taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
A884DA106BD150DAABC547E8FAF2D79D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460651	https://doi.org/10.3897/zookeys.1259.168419.figure20	Figure 20. Papuanatula (Papuafiliola) horvathrobi sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 20. Papuanatula (Papuafiliola) horvathrobi sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
A884DA106BD150DAABC547E8FAF2D79D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460652	https://doi.org/10.3897/zookeys.1259.168419.figure21	Figure 21. Papuanatula (Papuafiliola) horvathrobi sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Left mandible; d. Right mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h, i. Tergalius V; j. Head. Scale bars: 50 µm (j), 20 µm (c – h), 10 µm (a, b, i).	Figure 21. Papuanatula (Papuafiliola) horvathrobi sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Left mandible; d. Right mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h, i. Tergalius V; j. Head. Scale bars: 50 µm (j), 20 µm (c – h), 10 µm (a, b, i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
A884DA106BD150DAABC547E8FAF2D79D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460653	https://doi.org/10.3897/zookeys.1259.168419.figure22	Figure 22. Papuanatula (Papuafiliola) horvathrobi sp. nov., larva. a. Hind leg; b. Hind femur, outer margin; c. Hind tibia, anterior surface; d. Hind tarsus, outer margin; e. Abdomen; f. Abdominal terga; g. Paraproct and developing subimaginal gonostylus; h. Paracercus; i. Cercus; j. Abdominal tergum IV, surface. Scale bars: 100 µm (e), 50 µm (a), 20 µm (h), 10 µm (b – d, f, g, i).	Figure 22. Papuanatula (Papuafiliola) horvathrobi sp. nov., larva. a. Hind leg; b. Hind femur, outer margin; c. Hind tibia, anterior surface; d. Hind tarsus, outer margin; e. Abdomen; f. Abdominal terga; g. Paraproct and developing subimaginal gonostylus; h. Paracercus; i. Cercus; j. Abdominal tergum IV, surface. Scale bars: 100 µm (e), 50 µm (a), 20 µm (h), 10 µm (b – d, f, g, i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
A884DA106BD150DAABC547E8FAF2D79D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460660	https://doi.org/10.3897/zookeys.1259.168419.figure29	Figure 29. Habitats: P. batantaraja sp. nov. (a. Type locality, b); P. cukiclara sp. nov. (a, b, d. Type locality); P. cataracta sp. nov. (c. type locality); P. batanlenos sp. nov. (a, d. Type locality, e, f); P. horvathrobi sp. nov. (e); P. longabranchias sp. nov. (d. Type locality, f).	Figure 29. Habitats: P. batantaraja sp. nov. (a. Type locality, b); P. cukiclara sp. nov. (a, b, d. Type locality); P. cataracta sp. nov. (c. type locality); P. batanlenos sp. nov. (a, d. Type locality, e, f); P. horvathrobi sp. nov. (e); P. longabranchias sp. nov. (d. Type locality, f).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
A884DA106BD150DAABC547E8FAF2D79D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460661	https://doi.org/10.3897/zookeys.1259.168419.figure30	Figure 30. Distribution of Papuanatula species: yellow dots: known species (Kaltenbach et al. 2025); orange dot: Papuanatula arfak sp. nov. (present study); red dots: species from Batanta – P. batantaraja sp. nov., P. cukiclara sp. nov., P. cataracta sp. nov., P. batanlenos sp. nov., P. horvathrobi sp. nov., P. longabranchias sp. nov. (present study).	Figure 30. Distribution of Papuanatula species: yellow dots: known species (Kaltenbach et al. 2025); orange dot: Papuanatula arfak sp. nov. (present study); red dots: species from Batanta – P. batantaraja sp. nov., P. cukiclara sp. nov., P. cataracta sp. nov., P. batanlenos sp. nov., P. horvathrobi sp. nov., P. longabranchias sp. nov. (present study).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
DDF16181DA045D39AAEB357F74225338.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460651	https://doi.org/10.3897/zookeys.1259.168419.figure20	Figure 20. Papuanatula (Papuafiliola) horvathrobi sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 20. Papuanatula (Papuafiliola) horvathrobi sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
DDF16181DA045D39AAEB357F74225338.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460652	https://doi.org/10.3897/zookeys.1259.168419.figure21	Figure 21. Papuanatula (Papuafiliola) horvathrobi sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Left mandible; d. Right mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h, i. Tergalius V; j. Head. Scale bars: 50 µm (j), 20 µm (c – h), 10 µm (a, b, i).	Figure 21. Papuanatula (Papuafiliola) horvathrobi sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Left mandible; d. Right mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h, i. Tergalius V; j. Head. Scale bars: 50 µm (j), 20 µm (c – h), 10 µm (a, b, i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
DDF16181DA045D39AAEB357F74225338.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460653	https://doi.org/10.3897/zookeys.1259.168419.figure22	Figure 22. Papuanatula (Papuafiliola) horvathrobi sp. nov., larva. a. Hind leg; b. Hind femur, outer margin; c. Hind tibia, anterior surface; d. Hind tarsus, outer margin; e. Abdomen; f. Abdominal terga; g. Paraproct and developing subimaginal gonostylus; h. Paracercus; i. Cercus; j. Abdominal tergum IV, surface. Scale bars: 100 µm (e), 50 µm (a), 20 µm (h), 10 µm (b – d, f, g, i).	Figure 22. Papuanatula (Papuafiliola) horvathrobi sp. nov., larva. a. Hind leg; b. Hind femur, outer margin; c. Hind tibia, anterior surface; d. Hind tarsus, outer margin; e. Abdomen; f. Abdominal terga; g. Paraproct and developing subimaginal gonostylus; h. Paracercus; i. Cercus; j. Abdominal tergum IV, surface. Scale bars: 100 µm (e), 50 µm (a), 20 µm (h), 10 µm (b – d, f, g, i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
DDF16181DA045D39AAEB357F74225338.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460654	https://doi.org/10.3897/zookeys.1259.168419.figure23	Figure 23. Papuanatula (Papuafiliola) longabranchias sp. nov., larva, habitus. a. Dorsal view; b. lateral view; c. ventral view; d. legs. Scale bars: 1 mm.	Figure 23. Papuanatula (Papuafiliola) longabranchias sp. nov., larva, habitus. a. Dorsal view; b. lateral view; c. ventral view; d. legs. Scale bars: 1 mm.	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
