identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
E631879BBF68F025FF53FD51E1927A2F.text	E631879BBF68F025FF53FD51E1927A2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Benstonea odoardoi	<div><p>Benstonea odoardoi is a synonym of the large tree B. lauterbachii</p><p>Benstonea lauterbachii (Schumann &amp; Warburg in Warburg 1900: 81) Callmander &amp; Buerki in Callmander et al. (2012: 335).</p><p>Basionym: Pandanus lauterbachii Schumann &amp; Warburg in Warburg (1900: 81). Type:— PAPUA NEW GUINEA. Madang Prov.: Gogol, 28.X.1890, Lauterbach 863 (holo-: B [B 10 0216880]!).</p><p>Syn.: Pandanus odoardoi Martelli (1904: 304) . Benstonea odoardoi (Martelli) Callmander &amp; Buerki in Callmander et al. (2012: 336) [as “odoardii” correctable under ICN Art. 60.12 and Rec. 60C.1(a)]. Type:— PAPUA NEW GUINEA. Western Prov.: Fly river, 1877, d’Albertis s.n. (holotype, FI-B [FI003939, 3 sheets: Erbario Beccari 1526, 1526A, 1526B]!; isotype, FI [FI003940]!), syn. nov.</p><p>Notes:— Benstonea odoardoi was accepted by Callmander et al. (2012) following current taxonomic species circumscription (Stone 1978; Jebb 1992). New investigations of the type specimen (d’Albertis s.n.) deposited at FI showed a spicate infrutescence bearing massive oblong syncarps on a strong peduncle (Fig. 1A–B). It is considered here as a synonym of the earliest published B. lauterbachii, a well-known and widespread large tree species growing on wet soils in swampy lowland evergreen tropical forest often along river banks in New Guinea (Fig. 2B–C) and northern Australia (Jebb 1992; Wilson 2011).</p><p>Selected additional specimens seen: — INDONESIA [NEW GUINEA]. Papua Prov.: Nabire, river Boemi, 21.IV.1939, Inokuma 677 (FU) ; Mamberamo Raya, Aruh Creek, 19.XI.2008, Keim 1157 (BO) ; Mimika, Timika, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=136.96666&amp;materialsCitation.latitude=-4.3333335" title="Search Plazi for locations around (long 136.96666/lat -4.3333335)">Mt Jaya</a>, 4°20’S 136°58’E, 19.VIII.1998, Sands 7218 (A, BRI, K, L, MAN, SING) ; Jayapura, Cape Soeadja, 20.XII.1962, St. John 26162 (B carpo, BISH) . PAPUA NEW GUINEA. Central Prov.: Veiya river, 8.III.1935, Carr 1600 (K, L). East Sepik Prov.: Malu, 1.II.1913, Ledermann 10811 (B [B 010 0279960]) ; Arufufindi river, 29.VII.1967, Millar &amp; Dockrill NGF35188 (L) ; Brown river, 20 miles NE of Port Moresby, 26.I.1958, St. John 26231 (BISH, FI). Gulf Prov.: Lepokera, Vailala river, 16.II.1926, Brass 986 (FI). Morobe Prov.: Melahany [?] village, LAE, 4.I.1958, St. John 26145 (BISH, FI carpo, L) ; Margin of Sepik River between Mamber and Angoram, 31.XII.1962, St. John 26185 (BISH, P carpo). Western Prov.: Wuroi, Oriomo river, II–III.1934, Brass 5791 (FI) ; Lower Fly river, X.1936, Brass 8139 (A, L) .</p></div>	https://treatment.plazi.org/id/E631879BBF68F025FF53FD51E1927A2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Callmander, Martin W.;Keim, Ary P.;Heatubun, Charlie D.;Homot, Peter;Buerki, Sven	Callmander, Martin W., Keim, Ary P., Heatubun, Charlie D., Homot, Peter, Buerki, Sven (2016): Lifting the curtain on our knowledge of New Guinean Benstonea (Pandanaceae). Phytotaxa 275 (2): 168-174, DOI: 10.11646/phytotaxa.275.2.8, URL: http://dx.doi.org/10.11646/phytotaxa.275.2.8
