identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
E66B582DFFCEFF96FC47F940D15E532E.text	E66B582DFFCEFF96FC47F940D15E532E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sarametra A. H. Clark 1917	<div><p>Genus Sarametra A.H. Clark, 1917</p><p>Revised diagnosis: A genus of Zenometridae with a conical or truncated conical centrodorsal. Peripheral cirrus sockets separated interradially by narrow flat strip or weak interradial ridges, except in S. varians, which has more conspicuous, sinuous interradial ridges extending to the tip of the centrodorsal. Cirrus sockets with fulcral bowls surrounding central lumen; socket margins finely crenulate. Distal cirrals wider than long. Row of spines present on proximal brachials. First segment of P 1 with L/W = 1.1–1.8. Angle between the lateral edge of IBr 1 and the oral/aboral axis largely obtuse.</p><p>Type species: Zenometra triserialis A.H. Clark, 1908 (Fig. 6F).</p><p>Sarametra varians sp. nov. (Figs 7–10) ZooBankregistration: urn:lsid:zoobank.org:act: E9BAD48A-4D3D- 4AF5-BDBF-795804B707B2.</p><p>Diagnosis: Centrodorsal conical or truncated conical, taller than wide (H/D = 1.2–1.3), with two or three columns of cirrus sockets in each radial area. Cirrus sockets are separated interradially by weak to moderately developed interradial ridges, usually sinuous and extend to the aboral pole. Proximal brachials with conspicuous synarthrial and articular tubercles. Sparse and short spines on proximal brachials. P 1 6.7–10.0 mm, 13–15 pinnulars. The first pinnular of P</p><p>1</p><p>with L/W = 1.5–1.7.</p><p>Material examined: Holotype (RSIOCRI0292), Northwest Pacific, 12.5365°N, 149.6961°E, depth 2922 m, 26 July 2023, collected by HOV Jiaolong. Paratypes: RSIOCRI0007, Northwest Pacific, 15.9769°N, 155.2828°E, depth 2743m, 23 July 2014, collected by HOV Jiaolong; RSIOCRI0207, Northwest Pacific, 25.4525°N, 149.4811°E, depth 2749 m, 25 June 2023, collected by HOV Jiaolong; RSIOCRI0291, Northwest Pacific, 12.5365°N, 149.6961°E, depth 2922 m, 26 July 2023, collected by HOV Jiaolong. Other specimens: RSIOCRI0067 (broken arms only), Northwest Pacific, 15.4665°N, 161.7706°E, depth 2498 m, 21 September 2019, collected by ROV Hailong III.</p><p>Description of holotype RSIOCRI0292: Centrodorsal truncated conical and taller than wide (H = 5.1 mm, D = 3.9 mm, H/D = 1.3) (Fig. 8A). Cirrus sockets are separated interradially by elevated and sinuous interradial ridges that extends to the aboral pole. Aboral pole of centrodorsal excavated (Fig. 8B). Tegmen naked, domed, with mouth central or subcentral. Surface of tegmen level at IIBr 4–5 (Fig. 8D). Edge of lateral margins of radial articular facets visible interradially.</p><p>Cirri LII (functional sockets) in two or three columns per radial area. Each column with 4–6 functional sockets. There are very few obsolete sockets, only 1 or 2 per radial area. Fulcral bowl of cirrus sockets surrounding central lumen. Marginal crenulations of cirrus sockets fine. Cirri of 51–62 segments, 20–35 mm long. Longest cirral L/W = 2.0–2.5. Proximal cirrals cylindrical with distinctly concave aboral margins. c 1–4 very short. Cirrals increase in length from c 5 to c 10 (Fig. 8G). Cirrals at distal fourth of the cirrus shorter than wide. Terminal claw curved beyond aboral margin of penultimate cirral. Few cirri with opposing spine.</p><p>Basals visible externally in each interradial. Radials short, without spines. Arms 10. IBr 1 short, W/L = 2.7 (2.7/1.0 mm). The distal edge of IBr 1 is prominent at straight angles to the axillary (IBr 2). Distal margin broadly V-shaped, longer than the proximal margin. Axillary (IBr 2) rhombic, W/L= 1.3 (3.3/ 2.6 mm). The proximolateral and distal margins are slightly concave. The dorsal surfaces of IBr 1 and IBr 2 are very high and arched with conspicuous synarthrial tubercles (Fig. 8A, white arrow). IIBr 1 short, distal margin V-shaped, longer than proximal margin. W/L = 2.2 (2.0/ 0.9 mm). IIBr 2 cuneate, W = 1.6–2.2 mm, L = 1.6–1.8 mm, W/L = 1.0–1.3. First syzygy at IIBr 3+4, W = 1.9 mm. IIBr 5–8 square-shaped, W/L = 1.2–1.3. Second syzygy at IIBr 9 + 10, rectangle-shaped, W = 2.1 mm. Second syzygy occasionally at IIBr 10 + 11 or IIBr 12 +13. Subsequent interval 2–8. Proximal brachials equipped with sparse and short spines. From IIBr 5 onwards, brachials are smooth on the proximal margin, with fewer spines distally. Middle brachials cuneate or triangular, W/L = 1–1.2. Distal spines alternating on successive brachials. Distal brachials W/L = 0.9–1.2 (Fig. 8F). Distal brachials cuneate in shape and much slender as arm narrows nearer tip.