taxonID	type	description	language	source
E6209671FF8E050FFF07B2FEFAF566CF.taxon	diagnosis	Diagnosis. Medium to large, body 5.7 – 7.6 mm long, legs and halteres black. Hairs on thorax and abdomen whitish, scutum with long dense hairs leaving only inner stripes bare, acrostichals and dorsocentrals mostly white, humeral bristle distinct. Female scutum with few short hairs, legs without pennation or flattened hairs.	en	Çiftçi, Mustafa Cemal, Hasbenli, Abdullah, Canpolat, Derya (2012): Two new species of Empis (Diptera: Empididae) from Turkey. Zootaxa 3408: 47-53, DOI: 10.5281/zenodo.208769
E6209671FF8E050FFF07B2FEFAF566CF.taxon	description	Description. Male. Eyes meeting on frons for long distance, upper ommatidia enlarged. Face dull grey, lower edge shinning black. Pair of ocellar bristles as long as other occipital bristles. Occiput grey, upper occipital bristles black, lower part of occiput with thin whitish hairs. Antenna black, basal segments (scape and pedicel) with black bristly hairs as long as corresponding segments, stylus as long as half-length of 3 rd antennal segment (postpedicel). Palpus yellow, basally black with sparse bristly hairs. Labrum 1.5 X as long as height of head. Thorax grey, scutum with 4 dull black stripes, 2 narrow inner stripes separating acrostichals and dorsocentrals, outer stripes 2 X broader than inner stripes. Scutum with long dense whitish hairs as long as length of 1 st (scape) and 2 nd (pedicel) antennal segments leaving only inner stripes bare. Acrostichals and dorsocentrals whitish and as long as other hairs on scutum, acrostichals irregularly 3 – 5 serial, dorsocentrals multiserial also with row of black, slightly thicker bristles and ending with 2 pairs of long prescutellar bristles. Humeri (postpronotum) covered with whitish hairs, humeral bristle black anteriorly with 3 – 4 smaller bristles. Large thoracic bristles black, 1 intrahumeral, 3 notopleural, 1 supra-alar, 1 postalar, 2 – 4 scutellar bristles. Prothorax with whitish hairs. Metapleural fan (laterotergal bristles) long and whitish. Wing almost clear, all veins dark and distinct, only wing base yellowish. Costal bristle absent. Squama yellow with white fringes. Haltere with black knob and brownish stem. Legs long and slender, shinning black to blackish brown except for greyish dusted coxae. Coxae with long whitish hairs and 4 – 5 outer black bristles. Femora with short black hairs, mid and hind femora ventrally with row of spine-like bristles. Tibiae ventrally with short adpressed hairs, dorsal hairs bristle-like, slightly longer than ventral hairs. Mid tibia anteroventrally with row of short bristles. Tarsal segments with dense short hairs and apical circlet of bristles. Basitarsi (tarsomere 1) ventrally with spine-like bristles. Abdomen grey, terga much darker and slightly brownish pollinose in dorsal view. Terga 5 and 6 black and brownish pollinose, sternum 8 and genitalia shinning black with slightly brownish dusting. Abdominal hairs whitish, lateral hairs on anterior segments long. Terga 2 – 4 and 6 laterally with 1, tergum 5 laterally with 2 marginal bristles. Sternum 8 with sparse hairs and long and thick hind marginal bristles. Genitalia (terminalia) (Figs. 1 – 3) large and black with black marginal hairs, lower hind edge of lateral lamella (epandrium) brownish. Lateral lamella (epandrium) large, almost square, posterior edge oval (Fig. 1). Dorsal lamella (cercus) with short black hairs, anteriorly bilobed; inner lobe very narrowed and finger-like