E631879BBF68F024FF53F8FDE2F87C8F.text	E631879BBF68F024FF53F8FDE2F87C8F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Benstonea eumekes (H. St. John ex B. C. Stone 1978) Callm. & Buerki 2016	<div><p>Benstonea eumekes (H. St. John ex B.C. Stone) Callm. &amp; Buerki, comb. nov.</p><p>Basionym: Pandanus eumekes H. St. John ex B.C. Stone (1978: 43) . Type: — INDONESIA [NEW GUINEA]. West Papua Prov.: Sorong Dist., S of Radio Station Sorong, 7.IV.1954, P. Royen (holotype, L [L0050509]!).</p><p>Notes: — Stone (1978: 43) described this species based on its very long style ( “eumekes ” from Greek meaning “of a good height”). Benstonea eumekes differs from B. ihuana by its narrower and smaller leaves (125 × 1–2 cm) (vs. 250 × 6 cm) and by its drupes that have a two third connate pileus with the upper part not deciduous at maturity and a long,</p><p>fine and linear stigma (vs. half connate, the upper part deciduous and a pyramidal, angular stigma). This species is only known by the type collection from the West Papua Province in Indonesia.</p></div>	https://treatment.plazi.org/id/E631879BBF68F024FF53F8FDE2F87C8F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Callmander, Martin W.;Keim, Ary P.;Heatubun, Charlie D.;Homot, Peter;Buerki, Sven	Callmander, Martin W., Keim, Ary P., Heatubun, Charlie D., Homot, Peter, Buerki, Sven (2016): Lifting the curtain on our knowledge of New Guinean Benstonea (Pandanaceae). Phytotaxa 275 (2): 168-174, DOI: 10.11646/phytotaxa.275.2.8, URL: http://dx.doi.org/10.11646/phytotaxa.275.2.8
E631879BBF69F024FF53FEE1E0627E87.text	E631879BBF69F024FF53FEE1E0627E87.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Benstonea ihuana (Callm. & Buerki 2016) Callm. & Buerki 2016	<div><p>Benstonea ihuana (Martelli) Callm. &amp; Buerki, comb. nov.</p><p>Pandanus ihuanus Martelli (1929: 141) . Type:— PAPUA NEW GUINEA. Central Prov.: Vailala River, 14.II.1926, Brass 978 (holotype, A [A00020041, A00020042] images seen; isotypes, BRI [BRI-AQ0695558 carpo] image seen, FI [FI016602]!).</p><p>Notes:— Benstonea ihuana is easily recognized by its upper part of the pileus dehiscent at maturity and constricted at the base as well as its wide leaves (up to 6 cm). This species is most similar to Benstonea eumekes, but differs by the characters mentioned above. Furthermore, this taxon holds a short woody stipe, which seems to be unique in this group of species. Benstonea ihuana is known only by two collections from Papua New Guinea. Field observations have nevertheless been done in the Foja Mountains by T. Laman (Papua Prov., Indonesia) and near Wagau village (Morobe Prov., Papua New Guinea) by PH (Fig. 2A–B).</p><p>Additional specimens seen: — PAPUA NEW GUINEA. Central Prov.: Dieni, Ononge road, 1933, Brass 3924 (A, BO, BRI, FI, NY).</p></div>	https://treatment.plazi.org/id/E631879BBF69F024FF53FEE1E0627E87	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Callmander, Martin W.;Keim, Ary P.;Heatubun, Charlie D.;Homot, Peter;Buerki, Sven	Callmander, Martin W., Keim, Ary P., Heatubun, Charlie D., Homot, Peter, Buerki, Sven (2016): Lifting the curtain on our knowledge of New Guinean Benstonea (Pandanaceae). Phytotaxa 275 (2): 168-174, DOI: 10.11646/phytotaxa.275.2.8, URL: http://dx.doi.org/10.11646/phytotaxa.275.2.8