</p><p>Pinnules 10.0–14.0 mm in length, each with 9–20 pinnulars. Pinnular length 0.6–1.2mm. Proximal pinnulars compressed, following pinnulars gradually narrower, becoming extremely slender to tip. P 1 10 mm in length, with 15 segments. First pinnular L/W = 1.5 (Fig. 8E). Middle pinnular of P 1 L/W = 1.8–3. Distal pinnular of P 1 L/W = 3. P 2 8.9–12.0 mm with 11–13 pinnulars. First genital pinnule P 4. P 4 flat and thin, the length of which is about 7.5mm, genitals not conspicuous. The following pinnules have a broad base, with short first pinnular and extremely slender distal pinnulars (Fig.8F). The distal pinnules is about 14 mm long. P a 7.0– 8.7 mm, 11–13 pinnulars.The distal pinnules composed of oval-shaped and notably large pinnulars (Fig. 8C, white arrow).</p><p>Description of paratypes: RSIOCRI0007 (Fig. 9A–C), centrodorsal H = 4.1 mm, D = 3.1 mm, H/D = 1.3. Aboral pole of centrodorsal is convex. Interradial ridges slightly developed. Cirri XLVII (functional sockets). Functional cirrus sockets in two columns per radial area. Obsolete sockets 0–2 per radial area. 20–38mm long, with 38–60 segments. Longest cirral L/W = 2.4. c 1–4 very short. Cirrals increase in length from c 5 to c 8. IBr 1 short, W/L = 2.7 (2.5/ 0.9 mm). The distal edge of IBr 1 is prominent at straight angles to the axillary (IBr 2). Distal margin broadly V-shaped, longer than the proximal margin. Axillary (IBr 2) rhombic, W/L = 1.1 (3.2/3.0 mm). The proximolateral and distal margins are slightly concave.IIBr 1 short, distal margin V-shaped, longer than proximal margin. W/L = 2.4. IIBr 2 cuneate. W/L = 1.0 (1.6/ 1.6 mm). First syzygy at IIBr 3+ 4, W = 1.9 mm. Second syzygy at IIBr 9 +10, rectangle-shaped, W = 1.5 mm. IIBr 5–8 square-shaped, W/L = 1.1–1.2. Pinnules 7.0–15.0 mm in length, with 11–16 pinnulars. Pinnular length 0.6–1.1 mm. P 1 8.5 mm, 14 pinnulars. First pinnular L/W = 1.7. P 2 8.4 mm, 13 pinnulars. P a 7.0 mm, 12 pinnulars.</p><p>RSIOCRI0207 (Fig. 9E, G), centrodorsal half-broken, H = 5.3 mm, D = 4.4 mm, H/D = 1.2. Centrodorsal is completely hollow. Cirri XLVII (functional sockets).Functional cirrus sockets in two columns per radial area, separated by elevated interradial ridges extending to the aboral pole. Cirri of 60–71 segments. Longest cirral L/W = 2. IBr 1 short, W/L = 4.0 (2.9/ 0.7 mm). The distal edge of IBr 1 is prominent at straight angles to the axillary (IBr 2). Distal margin broadly V-shaped, longer than the proximal margin. Axillary (IBr 2) rhombic, W/L = 1.1 (2.6/ 2.2 mm). The proximolateral and distal margins are slightly concave. IIBr 1 short, distal margin V-shaped, longer than proximal margin. W/L = 1.9. IIBr 2 cuneate. W/L = 1.1 (1.9/ 1.8 mm). First syzygy at IIBr 3 + 4, W = 1.7 mm. Second syzygy at IIBr 9 + 10, rectangle-shaped, W = 1.8 mm. IIBr 5-8 square-shaped, W/L = 1.1–1.3. Pinnule length 8.0– 12.5 mm, each with 9–18 pinnulars. P 1 8.0 mm in length, with 13 pinnulars. First pinnular L/W = 1.7. P 2 7.3 mm with 13 pinnulars. P a 7.0 mm, 13 pinnulars.