in dorsal view; inner edge on apical half serrate in dorsal view (Fig. 2). Holotype male body length: 6.8 mm, wing length: 7.1 mm. Paratypes body length: male 5.7 – 7 mm; female 7 – 7.4 mm, wing length: male 6 – 7.2 mm; female 7.3 – 7.6 mm. Female. All main differential characters as in male but hairs and bristles shorter. Frons grey and wide. Whitish thin hairs on thorax and head considerably reduced. Dense whitish hairs on mesonotum in male reduced to few short hairs in female. Acrostichals and dorsocentral bristles black, very short and spine-like. Large thoracic bristles as in male but shorter. Wings clearly brownish. Legs long, slender and shinning black, without pennation and flattened hairs. Hairs and bristles on coxae short, ventral spine-like bristles on mid femur very shortened. Abdomen dull light grey with very short and sparse hairs, no hind marginal bristles. Last segments shinning black.	en	Çiftçi, Mustafa Cemal, Hasbenli, Abdullah, Canpolat, Derya (2012): Two new species of Empis (Diptera: Empididae) from Turkey. Zootaxa 3408: 47-53, DOI: 10.5281/zenodo.208769
E6209671FF8E050FFF07B2FEFAF566CF.taxon	discussion	Remarks. Empis nabucco is closely related to a small group of species (E. syriaca Collin, E. dedecor Loew and E. engeli Chvála) within E. haemorrhoica - group due to the densely haired mesonotum, black legs and halteres, whitish abdominal hairs and metapleural fan (laterotergal bristles). Empis nabucco mostly resembles E. dedecor and E. syriaca having long and dense pale hair-like acrostichals and dorsocentrals, but the latter two species have the humeral bristle not differentiated. Empis nabucco shares distinct humeral bristles as in E. engeli, but with additional 3 – 4 smaller anterior bristles. Female legs of E. nabucco are similar to E. dedecor in lacking pennate setae, but the female of E. nabucco is clearly differentiated from the female of E. dedecor by the short and reduced mesonotal hairs, spine-like acrostichals and dorsocentrals and completely grey abdomen except for the last shinning black segments.	en	Çiftçi, Mustafa Cemal, Hasbenli, Abdullah, Canpolat, Derya (2012): Two new species of Empis (Diptera: Empididae) from Turkey. Zootaxa 3408: 47-53, DOI: 10.5281/zenodo.208769
E6209671FF8E050FFF07B2FEFAF566CF.taxon	materials_examined	Type material. HOLOTYPE 3 (spn 13685): TURKEY. Ankara, Bala, northwest of Bahçekaradalak village, 1212 m, 04. vi. 2011. PARATYPES (spn 12648, 13649, 13686 - 13759): TURKEY. 3 Ƥ, Ankara, Bala, S of Abazlı village, 1133 m, 04. vi. 2011; 33, 20 Ƥ, Ankara, Bala, NW of Bahçekaradalak village, 1212 m, 04. vi. 2011; 4 Ƥ, Ankara, Bala, N of Bahçekaradalak village, 1216 m, 04. vi. 2011; 1 Ƥ, Yozgat, Akdaġmadeni, W of Bozhüyük village, 1785 m, 07. vi. 2011; 3 Ƥ, Sivas, Şarkışla, S of Hocabey village, 1453 m, 07. vi. 2011; 13, 3 Ƥ, Sivas, Şarkışla, NW of Gaziköy village, 1389 m, 07. vi. 2011; 23, 8 Ƥ, Sivas, Merkez, N of Büyükyüreġil village, 1441 m, 08. vi. 2011; 2 Ƥ, Sivas, Merkez, N of Kartalca village, 1366 m, 08. vi. 2011; 4 Ƥ, Sivas, Zara, W of Mahmutaġaçiftliġi village, 1412 m, 09. vi. 2011; 33, Sivas, Hafik, W of Topçuyeniköy village, 1438 m, 09. vi. 2011; 13, 2 Ƥ, Sivas, Hafik, N of Bakımlı village, 1374 m, 09. vi. 2011; 23, Sivas, Merkez, SE of Karayün village, 1477 m, 09.06.2011; 1 Ƥ, Sivas, Merkez, E of Yeniboġazkesen village, 1399 m, 09.06.2011; 1 Ƥ, Sivas, Hafik, E of Akpınar village, 1481 m, 10. vi. 2011; 2 Ƥ, Sivas, İmranlı, W of Kılıçköy, 1577 m, 10. vi. 2011; 1 Ƥ, Sivas, İmranlı, SE of Yaylacık village, 1623 m, 10. vi. 2011; 4 Ƥ, Sivas, İmranlı, W of Piredede village, 1727 m, 10. vi. 2011; 13, 2 Ƥ, Erzincan, Refahiye, NW of Kandilköy village, 2138 m, 11. vi. 2011; 13, Sivas, İmranlı, S of Toklucak village, 1700 m, 11. vi. 2011; 1 Ƥ, Erzincan, Refahiye, SE of Yurtbaşı village, 1728 m, 21. vi. 2011.	en	Çiftçi, Mustafa Cemal, Hasbenli, Abdullah, Canpolat, Derya (2012): Two new species of Empis (Diptera: Empididae) from Turkey. Zootaxa 3408: 47-53, DOI: 10.5281/zenodo.208769