E631879BBF69F024FF53FCE9E41079F7.text	E631879BBF69F024FF53FCE9E41079F7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Benstonea rostellata (Merr. & L. M. Perry 2016) Callm. & Buerki 2016	<div><p>Benstonea rostellata (Merr. &amp; L.M. Perry) Callm. &amp; Buerki, comb. nov.</p><p>Basionym: Pandanus rostellatus Merrill &amp; Perry (1939: 182) . Type:— PAPUA NEW GUINEA. Central Prov.: Kubuna, 15.III.1933, Brass 5675 (holotype, NY [NY00311360] image seen; isotype, A [A00020069] image seen, FI [FI015084]!) .</p><p>Syn.: Pandanus inokumae Kanehira (1940: 255) . Type: — INDONESIA [NEW GUINEA]. Papua Prov.: Nabire, Inokuma 574 (holotype, FU image seen; isotype, A [A00020043] image seen). Pandanus pentagonos St. John (1973: 50) . Type:— PAPUA NEW GUINEA. Morobe Prov.: Waret, M.S. Clemens 1537 (holotype, BR [BR0000005235674] image seen; isotype, L [L0050634]!)</p><p>Notes:— Benstonea rostellata is here circumscribed to include Pandanus inokumae and P. pentagonos as suggested by Stone (1978). This species can be recognized by its sub-elliptic syncarp disposed on a long peduncle and its pyramidal attenuate pileus ending into a linear stigma (Fig. 2C). Several collections represent immature fruits with a much shorter peduncle. More field-work and collections are required to further assess species delimitation within this taxon.</p><p>Additional specimens seen: — INDONESIA [NEW GUINEA]. Papua Prov.: Mimika, Timika, Mt Jaya, 12.III.1999, Willis 118 (BO, K, G, MAN). West Papua Province: Kabupaten Kaimana, Yamor Dist., Danau Yamor, Kampung Urubika (Gariau), forest behind Kampung, 6.VI.2015, Heatubun 1253 (MAN) . PAPUA NEW GUINEA. East Sepik Prov.: Aprilfluss, 19.XI.1912, Ledermann 9756 (B [B 010 0673216]; FI [FI014902]) ; Lordberg, 1.XII.1912, Ledermann 9953 (FI [FI014853]). Lordberg, 8.XII.1912, fr., Ledermann 10216 (B [B 010 0673215]). Morobe Prov.: Sattelberg, J. &amp; M. S. Clemens 861 (G [G00368998], L) ; Quembung mission, s.d., J. &amp; M. S. Clemens 1128 (L) ; Dallmann, 2.III.1940, Inokuma s.n. (FU) ; Bupu Village, Wampit, 13.VII.1967, Millar NGF 22970 (LAE) .</p></div>	https://treatment.plazi.org/id/E631879BBF69F024FF53FCE9E41079F7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Callmander, Martin W.;Keim, Ary P.;Heatubun, Charlie D.;Homot, Peter;Buerki, Sven	Callmander, Martin W., Keim, Ary P., Heatubun, Charlie D., Homot, Peter, Buerki, Sven (2016): Lifting the curtain on our knowledge of New Guinean Benstonea (Pandanaceae). Phytotaxa 275 (2): 168-174, DOI: 10.11646/phytotaxa.275.2.8, URL: http://dx.doi.org/10.11646/phytotaxa.275.2.8