</p><p>RSIOCRI0291 (Fig. 9D, F), centrodorsal H = 5.3 mm D = 4.1 mm, H/D = 1.3. Aboral pole of centrodorsal excavated. Cirrus sockets are separated interradially by elevated, sinuous interradial ridges that extends to the aboral pole. Cirri LV (functional sockets). Functional cirrus sockets in two or three columns per radial area. Obsolete sockets 1–4 per radial area. Cirri 20–45 mm long, with 49–70 segments. Longest cirral L/W = 2.5. c 1–4 very short. Cirrals increase in length from c 5 to c 10. IBr 1 short, W/L =2.6 (2.6/1.0 mm).The distal edge of IBr 1 isprominent at straight angles to the axillary (IBr 2). Distal margin broadly V-shaped, longer than the proximal margin. Axillary (IBr 2) rhombic, W/L = 1.3. The proximolateral and distal margins are slightly concave. IIBr 1 short, distal margin V-shaped, longer than proximal margin. W/L = 2.5. IIBr 2 cuneate. W/L = 1.3 (2.2/ 1.7 mm). First syzygy at IIBr 3 +4, W = 1.7 mm. Second syzygy at IIBr 9 + 10, rectangle-shaped, W = 1.3 mm. IIBr 5-8 square-shaped, W/L = 1.2–1.4. Pinnule length 8.0– 17.6 mm, each with 10–20 pinnulars. P 1 9.5 mm in length, with 13 pinnulars. First pinnular L/W = 1.5. P</p><p>2</p><p>8.5 mm with 11 pinnulars. P</p><p>a</p><p>8.7 mm, 11 pinnulars.</p><p>Ossicle morphology: Paratypes RSIOCRI0291, RSIOCRI0007, and RSIOCRI0207 were partially dissected for ossicle observation. Cirrus sockets are well separated by the sinuous interradial ridges (Fig. 10A). The proximal portion of the interradial ridges, where the basals are attached, exhibits a deep groove in both paratypes (Fig. 10A, white arrow). Socket margins finely crenulate; concave fulcral stereom restricted to central bowl surrounding socket lumen (Fig. 10A, B). The aboral pole shows a pentagonal-shaped regenerating skeleton with thin stereom (Fig. 10C). The five radials attached closely with the basals (Fig. 10D). Interradial processes of basal is broad, about as long as wide. The stereom on the inner side corresponds to a portion of the rosette (Fig. 10E). The articular facet of each radial to the first brachial bears five fossae, including a pair of adoral muscle fossae (mf), a pair of interarticular ligament fossae (il), and an aboral ligament fossae (al). Narrow ridges separate the two adoral muscle fossae and two interarticular ligament fossae. Aboral to the fulcral ridge is the unpaired aboral ligament fossa which houses the extensor ligament bundle (Hays 2016). IBr 1 W = 2.6 mm (Fig. 10F). Axillary (IBr 2) rhombic bears two articular faces distally, with thin marginal spines. W = 3.3 mm, L = 2.6 mm, W/L = 1.3 (Fig. 10G, H). IIBr 1 W = 2.8 mm (Fig. 10I). IIBr 2 with short and inconspicuous spines (Fig. 10J). First syzygy at IIBr 3+ 4. The syzygial articular surface of IIBr 3 oval shaped, with alternating radial ridges and depressions (Fig. 10K).</p><p>Colour: Purple brown coloured (Fig. 7A–C).</p><p>Etymology: The specific epithet ‘ varians ’ is a Latin adjective meaning ‘changing’ or ‘variable’, alluding to the morphological variability observed within this species, particularly in the shape of the centrodorsal and development of the interradial ridges.</p><p>Distribution: Known from RC seamount, Nazuna (HNN) seamount, RF seamount, Caiwei seamount, Northwest Pacific. Depth range 2498–2922 m. Attached to remnant coral stalk (RSIOCRI0291, RSIOCRI0292) or Primnoidae coral (RSIOCRI0207).</p><p>Remarks: The holotype RSIOCRI0292, and paratypes RSIOCRI0207 and RSIOCRI0291 demonstrated morphological similarities to the genus Zenometra, particularly in possessing a ‘cylindrical’ centrodorsal and elevated interradial ridges. However, molecular evidence suggests a close relationship of these specimens with Sarametra species (Fig. 2). Paratype RSIOCRI0007, on the other hand, presents traits that occurs in typical Sarametra, such as having a conical centrodorsal and less developed interradial ridges. With intraspecific COI distances less than 0.4% (Table 4), the polymorphism found in S. varians challenged the validity of the cylindrical centrodorsal and interradial ridge in defining genera, as noted in the assessment of generic diagnostic characters. Whether the observed similarity between S.varians and Zenometra columnaris suggests a close phylogenetic relationship is unknown as no DNA data of Z. columnaris is currently available.</p><p>In spite of the intraspecific variations, all specimens of S. varians share the same characters, in particular short and sparse spines on the proximal brachials, conspicuous synathrial and articular tubercles, as well as characters that fit with the genus, such as distal cirrals wider than long, first pinnular of P 1 less than twice as long as wide. It is attributed to Sarametra based on the current molecular and morphological evidence.The differences between S. varians and its congeners are listed in Table 6.