identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
9AA1FDCD8EC77EDF632AF957B51DBC37.text	9AA1FDCD8EC77EDF632AF957B51DBC37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mordellistena balearica Compte 1985	<div><p>Mordellistena (s. str.) balearica Compte, 1985 Fig. 8</p><p>Mordellistena balearica Compte, 1985: 63-64 (original description); Horák 2008: 96 (distribution).</p><p>Type locality.</p><p>Palma de Mallorca, Majorca.</p><p>Type depository.</p><p>According to the original description (Compte 1985), holotype should be deposited in MNCN. However, despite of the effort of the curator, the specimen was not found.</p><p>Diagnosis.</p><p>Mordellistena balearica was described based on a single male specimen from Mallorca. According to the original description, this species closely resembles M. pseudohirtipes and can be distinguished from this species by longer antennae (antennomeres V–X two times longer than wide) and different shape of parameres (Fig. 8) (Compte 1985). All characters mentioned in the original description suggest that this taxon is conspecific with M. pseudohirtipes; unfortunately, the authors did not have the opportunity to study the type.</p><p>Distribution.</p><p>Known only from type locality.</p><p>Remarks.</p><p>Compte (1985) mentioned following information: "This specimen, together with other specimens collected by P. López in Majorca, which current location I don’t know, was studied by the specialist Mr Ermisch, who considered it as a new species for science, called M. balearica, a name that seems to have remained in litteris". There are several specimens collected by P López in Mallorca in Ermisch’s collection (SNSD) labelled by Ermisch as M. balearica which in fact all belong to M. thuringiaca Ermisch, 1963.</p></div>	https://treatment.plazi.org/id/9AA1FDCD8EC77EDF632AF957B51DBC37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Selnekovic, David;Kodada, Jan	Selnekovic, David, Kodada, Jan (2019): Taxonomic revision of Mordellistenahirtipes species complex with new distribution records (Insecta, Coleoptera, Mordellidae). ZooKeys 854: 89-118, DOI: http://dx.doi.org/10.3897/zookeys.854.32299, URL: http://dx.doi.org/10.3897/zookeys.854.32299
11AC15046C2D472B592C5673521DBA79.text	11AC15046C2D472B592C5673521DBA79.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mordellistena hirtipes Schilsky 1895	<div><p>Mordellistena (s. str.) hirtipes Schilsky, 1895 Figs 1, 4A, B, 5 A–J</p><p>Mordellistena hirtipes Schilsky, 1895: 46 (original description); Heyden et al. 1906: 455 (catalogue); Csiki 1915: 35 (catalogue); Schaufuss 1916: 766 (distribution); Ermisch 1963: 62 (distribution); Ermisch 1965: 268 (distribution); Batten 1976: 168 (distribution); Batten 1977: 171-173 (distribution, figures, key); Horák 1990: 136 (lectotype and paralectotypes designation, figures); Odnosum 2003: 36-37, 40, 46 (key, figures, distribution); Horák 2008: 98 (catalogue, distribution); Odnosum 2010: 153, 192-194 (key, description, figures, distribution); Samin et al. 2016: 24 (distribution); Ruzzier et al. 2017: 152 (distribution).</p><p>Mordellistena aegea Franciscolo, 1949: 90, 93 syn. nov. (original description); Batten 1977: 169 (remarks); Horák 2008: 96 (catalogue).</p><p>Mordellistena podlussanyi Czető, 1990: 26-29 syn. nov. (original description); Horák 2008: 100 (catalogue).</p><p>Type locality.</p><p>Attalia [Turkey].</p><p>Type material examined.</p><p>M. hirtipes: LECTOTYPE (by designation of Horák (1990: 136)): 1 ♂, MNHU: "Attalia Reitter [hand written] / hirtipes Schils. [hand written] / Type [red label] / Zool. Mus. Berlin / [card with dissected genitalia] / LECTOTYPUS [red label] / Mordellistena hirtipes Schils. J. Horák det. 1985"; PARALECTOTYPES (by designation of Horák (1990: 136)): 4 ♂♂, 3 ♀♀, MNHU: "Attalia Reitter [hand written] / Coll. Schilsky / Type [red label] / Zool. Mus. Berlin / PARALECTOTYPUS [red label] / Mordellistena hirtipes Schils. J. Horák det. 1985"; 17 ♂♂, 9 ♀♀, MNHU: "♂ [or] ♀ / Coll. Schilsky / Type [red label] / Zool. Mus. Berlin / PARALECTOTYPUS [red label] / Mordellistena hirtipes Schils. J. Horák det. 1985"; 1 ♀, MNHU: "Syrien Kaifa. Reitter. / Coll. Schilsky / Type [red label] / Zool. Mus. Berlin / PARALECTOTYPUS [red label] / Mordellistena hirtipes Schils. J. Horák det. 1985"; 1 ♀, MNHU: "Morea Hagios Wlassis Brenske / hirtipes [hand written] / Type [red label] / Zool. Mus. Berlin / PARALECTOTYPUS [red label] / Mordellistena hirtipes Schils. J. Horák det. 1985". M. aegea: HOLOTYPE: 1 ♂, MCST: "Pod. Sper. Coo 7. VII.-VIII. [Podere sperimentale, Kos Island; hand written] / 7. VII.-VIII. Pod. Sper. Coo [hand written] / 19 [blue label] / [card with dissected median lobe] / Olo [hand written] Typus / [cover slides with dissected parameres and sternite VIII] / Mordellistena aegea n. sp. DET. FRANCSCOLO / HOLOTYPUS Mordellistena aegea Franciscolo, 1949 D. Selnekovič labelled 2018 / Mordellistena (s. str.) hirtipes Schilsky, 1895 D. Selnekovič det. 2018". M. podlussanyi: HOLOTYPE: 1 ♂, HNHM: "♂ / [transparent plastic board with dissected genitalia] / Krotos KRÉTA / 1981. V. 12. leg. Podlussány / Holotypus Mordellistena podlussanyi Czető, 1988 [red label, hand written] / Mordellistena (s. str.) hirtipes Schilsky, 1895 D. Selnekovič det. 2017".</p><p>Additional material examined.</p><p>Croatia: 1 ♂, HNHM: "Dalmatia leg. Endrödy-Younga / Dubrovnik Ins. Lokrum / 1958. VIII. 7. Kätscher / Mordellistena hirtipes Schils. det. R. Batten 1979 / Mordellistena (s. str.) hirtipes Schilsky, 1895 D. Selnekovič det. 2017". Cyprus: 1 ♂, HNHM: "Cyprus Laranka Glaszner / Mordellistena hirtipes Schils. det. R. Batten 1979 / Mordellistena (s. str.) hirtipes Schilsky, 1895 D. Selnekovič det. 2017"; 12 ♂♂, 2 ♀♀, CSB: "Cyprus W, Limassol env., Germasogeia Reservoir 34°45'19"N, 33°05'36"E, 27. IV. 2018 D. Selnekovič leg. / Mordellistena (s. str.) hirtipes Schilsky, 1895 D. Selnekovič det. 2018". Greece: 1 ♂, SNSD: "♂ / Insel Rhodos 24.5.-5.8.62 / Stadt Rhodos Umg. Dr Mand / Genitalpräparat / MORDELLISTENA pseudohirtipes Erm. K. Ermisch det. 19 / Mordellistena (s. str.) hirtipes Schilsky, 1895 D. Selnekovič det. 2017"; 1 ♂, HNHM: "Crete Biró / Ins. Dia 25.-29. V. / Mordellistena hirtipes Schils. det. R. Batten 1980 / Mordellistena (s. str.) hirtipes Schilsky, 1895 D. Selnekovič det. 2017". Montenegro: 3 ♂♂, 3 ♀♀, CSB: "Montenegro SW Bar city env. 42°07'56"N, 19°07'33"E, 22. VI. 2011 D. Selnekovič / Mordellistena (s. str.) hirtipes Schilsky, 1895 D. Selnekovič det. 2012". 2 ♂♂, 3 ♀♀, CSB: "Montenegro SW Bar city–Volujica hill 242°04'16"N, 19°06'10"E, 20. VI. 2011 D. Selnekovič / Mordellistena (s. str.) hirtipes Schilsky, 1895 D. Selnekovič det. 2012". 2 ♂♂, 2 ♀♀, CSB: "Montenegro SW Bar city–Stari Bar 42°05'31"N, 19°07'58"E, D. Selnekovič 19. VI. 2011 / Mordellistena (s. str.) hirtipes Schilsky, 1895 D. Selnekovič det. 2012". 1 ♂, CSB: "Montenegro SW Bar city, on Daucus 42°06'N, 19°06'E, 19. VI. 2011 D. Selnekovič 2011 / Mordellistena (s. str.) hirtipes Schilsky, 1895 D. Selnekovič det. 2012". Spain: 1 ♂, 1 ♂, SNSD: "Spanien, Prov. Gerona Tossa de mar A. Kampf, VII–VIII 35 / Paratypus Mordellistena geronensis Ermisch / Mordellistena (s. str.) hirtipes Schilsky, 1895 D. Selnekovič det. 2017". 