DDF16181DA045D39AAEB357F74225338.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460655	https://doi.org/10.3897/zookeys.1259.168419.figure24	Figure 24. Papuanatula (Papuafiliola) longabranchias sp. nov., larva. a. Labrum; b. Left mandible; c. Right mandible; d. Labium; e. Labial palp; f. Hypopharynx and superlinguae; g. Maxilla; h, i. Tergalius III; j. Head. Scale bars: 100 µm (j), 20 µm (b – d, f – h), 10 µm (a, e, i).	Figure 24. Papuanatula (Papuafiliola) longabranchias sp. nov., larva. a. Labrum; b. Left mandible; c. Right mandible; d. Labium; e. Labial palp; f. Hypopharynx and superlinguae; g. Maxilla; h, i. Tergalius III; j. Head. Scale bars: 100 µm (j), 20 µm (b – d, f – h), 10 µm (a, e, i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
DDF16181DA045D39AAEB357F74225338.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460656	https://doi.org/10.3897/zookeys.1259.168419.figure25	Figure 25. Papuanatula (Papuafiliola) longabranchias sp. nov., larva. a. Hind leg; b. Hind femur, inner margin; c. Hind femur, outer margin; d. Hind tibia, anterior surface; e. Hind tarsus, outer margin; f. Hind claw; g. Thorax, abdomen; h. Abdominal terga; i. Paraproct; j. Paracercus; k. Cercus. Scale bars: 100 µm (g), 50 µm (a), 20 µm (j), 10 µm (b – f, h, I, k).	Figure 25. Papuanatula (Papuafiliola) longabranchias sp. nov., larva. a. Hind leg; b. Hind femur, inner margin; c. Hind femur, outer margin; d. Hind tibia, anterior surface; e. Hind tarsus, outer margin; f. Hind claw; g. Thorax, abdomen; h. Abdominal terga; i. Paraproct; j. Paracercus; k. Cercus. Scale bars: 100 µm (g), 50 µm (a), 20 µm (j), 10 µm (b – f, h, I, k).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
DDF16181DA045D39AAEB357F74225338.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460657	https://doi.org/10.3897/zookeys.1259.168419.figure26	Figure 26. Papuanatula (Papuafiliola) longabranchias sp. nov, imago, male. a. Habitus, lateral view; b. Habitus, dorsal view; c. Fore leg; d. Hind leg. Scale bars: 1 mm (a, b), 100 µm (c, d).	Figure 26. Papuanatula (Papuafiliola) longabranchias sp. nov, imago, male. a. Habitus, lateral view; b. Habitus, dorsal view; c. Fore leg; d. Hind leg. Scale bars: 1 mm (a, b), 100 µm (c, d).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
DDF16181DA045D39AAEB357F74225338.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460658	https://doi.org/10.3897/zookeys.1259.168419.figure27	Figure 27. Papuanatula (Papuafiliola) longabranchias sp. nov. Imago, male. a. Fore wing; b. Genitalia. Subimagos. c, d. Habitus, male (lateral view, dorsal view); e, f. Habitus, female (lateral view, dorsal view); g. Hind leg; h. Fore tarsus; i. Hind tarsus. Scale bars: 1 mm (c – f), 100 µm (a, g), 20 µm (b, h), 10 µm (i).	Figure 27. Papuanatula (Papuafiliola) longabranchias sp. nov. Imago, male. a. Fore wing; b. Genitalia. Subimagos. c, d. Habitus, male (lateral view, dorsal view); e, f. Habitus, female (lateral view, dorsal view); g. Hind leg; h. Fore tarsus; i. Hind tarsus. Scale bars: 1 mm (c – f), 100 µm (a, g), 20 µm (b, h), 10 µm (i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
DDF16181DA045D39AAEB357F74225338.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460659	https://doi.org/10.3897/zookeys.1259.168419.figure28	Figure 28. Papuanatula (Papuafiliola) longabranchias sp. nov. a – c. Eggs; d. Micropyle. Scale bars: 10 µm (a – c), 5 µm (d).	Figure 28. Papuanatula (Papuafiliola) longabranchias sp. nov. a – c. Eggs; d. Micropyle. Scale bars: 10 µm (a – c), 5 µm (d).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
993AB6815BE75EF592FCECA16C6A659C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460654	https://doi.org/10.3897/zookeys.1259.168419.figure23	Figure 23. Papuanatula (Papuafiliola) longabranchias sp. nov., larva, habitus. a. Dorsal view; b. lateral view; c. ventral view; d. legs. Scale bars: 1 mm.	Figure 23. Papuanatula (Papuafiliola) longabranchias sp. nov., larva, habitus. a. Dorsal view; b. lateral view; c. ventral view; d. legs. Scale bars: 1 mm.	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
993AB6815BE75EF592FCECA16C6A659C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460655	https://doi.org/10.3897/zookeys.1259.168419.figure24	Figure 24. Papuanatula (Papuafiliola) longabranchias sp. nov., larva. a. Labrum; b. Left mandible; c. Right mandible; d. Labium; e. Labial palp; f. Hypopharynx and superlinguae; g. Maxilla; h, i. Tergalius III; j. Head. Scale bars: 100 µm (j), 20 µm (b – d, f – h), 10 µm (a, e, i).	Figure 24. Papuanatula (Papuafiliola) longabranchias sp. nov., larva. a. Labrum; b. Left mandible; c. Right mandible; d. Labium; e. Labial palp; f. Hypopharynx and superlinguae; g. Maxilla; h, i. Tergalius III; j. Head. Scale bars: 100 µm (j), 20 µm (b – d, f – h), 10 µm (a, e, i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
993AB6815BE75EF592FCECA16C6A659C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460656	https://doi.org/10.3897/zookeys.1259.168419.figure25	Figure 25. Papuanatula (Papuafiliola) longabranchias sp. nov., larva. a. Hind leg; b. Hind femur, inner margin; c. Hind femur, outer margin; d. Hind tibia, anterior surface; e. Hind tarsus, outer margin; f. Hind claw; g. Thorax, abdomen; h. Abdominal terga; i. Paraproct; j. Paracercus; k. Cercus. Scale bars: 100 µm (g), 50 µm (a), 20 µm (j), 10 µm (b – f, h, I, k).	Figure 25. Papuanatula (Papuafiliola) longabranchias sp. nov., larva. a. Hind leg; b. Hind femur, inner margin; c. Hind femur, outer margin; d. Hind tibia, anterior surface; e. Hind tarsus, outer margin; f. Hind claw; g. Thorax, abdomen; h. Abdominal terga; i. Paraproct; j. Paracercus; k. Cercus. Scale bars: 100 µm (g), 50 µm (a), 20 µm (j), 10 µm (b – f, h, I, k).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
