae from the Indo-Pacific: Elasmopus, Leeuwinella gen. nov., Maeropsis, Pseudelasmopus and Quadrimaera (Amphipoda: Crustacea). Zootaxa 4059 (2): 201-256, DOI: 10.11646/zootaxa.4059.2.1