1 ♂, SNSD: "Nordostspanien Costa brava 27. 7. 53 Dr David / Paratypus / PARATYPUS Mordellistena (s. str.) geronensis Ermisch, 1977 Selnekovič labelled 2017 [red label] / Mordellistena (s. str.) hirtipes Schilsky, 1895 D. Selnekovič det. 2017".</p><p>Differential diagnosis.</p><p>Parameres of M. hirtipes are shorter in proportion to the body dimensions than in M. pseudohirtipes and M. purpurascens (EL/LPrL ratio in M. hirtipes: 7.87-9.17 (8.48 ± 0.40, n = 14), M. pseudohirtipes: 4.65-7.17 (5.89 ± 0.71, n = 25) and M. purpurascens: 4.42-5.84 (4.98 ± 0.35, n = 19); EL/RPrL ratio in M. hirtipes: 10.07-11.89 (11.10 ± 0.50, n = 14), M. pseudohirtipes: 5.91-8.63 (7.42 ± 0.72, n = 25) and M. purpurascens: 5.57-6.94 (6.19 ± 0.41 n = 19). Ventral branch of the right paramere is in M. hirtipes (Fig. 5 D–G) usually distinctly shorter than the dorsal one whereas in M. pseudohirtipes (Fig. 6 E–H) and M. purpurascens (Fig. 7 G–J) it is subequal or longer. Basal part of the left paramere is in M. hirtipes (Fig. 5 D–G) distinctly shorter than in M. purpurascens (Fig. 7 G–J). Sides of elytra less convergent apically than in M. pseudohirtipes and M. purpurascens . Terminal maxillary palpomere in females shorter and broader, its inner angle is more acute (Fig. 4B) than in M. pseudohirtipes (Fig. 6D) and M. purpurascens (Fig. 7D).</p><p>Redescription.</p><p>Measurements: TL: ♂♂ 3.21-3.95 mm (3.51 ± 0.24 mm, n = 13), ♀♀ 2.79-4.68 mm (3.44 ± 0.52 mm, n = 9); HL: ♂♂ 0.72-0.93 mm (0.81 ± 0.06 mm, n = 14), ♀♀ 0.67-0.80 mm (0.74 ± 0.04 mm, n = 9); HW: ♂♂ 0.87-1.02 mm (0.94 ± 0.05 mm, n = 14), ♀♀ 0.77-0.96 mm (0.87 ± 0.06 mm, n = 9); PL: ♂♂ 1.04-1.33 mm (1.17 ± 0.11 mm, n = 14), ♀♀ 0.94-1.20 mm (1.09 ± 0.08 mm, n = 9); PW: ♂♂ 1.06-1.37 mm (1.23 ± 0.10 mm, n = 14), ♀♀ 0.98-1.29 mm (1.16 ± 0.10 mm, n = 9); EL: ♂♂ 2.34-2.96 mm (2.65 ± 0.19 mm, n = 14), ♀♀ 2.08-2.79 mm (2.48 ± 0.21 mm, n = 9); EW: ♂♂ 1.10-1.43 mm (1.25 ± 0.11 mm, n = 14), ♀♀ 1.10-1.36 mm (1.23 ± 0.09 mm, n = 9); PTiL: ♂♂ 0.70-0.87 mm (0.77 ± 0.05 mm, n = 14), ♀♀ 0.56-0.73 mm (0.65 ± 0.06 mm, n = 9); PTrL: ♂♂ 0.65-0.74 mm (0.71 ± 0.03 mm, n = 11), ♀♀ 0.57-0.66 mm (0.63 ± 0.03 mm, n = 7); MsTiL: ♂♂ 0.83-1.10 mm (0.97 ± 0.08 mm, n = 14), ♀♀ 0.78-1.01 mm (0.87 ± 0.07 mm, n = 9); MsTrL: ♂♂ 1.06-1.30 mm (1.16 ± 0.06 mm, n = 11), ♀♀ 0.91-1.13 mm (1.05 ± 0.07 mm, n = 9); MtTiL: ♂♂ 0.69-0.91 mm (0.81 ± 0.05 mm, n = 14), ♀♀ 0.66-0.86 mm (0.76 ± 0.06 mm, n = 9); MtTrL: ♂♂ 1.48-1.87 mm (1.67 ± 0.11 mm, n = 10), ♀♀ 1.33-1.69 mm (1.51 ± 0.11 mm, n = 9); PygL: ♂♂ 1.42-1.85 mm (1.58 ± 0.12 mm, n = 13), ♀♀ 1.17-1.56 mm (1.37 ± 0.12 mm, n = 9); TVtL: ♂♂ 0.54-0.87 mm (0.68 ± 0.10 mm, n = 13), ♀♀ 0.44-0.79 mm (0.63 ± 0.11 mm, n = 9); LPrL: 0.29-0.35 mm (0.31 ± 0.02 mm, n = 14); RPrL: 0.22-0.26 mm (0.24 ± 0.01 mm, n = 14); St8L: ♂♂ 0.57-0.63 mm (n = 2); St8W: ♂♂ 0.38-0.40 mm (n = 2).</p><p>Habitus illustrated in Fig. 1. Body slender, widest at the end of anterior third of elytra. Integument black. Head and pronotum covered with yellowish pubescence; pubescence on elytra yellowish in proximal half, gradually darkened towards apices, sometimes with reddish or violet metallic sheen; venter covered with yellowish pubescence, darkened along posterior margins of ventrites 3-5.</p><p>Head moderately convex dorsally, wider than long, widest before middle, HW/HL ratio: ♂♂ 1.10-1.23 (1.17 ± 0.04, n = 14), ♀♀ 1.13-1.23 (1.17 ± 0.03, n = 9). Dorsal surface with microreticulation and small round punctures bearing short setae; ventral surface with transverse microreticulation and sparse, small punctures bearing short setae; small medial triangular part before gula without punctures. Occipital margin rounded in dorsal aspect, straight or slightly concave if seen from behind. Eyes oval, finely faceted with short interfacetal setae. Anterior margin of clypeus straight. Labrum transverse, approximately two times as wide as long, anterior margin straight; surface with microreticulation and small, round punctures bearing short setae. Antennae rather long, slightly serrate (Fig. 5A, B); antennomeres I–IV subequal in length, slightly shorter and slenderer than following ones; antennomeres V–X in males 1.40 –1.60×, in females 1.20 –1.30× as long as wide; antennomere XI oval, ~2.30 × as long as wide. Mandibles symmetrical, bidentate, lateral portions microreticulated with short setae, outer distal portion with group of seven long sensilla; mola well developed, minutely dentate; prostheca well developed, setose. Galea gradually expanded toward apex, covered with apically expanded sensilla; lacinia setose medio-apically, reaching half of length of galea. Maxillary palpomere II distinctly expanded with long setae on ventral side in males (Fig. 4A); not expanded, without long setae in females (Fig. 4B); maxillary palpomere III short, ~1.50 × as long as wide; terminal maxillary palpomere broadly securiform, inner angle situated around middle, TPalL/TPalW ratio: ♂♂ 1.95-2.20 (2.09 ± 0.09, n = 14), ♀♀ 1.95-2.20 (2.10 ± 0.08, n = 9). Terminal labial palpomere fusiform, bearing sparse long sensilla on whole surface, and group of short sensilla at apex (Fig. 5C).</p><p>Pronotum moderately convex, approximately as long as wide, widest just behind middle, PW/PL ratio: ♂♂ 1.00-1.09 (1.05 ± 0.03, n = 14), ♀♀ 1.02-1.07 (1.06 ± 0.02, n = 9). Surface finely microreticulate, with small, rasp-like punctures bearing flat seta. Anterior margin rounded, slightly produced mesally, anterior angles broadly rounded; lateral carinae emarginated in lateral aspect; posterior margin forming short mesal lobe, emarginated before posterior angles; posterior angles rectangular in lateral aspect. Posterior marginal bead interrupted before posterior angles. Hypomeron triangular with round concavity posteriorly. Prosternum in front of procoxae narrow, expanded laterally; prosternal process incomplete, narrow, slightly constricted in the middle. Scutellar shield small, triangular, covered with small, round punctures bearing short setae. Mesoventral process ~0.50 × as wide as mesofemur, parallel-sided, truncate at apex. Metaventrite strongly convex in the middle; surface weakly microreticulated with small, transversally confluent, rasp-like punctures; posterior margin in the produced mesally; discrimen rather indistinct. Metanepisternum trapezoidal, narrowed posteriorly, dorsal margin emarginated, ventral margin straight.</p><p>Elytra long, narrow, widest in anterior 1/3, EL/EW ratio: ♂♂ 2.02-2.26 (2.12 ± 0.08, n = 14), ♀♀ 1.88-2.07 (2.01 ± 0.05, n = 9). Surface with weak transverse microreticulation and rasp-like punctures bearing flat setae. Lateral margins regularly rounded, apices separately rounded.