E6209671FF8E050FFF07B2FEFAF566CF.taxon	etymology	Derivatio nominis. This species is named after the Nabucco Gas Pipeline Project, the environmental baseline field studies for which provided the opportunity and financial support for the collection of the type materials.	en	Çiftçi, Mustafa Cemal, Hasbenli, Abdullah, Canpolat, Derya (2012): Two new species of Empis (Diptera: Empididae) from Turkey. Zootaxa 3408: 47-53, DOI: 10.5281/zenodo.208769
E6209671FF8C050EFF07B093FCBF6699.taxon	diagnosis	Diagnosis. Large (body 6 – 7 mm long), thoracic and abdominal hairs and bristles whitish yellow, legs and upper occipital hairs black. Halteres yellow with darker stem. Female legs without pennation or flattened hairs.	en	Çiftçi, Mustafa Cemal, Hasbenli, Abdullah, Canpolat, Derya (2012): Two new species of Empis (Diptera: Empididae) from Turkey. Zootaxa 3408: 47-53, DOI: 10.5281/zenodo.208769
E6209671FF8C050EFF07B093FCBF6699.taxon	description	Description. Male. Head black with grey dusting. Eyes with upper ommatidia enlarged. Face and frons grey dusted, face wide and broadened through mouth edge. Occiput grey, upper half of occiput with 2 rows of black occipital bristly-hairs, lower half around mouth opening with yellowish hairs. Antenna black, 1 st (scape) and 2 nd (pedicel) antennal segment with short black hairs, stylus as long as 3 rd antennal segment (postpedicel). Palpus black, with thin black hairs. Labrum twice as long as height of head. Thorax densely grey dusted, scutum also with slightly yellowish dusting. Scutum in frontal and dorsal view subshinning, in posterior and lateral view with 3 dull black stripes. Mesonotum densely clothed with yellowish hairs leaving only area between acrostichal and dorsocentral rows bare. Acrostichals and dorsocentrals yellowish, nearly as long as antennal stylus, acrostichals irregularly 3 – 4 serial, multiserial dorsocentral ending with 3 – 4 long yellowish prescutellar pairs. Large thoracic bristles yellowish, humeral bristle not differentiated because of long hairs on humeri (postpronotum), 4 notopleural, 2 supra-alars, 1 postalar and 2 pairs of scutellar yellowish and distinct. Prothorax with yellowish white hairs, pronotum with row of short hairs. Metapleural fan (laterotergal bristles) whitish. Wing very faintly brownish, all veins dark and distinct, only wing base yellowish. Costal bristle not differentiated, stigma black. Anal vein complete. Squama yellowish with pale fringes. Haltere yellow with darker stem. Legs long and slender, shinning black except for densely grey coxae. Hind tibia dilated towards tip, fore basitarsus slightly, hind basitarsus clearly swollen but not wider than tip of hind tibia. Coxae with whitish hairs. Fore femur dorsally with black, ventrally with sparse long yellowish hairs. Mid and hind femora with black dorsal hairs on apical half, other hairs pale, basally long shortened towards tip. Ventral hairs on mid and hind femora bristlelike. Hind femur anteroventrally with row of bristly hairs slightly