taxonID	type	description	language	source
A884DA106BD150DAABC547E8FAF2D79D.taxon	description	Figs 20, 21, 22	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
A884DA106BD150DAABC547E8FAF2D79D.taxon	diagnosis	Diagnosis. Larva. The following combination of characters distinguishes P. horvathrobi sp. nov. from other species of Papuanatula (Papuafiliola): thorax and abdomen dorsally without protuberances; femur with large blank in posterior ⅔; femur and abdomen without brown hypodermal maculae; tergalii oblong; paracercus with 15 segments.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
A884DA106BD150DAABC547E8FAF2D79D.taxon	description	Description. Larva (Fig. 20 – 22). Body length ~ 3.0 mm, cerci ~ 1.4 × body length. Cuticular coloration (Figs 20 a, b, 22 a). Head, thorax, and abdomen dorsally brown. Head, thorax and abdomen ventrally beige. Legs brown, femur with large blank in posterior ⅔, and distally with small blank area. Caudalii light brown. Hypodermal coloration (Fig. 20 a, b). Head, thorax, abdomen, and legs without hypodermal markings. Head. Antenna. Length ~ 2.5 × head length. Otherwise, as typical for the subgenus. Developing turbinate eyes in last instar male larva (Figs 20 a, 21 j) large, roundish, with some distance to each other. Labrum (Fig. 21 a, b). Length 0.6 × maximal width. With reduced, submarginal arc of three long, simple setae. Otherwise, as typical for subgenus. Right mandible (Fig. 21 d). Incisor with five denticles; kinetodontium with three denticles, inner lateral margin with row of small denticles, becoming smaller and finer toward base, and with row of short, fine setae; margin between prostheca and mola slightly convex, with few minute denticles. Otherwise, as typical for subgenus. Left mandible (Fig. 21 c). Incisor with five denticles, kinetodontium with three denticles; margin between prostheca and subtriangular process straight, with minute denticles. Otherwise, as typical for subgenus. Hypopharynx (Fig. 21 f) with well-developed tuft of long, spine-like setae. Otherwise, as typical for the genus. Maxilla (Fig. 21 g). Maxillary palp slightly longer than galea-lacinia; segment I 1.4 × length of segment II. Otherwise, as typical for the genus. Labium (Fig. 21 e) Paraglossa dorsally with three spine-like setae near inner, distolateral margin. Labial palp with segment I 0.7 × length of segments II and III combined. Segment II with narrow thumb-like distomedial protuberance, slightly bent distad, dorsally with one spine-like seta near outer, distolateral margin. Segment III oblique conical, apically rounded, approx. as long as segment II. Otherwise, as typical for the genus. Thorax. Sterna without protuberances. Terga. Without long setae on midline; without protuberance. Legs (Fig. 22 a – d). Ratio of leg segments: fore leg 1.1: 1.0: 0.3: 0.1, middle leg 1.2: 1.0: 0.3: 0.1 and hind leg 1.5: 1.0: 0.4: 0.2. Femur. Length ~ 3 × maximum width. Surface of brown areas with minute denticles. Tarsus. With regular row of medium, apically rounded setae along outer margin. Claw with one row of 11 denticles; one posterior seta. Otherwise, as typical for subgenus. Abdomen. Terga (Fig. 22 e, f, j). Abdominal terga without long setae on midline; abdominal terga without median or submedian elevations or protuberances; surface with minute denticles. Posterior margins of abdominal terga: I smooth, without denticles, II – IX with very small, triangular denticles. Tergalii (Fig. 21 h, i). Present on terga II – VII, oblong; tracheation well-developed, pigmentation not reaching margins. Each tergalius with anal rib longer than costal rib; ribs with minute, irregularly situated denticles on dorsal side. Tergalius II as long as terga III and ⅔ IV combined, tergalius IV as long as terga V and ⅔ VI combined, tergalius VII as long as terga VIII and ⅔ IX combined. Paraproct (Fig. 22 g) without posterior prolongation. Posterior margin with small denticles. Caudalii (Fig. 22 h, i). Cerci without swimming setae. Paracercus with 15 segments. Pose of subimaginal gonostyli under larval cuticle unknown. Subimago. Unknown. Imago. Unknown. Egg. Unknown.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
A884DA106BD150DAABC547E8FAF2D79D.taxon	etymology	Etymology. The new species is dedicated to Róbert Horváth, a Hungarian ornithologist, who initiated and organized the Hungarian research program on Biodiversity in Batanta in 2010. Since then, he gave continuous support in various aspects of the ten research trips conducted to date.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
A884DA106BD150DAABC547E8FAF2D79D.taxon	distribution	Distribution. New Guinea, Batanta Island (Fig. 30).	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
DDF16181DA045D39AAEB357F74225338.taxon	description	Figs 20, 21, 22, 23, 24, 25, 26, 27, 28	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