</p><p>Protibiae in males expanded basally, bearing fringe of long setae in basal 1/3; PTiL/PTrL ratio: ♂♂ 1.02-1.17 (1.08 ± 0.05, n = 11), ♀♀ 0.95-1.12 (1.03 ± 0.05, n = 7). Protarsomere I as long as two following tarsomeres combined; protarsomere IV simple, parallel-sided, shallowly emarginate at apex. Claws on protarsi with three, on meso and metatarsi with four denticles. Mesotibiae slightly bent inwards; mesotarsus longer than tibia, MsTiL/MsTrL ratio: ♂♂ 0.78-0.89 (0.82 ± 0.03, n = 11), ♀♀ 0.79-0.90 (0.83 ± 0.03, n = 9). Metacoxae large, anterior margin straight, posterior margin broadly rounded. Metatibiae bearing short subapical ridge and 3-4 lateral ridges parallel with apical margin of tibia, reaching 1/3 of tibial width. Metatibial spurs black, inner one ~1.30 × as long as outer one. Metatarsomere I bearing 4-5 short ridges, metatarsomere II bearing 2-3 ridges, metatarsomeres III and IV without ridges. Metatarsus ~2.00 × as long as metatibia, MtTrL/MtTiL ratio: ♂♂ 1.97-2.18 (2.07 ± 0.06, n = 10), ♀♀ 1.89-2.11 (1.98 ± 0.06, n = 9).</p><p>Pygidium long, slender, narrowly truncate at apex, PygL/TVtL ratio: ♂♂ 2.00-3.04 (2.36 ± 0.26, n = 13), ♀♀ 1.76-3.14 (2.22 ± 0.37, n = 9). Apical margin of terminal abdominal ventrite arcuate.</p><p>Male genitalia: sternite VIII with long setae in apical part, apical margin produced and weakly bilobed mesally (Fig. 5I), St8L/St8W ratio: ♂♂ 1.49-1.59 (n = 2). Sternite IX long, slender, arrow-shaped, with medial longitudinal keel at apex. Parameres (Fig. 5 D–G) rather short, EL/LPrL ratio: 7.87-9.17 (8.48 ± 0.40, n = 14); EL/RPrL ratio: 10.07-11.89 (11.10 ± 0.50, n = 14); LPrL/RPrL ratio: 1.24-1.37 (1.31 ± 0.04, n = 14). Left paramere with short basal part, LPrL/BLPr ratio: 1.88-2.12 (1.98 ± 0.08, n = 14); dorsal branch expanded and obliquely truncate apically; ventral branch slender, slightly bent medially, pointed at apex. Right paramere rather short, ventral branch distinctly shorter than dorsal one, dorsally bent; dorsal branch expanded and rounded apically. Median lobe long, slender, apical part slightly expanded (Fig. 5H). Phallobase with short tubular process (approximately 1/6 of total length) and long, slender furca.</p><p>Female genitalia: sternite VIII with apical protuberance and long setae alongside apical and lateral margins, spiculum ventrale short, broadly clavate (Fig. 5J); St8L/St8W ratio: ♀ 1.36 (n = 1).</p><p>Sexual dimorphism.</p><p>Females are usually more robust; with shorter antennae. Maxillary palpomere II is not expanded in females and without long setae on ventral side. Terminal maxillary palpomere is shorter in females, with angles more rounded. Protibiae are not expanded in females, without long setae in basal portion.</p><p>Distribution.</p><p>Croatia, Cyprus, France, Greece, Iran, Israel, Jordan, Macedonia, Montenegro, Romania, Spain, Syria, Turkey, Turkmenistan, Ukraine. Mordellistena hirtipes is reported here for the first time from Croatia, Montenegro, and Spain. Csiki (1915) mentioned also “Österreich” (Austria); however, this information is probably based on a misidentification.</p><p>Biology.</p><p>Adults were found on the flowers of Daucus sp. ( Apiaceae) and Helichrysum sp. ( Asteraceae) on dry grasslands and in urban environment.</p><p>Remarks.</p><p>Franciscolo (1949) described M. aegea based on three specimens from Kos island (Greece). Batten (1977) mentioned that this species does not belong to the micans group because the antennomere IV and V are equal in length. Examination of holotype revealed that this specimen belongs to M. hirtipes . We consider this taxon as a junior synonym of M. hirtipes .</p><p>Czető (1990) described M. podlussanyi based on a single male specimen from Crete. He mentioned in the original description that the maxillary palpomere II is not dilated. Examination of the holotype actually revealed, that the palpomere is expanded, and any other differences which could separate this taxon from M. hirtipes were found. This interpretation is also supported by the results of PCA analysis (Fig. 10A). We propose M. podlussanyi as a junior synonym of M. hirtipes .</p><p>In HNHM collections, there are three specimens of M. hirtipes, labelled by Reitter as holotype and paratypes. However, these specimens are not mentioned in original description and labels were probably added subsequently, after the description. These specimens are not parts of the type series.</p></div>	https://treatment.plazi.org/id/11AC15046C2D472B592C5673521DBA79	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Selnekovic, David;Kodada, Jan	Selnekovic, David, Kodada, Jan (2019): Taxonomic revision of Mordellistenahirtipes species complex with new distribution records (Insecta, Coleoptera, Mordellidae). ZooKeys 854: 89-118, DOI: http://dx.doi.org/10.3897/zookeys.854.32299, URL: http://dx.doi.org/10.3897/zookeys.854.32299
A538B4C37ADEC2D9152AC08F1C1FD675.text	A538B4C37ADEC2D9152AC08F1C1FD675.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mordellistena hirtipes species complex	<div><p>Mordellistena hirtipes species complex</p><p>Diagnosis.</p><p>Integument including legs and maxillary palpi completely black; metatibial spurs black; pubescence of dorsum yellowish, sometimes darkened in apical portions of elytra but never completely dark. Antennomeres I–IV shorter and narrower than following ones (Figs 5A, B, 6A, B, 7A, B). Maxillary palpomere II expanded in males, bearing very long setae on ventral surface (Figs 4A, 6C, 7C). Metatibiae at least with three lateral ridges, all parallel to apical margin of tibia. Metatarsomeres I and II with ridges.</p></div>	https://treatment.plazi.org/id/A538B4C37ADEC2D9152AC08F1C1FD675	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Selnekovic, David;Kodada, Jan	Selnekovic, David, Kodada, Jan (2019): Taxonomic revision of Mordellistenahirtipes species complex with new distribution records (Insecta, Coleoptera, Mordellidae). ZooKeys 854: 89-118, DOI: http://dx.doi.org/10.3897/zookeys.854.32299, URL: http://dx.doi.org/10.3897/zookeys.854.32299
521C5164BD67A916E12CFBD82F3B5096.text	521C5164BD67A916E12CFBD82F3B5096.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mordellistena irritans Franciscolo 1991	<div><p>Mordellistena (s. str.) irritans Franciscolo, 1991 Fig. 9</p><p>Mordellistena irritans Franciscolo, 1991: 168-173 (original description); Franciscolo 1995: 12 (distribution); Horák 2008: 98 (distribution).</p><p>Type locality.</p><p>Lampedusa Is., Italy.</p><p>Type depository.</p><p>Museo d’Aumale, Terrasini, Palermo, Italy: 1 ♀ holotype, 1 ♂ paratype (Franciscolo 1991). Not examined.</p><p>Diagnosis.</p><p>Mordellistena irritans can be assigned to M. hirtipes complex based on the expanded maxillary palpomere II in males and the shape of parameres. This species can be distinguished from all other species in the complex by the characteristic shape of the left paramere with dorsal branch parallel-sided and rounded at apex (Fig. 9).</p><p>Distribution.</p><p>Known only from the type locality.</p></div>	https://treatment.plazi.org/id/521C5164BD67A916E12CFBD82F3B5096	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Selnekovic, David;Kodada, Jan	Selnekovic, David, Kodada, Jan (2019): Taxonomic revision of Mordellistenahirtipes species complex with new distribution records (Insecta, Coleoptera, Mordellidae). ZooKeys 854: 89-118, DOI: http://dx.doi.org/10.3897/zookeys.854.32299, URL: http://dx.doi.org/10.3897/zookeys.854.32299