</p></div>	https://treatment.plazi.org/id/E66B582DFFCEFF96FC47F940D15E532E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mao, Jingwen;Zhang, Ruiyan;Eléaume, Marc;Zhou, Yadong;Zhang, Dongsheng;Sun, Shao’e;Sha, Zhongli;Wang, Chunsheng	Mao, Jingwen, Zhang, Ruiyan, Eléaume, Marc, Zhou, Yadong, Zhang, Dongsheng, Sun, Shao’e, Sha, Zhongli, Wang, Chunsheng (2025): Description of three new species of Zenometridae (Echinodermata, Crinoidea, Comatulida), with new insights on species and genus delimitation. Zoological Journal of the Linnean Society 204 (4), DOI: 10.1093/zoolinnean/zlaf089, URL: https://doi.org/10.1093/zoolinnean/zlaf089
E66B582DFFD7FF8AFEFAFB35D0275000.text	E66B582DFFD7FF8AFEFAFB35D0275000.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sarametra acuta Mao & Zhang & Eléaume & Zhou & Zhang & Sun & Sha & Wang 2025	<div><p>Sarametra acuta sp. nov. (Figs 7, 14)</p><p>ZooBank registration: urn:lsid:zoobank.org:act: BC37E273- 8CCE-4B56-A8BA-17951074FBFB.</p><p>Material examined: Holotype, RSIOCRI0272, Northwest Pacific, 12.6676°N, 156.5418°E, depth 1681 m, 22 July 2023, collected by HOV Jiaolong.</p><p>Diagnosis: A conical aboral pole on a conical centrodorsal with H/D = 1.8. Two or three columns of cirrus sockets per radial area separated by weak interradial ridges. The half of centrodorsal exhibits distinct spines, with some hook like, aligned along the interradial ridges and rims of obsolete sockets, forming continuous, spinose columns that extend to the apex. Radials and proximal brachials with short, protuberance-like spines.P 1 7.3–8.3 mm, 16 pinnulars. The first pinnular of P 1 with L/W = 1.3.</p><p>Description of holotype: Centrodorsal conical in shape. D = 3.9 mm, H = 7.1 mm, H/D = 1.8. Aboral pole is flat with a hole in the middle, probably eroded (Fig. 14A). Cirrus sockets separated interradially by weak interradial ridges. Longitudinal series of sharp spines present along the interradial ridges of centrodorsal and between columns of obsolete socket. These spines are very prominent, and some even form hooks (Fig. 14A, B).</p><p>Cirri XLIX (functional sockets). Functional cirrus sockets in two or three columns per radial area in basal half of centrodorsal. Each column of 4–6 functional sockets. Zero to four obsolete sockets per radial area. Fulcral bowl of cirrus sockets surrounding central lumen, with socket margins finely crenulate. Cirri of 49–60 segments, 30–40 mm long. Longest cirral L/W ratio is 2.9. c 1–4 are very short, measuring twice as wide as they are long. c 5–8 are the longest. Proximal cirrals cylindrical with distinctly concave aboral margins (Fig. 14H). Opposing spine exists in some cirri. The terminal claw is tapered and exhibits a weak curvature.</p><p>The external ends of the basal are visible interradially. Radials short, with a shallow U-shaped distal margin decorated by short spines. Arms 10, more than 150 mm in length. Proximal brachials with short, protuberance-like spines (Fig. 14C, E). IBr 1 short, W = 2.0 mm, L = 0.6 mm, W/L = 3.3. Distal margin of IBr 1 slightly raised from the axillary. Axillary (IBr 2) rhombic in shape, W = 2.6 mm, L = 1.7 mm, W/L = 1.5. The proximolateral and distal margins are slightly concave. IIBr 1 short, distal margin V-shaped, longer than proximal margin, W = 2.1 mm, L = 0.96mm, W/L = 2.1. IIBr 2 cuneate, W = 1.8 mm, L = 1.0 mm, W/L = 1.8.</p><p>First syzygy at IIBr 3 + 4, W = 1.5 mm. Second syzygy at IIBr 9 + 10, W = 1.8 mm. Syzygies at IIBr 3 + 4, IIBr 9 + 10 and IIBr 14 + 15; subsequent interval 4–6. IIBr 5–8 resembles a rectangle. Middle brachials W/L = 1.1–1.2. Brachials exhibit concave lateral margins and occasionally characteristic weakly alternating articular tubercles (Fig. 14F, G).