DDF16181DA045D39AAEB357F74225338.taxon	diagnosis	Diagnosis (larval characters; see Kaltenbach et al. 2025: 318). Antennal flagellum distally without brown dots; labrum widest at base, dorsally with few submarginal, simple setae (Fig. 24 b); both mandibles without prolonged, blade-like incisor (Fig. 24 c, d); labial palp with distomedian projection on segment II (Fig. 24 e); outer side of femur with regular row of long, slender, flattened, parallel-sided setae with blunt apex (Fig. 25 a, b); tibia with regular row of similar setae (Fig. 25 a, c); tarsus on inner margin with distalmost seta not longer or only slightly longer than other setae.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
993AB6815BE75EF592FCECA16C6A659C.taxon	description	Figs 23, 24, 25, 26, 27	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
993AB6815BE75EF592FCECA16C6A659C.taxon	diagnosis	Diagnosis. Larva. The following combination of characters distinguishes P. longabranchias sp. nov. from other species of Papuanatula (Papuafiliola): abdomen dorsally without long setae on midline; without median protuberances on abdominal terga; abdomen dorsally pale brown, with short, dark brown, lateral streaks along anterior margin of terga I – VII (VIII); femur with dark brown, shoe-shaped marking (marking longer than broad); tergalii very long and narrow; paracercus with 17 or 18 segments. The larva is hardly distinguished from P. stenophylla Kaltenbach, Kluge & Gattolliat, 2025.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
993AB6815BE75EF592FCECA16C6A659C.taxon	description	Description. Larva (Fig. 23 – 25). Body length 3.1 – 3.7 mm, cerci broken. Cuticular coloration (Figs 23 a – c, 25 g). Head, thorax and abdomen dorsally pale brown, mesonotum with some lighter areas, metanotum medially darker brownish. Legs pale brown, with large blank in basal part and small blank area in distal part. Head, thorax and abdomen ventrally beige. Cerci light brownish. Hypodermal coloration (Fig. 23 a, b). Abdomen dorsally with short, dark brown, lateral streaks along anterior margin of terga I – VII (VIII). Anterior side of femur with shoe-shaped, dark brown marking in blank area, longer than wide. Posterior side of femur with dark brown streak in outer distal area. Head. Antenna (Fig. 24 j). As typical for the subgenus, with each flagellomere symmetric, cylindrical. Developing turbinate eyes in last instar male larva (Figs 23 a, 24 j) large, roundish. Labrum (Fig. 24 a) Length 0.6 × maximal width. With reduced, submarginal arc of three long, simple setae. Otherwise, as typical for subgenus. Right mandible (Fig. 24 c) Incisor with five denticles; kinetodontium with three denticles, inner lateral margin with row of small denticles, becoming smaller and finer toward base, and with row of short, fine setae; margin between prostheca and mola slightly convex, smooth. Otherwise, as typical for subgenus. Left mandible (Fig. 24 b) Incisor with five denticles, kinetodontium with four denticles; margin between prostheca and subtriangular process straight, with few minute denticles. Otherwise, as typical for subgenus. Hypopharynx (Fig. 24 f) with tuft of long, straight, stout, spine-like setae. Otherwise, as typical for the genus. Maxilla (Fig. 24 g). Maxillary palp approx. as long as galea-lacinia; segment II ~ 1.5 × length of segment I. Otherwise, as typical for the genus. Labium (Fig. 24 d, e) Paraglossa dorsally with three spine-like setae near inner, distolateral margin. Labial palp with segment I 0.8 × length of segments II and III combined. Segment II with narrow thumb-like distomedial protuberance, slightly bent distad (protuberance somewhat varying), dorsally with one spine-like seta near outer, distolateral margin. Segment III oblong, apically rounded, ~ 0.9 × length of segment II. Otherwise, as typical for the genus. Thorax. Sterna without protuberances. Terga. Without long setae on midline; without protuberances. Legs (Fig. 25 a – f). Ratio of leg segments: fore leg 1.1: 1.0: 0.3: 0.1, middle leg 1.2: 1.0: 0.3: 0.1 and hind leg 1.2: 1.0: 0.3: 0.2. Femur. Length ~ 3.6 × maximum width. Surface rough in brown areas. Tarsus. With regular row of medium, apically rounded setae along outer margin, similar as on tibia, but much shorter. Claw with one row of 9 – 11 denticles; one posterior seta. Otherwise, as typical for subgenus. Abdomen. Terga (Fig. 25 g, h). Abdominal terga without long, fine setae on midline, without median or submedian elevations or protuberances. Surface of abdominal terga rough, with numerous short sensillae, without scattered fine simple setae. Posterior margins of abdominal terga: I smooth, without denticles, II – IX with very small, triangular, apically rounded, dark brown denticles. Tergalii (Fig. 24 h, i). Present on abdominal terga II – VII; long and narrow. Tracheae and pigmentation well-developed, reaching margins; ribs with very small, irregularly situated denticles on dorsal side. Tergalius II as long as terga III and IV combined; tergalius IV as long as terga V, VI, and ½ VII combined; tergalius VII as long as terga VIII, IX, and ½ X combined. Paraproct (Fig. 25 i). Without posterior prolongation; with small spines on posterior