993AB6815BE75EF592FCECA16C6A659C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460657	https://doi.org/10.3897/zookeys.1259.168419.figure26	Figure 26. Papuanatula (Papuafiliola) longabranchias sp. nov, imago, male. a. Habitus, lateral view; b. Habitus, dorsal view; c. Fore leg; d. Hind leg. Scale bars: 1 mm (a, b), 100 µm (c, d).	Figure 26. Papuanatula (Papuafiliola) longabranchias sp. nov, imago, male. a. Habitus, lateral view; b. Habitus, dorsal view; c. Fore leg; d. Hind leg. Scale bars: 1 mm (a, b), 100 µm (c, d).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
993AB6815BE75EF592FCECA16C6A659C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460658	https://doi.org/10.3897/zookeys.1259.168419.figure27	Figure 27. Papuanatula (Papuafiliola) longabranchias sp. nov. Imago, male. a. Fore wing; b. Genitalia. Subimagos. c, d. Habitus, male (lateral view, dorsal view); e, f. Habitus, female (lateral view, dorsal view); g. Hind leg; h. Fore tarsus; i. Hind tarsus. Scale bars: 1 mm (c – f), 100 µm (a, g), 20 µm (b, h), 10 µm (i).	Figure 27. Papuanatula (Papuafiliola) longabranchias sp. nov. Imago, male. a. Fore wing; b. Genitalia. Subimagos. c, d. Habitus, male (lateral view, dorsal view); e, f. Habitus, female (lateral view, dorsal view); g. Hind leg; h. Fore tarsus; i. Hind tarsus. Scale bars: 1 mm (c – f), 100 µm (a, g), 20 µm (b, h), 10 µm (i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
993AB6815BE75EF592FCECA16C6A659C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460659	https://doi.org/10.3897/zookeys.1259.168419.figure28	Figure 28. Papuanatula (Papuafiliola) longabranchias sp. nov. a – c. Eggs; d. Micropyle. Scale bars: 10 µm (a – c), 5 µm (d).	Figure 28. Papuanatula (Papuafiliola) longabranchias sp. nov. a – c. Eggs; d. Micropyle. Scale bars: 10 µm (a – c), 5 µm (d).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
993AB6815BE75EF592FCECA16C6A659C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460660	https://doi.org/10.3897/zookeys.1259.168419.figure29	Figure 29. Habitats: P. batantaraja sp. nov. (a. Type locality, b); P. cukiclara sp. nov. (a, b, d. Type locality); P. cataracta sp. nov. (c. type locality); P. batanlenos sp. nov. (a, d. Type locality, e, f); P. horvathrobi sp. nov. (e); P. longabranchias sp. nov. (d. Type locality, f).	Figure 29. Habitats: P. batantaraja sp. nov. (a. Type locality, b); P. cukiclara sp. nov. (a, b, d. Type locality); P. cataracta sp. nov. (c. type locality); P. batanlenos sp. nov. (a, d. Type locality, e, f); P. horvathrobi sp. nov. (e); P. longabranchias sp. nov. (d. Type locality, f).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
993AB6815BE75EF592FCECA16C6A659C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460661	https://doi.org/10.3897/zookeys.1259.168419.figure30	Figure 30. Distribution of Papuanatula species: yellow dots: known species (Kaltenbach et al. 2025); orange dot: Papuanatula arfak sp. nov. (present study); red dots: species from Batanta – P. batantaraja sp. nov., P. cukiclara sp. nov., P. cataracta sp. nov., P. batanlenos sp. nov., P. horvathrobi sp. nov., P. longabranchias sp. nov. (present study).	Figure 30. Distribution of Papuanatula species: yellow dots: known species (Kaltenbach et al. 2025); orange dot: Papuanatula arfak sp. nov. (present study); red dots: species from Batanta – P. batantaraja sp. nov., P. cukiclara sp. nov., P. cataracta sp. nov., P. batanlenos sp. nov., P. horvathrobi sp. nov., P. longabranchias sp. nov. (present study).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
79AAA1BCD8D859788D234314421AF9E2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460632	https://doi.org/10.3897/zookeys.1259.168419.figure1	Figure 1. Papuanatula (Papuanatula) arfak sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 1. Papuanatula (Papuanatula) arfak sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
79AAA1BCD8D859788D234314421AF9E2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460633	https://doi.org/10.3897/zookeys.1259.168419.figure2	Figure 2. Papuanatula (Papuanatula) arfak sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Right mandible; d. Left mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h. Maxillary palp; i. Developing subimaginal gonostyli. Scale bars: 50 µm (e), 20 µm (a, c, d, f, g, i), 10 µm (b, h).	Figure 2. Papuanatula (Papuanatula) arfak sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Right mandible; d. Left mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h. Maxillary palp; i. Developing subimaginal gonostyli. Scale bars: 50 µm (e), 20 µm (a, c, d, f, g, i), 10 µm (b, h).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
79AAA1BCD8D859788D234314421AF9E2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460634	https://doi.org/10.3897/zookeys.1259.168419.figure3	Figure 3. Papuanatula (Papuanatula) arfak sp. nov., larva. a. Middle leg; b. Fore tarsus; c. Fore claw; d. Fore femur, posterior apex; e. Fore tibia, posterior surface; f. Middle tibia, inner margin; g. Fore femur, ventral margin; h. Head, mature male larva. Scale bars: 100 µm (h), 50 µm (a), 20 µm (b), 10 µm (c – g).	Figure 3. Papuanatula (Papuanatula) arfak sp. nov., larva. a. Middle leg; b. Fore tarsus; c. Fore claw; d. Fore femur, posterior apex; e. Fore tibia, posterior surface; f. Middle tibia, inner margin; g. Fore femur, ventral margin; h. Head, mature male larva. Scale bars: 100 µm (h), 50 µm (a), 20 µm (b), 10 µm (c – g).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
79AAA1BCD8D859788D234314421AF9E2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460635	https://doi.org/10.3897/zookeys.1259.168419.figure4	Figure 4. Papuanatula (Papuanatula) arfak sp. nov., larva. a. Abdominal terga, posterior margins; b. Paracercus; c. Cercus, basal part; d. Cercus, distal part; e, f. Tergalius IV; g – i. Paraproct. Scale bars: 50 µm (h), 20 µm (a, b, g), 10 µm (c, d, f, h, i).	Figure 4. Papuanatula (Papuanatula) arfak sp. nov., larva. a. Abdominal terga, posterior margins; b. Paracercus; c. Cercus, basal part; d. Cercus, distal part; e, f. Tergalius IV; g – i. Paraproct. Scale bars: 50 µm (h), 20 µm (a, b, g), 10 µm (c, d, f, h, i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