</p><p>Proximal pinnulars compressed, following pinnulars gradually narrower, becoming extremely slender to tip. Pinnule length 8.3–14.2 mm, 9–15 pinnulars. Pinnulars length 0.4–1.5 mm. P 1 7.3–8.3 mm 16 pinnulars. The first pinnular of P 1 with L/W = 1.3. P 1 wide at the base, getting thinner. Middle pinnular of P 1 L/W = 3.73. Distal pinnular of P 1 L/W = 4.0. P 2 up to 11.0 mm long with 19 pinnulars. Middle pinnular of P</p><p>2</p><p>L/W = 2.0. Distal pinnular of P 2 L/W = 3.8 (Fig. 14F). The following pinnules have a broad base, the first pinnular is short, and the distal pinnulars are extremely slender (Fig.14G).Pa 6.5–7.0mm, with11–14 pp.The distal pinnules are composed of oval-shaped and notably large pinnulars (Fig. 14D).</p><p>Colour: Deep purple brown coloured (Fig. 7F).</p><p>Etymology: The specific epithet ‘ acuta ’ is derived from the Latin word for ‘sharp’ or ‘pointed’, reflecting the sharp, prominent spine present on the centrodorsal in this new species.</p><p>Distribution: Known from Weijia seamount, Northwest Pacific. At 1681 m depth. Attached to Caulophacus sponge.</p><p>Remarks: A defining characteristic of this new species is the development of prominent spines along the interradial ridges and obsolete socket rims of the centrodorsal. In S. acuta, these spines are not only more pronounced than the cirrus sockets but also extend from the middle of the centrodorsal to the aboral pole, with some even developing into a hook. This distinctive morphology of the centrodorsal in Sarametra acuta provides a diagnostic characteristic that differentiates it from other species within the genus. In addition, the characters of P 1 distinguish it from Sarametra triserialis . The spines on the proximal brachials are also of a different shape as compared to what is observed in other species, as they form rows of small protuberance instead of long and sharp spines.</p></div>	https://treatment.plazi.org/id/E66B582DFFD7FF8AFEFAFB35D0275000	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mao, Jingwen;Zhang, Ruiyan;Eléaume, Marc;Zhou, Yadong;Zhang, Dongsheng;Sun, Shao’e;Sha, Zhongli;Wang, Chunsheng	Mao, Jingwen, Zhang, Ruiyan, Eléaume, Marc, Zhou, Yadong, Zhang, Dongsheng, Sun, Shao’e, Sha, Zhongli, Wang, Chunsheng (2025): Description of three new species of Zenometridae (Echinodermata, Crinoidea, Comatulida), with new insights on species and genus delimitation. Zoological Journal of the Linnean Society 204 (4), DOI: 10.1093/zoolinnean/zlaf089, URL: https://doi.org/10.1093/zoolinnean/zlaf089
E66B582DFFCAFF8BFF1EF950D1C2518A.text	E66B582DFFCAFF8BFF1EF950D1C2518A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sarametra subtilis Mao & Zhang & Eléaume & Zhou & Zhang & Sun & Sha & Wang 2025	<div><p>Sarametra subtilis sp. nov. (Figs 7, 11–13)</p><p>ZooBank registration: urn:lsid:zoobank.org:act: 7035258B-7689- 46E3-BBCD-D74AA03F09BF.</p><p>Material examined: Holotype RSIOCRI0108, Northwest Pacific, 13.3520°N, 134.5292°E, depth 1807 m, 6 August 2020, collected by ROV Hailong IV. Paratypes, RSIOCRI0016, Northwest Pacific, 12.6948°N, 156.5382°E, depth 1670 m, 29 September 2017, collected by ROV Haima; RSIOCRI0102, Northwest Pacific, 13.3316°N, 134.5094°E, depth 2097m, 3 August 2020, collected by ROV Hailong IV. RSIOCRI0253, Northwest Pacific, 12.8623°N, 157.0395°E, depth 1947 m, 19 July 2023, collected by HOV Jiaolong.</p><p>Diagnosis: A rounded, conical aboral pole on a conical centrodorsal with a height-to-diameter ratio (H/D) ranging from 1.3 to 1.5. Two or three columns of cirrus sockets per radial area separated by weak interradial ridges. Long, fine spines webbed with tissues densely distributed on the radials and proximal brachials. P 1 7.0–10.0 mm, 11–17 pinnulars. The first pinnular of P 1 with L/W = 1.7–1.8.</p><p>Description of holotype RSIOCRI0108: Centrodorsal conical in shape, H/D = 1.4 (5 mm / 3.6 mm). The aboral pole rather flat. The interradial ridges weakly developed restricted to the centrodorsal periphery (Fig. 11A). Tegmen naked, domed, with mouth central or subcentral. Surface of tegmen level at IIBr 5–6 (Fig. 11B). Radials spiny, with a shallow U-shaped distal margin.</p><p>Cirri XLVII (functional sockets), arranged in two or three columns per radial area in basal half of centrodorsal. Each column with 3–5 functional sockets. 