79A3E50824AB3DAEF2135DE2BF0B11C8.text	79A3E50824AB3DAEF2135DE2BF0B11C8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mordellistena pseudohirtipes Ermisch 1965	<div><p>Mordellistena (s. str.) pseudohirtipes Ermisch, 1965 Figs 2, 6 A–J</p><p>Mordellistena pseudohirtipes Ermisch, 1965: 268 (original description); Batten 1976: 168 (distribution); Batten 1977: 171-173 (distribution, figures, key); Plaza 1983: 574 (distribution, biology); Czető 1990: 28-29 (description, figure); Franciscolo 1995: 12 (distribution); Horák 2008: 100 (distribution); Odnosum 2010: 153, 194-195 (key, description, distribution); Ruzzier 2013: 109 (distribution).</p><p>Mordellistena fageli Ermisch, 1969: 112 syn. nov. (original description); Batten 1976: 168 (distribution); Plaza 1983: 575-576 (distribution, biology); Horák 1983: 13 (remarks); 2008: 97 (distribution).</p><p>Mordellistena pseudohirtipes krotosensis Czető, 1990: 28 syn. nov. (original description); Horák 2008: 100 (distribution).</p><p>Type locality.</p><p>Nessebar env., Bulgaria.</p><p>Type material examined.</p><p>M. pseudohirtipes pseudohirtipes: HOLOTYPE: 1 ♂, SNSD: "♂ / Genitalpräparat / Bulgaria Umg. Nessebar Juli 1961 leg. BECH / Holotypus [red label] / MORDELLISTENA pseudohirtipes Erm. K Ermisch det. 19 / Coll. ERMISCH Leipzig Ankauf 1970 / Staatl. Museum für Tierkunde Dresden"; PARATYPE: 1 ♀, SNSD: "♀ / Bulgaria Umg. Nessebar Juli 1961 leg. BECH / Allotypus [red label] / MORDELLISTENA pseudohirtipes Erm. K Ermisch det. 19 / Coll. ERMISCH Leipzig Ankauf 1970 / Staatl. Museum für Tierkunde Dresden". M. pseudohirtipes krotosensis: HOLOTYPE: 1 ♂, HNHM: "[transparent plastic board with dissected genitalia] / ♂ / Krotos KRÉTA / 1981. V. 12. leg. Podlussány / Holotypus Mordellistena pseudohirtipes Ermisch, 1965 ssp. krotosensis, Czető 1988 [red label] / Mordellistena (s. str.) pseudohirtipes Ermisch, 1965 D. Selnekovič det. 2017". M. fageli: HOLOTYPE: 1 ♂, SNSD: "♂ / Genitalpräparat / Portugal: Algrave Caldas de Monchique V– 1960 G. Fagel / R. I. Sc. N. B. I. G. 22.145 / Holotypus [red label] / coll. ERMISCH, Leipzig, Ankauf 1970 / Staatl. Museum für Tierkunde Dresden / HOLOTYPUS Mordellistena (s. str.) fageli Ermisch, 1969 D. Selnekovič labelled 2017 [red label] / Mordellistena (s. str.) pseudohirtipes Ermisch, 1965 D. Selnekovč det. 2017"; PARATYPE: 1 ♀, SNSD: "♀ / Portugal: Algarve Caldas de Monchique V– 1960 G. Fagel / R. I. Sc. N. B. I. G. 22.145 / Allotypus [red label] / coll. ERMISCH, Leipzig, Ankauf 1970 / Staatl. Museum für Tierkunde Dresden / ALLOTYPUS (PARATYPUS) Mordellistena (s. str.) fageli Ermisch, 1969 D. Selnekovič labelled 2017 [red label] / Mordellistena (s. str.) pseudohirtipes Ermisch, 1965 D. Selnekovič det. 2017".</p><p>Additional material examined.</p><p>Algeria. 1 ♂, SNSD: “Algérie: Algérois, Kaddous 3 –V– 1954 G. Fagel / R. I. Sc. N. B. I. G. 19.867 / coll. ERMISCH, Leipzig, Ankauf 1970 / Staatl. Museum für Tierkunde Dresden / " Mordellistena (s. str.) pseudohirtipes Ermisch, 1965 D. Selnekovič det. 2017" [in collection as M. fageli]. Bulgaria. 1 ♂, SNSD: "♂ / Nessebar, Bulgaria 28. 5. - 10. 6. 1963 Karl Bleyl / MORDELLISTENA pseudohirtipes Erm. K. Ermisch det. 19"; 1 ♂, CSB: "Bulgaria mer. occ. Sandanski (→ Liljanovo) 5. - 10. 1976 Karel Majer lgt. / Mordellistena (s. str.) pseudohirtipes Ermisch, 1965 D. Selnekovič det. 2016". France. 2 ♂♂, SNSD: "♂ / Genitalpräparat / France Basses Alpes St. Michel l’Observat . 24. 7 - 10. 8. 63 Rudkjöb . / MORDELLISTENA pseudohirtipes Erm. K. Ermisch det. 19"; 1 ♂, SNSD: "♂ / Ardêche 10. 7. 65 Banne Balazuc / MORDELLISTENA pseudohirtipes Erm. K. Ermisch det. 19; 1 ♂ SNSD: "♂ / Genitalpräparat / Südfrankreich Camargue, 13. 6. 1952, leg. Freude / Mordellistena Lopezi Ermisch det. K. Ermisch 63" [in collection as M. lopezi]; 1 ♂, SNSD: "♂ / Genitalpräparat / Pyrenées or. Umg. Banyuls 30. 5.-10. 6. 53 / Mordellistena Lopezi Ermisch det. K. Ermisch 63" [in collection as M. lopezi]; 1 ♂, SNSD: "Banyuls Pyr. or. VI. 53 J. u. B. Bechyne / Museum Frey München” [in collection as M. lopezi]; 1 ♂, SNSD: "♂ / Gall. mer. Agay (Var) 18. 7. 58 W. Liebmann / Genitalpräparat” [in collection as M. lopezi]; 1 ♂, SNSD: "♂ / Genitalpräparat / Fr. Ardêche Bois de Paiolive 1. 7. 66 Balazuc [hand written] / Paratypus / Staatl. Museum für Tierkunde Dresden / PARATYPUS Mordellistena (s. str.) geronensis Ermisch, 1977, Selnekovič labelled 2017 [red label] / Mordellistena (s. str.) pseudohirtipes Ermisch, 1965 D. Selnekovič det. 2017" [in collection as M. geronensis]. Georgia. 1 ♂, SNSD: "♂ / Genitalpräparat / SSSR–Gruzie Tbilisi 7.57 R. Dvořák / MORDELLISTENA pseudohirtipes Erm. K. Ermisch det. 19 / Coll. ERMISCH Leipzig Ankauf 1970 / Staatl. Museum für Tierkunde Dresden"; 2 ♂♂, SNSD: "♂ / SSSR–Gruzie Tbilisi 7.57 R. Dvořák / MORDELLISTENA pseudohirtipes Erm. K. Ermisch det. 19". Greece. 1 ♂, SNSD: "♂ / Genitalpräparat / Athos Daphni A. Schatzmayr / MORDELLISTENA pseudohirtipes Erm. K Ermisch det. 19"; 1 ♂, SNSD: "♂ / Genitalpräparat / Ephesus / J. Sahlb. / MORDELLISTENA pseudohirtipes Erm. K. Ermisch det. 19"; 1 ♂, HNHM: "Creta Biró / Amari 4. VII. 06 / Mordellistena pseudohirtipes Erm. det. R. Batten 1980 / Mordellistena (s. str.) pseudohirtipes Ermisch, 1965 D. Selnekovič det. 2017". Israel. 1 ♂, CSB: "Israel Jerusalem 25. III. 2001, ??? leg. / Mordellistena s. str. pseudohirtipes Ermisch, 1965 D. Selnekovič det. 2014"; 1 ♂, HNHM: "Izrael Rehovot 1965. V. 20. Dr. Erdös / coll. Dr. J. Erdös / Mordellistena pseudohirtipes Erm. det. R. Batten 1980 / Mordellistena (s. str.) pseudohirtipes Ermisch, 1965 D. Selnekovič det. 2017". Italy. 1 ♂, SNSD: "♂ / Genitalpräparat / Sicilia Magara d. V. 16. 5. 61 W. Liebmann" [in collection as M. lopezi]. Macedonia. 1 ♂, SNSD: "♂ / Veles, Mac. 23. - 25. 5. 55. leg. F. Schubert / Genitalpräparat / MORDELLISTENA pseudohirtipes Erm. K Ermisch det. 19". Montenegro. 2 ♂♂, 1 ♀, CSB: "Montenegro S Skadarske jazero lake N, Virpazar village env. D. Selnekovič 21. VI. 2011 / Mordellistena pseudohirtipes Ermisch, 1965 D. Selnekovič det. 2012". Spain. 1 ♂, SNSD: "♂ / Genitalpräparat / Son Españolet 1 –VI– 1958 R. López / Paratypus [red label] / Mordellistena Lopezi Ermisch det. K. Ermisch / Mordellistena (s. str.) pseudohirtipes Ermisch, 1965 D. Selnekovič det. 2017" [in collection as M. lopezi]; 1 ♂, SNSD: "♂ / Genitalpräparat / SEVILLA Hi. m. Marismas, V. 1943 G. Frey, C. Koch / Mordellistena Lopezi Ermisch det. K. Ermisch 63 / Mordellistena (s. str.) pseudohirtipes Ermisch, 1965 D. Selnekovič det. 2017" [in collection as M. lopezi]; 1 ♂, CSB: "Spain, Málaga, Lagunas de Archidona, 800m 37°06'N, 04°18'40"W, 12.-14. V. 2018 E. Jendek / Mordellistena (s. str.) pseudohirtipes Ermisch, 1965 D. Selnekovič det. 2018". Ukraine. 