margin. Caudalii (Fig. 25 j, k). Cerci without swimming setae. Paracercus with 17 or 18 segments. Pose of subimaginal gonostyli under larval cuticle. Unknown. Subimago (Fig. 27 c – i). Body length 3.6 – 4.1 mm. Turbinate eyes of male ochre. Thorax pale brown. Wing membrane colourless, veins pale brown, microtrichia pale brown. Legs very pale brown; femur anteriorly with hypodermal, shoe-shaped, dark brown marking in basal ½; femur posteriorly with hypodermal, dark brown distal streak. Abdomen of female pale brown; male with abdominal segments I and VII – X pale brown, II – VI nearly colourless; both sexes with hypodermal, sublateral, transvers, dark brown short streaks on posterior margin of terga I – VII. Cerci colourless. Texture (Fig. 27 h, i). On all legs of male and female subimagos, terminal tarsomere covered with pointed microlepides only; other tarsomeres covered mostly with blunt microlepides, with pointed microlepides near apex. Imago, male (Figs 26 a – d, 27 a, b). Body length ~ 3.8 mm. Head pale brown, antennae pale brown, turbinate eyes pale brown, wide, with facetted surfaces round. Thorax pale brown to brown. Fore wing with membrane colourless, veins pale brown, base of RA and costal brace brownish. Pterostigma with two or three oblique crossveins, basalmost crossvein nearly complete, others incomplete. Legs brownish; femur with hypodermal, dark brown, shoe-shaped macula in basal ½ of anterior side; femur on posterior side distally with hypodermal, dark brown streak. Abdominal segments I and VII – X pale brown, II – VI nearly colourless; with hypodermal, dark brown, short, sublateral, transvers streaks on posterior margin of terga I – VII. Cerci colourless. Genitalia (Fig. 27 b). Unistyliger slightly widened apically, with median and lateral margins slightly convex. Gonostylus with segment I on lateral and median side convex and separated from segment II by concavity. Segment II almost equally wide and straight all over its length. Terminal segment III slightly wider than segment II, cube-like. Penial bridge with poorly expressed membranous projection between unistyligers. Imago, female. Unknown. Egg (Fig. 28 a – d). Elongate ovoid. Chorion entirely covered with ridges forming a net-like relief.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
993AB6815BE75EF592FCECA16C6A659C.taxon	etymology	Etymology. The species name longabranchias, meaning “ long gills ” in Latin, refers to the very long and slender shape of the tergalii.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
993AB6815BE75EF592FCECA16C6A659C.taxon	distribution	Distribution. New Guinea, Batanta Island (Fig. 30).	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
79AAA1BCD8D859788D234314421AF9E2.taxon	description	Figs 1, 2, 3, 4, 5	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
79AAA1BCD8D859788D234314421AF9E2.taxon	diagnosis	Diagnosis. Larva. The following combination of characters distinguishes P. arfak sp. nov. from other species of Papuanatula s. str.: body dorsally with irregular row of long, fine, simple setae along midline; abdominal terga without distinct protuberances; abdominal terga I – VI with hypodermal wide dark brown transverse band close to anterior margin, I – IX with medial narrow, dark brown, longitudinal streak; femur proximally with wedge-shaped blank; abdominal terga with triangular, apically rounded denticles on posterior margin; small scattered scales on abdominal terga oblong, striated, apically denticulate; paracercus with nine segments.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
79AAA1BCD8D859788D234314421AF9E2.taxon	description	Description. Larva (Figs 1 – 5). Body length 4.5 – 5.7 mm, cerci much longer than body length (~ 1.4 ×). Cuticular coloration (Figs 1 a – c, 3 a). Head, thorax and abdomen dorsally brown; thorax with indistinct, complex pattern. Femur proximally with wedge-shaped blank, surrounded by grey area, distally yellow-brown to grey-brown; tibia grey; tarsus grey-brown, distally darker. Head, thorax and abdominal segment I ventrally ecru, protuberances of thoracic sterna brown; abdominal segments II – X ventrally brown, laterally paler. Cerci yellow-brown. Hypodermal coloration (Fig. 1 a). Abdominal terga I – VI (VII) with wide dark brown transverse band close to anterior margin, I – IX with narrow dark brown transverse band close to posterior margin and medially narrow dark brown, longitudinal streak. Head (Figs 1 b, 3 h). Dorsally with irregular row of long, fine, simple setae along midline. Antenna. Length ~ 1.5 × head length. As typical for subgenus. Developing turbinate eyes in last instar male larva (Fig. 3 h) large, subquadrangular, touching each other in the middle. Labrum (Fig. 2 a, b). Length 0.5 × maximum width, laterally convex. Dorsal, sub-marginal arc with ~ 30 feathered setae. Right mandible (Fig. 2 c). Margin between prostheca and mola straight, with row of minute denticles. Otherwise, as typical for subgenus. Left mandible (Fig. 2 d). Margin between prostheca and mola straight, with