79AAA1BCD8D859788D234314421AF9E2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460636	https://doi.org/10.3897/zookeys.1259.168419.figure5	Figure 5. Papuanatula (Papuanatula) arfak sp. nov., larva, SEM pictures. a. Abdominal terga III – VII; b. Abdominal tergum VII, posterior margin. Scale bars: 300 µm (a), 40 µm (b).	Figure 5. Papuanatula (Papuanatula) arfak sp. nov., larva, SEM pictures. a. Abdominal terga III – VII; b. Abdominal tergum VII, posterior margin. Scale bars: 300 µm (a), 40 µm (b).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
79AAA1BCD8D859788D234314421AF9E2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460661	https://doi.org/10.3897/zookeys.1259.168419.figure30	Figure 30. Distribution of Papuanatula species: yellow dots: known species (Kaltenbach et al. 2025); orange dot: Papuanatula arfak sp. nov. (present study); red dots: species from Batanta – P. batantaraja sp. nov., P. cukiclara sp. nov., P. cataracta sp. nov., P. batanlenos sp. nov., P. horvathrobi sp. nov., P. longabranchias sp. nov. (present study).	Figure 30. Distribution of Papuanatula species: yellow dots: known species (Kaltenbach et al. 2025); orange dot: Papuanatula arfak sp. nov. (present study); red dots: species from Batanta – P. batantaraja sp. nov., P. cukiclara sp. nov., P. cataracta sp. nov., P. batanlenos sp. nov., P. horvathrobi sp. nov., P. longabranchias sp. nov. (present study).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
A8328AE5F38E58CEA5CA64B11904DDA9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460646	https://doi.org/10.3897/zookeys.1259.168419.figure15	Figure 15. Papuanatula (Papuanatula) batanlenos sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 15. Papuanatula (Papuanatula) batanlenos sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
A8328AE5F38E58CEA5CA64B11904DDA9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460647	https://doi.org/10.3897/zookeys.1259.168419.figure16	Figure 16. Papuanatula (Papuanatula) batanlenos sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Left mandible; d. Right mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h, i. Tergalius IV; j. Head; k. Developing subimaginal gonostyli. Scale bars: 100 µm (j), 20 µm (a, c – h, k), 10 µm (b, i).	Figure 16. Papuanatula (Papuanatula) batanlenos sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Left mandible; d. Right mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h, i. Tergalius IV; j. Head; k. Developing subimaginal gonostyli. Scale bars: 100 µm (j), 20 µm (a, c – h, k), 10 µm (b, i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
A8328AE5F38E58CEA5CA64B11904DDA9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460648	https://doi.org/10.3897/zookeys.1259.168419.figure17	Figure 17. Papuanatula (Papuanatula) batanlenos sp. nov., larva. a. Middle leg; b. Middle femur, outer margin; c. Middle tarsus, outer margin; d. Middle claw; e. Abdomen; f. Abdominal terga; g. Paraproct; h. Paracercus; i. Cercus. Scale bars: 100 µm (a, e), 20 µm (h), 10 µm (b – d, f, g, i).	Figure 17. Papuanatula (Papuanatula) batanlenos sp. nov., larva. a. Middle leg; b. Middle femur, outer margin; c. Middle tarsus, outer margin; d. Middle claw; e. Abdomen; f. Abdominal terga; g. Paraproct; h. Paracercus; i. Cercus. Scale bars: 100 µm (a, e), 20 µm (h), 10 µm (b – d, f, g, i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
A8328AE5F38E58CEA5CA64B11904DDA9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460649	https://doi.org/10.3897/zookeys.1259.168419.figure18	Figure 18. Papuanatula (Papuanatula) batanlenos sp. nov, imago, male. a. Habitus, lateral view; b. Habitus, dorsal view; c. Middle leg; d. Genitalia. Scale bars: 1 mm (a, b), 100 µm (c), 50 µm (d).	Figure 18. Papuanatula (Papuanatula) batanlenos sp. nov, imago, male. a. Habitus, lateral view; b. Habitus, dorsal view; c. Middle leg; d. Genitalia. Scale bars: 1 mm (a, b), 100 µm (c), 50 µm (d).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
A8328AE5F38E58CEA5CA64B11904DDA9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460650	https://doi.org/10.3897/zookeys.1259.168419.figure19	Figure 19. Papuanatula (Papuanatula) batanlenos sp. nov. Imago, male. a. Fore wing. Subimagos. b. Habitus, male; c. Habitus, female; d. Middle leg; e. Fore tarsus; f. Middle tarsus. Scale bars: 1 mm (a – c), 100 µm (d), 20 µm (e), 10 µm (f).	Figure 19. Papuanatula (Papuanatula) batanlenos sp. nov. Imago, male. a. Fore wing. Subimagos. b. Habitus, male; c. Habitus, female; d. Middle leg; e. Fore tarsus; f. Middle tarsus. Scale bars: 1 mm (a – c), 100 µm (d), 20 µm (e), 10 µm (f).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
A8328AE5F38E58CEA5CA64B11904DDA9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460660	https://doi.org/10.3897/zookeys.1259.168419.figure29	Figure 29. Habitats: P. batantaraja sp. nov. (a. Type locality, b); P. cukiclara sp. nov. (a, b, d. Type locality); P. cataracta sp. nov. (c. type locality); P. batanlenos sp. nov. (a, d. Type locality, e, f); P. horvathrobi sp. nov. (e); P. longabranchias sp. nov. (d. Type locality, f).	Figure 29. Habitats: P. batantaraja sp. nov. (a. Type locality, b); P. cukiclara sp. nov. (a, b, d. Type locality); P. cataracta sp. nov. (c. type locality); P. batanlenos sp. nov. (a, d. Type locality, e, f); P. horvathrobi sp. nov. (e); P. longabranchias sp. nov. (d. Type locality, f).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
A8328AE5F38E58CEA5CA64B11904DDA9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460661	https://doi.org/10.3897/zookeys.1259.168419.figure30	Figure 30. Distribution of Papuanatula species: yellow dots: known species (Kaltenbach et al. 2025); orange dot: Papuanatula arfak sp. nov. (present study); red dots: species from Batanta – P. batantaraja sp. nov., P. cukiclara sp. nov., P. cataracta sp. nov., P. batanlenos sp. nov., P. horvathrobi sp. nov., P. longabranchias sp. nov. (present study).	Figure 30. Distribution of Papuanatula species: yellow dots: known species (Kaltenbach et al. 2025); orange dot: Papuanatula arfak sp. nov. (present study); red dots: species from Batanta – P. batantaraja sp. nov., P. cukiclara sp. nov., P. cataracta sp. nov., P. batanlenos sp. nov., P. horvathrobi sp. nov., P. longabranchias sp. nov. (present study).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