2 ♂♂, SNSD: "♂ / Genitalpräparat / Umgeb. Jalta Krim, Ende Juli 1965 leg. F. Hieke / MORDELLISTENA pseudohirtipes Erm. K Ermisch det. 19"; 1 ♂, HNHM: "Krim Jaila 17. VI. 1956 leg. L. Horváth / Mordellistena pseudohirtipes Erm. det. R. Batten 1980 / Mordellistena (s. str.) pseudohirtipes Ermisch, 1965 D. Selnekovič det. 2017".</p><p>Differential diagnosis.</p><p>From M. purpurascens it differs in shorter parameres (EL/LPrL ratio: M. pseudohirtipes: 4.65-7.17 (5.89 ± 0.71, n = 25), M. purpurascens: 4.42-5.84 (4.98 ± 0.35, n = 19); EL/RPrL ratio: M. pseudohirtipes: 5.91-8.63 (7.42 ± 0.72, n = 25), M. purpurascens: 5.57-6.94 (6.19 ± 0.41 n = 19)). Basal part of the left paramere (Fig. 6 E–H) is shorter than in M. purpurascens (Fig. 7 G–J). Body usually smaller (TL: M. pseudohirtipes: ♂♂ 2.47-4.05 mm (3.20 ± 0.43 mm, n = 25) ♀♀ 3.26-4.47 mm (3.68 ± 0.56 mm, n = 3), M. purpurascens: ♂♂ 3.10-4.42 mm (3.75 ± 0.35 mm, n = 19), ♀♀ 3.31-4.42 mm (3.82 ± 0.36 mm, n = 14)). Differences between M. hirtipes are mentioned above.</p><p>Redescription.</p><p>Measurements: TL: ♂♂ 2.47-4.05 mm (3.20 ± 0.43 mm, n = 25) ♀♀ 3.26-4.47 mm (3.68 ± 0.56 mm, n = 3); HL: ♂♂ 0.64-0.93 mm (0.75 ± 0.09 mm, n = 25), ♀♀ 0.77-1.04 mm (0.87 ± 0.12 mm, n = 3); HW: ♂♂ 0.70-1.06 mm (0.87 ± 0.11 mm, n = 29), ♀♀ 0.87-1.20 mm (0.99 ± 0.15 mm, n = 3); PL: ♂♂ 0.83-1.29 mm (1.05 ± 0.14 mm, n = 25), ♀♀ 1.06-1.42 mm (1.19 ± 0.16 mm, n = 3); PW: ♂♂ 0.83-1.39 mm (1.07 ± 0.17 mm, n = 29), ♀♀ 1.12-1.67 mm (1.31 ± 0.25 mm, n = 3); EL: ♂♂ 1.88-3.02 mm (2.37 ± 0.31 mm, n = 25), ♀♀ 2.44-3.35 mm (2.75 ± 0.42 mm, n = 3); EW: ♂♂ 0.81-1.46 mm (1.09 ± 0.18 mm, n = 25), ♀♀ 1.17-1.69 mm (1.35 ± 0.24 mm, n = 6); PTiL: ♂♂ 0.57-0.91 mm (0.70 ± 0.10 mm, n = 25), ♀♀ 0.65-0.90 mm (0.74 ± 0.11 mm, n = 6); PTrL: ♂♂ 0.52-0.84 mm (0.63 ± 0.08 mm, n = 24), ♀♀ 0.60-0.80 mm (0.68 ± 0.09, n = 3); MsTiL: 0.71-1.17 mm (0.87 ± 0.12 mm, n = 25), ♀♀ 0.86-1.25 mm (0.99 ± 0.18 mm, n = 3); MsTrL: ♂♂ 0.83-1.34 mm (1.02 ± 0.15 mm, n = 13), ♀♀ 1.04-1.35 mm (n = 2); MtTiL: ♂♂ 0.60-0.92 mm (0.74 ± 0.09 mm, n = 25), ♀♀ 0.74-1.04 mm (0.84 ± 0.14 mm, n = 3); MtTrL: ♂♂ 1.27-1.98 mm (1.59 ± 0.20 mm, n = 17), ♀♀ 1.51-2.05 mm (n = 2); PygL: ♂♂ 1.29-1.96 mm (1.54 ± 0.18 mm, n = 29), ♀♀ 1.39-1.75 mm (1.57 ± 0.14 mm, n = 3); TVtL: ♂♂ 0.56-1.25 mm (0.73 ± 0.13 mm, n = 25), ♀♀ 0.50-0.71 mm (0.64 ± 0.10 mm, n = 3); LPrL: 0.33-0.46 mm (0.40 ± 0.03 mm, n = 25); RPrL: 0.26-0.37 mm (0.32 ± 0.03 mm, n = 25); St8L: ♂ 0.47 mm (n = 1); St8W: ♂ 0.31 mm (n = 1).</p><p>Habitus given in Fig. 2. Body strongly elongate, slender, widest just behind humeri. Integument black, mouthparts sometimes paler. Pubescence on head pale yellowish, on pronotum yellowish to dark grey, on elytra yellowish in anterior 1/2, darkened towards apices, or completely dark grey, sometimes with reddish or purplish metallic sheen, on pygidium dark grey, on venter yellowish, darkened along posterior margins of abdominal ventrites.</p><p>Head moderately convex dorsally, wider than long, widest just before middle, HW/HL ratio: ♂♂ 1.08-1.23 (1.15 ± 0.03, n = 25), ♀♀ 1.11-1.16 (1.13 ± 0.02, n = 3). Dorsal surface weakly microreticulated with small, round punctures bearing short setae. Occipital margin rounded in dorsal aspect, straight, or slightly concave seen from behind. Eyes oval, completely reaching occiput, not expanded onto ventral surface, finely faceted, with short interfacetal setae. Anterior margin of clypeus straight. Labrum transverse, anterior margin straight or very slightly emarginate; surface microreticulation with small, round punctures and setae. Antennae slightly serrate (Fig. 6A, B); antennomeres I–IV subequal in length; antennomeres V–X longer and wider, in males ~1.30 ×, in females ~1.20 × as long as wide; terminal antennomere elongate oval, ~1.90 × as long as wide. Galea gradually expanded toward apex, covered with apically expanded sensilla. Maxillary palpi (Fig. 6 C–D) black; palpomere I very short; palpomere II distinctly expanded, with long setae on ventral side in males, not expanded, rather long and narrow in females; palpomere III short, ~1.50 × as long as wide; terminal palpomere in males broadly securiform, with inner angle situated around the middle, in females slenderer, with inner angle situated in apical 1/3; TPalL/TPalW ratio: ♂♂ 1.80-2.30 (2.07 ± 0.11, n = 25), ♀♀ 2.05-2.31 (2.21 ± 0.12, n = 3).</p><p>Pronotum moderately convex, approximately as long as wide, PW/PL ratio: ♂♂ 0.97-1.12 (1.02 ± 0.03, n = 25), ♀♀ 1.04-1.18 (1.10 ± 0.06, n = 3). Surface finely transversally microreticulate, covered with rasp-like punctures, distance between punctures 2.00-4.00 times as long as the diameter, each puncture bears flat, pointed seta. Anterior margin rounded, slightly produced mesally, anterior angles broadly rounded; lateral carinae emarginated in lateral aspect; posterior margin forming short mesal lobe, emarginated before posterior angles; posterior angles in lateral aspect rectangular, acute. Posterior marginal bead interrupted before posterior angles. Scutellar shield small, triangular, with small, rasp-like punctures bearing setae. Metanepisternum trapezoidal, narrowed posteriorly, ventral margin straight, dorsal margin emarginate.</p><p>Elytra long and narrow, moderately convex, widest at the end of anterior 1/4, EL/EW ratio: ♂♂ 2.02-2.40 (2.18 ± 0.10, n = 25), ♀♀ 1.99-2.08 (2.04 ± 0.04, n = 3). Surface with weak transverse microreticulation and rasp-like punctures, these are larger and more densely arranged than those on pronotum, each puncture bears flat seta. Lateral margins rather strongly convergent, regularly rounded; apices separately rounded.</p><p>Profemora slender, in males somewhat stouter than in females. Protibiae straight, in males distinctly expanded in basal half, here with fringe of long, thick setae; PTiL/PTrL ratio: ♂♂ 1.02-1.24 (1.11 ± 0.05, n = 25), ♀♀ 1.02-1.13 (1.09 ± 0.05, n = 3). Protarsomere I in females as long as two following tarsomeres combined, in males slightly longer; protarsomere IV simple, slightly shorter than previous one, shallowly emarginate at apex; terminal protarsomere slightly shorter than previous two tarsomeres combined. Claws on protarsi with three denticles, on meso- and metatarsi with four denticles. Mesotibiae slightly bent medially; mesotarsus longer than tibia, MsTiL/MsTrL ratio: ♂♂ 0.79-0.90 (0.83 ± 0.03, n = 13), ♀♀ 0.85-0.92 (n = 2). Metacoxae large, anterior margin slightly emarginated, posterior margin broadly rounded. Metatibiae bearing short subapical ridge and 3-4 lateral ridges parallel to apical margin of tibia, reaching 1/3 of tibial width. Metatibial spurs black, long, inner one ~1.30 × as long as outer one. Metatarsomere I bearing 3-5 short ridges, metatarsomere II bearing 2-3 ridges, metatarsomere III without ridges. Metatarsus ~2.00 × as long as metatibia, MtTrL/MtTiL ratio: ♂♂ 2.02-2.32 (2.18 ± 0.07, n = 17), ♀♀ 1.98-2.00 (n = 2).</p><p>Pygidium long and slender, PygL/TVtL ratio: ♂♂ 1.82-2.47 (2.19 ± 0.15, n = 23), ♀♀ 1.88-2.79 (2.22 ± 0.32, n = 6). Apical margin of terminal ventrite arcuate.