row of minute denticles. Otherwise, as typical for subgenus. Hypopharynx (Fig. 2 f). As typical for genus. Maxilla (Fig. 2 g, h). Maxillary palp subequal in length to galea-lacinia; palp segment II slightly longer than segment I. Otherwise, as typical for the genus. Labium (Fig. 2 e). Paraglossa dorsally with two spine-like setae near inner, distolateral margin. Labial palp with segment I subequal in length to segments II and III combined. Segment II with minute distomedial protuberance, dorsally with row of five or six spine-like setae near outer, distolateral margin. Segment III slightly pentagonal, pointed; 0.8 × length of segment II. Otherwise, as typical for the genus. Thorax. Sterna (Fig. 1 c). With small protuberances on sides of prosternum and close to openings of mesothoracic and metathoracic sternal apodemes. Terga (Fig. 1 b) without protuberance; with irregular row of long, fine, simple setae along midline. Legs (Fig. 3 a – g). Ratio of leg segments: fore leg 0.9: 1.0: 0.3: 0.2, middle leg 1.0: 1.0: 0.3: 0.2 and hind leg 1.1: 1.0: 0.3: 0.2. Femur. Length ~ 4 × maximum width. Claw with one row of 5 – 8 denticles and one posterior setae. Otherwise, as typical for subgenus. Abdomen. Terga (Figs 1 b, 4 a, 5 a, b) with irregular row of long, fine, simple setae along midline. Terga without protuberances, terga I – IV with slight, paired medioposterior elevations. Posterior margin of terga: I smooth, without denticles, II – IX with triangular, partly apically rounded denticles, becoming longer toward end of abdomen. Surface with scattered small, oblong, striated, apically serrate scales. Tergalii (Fig. 4 e, f). Present on terga II – VII. Broad ovoid, tracheation well developed and pigmented; margin smooth, with short, fine, simple setae. Tergalius II nearly reaching end of tergum IV, tergalius IV as long as length of terga V and VI combined, tergalius VII nearly reaching end of tergum IX. Paraproct (Fig. 4 g – i). Posterior margin with prolongation and dense row of minute denticles. Caudalii (Fig. 4 b – d). Cerci apart from basal part with 1 – 6 swimming setae per segment, increasing toward distal part. Paracercus with nine segments. Pose of subimaginal gonostyli under larval cuticle (Fig. 2 i) as typical for the subgenus. Subimago. Unknown. Imago. Unknown. Egg. Unknown.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
79AAA1BCD8D859788D234314421AF9E2.taxon	etymology	Etymology. The species name refers to the locality Arfak, the type locality of the species.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
79AAA1BCD8D859788D234314421AF9E2.taxon	distribution	Distribution. New Guinea (Fig. 30).	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
A8328AE5F38E58CEA5CA64B11904DDA9.taxon	description	Figs 15, 16, 17, 18, 19	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
A8328AE5F38E58CEA5CA64B11904DDA9.taxon	diagnosis	Diagnosis. Larva. The following combination of characters distinguishes P. batanlenos sp. nov. from other species of Papuanatula s. str.: body dorsally without row of long, fine, simple setae along midline; body dorsally without protuberances; abdomen dorsally grey-brown, abdominal terga IV, VII, and VIII with trough-like, dark grey-brown marking, abdominal terga III, V, and VI with dark grey-brown, oblique, lateral streaks; femur with irregular, shoe-shaped marking in large, basal blank; paracercus with 14 – 16 segments; paraproct without posterior prolongation.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
A8328AE5F38E58CEA5CA64B11904DDA9.taxon	description	Description. Larva (Figs 15 – 17). Body length 3.5 – 4.0 mm, cerci ~ 1.6 × body length. Cuticular coloration (Figs 15 a – c, 17 e). Head, thorax and abdomen dorsally grey-brown, thorax darker; abdominal terga IV, VII, and VIII with trough-like, dark grey-brown marking; abdominal terga III, V, and VI with dark grey-brown, oblique, lateral streaks (dorsal abdominal pattern slightly varying). Head, thorax and abdomen ventrally grey-brown. Legs grey-brown, with large blank in basal area. Caudalii yellow-brown. Hypodermal coloration (Fig. 15 a – c). Abdominal terga I – IX with narrow, dark brown, transverse band along posterior margin. Femur anteriorly with dark brown, irregular, shoe-shaped marking in basal ½ (somewhat variable in shape; sometimes poorly developed); posteriorly with dark brown streak distally close to outer margin. Head. Antenna (Fig. 16 j). Length ~ 2.5 × head length. As typical for the subgenus. Developing turbinate eyes in last instar male larva (Fig. 15 a) large, round, nearly touching each other in the middle. Labrum (Fig. 16 a, b). Length 0.5 × maximum width, laterally convex. Dorsal, sub-marginal arc with ~ 15 feathered setae. Right mandible (Fig. 16 d). Margin between prostheca and mola straight, with few minute denticles. Otherwise, as typical for subgenus. Left mandible (Fig. 16 c). Margin between prostheca and mola straight, with few minute denticles. Subtriangular process with undulating margins, outer margin with denticles. Otherwise, as