13E61C5AA69F58A6B76B6E98436A730A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460637	https://doi.org/10.3897/zookeys.1259.168419.figure6	Figure 6. Papuanatula (Papuanatula) batantaraja sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 6. Papuanatula (Papuanatula) batantaraja sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
13E61C5AA69F58A6B76B6E98436A730A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460638	https://doi.org/10.3897/zookeys.1259.168419.figure7	Figure 7. Papuanatula (Papuanatula) batantaraja sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Right mandible; d. Left mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h, i. Tergalius IV; j. Head. Scale bars: 100 µm (j), 20 µm (a, c – h), 10 µm (b, i).	Figure 7. Papuanatula (Papuanatula) batantaraja sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Right mandible; d. Left mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h, i. Tergalius IV; j. Head. Scale bars: 100 µm (j), 20 µm (a, c – h), 10 µm (b, i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
13E61C5AA69F58A6B76B6E98436A730A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460639	https://doi.org/10.3897/zookeys.1259.168419.figure8	Figure 8. Papuanatula (Papuanatula) batantaraja sp. nov., larva. a. Hind leg; b. Hind femur, outer margin; c. Hind femur, surface; d. Hind tarsus, claw (arrow: posterior seta); e. Abdomen (arrow: protuberance on fore protoptera); f. Abdominal tergum V, surface; g. Abdominal terga; h. Paracercus; i. Cercus; j. Paraproct. Scale bars: 100 µm (e), 50 µm (a), 20 µm (b, d, h, j), 10 µm (c, f, g, i).	Figure 8. Papuanatula (Papuanatula) batantaraja sp. nov., larva. a. Hind leg; b. Hind femur, outer margin; c. Hind femur, surface; d. Hind tarsus, claw (arrow: posterior seta); e. Abdomen (arrow: protuberance on fore protoptera); f. Abdominal tergum V, surface; g. Abdominal terga; h. Paracercus; i. Cercus; j. Paraproct. Scale bars: 100 µm (e), 50 µm (a), 20 µm (b, d, h, j), 10 µm (c, f, g, i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
13E61C5AA69F58A6B76B6E98436A730A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460660	https://doi.org/10.3897/zookeys.1259.168419.figure29	Figure 29. Habitats: P. batantaraja sp. nov. (a. Type locality, b); P. cukiclara sp. nov. (a, b, d. Type locality); P. cataracta sp. nov. (c. type locality); P. batanlenos sp. nov. (a, d. Type locality, e, f); P. horvathrobi sp. nov. (e); P. longabranchias sp. nov. (d. Type locality, f).	Figure 29. Habitats: P. batantaraja sp. nov. (a. Type locality, b); P. cukiclara sp. nov. (a, b, d. Type locality); P. cataracta sp. nov. (c. type locality); P. batanlenos sp. nov. (a, d. Type locality, e, f); P. horvathrobi sp. nov. (e); P. longabranchias sp. nov. (d. Type locality, f).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
13E61C5AA69F58A6B76B6E98436A730A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460661	https://doi.org/10.3897/zookeys.1259.168419.figure30	Figure 30. Distribution of Papuanatula species: yellow dots: known species (Kaltenbach et al. 2025); orange dot: Papuanatula arfak sp. nov. (present study); red dots: species from Batanta – P. batantaraja sp. nov., P. cukiclara sp. nov., P. cataracta sp. nov., P. batanlenos sp. nov., P. horvathrobi sp. nov., P. longabranchias sp. nov. (present study).	Figure 30. Distribution of Papuanatula species: yellow dots: known species (Kaltenbach et al. 2025); orange dot: Papuanatula arfak sp. nov. (present study); red dots: species from Batanta – P. batantaraja sp. nov., P. cukiclara sp. nov., P. cataracta sp. nov., P. batanlenos sp. nov., P. horvathrobi sp. nov., P. longabranchias sp. nov. (present study).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
560DB81217BB52A5B0B34CC75F8DB52D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460643	https://doi.org/10.3897/zookeys.1259.168419.figure12	Figure 12. Papuanatula (Papuanatula) cataracta sp. nov., larva, habitus. a. Dorsal view; b, c. Lateral view; d. Ventral view. Scale bars: 1 mm.	Figure 12. Papuanatula (Papuanatula) cataracta sp. nov., larva, habitus. a. Dorsal view; b, c. Lateral view; d. Ventral view. Scale bars: 1 mm.	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
560DB81217BB52A5B0B34CC75F8DB52D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460644	https://doi.org/10.3897/zookeys.1259.168419.figure13	Figure 13. Papuanatula (Papuanatula) cataracta sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Right mandible; d. Left mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h. Tergalius IV; i. Head. Scale bars: 100 µm (i), 20 µm (a, c – g), 10 µm (b, h).	Figure 13. Papuanatula (Papuanatula) cataracta sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Right mandible; d. Left mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h. Tergalius IV; i. Head. Scale bars: 100 µm (i), 20 µm (a, c – g), 10 µm (b, h).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
560DB81217BB52A5B0B34CC75F8DB52D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460645	https://doi.org/10.3897/zookeys.1259.168419.figure14	Figure 14. Papuanatula (Papuanatula) cataracta sp. nov., larva. a. Hind leg; b. Hind tarsus and claw; c. Abdomen; d. Abdominal tergum V; e. Abdominal terga V and VI; f. Cercus; g. Paracercus; h. Paraproct; i. Hind femur, surface. Scale bars: 50 µm (a, c), 10 µm (b, d – i).	Figure 14. Papuanatula (Papuanatula) cataracta sp. nov., larva. a. Hind leg; b. Hind tarsus and claw; c. Abdomen; d. Abdominal tergum V; e. Abdominal terga V and VI; f. Cercus; g. Paracercus; h. Paraproct; i. Hind femur, surface. Scale bars: 50 µm (a, c), 10 µm (b, d – i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