2–4 obsolete sockets per radial area. Fulcral bowl of cirrus sockets surrounding central lumen, with fine marginal crenulations of sockets. Cirri of 49–63 segments, 23–40 mm long. The longest cirral with L/W = 2.2. Proximal several cirrals cylindrical with distinctly concave aboral margins. c 1–4 are very short, measuring twice as wide as long (Fig. 11F). c 5–7 are the longest. The following cirrals become shorter, with distal cirrals wider than long. Few cirri with opposing spine. The terminal claw is tapered and exhibits a weak curvature.</p><p>Basals visible from each interradius. The radials are short with spines. Arms 10, all broken. IBr 1 short, W = 2.6 mm, L = 0.7– 1.0 mm, W/L = 2.6. The distal margin of IBr 1 is longer than the proximal margin. Distal margin of IBr 1 slightly raised from the axillary. Axillary (IBr 2) rhombic in shape, W/L = 1.1 (3.0 mm/ 2.8 mm). The proximolateral and distal margins are slightly concave. IIBr 1 short, W = 1.8–2.0 mm, L = 0.7–0.8 mm, W/L = 2.5–2.8. Distal margin V-shaped, longer than proximal margin, IIBr 2 cuneate, W = 1.9–2.2 mm, L = 1.0– 1.3 mm, W/L = 1.5–2.0.</p><p>Syzygies at IIBr 3+ 4, IIBr 9+ 10 and IIBr 14 +15 (or IIBr 15+ 16, IIBr 17 +18); subsequent interval 2–8. First syzygy W = 1.5 mm. Second syzygy W = 2.1 mm. In one case the second syzygy is at IIBr 7 +8, and the third syzygy at IIBr 10 +11, and in a second case the second syzygy at IIBr 8 +9 (Fig. 11B). Long, fine spines webbed with tissues, densely distributed on both the proximal and distal edges of the proximal brachials (Fig. 11B). IIBr 5–8 rectangular, with W/L ranging from 1.3 to 1.4. Middle brachials W/L = 1.3–1.5. Distal brachials W/L = 0.9–1.2. Brachials exhibit concave lateral margins and occasionally with weakly alternating articular tubercles (Fig. 11E).</p><p>Proximal pinnulars compressed, following pinnulars gradually narrower, becoming extremely slender to tip. Pinnules 7.1– 14.0 mm in length, with 10–14 pinnulars. Pinnulars 0.9–1.2 mm in length. P 1 8.3–10.0 mm long with 15–17 pinnulars. The first pinnular of P 1 with L/W = 1.7–1.8 (Fig. 11C). Middle pinnular of P 1 L/W = 2.7–4.0. Distal pinnular of P 1 L/W = 4.0. P 2 9.1–10.2mm long with 16 pinnulars. The following pinnules have a broad base, the first pinnular is short, and the distal pinnulars are extremely slender (Fig. 11D). P a 5.0– 8.7 mm, 6–14 pinnulars.</p><p>Description of paratypes: RSIOCRI0016, centrodorsal H/D = 1.3 (5.1 mm / 3.8 mm). Cirri XLIX (functional sockets). Obsolete sockets 1–3 per radial area. Functional cirrus sockets in one or three columns per radial area. Spines conspicuous, webbed with tissues, densely distributed on proximal brachials (Fig.12C). IBr 1 short, W/L = 3.8 (2.9/ 0.7 mm). Distal margin broadly V-shaped, longer than the proximal margin. Axillary (IBr 2) rhombic, W/L = 1.1. The proximolateral and distal margins are slightly concave. IIBr 1 short, distal margin V-shaped, longer than proximal margin. W/L = 2.8. IIBr 2 cuneate. W/L = 1.5. First syzygy at IIBr 3 +4, W = 1.8 mm. Second syzygy at IIBr 9+ 10, rectangle-shaped, W = 1.7–2.1 mm. IIBr 5–8 square-shaped, W/L = 1.4. Pinnule length 7.1–11.4 mm, 10–13 pinnulars. Pinnular length 0.4– 1.0 mm.P 1 7.8mm, 14 pinnulars.ThefirstpinnularofP 1 L/W = 1.8. P 2 6.7 mm, 8 pinnulars. P a 7.5 mm, 13 pinnulars.</p><p>RSIOCRI0102, centrodorsal H/D = 1.5 (5.9 mm / 3.8 mm). Cirri XL (functional sockets). Four to eight obsolete sockets per radial area. Cirri of 40–54 segments, 20–30 mm long. Longest cirral L/W = 2.8. Few cirri with opposing spine. IBr 1 short, W/L = 3.5 (2.1/ 0.6 mm). Distal margin broadly V-shaped, longer than the proximal margin. Axillary (IBr 2) rhombic, W/L = 1.1 (2.6/ 2.4 mm). The proximolateral and distal margins are slightly concave.IIBr 1 short, distal margin V-shaped, longer than proximal margin. W/L = 2.5. IIBr 2 cuneate. W/L = 1.4. First syzygy at IIBr 3 +4, W = 1.7 mm. Second syzygy at IIBr 9 +10, rectangle shaped, W = 1.7 mm. IIBr 5–8 square shaped, W/L = 1.4. Pinnule length 10.0– 17.6 mm, 10–20 pinnulars. Pinnular length 0.4–1.1 mm. P 1 7 mm, 11 pinnulars. The first pinnular of P 1 with L/W = 1.8. P 2 7 mm, 9 pinnulars. P</p><p>a</p><p>6.4 mm, 8 pinnulars.