typical for subgenus. Hypopharynx (Fig. 16 f). Medial tuft of spine-like setae laterally denser, giving the impression of a pair of tufts. Otherwise, as typical for the genus. Maxilla (Fig. 16 g). Maxillary palp slightly longer than galea-lacinia; palp segment II ~ 1.2 × length of segment I. Otherwise, as typical for the genus. Labium (Fig. 16 e). Paraglossa with two spine-like setae on inner, distolateral margin. Labial palp with segment I ~ 0.8 × length of segments II and III combined. Segment II without distomedial protuberance, dorsally with row of four spine-like setae near outer, distolateral margin. Segment III broad pentagonal, pointed, 0.8 × length of segment II. Otherwise, as typical for the genus. Thorax. Sterna without protuberances. Terga without protuberances. Legs (Fig. 17 a – d). Ratio of leg segments: fore leg 1.0: 1.0: 0.2: 0.1, middle leg 1.0: 1.0: 0.2: 0.1 and hind leg 1.1: 1.0: 0.2: 0.1. Femur. Length 3 × maximum width. Many short, spine-like setae along ventral margin. Claw with one row of six or seven denticles, distalmost denticle with distance to other denticles; one or two posterior setae. Otherwise, as typical for subgenus. Abdomen. Terga (Fig. 17 f). Abdominal terga without protuberances. Posterior margin of terga: I smooth, without denticles; II – IX with triangular, pointed denticles. Tergalii (Fig. 16 h, i). Ovoid, tracheation rather well developed; margin with minute serration and short, fine, simple setae. Tergalius II as long as abdominal terga III, IV, and ¼ V combined, tergalius IV as long as terga V, VI and ⅓ VII combined, tergalius VII as long as terga VIII and ⅓ IX combined. Paraproct (Fig. 17 g). Posterior margin without prolongation, smooth, without denticles. Caudalii (Fig. 17 h, i). Cerci without swimming setae. Paracercus with 14 – 16 segments. Pose of subimaginal gonostyli under larval cuticle (Fig. 16 k). Segment III conical. Otherwise, as typical for the subgenus. Subimago (Fig. 19 b – f). Body length 4.2 – 4.5 mm. Turbinate eyes of male ochre. Thorax brown. Wing membrane colourless, veins pale yellow-brown, base of wings brown, microtrichia pale yellow-brown. Legs very pale brown; coxa with hypodermal, dark brown fleck; femur anteriorly with hypodermal, shoe-shaped, dark brown marking in basal ½; femur posteriorly with hypodermal, dark brown distal streak and fleck in basal ½. Abdomen yellow-brown; male with hypodermal, dark brown, trough-shaped marking on terga III, IV, VII, and VIII, and sublateral, oblique, dark brown streaks on terga II, V and VI; female with same hypodermal markings, but more diffuse; all terga of both sexes with hypodermal, narrow, transvers, dark brown band on posterior margin. Cerci colourless. Texture (Fig. 19 e, f). On all legs of male and female subimagos, terminal tarsomere covered with pointed microlepides; other tarsomeres covered mostly with blunt microlepides, with pointed microlepides near apex. Imago, male (Figs 18 a – d, 19 a). Body length ~ 3.8 mm. Head and antennae pale brown. Turbinate eyes pale brown, wide, with facetted surfaces round. Thorax pale brown. Fore wing with membrane colourless, base of wings brownish. Pterostigma with three or four oblique crossveins, basal two crossveins nearly complete. Legs light brownish; coxa with hypodermal, dark brown fleck; femur anteriorly with hypodermal, shoe-shaped, dark brown marking in basal ½; femur posteriorly with hypodermal, dark brown distal streak and fleck in basal ½. Abdominal segments I and VII – X pale brown, II – VI transparent, giving a blueish impression; with hypodermal, dark brown, trough-shaped marking on terga III, IV, VII, and VIII, and sublateral, oblique, dark brown streaks on terga II, V, and VI. Cerci colourless. Genitalia (Fig. 18 d). Unistyliger slightly widened apically, with lateral margin slightly concave. Gonostylus with segment I conical and separated from segment II by concavity. Segment II slightly wider in distal ½, bent inward. Terminal segment III narrower than segment II, ovoid, length ~ 1.5 × maximal width. Penial bridge with poorly expressed membranous projection between unistyligers. Imago, female. Unknown. Egg. Unknown. Eggs extracted from female subimagos are undeveloped.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
A8328AE5F38E58CEA5CA64B11904DDA9.taxon	etymology	Etymology. The species name batanlenos refers to the type locality on Batanta island and its belonging to the lenos species group.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
A8328AE5F38E58CEA5CA64B11904DDA9.taxon	distribution	Distribution. New Guinea, Batanta Island (Fig. 30). Key to the species of P. lenos species group from New Guinea (larvae) Based on Kaltenbach et al. 2025.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
13E61C5AA69F58A6B76B6E98436A730A.taxon	description	Figs 6, 7, 8	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