E631879BBF69F022FF53FA39E3EF7DAF.text	E631879BBF69F022FF53FA39E3EF7DAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Benstonea papuana Callm. & Buerki 2016	<div><p>Benstonea papuana Callm. &amp; Buerki, nom. nov.</p><p>Basionym: Pandanus papuanus Ridley (1916:237), nom.illeg. [non Pandanus papuanus Solms-Laubach (1883: 93)]. Type:— INDONESIA [NEW GUINEA]. Papua Prov.: Mimika, Mt. Carstensz [now Mt Jaya], XI.1912, Kloss s.n. (holotype, BM [BM000958474]!).</p><p>Notes:— Benstonea papuana is based on a very poor collection made by Cecil Boden Kloss (1877–1949) while he took part in the Second Wollaston Expedition in Dutch New Guinea in 1912–1913 (Ridley 1916). This species was the taxonomic entity accepted under the epithet “ odoardoi ” in the genus Pandanus Parkinson (1773: 76) by Merrill &amp; Perry (1939) and Stone (1978). Benstonea papuana is a poorly known species differing from the other species of this group by its rounded free pileus at the base of the style (Fig. 2D). Brass 7190 represents a better-preserved collection than the type specimen and is a good representative of the morphology of this species.</p><p>Additional specimens seen: — INDONESIA [NEW GUINEA]. West Papua Prov.: Teluk Wondama, Windesi, VII.1909, von Balen s.n. (FI) ; Kota Sorong, Klasaman, Taman Wisata Alam, 29. V.2010, Heatubun et al. 1017 (MAN). Papua Prov.: Mamberamo Raya, Idenburg river, III.1939, Brass 13620 (A, BO, L) . PAPUA NEW GUINEA. Western Prov.: Palmer river, 2 mi below black river junction, VI.1936, Brass 7190 (A, BM, BO, L, LAE) .</p><p>Benstonea permicron (Kanehira 1940: 258) Callmander &amp; Buerki in Callmander et al. (2012: 336). Type:— INDONESIA [NEW GUINEA]. Papua Prov.: Nabire, Dallman river, 2.III.1940, Inokuma 603 (holotype, FU image seen: isotype, A [A00020067] image seen).</p><p>Syn.; Pandanus bintuniensis Wiriadinata (2009: 443) . Type:— INDONESIA [NEW GUINEA]. West Papua Prov.: Bintuni Bay, Tanah Merah [from cult. in Bogor], XII.2005, Wiriadinata HW 13274 (holotype, BO!), syn. nov .</p><p>Notes: — Pandanus bintuniensis has been described based on a plant cultivated at the Research Centre for Biology of LIPI (Indonesia Institute of Sciences) in Bogor. Its small rounded syncarp (3–5 cm in diam.) with its long pyramidal pileus ending with an acute somewhat beak-shaped stigma is highly similar to Benstonea permicron (Fig. 2E). The only morphological difference between Pandanus bintuniensis and Benstonea permicron relates with the width of the leaves (3.5–5 cm vs. 1–1.5 cm), however the authors do not consider this character to be sufficient to confidently discriminate these two species. We therefore consider Pandanus bintuniensis as a synonym of Benstonea permicron .</p><p>Additional specimens seen: — INDONESIA [NEW GUINEA]. West Papua Prov.: Kabupaten Teluk Bintuni, Kampung Yakati, forest behind Kampung, 26.VII.2010, fr., Heatubun et al. 1063 (MAN) ; Kabupaten Manokwari Selatan, Oransbari Distr., Muari River, 24.II.2016, W. A. Mustaqim et al. 1529 (BO). Papua Prov.: Kabupaten Paniai, Wissel Lake region, 23.II.1939, Eyma 4580 (BO, L) .</p></div>	https://treatment.plazi.org/id/E631879BBF69F022FF53FA39E3EF7DAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Callmander, Martin W.;Keim, Ary P.;Heatubun, Charlie D.;Homot, Peter;Buerki, Sven	Callmander, Martin W., Keim, Ary P., Heatubun, Charlie D., Homot, Peter, Buerki, Sven (2016): Lifting the curtain on our knowledge of New Guinean Benstonea (Pandanaceae). Phytotaxa 275 (2): 168-174, DOI: 10.11646/phytotaxa.275.2.8, URL: http://dx.doi.org/10.11646/phytotaxa.275.2.8