</p><p>RSIOCRI00253, centrodorsal H/D = 1.4 (5.0 mm/ 3.6 mm). Tegmen naked, domed, with mouth central (Fig. 12B); surface of tegmen level with IIBr 3–4. Middle brachials W/L = 1.3. Distal brachials W/L = 1.4. Syzygies at IIBr 3 +4, IIBr 9+ 10, IIBr 14 + 15. Second syzygy appears in IIBr 11 + 12 in one arm. Cirri L (functional sockets). There are very few obsolete sockets, 2–3 per radial area. Functional cirrus sockets in two or three columns per radial area. There are spines on both the proximal and distal edges of the proximal brachials (Fig. 12A). IBr 1 short, W/L = 2.7 (1.9/ 0.7 mm). Distal margin broadly V-shaped, longer than the proximal margin. Axillary (IBr 2) rhombic, W/L = 1.2 (2.4/ 2.1 mm). The proximolateral and distal margins are slightly concave. IIBr 1 short, distal margin V-shaped, longer than proximal margin. W/L = 2.6. IIBr</p><p>2</p><p>cuneate. W/L = 1.5. First syzygy at IIBr 3+ 4, W = 1.7 mm. Second syzygy at IIBr 9 +10, rectangle shaped, W = 1.8 mm. IIBr 5–8 square shaped, W/L = 1.1. Pinnule length 6.0– 12.3 mm, 6–15 pinnulars. Pinnular length 0.6–0.9mm. P 1 8 mm, 14 pinnulars. The first pinnular of P 1 L/W = 1.8. P 2 6.3 mm, 9 pp. The distal pinnules have oval-shaped and notably large plates (Fig. 12D). Delicate side and covering plates present.</p><p>Ossicle morphology of RSIOCRI0016: Cirrus sockets are well separated by the weak interradial ridges. Concave fulcral stereom restricted to central bowl surrounding socket lumen; socket margins finely crenulate (Fig. 13A). The centrodorsal ossicle exhibits a shallow depression on its edge in each interradius, designed for adhesion to the basal surface (Fig. 13B, white arrow). The aboral pole at the centrodorsal was broken, possibly by erosion (Fig. 13C). Basal circlet surrounding centrodorsal central cavity, with a fused rosette in the centre (Fig. 5D), broken off during ossicle preparation (Fig. 13D). Interradial processes of basal is broad, about as long as wide (Fig. 13D). External end of basal pentagonal in shape, with a curved aboral margin (Fig. 13E). The articular facet of each radial to the first brachial bears five fossae. The two adoral muscle fossae and two interarticular ligament fossae separated by a narrow ridge. Radial W = 2.0 mm with spines (Fig. 13F). IBr 1 W = 2.8 mm with long spine (Fig. 13G). Axillary (IBr 2) rhombic in shape, bears two articular faces distally (Fig. 13H). In external view, axillary (IBr 2) bears long, lamellate spines densely distributed in several rows (Fig. 13I). L = 2.8 mm, W = 3.0 mm, W/L =1.1. IIBr 1 W = 2 mm (Fig.13J). IIBr 2 W = 2 mm with dense spines (Fig. 13K). First syzygy IIBr 3+ 4. The syzygial articular surface on IIBr 3 has alternating radial ridges and depressions (Fig. 13L), which are nearly circular.</p><p>Colour: Deep purple brown coloured (Fig. 7D, E).</p><p>Etymology: The specific epithet ‘ subtilis ’ is a Latin adjective meaning ‘fine, thin, finely woven’, reflecting the long, fine spines webbed by tissues densely distributed on proximal brachials.</p><p>Distribution: Known from Weijia seamount, Roischesar Peak, Northwest Pacific. Depth range 1670–2097m. Attached to remnant coral stalk (RSIOCRI0108) or Caulophacus sponge (RSIOCRI0102, RSIOCRI0253).</p><p>Remarks: The new species Sarametra subtilis presented morphological similarities to Sarametra triserialis, the type species of the genus. In Sarametra triserialis, the P 1 measures 9.6 mm in length with 22 pinnulars, with the first segment as long as broad. P 1 of Sarametra subtilis is often longer than those in Sarametra triserialis, yet the number of pinnulars is less numerous. These differences in the P 1 dimensions, although minor, are consistent across all specimens. Furthermore, Messing and White (2001) described S. triserialis as having long, slender, webbed spines on proximal brachials. Here, webbed spines are less prominent and more or less restricted to IBr 1 and axillary (Fig. 6F).