13E61C5AA69F58A6B76B6E98436A730A.taxon	diagnosis	Diagnosis. Larva. The following combination of characters distinguishes P. batantaraja sp. nov. from other species of Papuanatula s. str.: large species (> 7 mm body length); body dorsally without row of long, fine, simple setae along midline; fore protoptera posteromedially with pair of minute, broad protuberances and one minute, rather pointed protuberance between them; metanotum and abdominal terga I – X medially with conspicuous, long, pointed protuberance, slightly bent posteriad; left mandible without subtriangular process; femur anteriorly with angulate blank in basal ½; transparent, semicircular scales on anterior surface of femur and dorsal surface of abdominal terga; paracercus vestigial.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
13E61C5AA69F58A6B76B6E98436A730A.taxon	description	Description. Larva (Figs 6 – 8). Body length 7.0 – 7.5 mm, largest Papuanatula species known so far. Cerci ~ 2 × as long as body length. Cuticular coloration (Figs 6 a – c, 8 e). Head, thorax and abdomen dorsally uniform grey-brown. Head, thorax and abdomen ventrally beige. Legs yellow-brown, femur basally with angulate blank, apically blank. Caudalii yellow-brown. Hypodermal coloration (Fig. 6 a, b). Thorax dorsally with few blackish markings. Abdominal terga with narrow dark brown to blackish transverse band along posterior margins; sigillae blackish. Head. Antenna (Fig. 7 j). Length ~ 1.5 × head length. Flagellum distally with brown dots. Developing turbinate eyes in last instar male larva unknown. Labrum (Fig. 7 a, b). Very wide, length 0.5 × maximum width, laterally angulate. Dorsal, sub-marginal arc with> 40 densely articulated, feathered setae. Right mandible (Fig. 7 d). Margin between prostheca and mola straight, smooth. Otherwise, as typical for the subgenus. Left mandible (Fig. 7 c). Margin between prostheca and mola straight, smooth; subtriangular process not developed. Otherwise, as typical for the subgenus. Hypopharynx (Fig. 7 f). Apical tuft of spine-like setae laterally denser, giving the impression of a pair of tufts. Otherwise, as typical for the genus. Maxilla (Fig. 7 g). Maxillary palp slightly longer than galea-lacinia; palp segment II approx. as long as segment I; segment I thicker than segment II. Otherwise, as typical for the genus. Labium (Fig. 7 e). Paraglossa dorsally with one spine-like seta near inner, distolateral margin. Labial palp with segment I 0.9 × length of segments II and III combined. Segment II with very small, rounded, distomedial protuberance, dorsally with row of five spine-like setae near outer, distolateral margin. Segment III slightly pentagonal, pointed, 0.8 × length of segment II. Otherwise, as typical for the genus. Thorax. Sterna. Protuberances not developed. Terga (Fig. 8 e). Metanotum posteromedially with long, pointed protuberance, slightly bent posteriad. Fore protoptera posteromedially with pair of minute, broad protuberances and one minute, rather pointed protuberance between them. Legs (Fig. 8 a – d). Ratio of leg segments: fore leg 1.1: 1.0: 0.3: 0.1, middle leg 1.0: 1.0: 0.3: 0.1 and hind leg 1.1: 1.0: 0.3: 0.1. Femur. Length ~ 3 × maximum width. Anterior surface with transparent, semicircular scales. Claw with one row of 7 – 9 denticles, apical denticle larger than other ones, and one or two posterior setae, and one or two short, reduced setae opposite to posterior setae. Otherwise, as typical for the subgenus. Abdomen. Terga (Figs 6 a, b, 8 e – g). Abdominal terga I – X posteromedially with conspicuous, long, pointed protuberance, slightly bent posteriad; Posterior margin of terga: I – IX with minute, pointed denticles, slightly increasing in size toward IX. Surface with scattered transparent, semicircular scales. Tergalii (Fig. 7 h, i). Present on terga II – VII. Broad oblique ovoid; tracheation strongly developed; with grey pigmentation, especially in anal ½; margins smooth, with many short, fine, simple setae. Tergalius II as long as abdominal terga III and IV combined, tergalius IV as long as terga V, VI and ⅓ VII combined, tergalius VII reaching beginning of tergum X. Paraproct (Fig. 8 j). Posterior margin expanded, smooth. Caudalii (Fig. 8 h, i) Cerci without swimming setae. Paracercus vestigial. Pose of subimaginal gonostyli under larval cuticle. Unknown. Subimago. Unknown. Imago. Unknown. Egg. Unknown.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
13E61C5AA69F58A6B76B6E98436A730A.taxon	etymology	Etymology. The species name batantaraja refers to the Indonesian island Batanta, where the species was found, and the Indonesian word “ raja ” meaning king, because it is the largest species known in the genus Papuanatula.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
13E61C5AA69F58A6B76B6E98436A730A.taxon	distribution	Distribution. New Guinea, Batanta Island (Fig. 30).	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
560DB81217BB52A5B0B34CC75F8DB52D.taxon	description	Figs 12, 13, 14	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