longer than width of hind femora. Fore tibia slen- der, covered with short black hairs, dorsal hairs slightly longer. Mid tibia (Fig. 4) with short black hairs, anterodorsally and ventrally with row of bristles, apically and basally shortened ventral row long at middle. Hind tibia dorsally with 2 rows of bristles, ventrally with short adpressed bristly hairs. Tarsal segments with short hairs and apical circlets of bristly-hairs. Fore and mid basitarsi and hind tarsal segments ventrally with short spine-like hairs. Hind basitarsus dorsally with long hairs. Abdomen black, dorsum black, subshinning and slightly grey dusted, sterna densely grey dusted. Abdominal hairs whitish, long on anterior segments and shortened towards tip. Sternum 8 with long yellowish bristles on hind margin. Genitalia (terminalia) (Fig. 5) large and distinct. Aedeagus (phallus) (Fig. 6) brownish yellow, long and bow-shaped, sligthly thickened at base. Lateral lamella (epandrium) triangular, hind margin with long pale hairs. Dorsal lamella (cercus) with short dark hairs, anteriorly oval, suddenly narrowed and finger-like at tip. Holotype male body length: 6.2 mm, wing length: 6.8 mm. Paratypes body length: male 5.6 – 6.4 mm; female 6.5 – 7.1 mm, wing length: male 6.2 – 7 mm; female 6.3 – 6.9 mm. Female. Main differential characters and general appearance as in male with shorter hairs. Wings distinctly brownish. Abdomen densely grey dusting on terga and sterna. Legs shinning black but especially on femora with slightly grey dusting. Fore basitarsus (tarsomere 1) simple, hind basitarsus and tip of hind tibia little swollen as in male. Legs without pennation and flattened hairs. Other characters on legs as in male with shorter bristles.	en	Çiftçi, Mustafa Cemal, Hasbenli, Abdullah, Canpolat, Derya (2012): Two new species of Empis (Diptera: Empididae) from Turkey. Zootaxa 3408: 47-53, DOI: 10.5281/zenodo.208769
E6209671FF8C050EFF07B093FCBF6699.taxon	discussion	Remarks. Considering the dense, long and soft yellowish white hairs on the mesonotum, E. doronicola is very much like E. decora Meigen, but in the former species the mesonotum has only yellowish white hairs including strong notopleurals, supra-alars, postalar and also scutellar setae. Empis decora have black bristling and hairing on posterior part of mesonotum. Female legs not pennate in E. doronicola but in E. decora female legs are pennate.	en	Çiftçi, Mustafa Cemal, Hasbenli, Abdullah, Canpolat, Derya (2012): Two new species of Empis (Diptera: Empididae) from Turkey. Zootaxa 3408: 47-53, DOI: 10.5281/zenodo.208769
E6209671FF8C050EFF07B093FCBF6699.taxon	materials_examined	Type material. HOLOTYPE 3 (spn 13760): TURKEY. Yozgat, Akdaġmadeni, north of Davulbaz village, 1737 m, 06. vi. 2011. PARATYPES (spn 13761 - 13802): TURKEY. 83, 10 Ƥ, Yozgat, Akdaġmadeni, SE of Özer village, 1839 m, 06. vi. 2011; 133, 11 Ƥ, Yozgat, Akdaġmadeni, N of Davulbaz village, 1737 m, 06. vi. 2011.	en	Çiftçi, Mustafa Cemal, Hasbenli, Abdullah, Canpolat, Derya (2012): Two new species of Empis (Diptera: Empididae) from Turkey. Zootaxa 3408: 47-53, DOI: 10.5281/zenodo.208769
E6209671FF8C050EFF07B093FCBF6699.taxon	etymology	Derivatio nominis. The species is named after the plant Doronicum orientale Hoffmann, 1808 from the family Asteraceae, upon which the type material were collected.	en	Çiftçi, Mustafa Cemal, Hasbenli, Abdullah, Canpolat, Derya (2012): Two new species of Empis (Diptera: Empididae) from Turkey. Zootaxa 3408: 47-53, DOI: 10.5281/zenodo.208769