</p><p>Male genitalia: sternite VIII rather short, setae present in apical 1/3, apical protuberance short, slightly bilobed at apex (Fig. 6J); St8L/St8W ratio: ♂♂ 1.50-1.53 (n = 2). Sternite IX long, slender, arrow-shaped, with medial longitudinal keel in apical part. Parameres (Fig. 6 E–H) rather long, EL/LPrL ratio: 4.65-7.17 (5.89 ± 0.71, n = 25), EL/RPrL ratio: 5.91-8.63 (7.42 ± 0.72, n = 25), LPrL/RPrL ratio: 1.12-1.39 (1.26 ± 0.06, n = 25). Left paramere: dorsal branch expanded apically, obliquely truncate at apex; ventral branch narrow, slightly bent medially, pointed at apex, LPrL/BLPr ratio: 1.73-2.24 (1.95 ± 0.11, n = 25). Right paramere: dorsal branch rather narrow, slightly expanded, rounded at apex; ventral branch as long as or slightly shorter than dorsal branch, bent dorsally in apical part, pointed at apex. Median lobe long, slender, slightly expanded in apical part (Fig. 6I). Phallobase with short tubular process (approximately 1/6 of total length) and long, slender furca.</p><p>Sexual dimorphism.</p><p>Females are more robust than males, their protibiae are not expanded in basal 1/3 and without fringe of long setae. Maxillary palpomere II is not expanded in females and without long setae on ventral side. Terminal maxillary palpomere is slenderer in females and its inner angle is situated more distally than in males. Antennae are somewhat shorter in females.</p><p>Distribution.</p><p>Algeria, Azerbaijan, Bulgaria, Croatia, Cyprus, France, Georgia, Greece, Israel, Italy, Macedonia, Montenegro, Morocco, Portugal, Spain, Turkey, Ukraine. Mordellistena pseudohirtipes is reported here for the first time from Israel and Montenegro.</p><p>Biology.</p><p>Adults were collected on the flowers of Apiaceae plants on dry grasslands. Plaza (1983) mentioned that M. pseudohirtipes was collected in Spain on following plant species: Thapsia villosa, Daucus carota (both Apiaceae) and Ruta montana ( Rutaceae).</p><p>Remarks.</p><p>Mordellistena fageli was placed in the pentas–group in the original description, based on the dark pubescence and three ridges on the metatarsomere II. In fact, as Horák (1983) already mentioned, this species belongs to M. hirtipes complex, based on the strongly convex body and expanded maxillary palpomere II. Examination of type material did not reveal any characters which could separate this taxon from M. pseudohirtipes . In the plot from PCA analysis (Fig. 10A), M. fageli is placed just next to the cluster of M. pseudohirtipes, in the same plane along the PC 1 axis. We consider these taxa as conspecific and propose M. fageli as a junior synonym of M. pseudohirtipes .</p><p>Czető (1990) described M. pseudohirtipes krotosensis based on two male specimens from Crete island. Characters such as length of the body, length of pygidium and colouration of pubescence, that he used for differentiation of the subspecies, are subjects of individual variability. Results of PCA (Fig. 10A) show, that holotype of M. pseudohirtipes krotosensis is placed within the cluster of the nominotypical subspecies. After examination of holotype we consider this subspecies as a junior synonym of M. pseudohirtipes pseudohirtipes .</p><p>In Ermisch’s collection, there is a series of specimens named Mordellistena lopezi . Such species has not been described, and in fact, all the specimens belong to M. pseudohirtipes, except the one labelled as “Type”, which belongs to M. purpurascens .</p></div>	https://treatment.plazi.org/id/79A3E50824AB3DAEF2135DE2BF0B11C8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Selnekovic, David;Kodada, Jan	Selnekovic, David, Kodada, Jan (2019): Taxonomic revision of Mordellistenahirtipes species complex with new distribution records (Insecta, Coleoptera, Mordellidae). ZooKeys 854: 89-118, DOI: http://dx.doi.org/10.3897/zookeys.854.32299, URL: http://dx.doi.org/10.3897/zookeys.854.32299
858198DCA030C5EE35BEF78642EB316A.text	858198DCA030C5EE35BEF78642EB316A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mordellistena purpurascens Costa 1854	<div><p>Mordellistena (s. str.) purpurascens Costa, 1854 Figs 3, 7 A–K</p><p>Mordellistena purpurascens Costa, 1854: 17 + Plate XXI (original description, figure); Gemminger and Harold 1870: 2113 (catalogue, as syn. of Mordellistena pumila (Gyllenhal, 1810)); Emery 1876: 95 (as syn. of Mordellistena micans (Germar, 1817)); Baudi di Selve 1877: 827 (as syn. of M. micans); Heyden et al. 1883: 142 (catalogue, as syn. of M. micans); Schilsky 1898: 77 (as syn. of M. micans); Heyden et al. 1906: 456 (catalogue, as syn. of M. micans); Csiki 1915: 37 (catalogue, as syn. of M. micans); Ermisch 1977: 169 (misidentification); Batten 1977 (misidentification); Kaszab 1979: 72, 74 (misidentification); Franciscolo 1991: 172-173 (remarks); Odnosum 2003: 36-46 (misidentification); Odnosum 2005: 100-108 (misidentification); Horák 2008: 100 (misidentification); Odnosum 2010: 199 (misidentification); Ruzzier 2013: 110 (misidentification).</p><p>Mordellistena geronensis Ermisch, 1977: 169 syn. nov. (original description in the key); Kaszab 1979: 72-73 (figures, key); Franciscolo 1991: 171-172 (key, figures); Horák 2008: 98 (distribution).</p><p>Mordellistena istrica Ermisch, 1977: 169 syn. nov. (original description in the key); Kaszab 1979: 73 (key); Horák 2008: 98 (distribution).</p><p>Type locality.</p><p>Naples, Italy.</p><p>Type material examined.</p><p>M. purpurascens: LECTOTYPE, here designated, glued, genitalia in separate microvial, right metatarsus missing: 1 ♂, MZFN: " Mordellistena purpurascens n. Napoli [hand written by Costa] / LECTOTYPE Mordellistena purpurascens Costa, 1854 D. Selnekovič des. 2019" [red label]. M. geronensis: HOLOTYPE: 1 ♂, SNSD: "♂ / Genitalpräparat / [card with dissected genitalia, right antenna and right protarsus] / Spanien, Prov. Gerona, Tossa de mar, A. Kampf. VII–VIII 35 / Holotypus [red label] / Holotypus Mordellistena geronensis Ermisch / Staatl. Museum für Tierkunde Dresden"; PARATYPES: 1 ♀, SNSD: "Spanien, Prov. Gerona, Tossa de mar, A. Kampf, V–VI 35 / sp.? grupe micans, det. Ermisch 1940 / Paratypus / PARATYPUS Mordellistena (s. str.) geronensis Ermisch, 1977, Selnekovič labelled 2017 [red label] / Mordellistena (s. str.) purpurascens Costa, 1854, D. Selnekovič det. 2019"; 2 ♂♂, 2 ♀♀, SNSD: "♂ [or] ♀ / Ardêche, 10. 7. 65, Banne, J. Balazuc / Paratypus / PARATYPUS Mordellistena (s. str.) geronensis Ermisch, 1977, Selnekovič labelled 2017 [red label] / Mordellistena (s. str.) purpurascens Costa, 1854, D. Selnekovič det. 2017"; 1 ♂, SNSD: "♂ / Ardêche, Sammzon, 8. 7. 65, Balazuc / Paratypus / PARATYPUS Mordellistena (s. str.) geronensis Ermisch, 1977, Selnekovič labelled 2017 [red label] / Mordellistena (s. str.) purpurascens Costa, 1854, D. Selnekovič det. 2017". M. istrica: HOLOTYPE: 1 ♂, SNSD: "♂ / [card with dissected genitalia] / Pola, Istr. F. Lang / [blank red circular label] / MORDELLISTENA istrian [illegible handwriting] det. Ermisch 1952 / Type [red label] / Holotypus Mordellistena istrica Ermisch / Mordellistena ( Mordellistena) geronensis Ermisch det. P. Leblanc 2007 / Mordellistena (s. str.) purpurascens Costa, 1854 D. Selnekovič det. 2019"; PARATYPES: 1 ♀, SNSD: "Plomin, Warmehang 14. 