560DB81217BB52A5B0B34CC75F8DB52D.taxon	diagnosis	Diagnosis. Larva. The following combination of characters distinguishes P. cataracta sp. nov. from other species of Papuanatula s. str.: body dorsally without row of long, fine, simple setae along midline; metanotum and abdominal terga I – VIII posteromedially with cone-like, pointed protuberance, longest on terga III – V, small on VII and VIII; femur anteriorly with irregular blank in basal part; anterior surface of femur and abdominal terga covered with minute, pointed spines; paracercus with five segments.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
560DB81217BB52A5B0B34CC75F8DB52D.taxon	description	Description. Larva (Figs 12 – 14). Body length 2.5 – 3.4 mm, cerci ~ 1.6 × as long as body length. Cuticular coloration (Figs 12 a – d, 14 c). Head, thorax and abdomen dorsally pale brown. Head, thorax and abdomen ventrally pale brown. Legs pale brown, femur anteriorly with irregular blank areas in basal and distal part. Caudalii pale brown. Hypodermal coloration (Fig. 12 a). Abdominal terga with narrow dark brown, transverse band along posterior margins. Head. Antenna (Fig. 13 i). Length ~ 1.5 × head length. Flagellum distally without brown dots. Developing turbinate eyes in last instar male larva (Fig. 13 i) rather small, ovoid, widely spaced. Labrum (Fig. 13 a, b). Length ~ 0.5 × maximum width, laterally convex. Dorsal, sub-marginal arc with ~ 22 feathered setae. Right mandible (Fig. 13 c). Margin between prostheca and mola straight, smooth. Otherwise, as typical for the subgenus. Left mandible (Fig. 13 d). Subtriangular process poorly developed, transparent. Margin between prostheca and mola straight, smooth. Otherwise, as typical for the subgenus. Hypopharynx (Fig. 13 f). As typical for the genus. Maxilla (Fig. 13 g). Maxillary palp approx. as long as galea-lacinia; palp segment II ~ 1.3 × as long as segment I. Otherwise, as typical for the genus. Labium (Fig. 13 e). Paraglossa dorsally with two spine-like setae near inner, distolateral margin. Labial palp with segment I 0.7 × length of segments II and III combined. Segment II without distomedial protuberance, dorsally with row of four or five spine-like setae near outer, distolateral margin. Segment III slightly pentagonal, pointed, 0.8 × length of segment II. Otherwise, as typical for the genus. Thorax. Sterna. With small protuberances on sides of prosternum and close to openings of mesothoracic and metathoracic sternal apodemes (as typical for the subgenus). Terga (Figs 12 c, 14 c). Metanotum posteromedially with cone-like protuberance. Legs (Fig. 14 a, b, i). Ratio of leg segments: fore leg 1.0: 1.0: 0.4: 0.2, middle leg 1.0: 1.0: 0.4: 0.2, and hind leg 1.1: 1.0: 0.4: 0.2. Femur. Length ~ 3 × maximum width. Surface covered with minute, pointed denticles. Tarsus. Setae along outer margin not similar as on tibia as usually, but much shorter, fine, and not ciliate. Claw with one row of six or seven denticles, and one posterior seta. Otherwise, as typical for the subgenus. Abdomen. Terga (Fig. 14 c – e). Abdominal terga I – VIII posteromedially with cone-like, pointed protuberance, longest on terga III – V, small on VII and VIII. Posterior margin of terga: I – IX with short, rounded spines, apically carrying needle-like denticles; partly also triangular, pointed spines toward end of abdomen. Tergalii (Fig. 13 h). Present on terga II – VII. Ovoid; tracheation poorly developed, pigmentation of tracheae mainly limited to main trunk; margins smooth, with short, fine, simple setae. Tergalius II as long as abdominal terga III and ½ IV combined, tergalius IV as long as terga V and ⅓ VI combined, tergalius VII reaching anterior margin of tergum IX. Paraproct (Fig. 14 h). Posterior margin expanded, with denticles in distal part. Caudalii (Fig. 14 f, g) Cerci without swimming setae. Paracercus with five segments. Pose of subimaginal gonostyli under larval cuticle. As typical for the subgenus. Subimago. Unknown. Imago. Unknown. Egg. Unknown.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
560DB81217BB52A5B0B34CC75F8DB52D.taxon	etymology	Etymology. The species name cataracta, meaning waterfall in Latin, refers to the habitat of the larva in the water-film on rocks in fast flowing water.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
560DB81217BB52A5B0B34CC75F8DB52D.taxon	distribution	Distribution. New Guinea, Batanta Island (Fig. 30). Key to the species of P. copis species group from New Guinea (larvae) Based on Kaltenbach et al. 2025.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
77142C5DC41F5B859C334948BF519B49.taxon	description	Figs 9, 10, 11	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
77142C5DC41F5B859C334948BF519B49.taxon	diagnosis	Diagnosis. Larva. The following combination of characters distinguishes P. cukiclara sp. nov. from other species of Papuanatula s. str.: body dorsally without row of long, fine, simple setae along midline; metanotum and abdominal terga I – VIII posteromedially with conspicuous, long, pointed protuberance, slightly bent posteriad; femur anteriorly with irregular wedge-shaped blank in basal part, blank area in distal part, and blank streak along outer margin; paracercus with five or six segments.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