</p></div>	https://treatment.plazi.org/id/E66B582DFFCAFF8BFF1EF950D1C2518A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mao, Jingwen;Zhang, Ruiyan;Eléaume, Marc;Zhou, Yadong;Zhang, Dongsheng;Sun, Shao’e;Sha, Zhongli;Wang, Chunsheng	Mao, Jingwen, Zhang, Ruiyan, Eléaume, Marc, Zhou, Yadong, Zhang, Dongsheng, Sun, Shao’e, Sha, Zhongli, Wang, Chunsheng (2025): Description of three new species of Zenometridae (Echinodermata, Crinoidea, Comatulida), with new insights on species and genus delimitation. Zoological Journal of the Linnean Society 204 (4), DOI: 10.1093/zoolinnean/zlaf089, URL: https://doi.org/10.1093/zoolinnean/zlaf089
E66B582DFFD6FF89FC63FABFD0BE5065.text	E66B582DFFD6FF89FC63FABFD0BE5065.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zenometridae A. H. Clark 1909	<div><p>Zenometridae gen.et sp. indet. (Fig. 15)</p><p>Material examined: MNHN-IE-2016-1652, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-147.1447&amp;materialsCitation.latitude=-15.965" title="Search Plazi for locations around (long -147.1447/lat -15.965)">South Pacific</a>, 15.9650°S, 147.1447°W, depth 462–980 m, 1 October 2009 . MNHN-IE-2023-89, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-140.1333&amp;materialsCitation.latitude=-9.3167" title="Search Plazi for locations around (long -140.1333/lat -9.3167)">South Pacific</a>, 9.3167°S, 140.1333°W, depth 1107 m, 3 September 2009 .</p><p>Description: MNHN-IE-2016-1652. Centrodorsal long conical in shape. D = 3.7 mm, H = 7.0 mm, H/D = 1.9. The aboral pole is conical in form. Cirrus sockets separated interradially by narrow flat strip (Fig. 15A).</p><p>Cirri LXI (functional sockets). Functional cirrus sockets in three columns per radial area in centrodorsal. Each column with 3–5 functional sockets. Six to nine obsolete sockets per radial area. Fulcral bowl of cirrus sockets surrounding central lumen. All cirri lost, except one with only 9 cirrals preserved (Fig. 15D). The first cirral retained (probably c 2) compressed, L/W = 0.83. Following 8 cirrals longer than wide. The longest has L/W ratio of 2.3. Cirrals with concave aboral margins.</p><p>Arms 10, all broken. Thin, short spines sparsely distributed on proximal brachials. IBr 1 short, W = 2.1 mm, L = 0.56 mm, W/L = 3.8. Axillary (IBr 2) rhombic in shape, W = 2.5 mm, L = 2.6 mm, W/L = 0.96. IIBr 1 short, W = 2.0– 2.1 mm, L = 0.5– 0.6mm, W/L = 3.2–4.2. IIBr 2 cuneate, W = 2.2 mm, L = 1.9 mm, W/L = 1.2. First syzygy at IIBr 3 +4, W = 2.0 mm. Pinnules broken. The first pinnular of P 1 with L/W = 2.4.</p><p>MNHN-IE-2023-89. Centrodorsal conical in shape. D = 3.9 mm, H = 9.6 mm, H/D = 2.5. The aboral pole is conical in form. Cirrus sockets separated interradially by narrow flat strip (Fig. 15B).</p><p>Cirri LVI (functional sockets). Cirri all lost. Functional cirrus sockets in three columns per radial area in centrodorsal. Each column with 3–5 functional sockets. Nine to thirteen obsolete sockets per radial area. Fulcral bowl of cirrus sockets surrounding central lumen.</p><p>Arms 10, all broken. Sparse and short spines on proximal brachials. IBr 1 short, W = 2.3 mm, L = 0.8 mm, W/L = 2.9. Axillary (IBr 2) rhombic in shape (Fig. 15C), W = 2.8 mm, L = 3.1 mm, W/L = 0.9. IIBr 1 short, W = 2.2 mm, L = 0.68 mm, W/L = 3.2. IIBr 2 cuneate, W = 2.6 mm, L = 2.1 mm, W/L = 1.2. First syzygy at IIBr 3+ 4, W = 2.1–2.3 mm. Pinnules broken. The first pinnular of P 1 with L/W = 2.2.</p></div>	https://treatment.plazi.org/id/E66B582DFFD6FF89FC63FABFD0BE5065	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mao, Jingwen;Zhang, Ruiyan;Eléaume, Marc;Zhou, Yadong;Zhang, Dongsheng;Sun, Shao’e;Sha, Zhongli;Wang, Chunsheng	Mao, Jingwen, Zhang, Ruiyan, Eléaume, Marc, Zhou, Yadong, Zhang, Dongsheng, Sun, Shao’e, Sha, Zhongli, Wang, Chunsheng (2025): Description of three new species of Zenometridae (Echinodermata, Crinoidea, Comatulida), with new insights on species and genus delimitation. Zoological Journal of the Linnean Society 204 (4), DOI: 10.1093/zoolinnean/zlaf089, URL: https://doi.org/10.1093/zoolinnean/zlaf089