560DB81217BB52A5B0B34CC75F8DB52D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460660	https://doi.org/10.3897/zookeys.1259.168419.figure29	Figure 29. Habitats: P. batantaraja sp. nov. (a. Type locality, b); P. cukiclara sp. nov. (a, b, d. Type locality); P. cataracta sp. nov. (c. type locality); P. batanlenos sp. nov. (a, d. Type locality, e, f); P. horvathrobi sp. nov. (e); P. longabranchias sp. nov. (d. Type locality, f).	Figure 29. Habitats: P. batantaraja sp. nov. (a. Type locality, b); P. cukiclara sp. nov. (a, b, d. Type locality); P. cataracta sp. nov. (c. type locality); P. batanlenos sp. nov. (a, d. Type locality, e, f); P. horvathrobi sp. nov. (e); P. longabranchias sp. nov. (d. Type locality, f).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
560DB81217BB52A5B0B34CC75F8DB52D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460661	https://doi.org/10.3897/zookeys.1259.168419.figure30	Figure 30. Distribution of Papuanatula species: yellow dots: known species (Kaltenbach et al. 2025); orange dot: Papuanatula arfak sp. nov. (present study); red dots: species from Batanta – P. batantaraja sp. nov., P. cukiclara sp. nov., P. cataracta sp. nov., P. batanlenos sp. nov., P. horvathrobi sp. nov., P. longabranchias sp. nov. (present study).	Figure 30. Distribution of Papuanatula species: yellow dots: known species (Kaltenbach et al. 2025); orange dot: Papuanatula arfak sp. nov. (present study); red dots: species from Batanta – P. batantaraja sp. nov., P. cukiclara sp. nov., P. cataracta sp. nov., P. batanlenos sp. nov., P. horvathrobi sp. nov., P. longabranchias sp. nov. (present study).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
560DB81217BB52A5B0B34CC75F8DB52D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460642	https://doi.org/10.3897/zookeys.1259.168419.figure11	Figure 11. Papuanatula (Papuanatula) cukiclara sp. nov., larva. a. Hind leg; b. Fore tarsus, claw; c. Fore femur, anterior surface (brown areas); d. Abdominal terga; e. Paraproct; f. Paracercus; g. Cercus; h. Abdomen. Scale bars: 100 µm (h), 50 µm (a), 20 µm (f), 10 µm (b – e, g).	Figure 11. Papuanatula (Papuanatula) cukiclara sp. nov., larva. a. Hind leg; b. Fore tarsus, claw; c. Fore femur, anterior surface (brown areas); d. Abdominal terga; e. Paraproct; f. Paracercus; g. Cercus; h. Abdomen. Scale bars: 100 µm (h), 50 µm (a), 20 µm (f), 10 µm (b – e, g).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
77142C5DC41F5B859C334948BF519B49.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460640	https://doi.org/10.3897/zookeys.1259.168419.figure9	Figure 9. Papuanatula (Papuanatula) cukiclara sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 9. Papuanatula (Papuanatula) cukiclara sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
77142C5DC41F5B859C334948BF519B49.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460641	https://doi.org/10.3897/zookeys.1259.168419.figure10	Figure 10. Papuanatula (Papuanatula) cukiclara sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Left mandible; d. Right mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h. Tergalius IV; i. Head. Scale bars: 100 µm (i), 20 µm (a, c – g), 10 µm (b, h).	Figure 10. Papuanatula (Papuanatula) cukiclara sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Left mandible; d. Right mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h. Tergalius IV; i. Head. Scale bars: 100 µm (i), 20 µm (a, c – g), 10 µm (b, h).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
77142C5DC41F5B859C334948BF519B49.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460642	https://doi.org/10.3897/zookeys.1259.168419.figure11	Figure 11. Papuanatula (Papuanatula) cukiclara sp. nov., larva. a. Hind leg; b. Fore tarsus, claw; c. Fore femur, anterior surface (brown areas); d. Abdominal terga; e. Paraproct; f. Paracercus; g. Cercus; h. Abdomen. Scale bars: 100 µm (h), 50 µm (a), 20 µm (f), 10 µm (b – e, g).	Figure 11. Papuanatula (Papuanatula) cukiclara sp. nov., larva. a. Hind leg; b. Fore tarsus, claw; c. Fore femur, anterior surface (brown areas); d. Abdominal terga; e. Paraproct; f. Paracercus; g. Cercus; h. Abdomen. Scale bars: 100 µm (h), 50 µm (a), 20 µm (f), 10 µm (b – e, g).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
77142C5DC41F5B859C334948BF519B49.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460660	https://doi.org/10.3897/zookeys.1259.168419.figure29	Figure 29. Habitats: P. batantaraja sp. nov. (a. Type locality, b); P. cukiclara sp. nov. (a, b, d. Type locality); P. cataracta sp. nov. (c. type locality); P. batanlenos sp. nov. (a, d. Type locality, e, f); P. horvathrobi sp. nov. (e); P. longabranchias sp. nov. (d. Type locality, f).	Figure 29. Habitats: P. batantaraja sp. nov. (a. Type locality, b); P. cukiclara sp. nov. (a, b, d. Type locality); P. cataracta sp. nov. (c. type locality); P. batanlenos sp. nov. (a, d. Type locality, e, f); P. horvathrobi sp. nov. (e); P. longabranchias sp. nov. (d. Type locality, f).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
77142C5DC41F5B859C334948BF519B49.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460661	https://doi.org/10.3897/zookeys.1259.168419.figure30	Figure 30. Distribution of Papuanatula species: yellow dots: known species (Kaltenbach et al. 2025); orange dot: Papuanatula arfak sp. nov. (present study); red dots: species from Batanta – P. batantaraja sp. nov., P. cukiclara sp. nov., P. cataracta sp. nov., P. batanlenos sp. nov., P. horvathrobi sp. nov., P. longabranchias sp. nov. (present study).	Figure 30. Distribution of Papuanatula species: yellow dots: known species (Kaltenbach et al. 2025); orange dot: Papuanatula arfak sp. nov. (present study); red dots: species from Batanta – P. batantaraja sp. nov., P. cukiclara sp. nov., P. cataracta sp. nov., P. batanlenos sp. nov., P. horvathrobi sp. nov., P. longabranchias sp. nov. (present study).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
7932E4979B925620A02E574FA696DA9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460632	https://doi.org/10.3897/zookeys.1259.168419.figure1	Figure 1. Papuanatula (Papuanatula) arfak sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 1. Papuanatula (Papuanatula) arfak sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