77142C5DC41F5B859C334948BF519B49.taxon	description	Description. Larva (Figs 9 – 11). Body length 2.7 – 3.9 mm, cerci ~ 2 × body length. Cuticular coloration (Figs 9 a – c, 11 c). Head, thorax and abdomen dorsally pale brown; thorax with indistinct pattern; abdominal terga V, VI, and X dorsally and laterally brighter, terga III and IV laterally brighter. Head, thorax and abdomen ventrally pale brown. Legs pale brown, femur anteriorly with irregular wedge-shaped blank in basal part, blank area in distal part, and blank streak along outer margin. Caudalii ecru. Hypodermal coloration (Fig. 9 a, b). Abdominal terga with fine dark brown transverse band along posterior margins. Head. Antenna (Fig. 10 i). Length ~ 1.5 × head length. Flagellum distally with brown dots. Developing turbinate eyes in last instar male larva unknown. Labrum (Fig. 10 a, b). Length ~ 0.5 × maximum width, laterally convex. Dorsal, sub-marginal arc with ~ 22 feathered setae. Right mandible (Fig. 10 d). Margin between prostheca and mola straight, smooth. Otherwise, as typical for the subgenus. Left mandible (Fig. 10 c). Margin between prostheca and mola straight, smooth. Otherwise, as typical for the subgenus. Hypopharynx (Fig. 10 f). As typical for the genus. Maxilla (Fig. 10 g). Maxillary palp approx. as long as galea-lacinia; palp segment II approx. as long as segment I. Otherwise, as typical for the genus. Labium (Fig. 10 e). Paraglossa dorsally with two spine-like setae near inner, distolateral margin. Labial palp with segment I 0.7 × length of segments II and III combined. Segment II without distomedial protuberance, dorsally with row of four spine-like setae near outer, distolateral margin. Segment III slightly pentagonal, pointed, 0.8 × length of segment II. Otherwise, as typical for the genus. Thorax. Sterna. Protuberances not developed. Terga (Figs 9 a, 11 c). Metanotum posteromedially with stout, pointed protuberance. Legs (Fig. 11 a – c). Ratio of leg segments: fore leg 0.9: 1.0: 0.3: 0.2, middle leg 1.0: 1.0: 0.3: 0.2, and hind leg 1.0: 1.0: 0.3: 0.2. Femur. Length ~ 3 × maximum width; surface rough, denticulate on brown areas. Claw with one row of seven or eight denticles, and one posterior seta. Otherwise, as typical for the subgenus. Abdomen. Terga (Figs 9 a, b, 11 h). Abdominal terga I – VIII posteromedially with conspicuous, long, pointed protuberance, slightly bent posteriad; Posterior margin of terga: I smooth, without spines, II – IX with short, rounded spines, apically carrying needle-like denticles. Tergalii (Fig. 10 h). Present on terga II – VII. Oblique ovoid; tracheation poorly developed, pigmentation of tracheae limited to main trunk; margins smooth, with short, fine, simple setae. Tergalius II as long as abdominal terga III and IV combined, tergalius IV as long as terga V and VI combined, tergalius VII reaching middle of tergum IX. Paraproct (Fig. 11 e). Posterior margin expanded, with minute denticles in distal part. Caudalii (Fig. 11 f, g) Cerci without swimming setae. Paracercus with six segments. Pose of subimaginal gonostyli under larval cuticle. As typical for the subgenus (folding in “ Labiobaetis ” - type). Subimago. Unknown. Imago. Unknown. Egg. Unknown.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
77142C5DC41F5B859C334948BF519B49.taxon	etymology	Etymology. The species name is composed of “ cuki ”, meaning cute in Hungarian, and “ clara ”, meaning “ bright ” in Latin. The latter refers to the bright colour of the larva, especially on parts of the abdomen.	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
77142C5DC41F5B859C334948BF519B49.taxon	distribution	Distribution. New Guinea, Batanta Island (Fig. 30).	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
7932E4979B925620A02E574FA696DA9C.taxon	description	Figs 1, 2, 3, 4, 5, 6, 7, 8, 9, 10, 11, 12, 13, 14, 15, 16, 17, 18, 19	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
7932E4979B925620A02E574FA696DA9C.taxon	diagnosis	Diagnosis (larval characters; according to Kaltenbach et al. 2025: 166). Antennal flagellum distally with brown dots; labrum wide, widest in medial area, dorsally with submarginal row of long, feathered setae (Fig. 2 b); both mandibles with incisor strongly elongated, blade-like (Fig. 7 c, d; full-length incisors present only at the beginning of each instar, mostly worn at the end of instar); labial palp without clear distomedial projection at segment II (Fig. 2 e); outer side of femur usually with single regular row of long, hair-like setae bearing numerous fine, short branches on all sides (Fig. 8 a, b); anterior side of tibia usually with regular row of setae similar to that on femur (Fig. 8 a).	en	Kaltenbach, Thomas, Kovács, Tibor, Gattolliat, Jean-Luc (2025): New species of Papuanatula Lugo-Ortiz & McCafferty, 1999 from New Guinea (Ephemeroptera, Baetidae) with focus on Batanta Island. ZooKeys 1259: 1-55, DOI: 10.3897/zookeys.1259.168419