6. 1965 / Istrien K. Wellschmeld / Paratypus / PARATYPUS Mordellistena (s. str.) istrica Ermisch, 1977, Selnekovič labelled 2017" [red label]; 1 ♂, SNSD: "♂ / Corsica / Paratypus / PARATYPUS Mordellistena (s. str.) istrica Ermisch, 1977, Selnekovič labelled 2017 [red label] / Mordellistena ( Mordellistena) geronensis Ermisch det. P. Leblanc 2007 / Mordellistena (s. str.) purpurascens Costa, 1854 D. Selnekovič det. 2019"; 1 ♂, 1 ♀, SNSD: "12. 7. 14 Gallia m. Agay Rapp / Paratypus / PARATYPUS Mordellistena (s. str.) istrica Ermisch, 1977, Selnekovič labelled 2017" [red label].</p><p>Additional material examined.</p><p>Greece: 1 ♂, 2 ♀♀, CSB: "Greece N, Corfu - Kavos, 39°24'16"N, 20°05'53"E, F. Repta leg., 28. VIII. 2011 / Mordellistena purpurascens Costa, 1854, D. Selnekovič det. 2019". Italy: 1 ♂, CSB: “IT–Sicilia, Madonia, Termini, Sciara, M San Calogero, ex. l, 2.-3. 6. 2011, M. Šárovec, 3. 8 / Mordellistena purpurascens Costa, 1854, D. Selnekovič det. 2019"; 1 ♂, SNSD: "♂ / Gavoi Sard. 750m 21.-26. 8. 55 J. Kless 78 / Mordellistena Lopezi Ermisch det. K. Ermisch / Mordellistena (s. str.) purpurascens Costa, 1854 D. Selnekovič det. 2019" [in collection as M. lopezi]; 1 ♂, SNSD: "♂ / Genitalpräparat / ITALIA mer. Capaccio Hüdepohl VI. 64 / Mordellistena (s. str.) purpurascens Costa, 1854 D. Selnekovič det. 2019" [in collection as M. lopezi]. Montenegro: 4 ♂♂, 4 ♀♀ CSB: "Montenegro SE, 42°06'N, 19°06'E, Bar–centrum, on Daucus sp., D. Selnekovič 19. VI. 2011 / Mordellistena purpurascens Costa, 1854, D. Selnekovič det. 2019"; 1 ♀ CSB: "Montenegro SE, BAR env., 42°07'56"N, 19°07'33"E, 22. VI. 2011 / Mordellistena (s. str.) purpurascens Costa, 1854, D. Selnekovič det. 2019"; 1 ♂ HNHM: "Dalmatia Horváth / Zelenika 906. VIII. / Mordellistena pseudohirtipes Erm. det. R. Batten 1979 / Mordellistena (s. str.) purpurascens, Costa, 1854 D. Selnekovič det. 2019". Morocco: 1 ♂, CSB: Morocco Moyen Atl, Khenifra 15km E M. Šárovec 11. VII. 2007 / Mordellistena purpurascens Costa, 1854 D. Selnekovič det. 2019"; 4 ♂♂, 3 ♀♀, CSB: "Morocco Moyen Atl, Khenifra 10km I M. Šárovec 30. V. 2007 / Mordellistena purpurascens Costa, 1854 D. Selnekovič det. 2019". Spain: 1 ♂, SNSD: "♂ / Genitalpräparat / Son Españolet 1 –VI– 1958 R. López / Typus [red label] / Mordellistena Lopezi Ermisch det. K. Ermisch / Mordellistena (s. str.) purpurascens Costa, 1854 D. Selnekovič det. 2019" [in collection as M. lopezi].</p><p>Differential diagnosis.</p><p>M. purpurascens closely resembles M. hirtipes and M. pseudohirtipes . The differences are described under these species.</p><p>Redescription.</p><p>Measurements: TL: ♂♂ 3.10-4.42 mm (3.75 ± 0.35 mm, n = 19), ♀♀ 3.31-4.42 mm (3.82 ± 0.36 mm, n = 14); HL: ♂♂ 0.77-0.97 mm (0.85 ± 0.06 mm, n = 19), ♀♀ 0.78-0.96 mm (0.86 ± 0.06 mm, n = 14); HW: ♂♂ 0.91-1.17 mm (1.01 ± 0.07 mm, n = 19), ♀♀ 0.84-1.12 mm (0.98 ± 0.09 mm, n = 14); PL: ♂♂ 1.06-1.44 mm (1.23 ± 0.10 mm, n = 19), ♀♀ 1.04-1.44 mm (1.25 ± 0.11 mm, n = 14); PW: 1.10-1.56 mm (1.30 ± 0.13 mm, n = 19), ♀♀ 1.13-1.58 mm (1.35 ± 0.14 mm, n = 13); EL: ♂♂ 2.44-3.35 mm (2.81 ± 0.27 mm, n = 19), ♀♀ 2.50-3.38 mm (2.88 ± 0.27 mm, n = 14); EW: ♂♂ 1.15-1.59 mm (1.35 ± 0.13 mm, n = 19), ♀♀ 1.19-1.66 mm (1.43 ± 0.15 mm, n = 14); ATiL: ♂♂ 0.71-0.93 mm (0.81 ± 0.07 mm, n = 19), ♀♀ 0.65-0.91 mm (0.75 ± 0.08 mm, n = 14); ATrL: ♂♂ 0.64-0.84 mm (0.72 ± 0.07 mm, n = 15), ♀♀ 0.62-0.80 mm (0.69 ± 0.06 mm, n = 13); ITiL: ♂♂ 0.91-1.23 mm (1.03 ± 0.10 mm, n = 19), ♀♀ 0.86-1.23 mm (1.01 ± 0.12 mm, n = 14); ITrL: ♂♂ 1.12-1.64 mm (1.27 ± 0.15 mm, n = 9), ♀♀ 1.02-1.34 mm (1.16 ± 0.10 mm, n = 14); PTiL: 0.78-1.08 mm (0.88 ± 0.08 mm, n = 19), ♀♀ 0.75-1.05 mm (0.88 ± 0.08 mm, n = 14); PTrL: 1.64-2.18 mm (1.87 ± 0.19 mm, n = 9), ♀♀ 1.48-2.16 mm (1.77 ± 0.21 mm, n = 11); PygL: ♂♂ 1.50-2.12 mm (1.86 ± 0.18 mm, n = 19), ♀♀ 1.35-1.98 mm (1.67 ± 0.20 mm, n = 14); TVtL: ♂♂ 0.58-0.87 mm (0.77 ± 0.10 mm, n = 18), ♀♀ 0.60-0.92 mm (0.77 ± 0.08 mm, n = 14); RPrL: 0.52-0.64 mm (0.56 ± 0.03 mm, n = 19); LPrL: 0.41-0.51 mm (0.45 ± 0.03 mm, n = 19); St8L: ♂♂ 0.65-0.80 mm (0.70 ± 0.07 mm, n = 3); St8W: ♂♂ 0.49-0.52 mm (0.50 ± 0.01 mm, n = 3).</p><p>Habitus illustrated in Fig. 3. Body strongly elongate, slender, widest behind anterior 1/4 of elytra. Integument black, anterior margin of clypeus and mandibles somewhat paler. Pubescence on head and thorax yellowish; on elytra yellowish in anterior half, gradually darkened apically; on venter yellowish, darkened along posterior margins of ventrites 3 and 4 and completely dark grey on terminal ventrite and pygidium.</p><p>Head convex dorsally, wider than long, widest about middle, HW/HL ratio: ♂♂ 1.11-1.23 (1.19 ± 0.03, n = 19), ♀♀ 1.06-1.19 (1.14 ± 0.03, n = 14). Dorsal surface weakly microreticulated, with small, round punctures, each bearing short seta. Ventral surface with weak transverse microreticulation and sparsely arranged, round punctures, each bearing short seta. Occipital margin rounded in dorsal aspect, straight if seen from behind. Eyes oval, completely reaching occiput, not expanded onto ventral surface, finely faceted, with short interfacetal setae. Anterior margin of clypeus straight. Labrum transverse, LabW/LabL: ♂♂ 2.04-2.27 (2.15 ± 0.10, n = 5), ♀♀ 1.88-2.38 (2.21 ± 0.20, n = 5), anterior margin straight or very shallowly emarginate mesally; surface covered with small, round punctures, each bearing seta. Antennae slightly serrate, expanded from antennomere V (Fig. 7A, B); antennomeres I and II short, subequal in length and width; antennomere III equal in length and slightly slenderer than previous two; antennomere IV slightly longer and wider than previous one; antennomeres V–X wider than previous four, in males ~1.60 ×, in females ~1.30 × as long as wide; antennomere XI elongate oval, ~2.20 × as long as wide. Galea gradually expanded apically, covered with apically expanded sensilla. Maxillary palpi (Fig. 7 C–D) black; palpomere I very short; palpomere II in males expanded with long setae on ventral side, in females slenderer, without long setae; palpomere III short, ~1.80 × as long as wide, in males with long setae on ventral side; terminal palpomere securiform, in males wider than in females, inner angle situated around middle in males, in terminal 1/3 in females; TPalL/TPalW ratio: ♂♂ 1.72-2.16 (1.96 ± 0.10, n = 17), ♀♀ 2.08-2.34 (2.21 ± 0.09, n = 14).</p><p>Pronotum moderately convex, slightly wider than long, PW/PL ratio: ♂♂ 0.97-1.10 (1.05 ± 0.03, n = 19), ♀♀ 1.00-1.15 (1.07 ± 0.04, n = 13). Surface weakly microreticulated with small, rasp-like punctures, distance between punctures 1.50 –2.00× as long as puncture diameter, each puncture bearing flat seta. Anterior margin rounded, slightly produced mesally, anterior angles broadly rounded; lateral carinae rounded in dorsal aspect, shallowly but distinctly emarginate in lateral aspect; posterior margin forming short mesal lobe, emarginated laterally before posterior angles; posterior angles rectangular, pointed in lateral aspect. Posterior marginal bead interrupted before posterior angles. Prosternum in front of procoxae narrow, laterally expanded. Scutellar shield small, triangular, with small punctures bearing short setae. Mesoventral process ca. half as wide as mesofemora. Metaventrite large, posterior margin produced mesally between metacoxae; longitudinal discrimen rather indistinct. Metanepisternum trapezoidal, slightly narrowed posteriorly, dorsal margin emarginate, ventral margin straight.