7932E4979B925620A02E574FA696DA9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460633	https://doi.org/10.3897/zookeys.1259.168419.figure2	Figure 2. Papuanatula (Papuanatula) arfak sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Right mandible; d. Left mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h. Maxillary palp; i. Developing subimaginal gonostyli. Scale bars: 50 µm (e), 20 µm (a, c, d, f, g, i), 10 µm (b, h).	Figure 2. Papuanatula (Papuanatula) arfak sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Right mandible; d. Left mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h. Maxillary palp; i. Developing subimaginal gonostyli. Scale bars: 50 µm (e), 20 µm (a, c, d, f, g, i), 10 µm (b, h).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
7932E4979B925620A02E574FA696DA9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460634	https://doi.org/10.3897/zookeys.1259.168419.figure3	Figure 3. Papuanatula (Papuanatula) arfak sp. nov., larva. a. Middle leg; b. Fore tarsus; c. Fore claw; d. Fore femur, posterior apex; e. Fore tibia, posterior surface; f. Middle tibia, inner margin; g. Fore femur, ventral margin; h. Head, mature male larva. Scale bars: 100 µm (h), 50 µm (a), 20 µm (b), 10 µm (c – g).	Figure 3. Papuanatula (Papuanatula) arfak sp. nov., larva. a. Middle leg; b. Fore tarsus; c. Fore claw; d. Fore femur, posterior apex; e. Fore tibia, posterior surface; f. Middle tibia, inner margin; g. Fore femur, ventral margin; h. Head, mature male larva. Scale bars: 100 µm (h), 50 µm (a), 20 µm (b), 10 µm (c – g).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
7932E4979B925620A02E574FA696DA9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460635	https://doi.org/10.3897/zookeys.1259.168419.figure4	Figure 4. Papuanatula (Papuanatula) arfak sp. nov., larva. a. Abdominal terga, posterior margins; b. Paracercus; c. Cercus, basal part; d. Cercus, distal part; e, f. Tergalius IV; g – i. Paraproct. Scale bars: 50 µm (h), 20 µm (a, b, g), 10 µm (c, d, f, h, i).	Figure 4. Papuanatula (Papuanatula) arfak sp. nov., larva. a. Abdominal terga, posterior margins; b. Paracercus; c. Cercus, basal part; d. Cercus, distal part; e, f. Tergalius IV; g – i. Paraproct. Scale bars: 50 µm (h), 20 µm (a, b, g), 10 µm (c, d, f, h, i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
7932E4979B925620A02E574FA696DA9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460636	https://doi.org/10.3897/zookeys.1259.168419.figure5	Figure 5. Papuanatula (Papuanatula) arfak sp. nov., larva, SEM pictures. a. Abdominal terga III – VII; b. Abdominal tergum VII, posterior margin. Scale bars: 300 µm (a), 40 µm (b).	Figure 5. Papuanatula (Papuanatula) arfak sp. nov., larva, SEM pictures. a. Abdominal terga III – VII; b. Abdominal tergum VII, posterior margin. Scale bars: 300 µm (a), 40 µm (b).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
7932E4979B925620A02E574FA696DA9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460637	https://doi.org/10.3897/zookeys.1259.168419.figure6	Figure 6. Papuanatula (Papuanatula) batantaraja sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 6. Papuanatula (Papuanatula) batantaraja sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
7932E4979B925620A02E574FA696DA9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460638	https://doi.org/10.3897/zookeys.1259.168419.figure7	Figure 7. Papuanatula (Papuanatula) batantaraja sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Right mandible; d. Left mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h, i. Tergalius IV; j. Head. Scale bars: 100 µm (j), 20 µm (a, c – h), 10 µm (b, i).	Figure 7. Papuanatula (Papuanatula) batantaraja sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Right mandible; d. Left mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h, i. Tergalius IV; j. Head. Scale bars: 100 µm (j), 20 µm (a, c – h), 10 µm (b, i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
7932E4979B925620A02E574FA696DA9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460639	https://doi.org/10.3897/zookeys.1259.168419.figure8	Figure 8. Papuanatula (Papuanatula) batantaraja sp. nov., larva. a. Hind leg; b. Hind femur, outer margin; c. Hind femur, surface; d. Hind tarsus, claw (arrow: posterior seta); e. Abdomen (arrow: protuberance on fore protoptera); f. Abdominal tergum V, surface; g. Abdominal terga; h. Paracercus; i. Cercus; j. Paraproct. Scale bars: 100 µm (e), 50 µm (a), 20 µm (b, d, h, j), 10 µm (c, f, g, i).	Figure 8. Papuanatula (Papuanatula) batantaraja sp. nov., larva. a. Hind leg; b. Hind femur, outer margin; c. Hind femur, surface; d. Hind tarsus, claw (arrow: posterior seta); e. Abdomen (arrow: protuberance on fore protoptera); f. Abdominal tergum V, surface; g. Abdominal terga; h. Paracercus; i. Cercus; j. Paraproct. Scale bars: 100 µm (e), 50 µm (a), 20 µm (b, d, h, j), 10 µm (c, f, g, i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
7932E4979B925620A02E574FA696DA9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460640	https://doi.org/10.3897/zookeys.1259.168419.figure9	Figure 9. Papuanatula (Papuanatula) cukiclara sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 9. Papuanatula (Papuanatula) cukiclara sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
7932E4979B925620A02E574FA696DA9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460641	https://doi.org/10.3897/zookeys.1259.168419.figure10	Figure 10. Papuanatula (Papuanatula) cukiclara sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Left mandible; d. Right mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h. Tergalius IV; i. Head. Scale bars: 100 µm (i), 20 µm (a, c – g), 10 µm (b, h).	Figure 10. Papuanatula (Papuanatula) cukiclara sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Left mandible; d. Right mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h. Tergalius IV; i. Head. Scale bars: 100 µm (i), 20 µm (a, c – g), 10 µm (b, h).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