</p><p>Elytra long, narrow, widest at end of anterior 1/4, EL/EW ratio: ♂♂ 1.97-2.23 (2.08 ± 0.07, n = 19), ♀♀ 1.83-2.15 (2.02 ± 0.07, n = 14). Dorsal surface covered with weak transverse microreticulation and rasp-like punctures, distance between punctures ~1,50 × as long as puncture diameter; each puncture bearing flat seta. Lateral margins regularly rounded, apices separately rounded.</p><p>Protibiae straight, basal part in males slightly expanded and bearing distinct fringe of long setae; PTiL/PTrL ratio: ♂♂ 0.98-1.24 (1.13 ± 0.06, n = 15), ♀♀ 0.98-1.17 (1.10 ± 0.05, n = 13). Protarsomere I in females as long as two following tarsomeres combined, in males slightly longer; protarsomere IV simple, parallel-sided, very shallowly emarginated at apex. Claws on protarsi rather long, slender, with three denticles, on meso- and metatarsi with four denticles. Mesotibiae slightly bent medially; mesotarsus longer than tibia, MsTiL/MsTrL ratio: ♂♂ 0.75-0.90 (0.84 ± 0.04, n = 9), ♀♀ 0.82-0.95 (0.88 ± 0.04, n = 14). Metacoxae large, anterior margin straight, posterior margin broadly rounded. Metatibiae bearing short subapical ridge and 3-4 lateral ridges parallel to apical tibial margin, reaching 1/3 of tibial width. Metatibial spurs black, inner one ~1.30 × as long as outer one. Metatarsomere I bearing in males 5, in females 3-4 short lateral ridges; metatarsomere II bearing 2-3 ridges; metatarsomeres III and IV without ridges. Metatarsus ~2.00 × as long as metatibia, MtTrL/MtTiL ratio: ♂♂ 2.00-2.33 (2.13 ± 0.10, n = 10), ♀♀ 1.90-2.18 (2.01 ± 0.07, n = 11).</p><p>Pygidium long, slender, PygL/TVtL ratio: ♂♂ 1.88-3.32 (2.43 ± 0.35, n = 18), ♀♀ 1.88-2.64 (2.18 ± 0.17, n = 14). Apical margin of terminal abdominal ventrite arcuate.</p><p>Male genitalia: sternite VIII rather short, with long setae in apical part, apical protuberance rather short, slightly bilobed at apex (Fig. 7E); St8L/St8W ratio: ♂♂ 1.26-1.32 (1.28 ± 0.02, n = 4). Sternite IX long, slender, arrow-shaped, with medial longitudinal keel apically. Parameres (Fig. 7 G–J) rather long, EL/LPrL ratio: 4.42-5.84 (4.98 ± 0.35, n = 19), EL/RPrL ratio: 5.57-6.94 (6.19 ± 0.41 n = 19); LPrL/RPrL ratio: 1.16-1.30 (1.24 ± 0.03, n = 19). Left paramere with very long basal part, LPrL/BLPr ratio: 1.50-1.91 (1.76 ± 0.09, n = 19); dorsal branch strongly expanded apically, obliquely truncate at apex; ventral branch slender, slightly bent medially. Right paramere rather long with long branches; ventral branch longer than dorsal one, pointed at apex; dorsal branch rather narrow, slightly expanded apically, rounded at apex. Median lobe (Fig. 7K) long, slender, apical part narrow or slightly expanded. Phallobase with short tubular process (approximately 1/6 of total length) and long, slender furca.</p><p>Female genitalia: sternite VIII (Fig. 7F) with slightly bilobed apical protuberance, long setae situated at apex and alongside lateral margins; spiculum ventrale short, broadly clavate; St8L/St8W ratio: 1.62 (n = 1).</p><p>Sexual dimorphism.</p><p>Females are more robust, with protibiae not expanded and without fringe of long setae in basal part. Maxillary palpomere II not expanded in females and without long setae on ventral side. Terminal maxillary palpomere is wider in males, with its inner angle situated approximately in the middle (Fig. 7C), in females it is generally slenderer, with its inner angle situated in terminal 1/3 (Fig. 7D). Antennae are shorter in females; antennomeres V–X ~1.60 × as long as wide in males, ~1.30 × in females.</p><p>Distribution.</p><p>Croatia, France, Greece, Italy, Montenegro, Morocco, Spain. Mordellistena purpurascens is reported here for the first time from Greece and Montenegro. Odnosum (2003, 2005, 2010) reported M. purpurascens from Kazakhstan, Turkmenistan, Tajikistan, and Ukraine. However, based on the figures of parameres provided in all the three mentioned studies, it is obvious that he referred to a different species (see Discussion).</p><p>Biology.</p><p>Adults were collected by the first author in Montenegro, in urban environment of Bar on the flowers of Daucus sp. ( Apiaceae).</p><p>Remarks.</p><p>Mordellistena purpurascens was described by Costa (1854) and referred to be found in several localities in former "Regno di Napoli" (southern parts of present Italy). Series of M. purpurascens in Costa’s collection in MZFN contains only two specimens. One of them with the original label " Mordellistena purpurascens n. Napoli" is designed here as a lectotype. The other specimen labelled "S. Severina" without identification label belongs to a different species from the gemellata-group (sensu Ermisch 1956). Genitalia of the lectotype were examined for the first time for the purposes of the present study (Fig. 7G).</p><p>Ermisch (1977) briefly described two new species M. geronensis and M. istrica as a part of an identification key. He differentiated these species from each other based on the shape of the apical part of the median lobe (expanded in M. istrica, not expanded in M. geronensis). Shape of the apical part of median lobe depends on the observation method (dry/wet, card mounted/slide mounted). After examining the series of slide mounted median lobes of both taxa, we did not find any differences in the shape. Examination of the male genitalia from type specimens of M. purpurascens (Fig. 7G), M. geronensis (Fig. 7H) and M. istrica (Fig. 7I) revealed that these taxa are conspecific. We thus propose M. istrica and M. geronensis as the junior synonyms of M. purpurascens .</p><p>Type series of M. istrica includes a female paratype (Pola, Croatia), which we were not able to assign to M. purpurascens or M. pseudohirtipes . Type series of M. geronensis includes a male paratype (Bois de Paiolive, Ardêche, France), which in fact belongs to M. pseudohirtipes, and three paratypes (Tossa de mar, Spain; Costa Brava, Spain), which belong to M. hirtipes . In Ermisch’s collection, there is a series of specimens named Mordellistena lopezi . Such species has not been described. Specimen labelled as “Typus”, in fact, belongs to M. purpurascens, the rest of the specimens belongs to M. pseudohirtipes .</p></div>	https://treatment.plazi.org/id/858198DCA030C5EE35BEF78642EB316A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Selnekovic, David;Kodada, Jan	Selnekovic, David, Kodada, Jan (2019): Taxonomic revision of Mordellistenahirtipes species complex with new distribution records (Insecta, Coleoptera, Mordellidae). ZooKeys 854: 89-118, DOI: http://dx.doi.org/10.3897/zookeys.854.32299, URL: http://dx.doi.org/10.3897/zookeys.854.32299