7932E4979B925620A02E574FA696DA9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460642	https://doi.org/10.3897/zookeys.1259.168419.figure11	Figure 11. Papuanatula (Papuanatula) cukiclara sp. nov., larva. a. Hind leg; b. Fore tarsus, claw; c. Fore femur, anterior surface (brown areas); d. Abdominal terga; e. Paraproct; f. Paracercus; g. Cercus; h. Abdomen. Scale bars: 100 µm (h), 50 µm (a), 20 µm (f), 10 µm (b – e, g).	Figure 11. Papuanatula (Papuanatula) cukiclara sp. nov., larva. a. Hind leg; b. Fore tarsus, claw; c. Fore femur, anterior surface (brown areas); d. Abdominal terga; e. Paraproct; f. Paracercus; g. Cercus; h. Abdomen. Scale bars: 100 µm (h), 50 µm (a), 20 µm (f), 10 µm (b – e, g).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
7932E4979B925620A02E574FA696DA9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460643	https://doi.org/10.3897/zookeys.1259.168419.figure12	Figure 12. Papuanatula (Papuanatula) cataracta sp. nov., larva, habitus. a. Dorsal view; b, c. Lateral view; d. Ventral view. Scale bars: 1 mm.	Figure 12. Papuanatula (Papuanatula) cataracta sp. nov., larva, habitus. a. Dorsal view; b, c. Lateral view; d. Ventral view. Scale bars: 1 mm.	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
7932E4979B925620A02E574FA696DA9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460644	https://doi.org/10.3897/zookeys.1259.168419.figure13	Figure 13. Papuanatula (Papuanatula) cataracta sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Right mandible; d. Left mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h. Tergalius IV; i. Head. Scale bars: 100 µm (i), 20 µm (a, c – g), 10 µm (b, h).	Figure 13. Papuanatula (Papuanatula) cataracta sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Right mandible; d. Left mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h. Tergalius IV; i. Head. Scale bars: 100 µm (i), 20 µm (a, c – g), 10 µm (b, h).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
7932E4979B925620A02E574FA696DA9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460645	https://doi.org/10.3897/zookeys.1259.168419.figure14	Figure 14. Papuanatula (Papuanatula) cataracta sp. nov., larva. a. Hind leg; b. Hind tarsus and claw; c. Abdomen; d. Abdominal tergum V; e. Abdominal terga V and VI; f. Cercus; g. Paracercus; h. Paraproct; i. Hind femur, surface. Scale bars: 50 µm (a, c), 10 µm (b, d – i).	Figure 14. Papuanatula (Papuanatula) cataracta sp. nov., larva. a. Hind leg; b. Hind tarsus and claw; c. Abdomen; d. Abdominal tergum V; e. Abdominal terga V and VI; f. Cercus; g. Paracercus; h. Paraproct; i. Hind femur, surface. Scale bars: 50 µm (a, c), 10 µm (b, d – i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
7932E4979B925620A02E574FA696DA9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460646	https://doi.org/10.3897/zookeys.1259.168419.figure15	Figure 15. Papuanatula (Papuanatula) batanlenos sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 15. Papuanatula (Papuanatula) batanlenos sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
7932E4979B925620A02E574FA696DA9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460647	https://doi.org/10.3897/zookeys.1259.168419.figure16	Figure 16. Papuanatula (Papuanatula) batanlenos sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Left mandible; d. Right mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h, i. Tergalius IV; j. Head; k. Developing subimaginal gonostyli. Scale bars: 100 µm (j), 20 µm (a, c – h, k), 10 µm (b, i).	Figure 16. Papuanatula (Papuanatula) batanlenos sp. nov., larva. a. Labrum; b. Labrum, submarginal arc of setae; c. Left mandible; d. Right mandible; e. Labium; f. Hypopharynx and superlinguae; g. Maxilla; h, i. Tergalius IV; j. Head; k. Developing subimaginal gonostyli. Scale bars: 100 µm (j), 20 µm (a, c – h, k), 10 µm (b, i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
7932E4979B925620A02E574FA696DA9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460648	https://doi.org/10.3897/zookeys.1259.168419.figure17	Figure 17. Papuanatula (Papuanatula) batanlenos sp. nov., larva. a. Middle leg; b. Middle femur, outer margin; c. Middle tarsus, outer margin; d. Middle claw; e. Abdomen; f. Abdominal terga; g. Paraproct; h. Paracercus; i. Cercus. Scale bars: 100 µm (a, e), 20 µm (h), 10 µm (b – d, f, g, i).	Figure 17. Papuanatula (Papuanatula) batanlenos sp. nov., larva. a. Middle leg; b. Middle femur, outer margin; c. Middle tarsus, outer margin; d. Middle claw; e. Abdomen; f. Abdominal terga; g. Paraproct; h. Paracercus; i. Cercus. Scale bars: 100 µm (a, e), 20 µm (h), 10 µm (b – d, f, g, i).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
7932E4979B925620A02E574FA696DA9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460649	https://doi.org/10.3897/zookeys.1259.168419.figure18	Figure 18. Papuanatula (Papuanatula) batanlenos sp. nov, imago, male. a. Habitus, lateral view; b. Habitus, dorsal view; c. Middle leg; d. Genitalia. Scale bars: 1 mm (a, b), 100 µm (c), 50 µm (d).	Figure 18. Papuanatula (Papuanatula) batanlenos sp. nov, imago, male. a. Habitus, lateral view; b. Habitus, dorsal view; c. Middle leg; d. Genitalia. Scale bars: 1 mm (a, b), 100 µm (c), 50 µm (d).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
7932E4979B925620A02E574FA696DA9C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1460650	https://doi.org/10.3897/zookeys.1259.168419.figure19	Figure 19. Papuanatula (Papuanatula) batanlenos sp. nov. Imago, male. a. Fore wing. Subimagos. b. Habitus, male; c. Habitus, female; d. Middle leg; e. Fore tarsus; f. Middle tarsus. Scale bars: 1 mm (a – c), 100 µm (d), 20 µm (e), 10 µm (f).	Figure 19. Papuanatula (Papuanatula) batanlenos sp. nov. Imago, male. a. Fore wing. Subimagos. b. Habitus, male; c. Habitus, female; d. Middle leg; e. Fore tarsus; f. Middle tarsus. Scale bars: 1 mm (a – c), 100 µm (d), 20 µm (e), 10 µm (f).	2025-11-10	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc		Zenodo	biologists	Kaltenbach, Thomas;Kovács, Tibor;Gattolliat, Jean-Luc			
