identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
E4294C700C395C53AEEBC596FBB8812B.text	E4294C700C395C53AEEBC596FBB8812B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ononis tridentata subsp. tridentata	<div><p>Ononis tridentata Linnaeus (1753: 718) subsp. tridentata</p><p>Ononis tridentata Linnaeus (1753: 718) was described in the protologue with a phrase-name “Ononis foliis ternatis carnosis sublinearibus tridentatis”. Linnaeus (1753) stated about the origin of the plant “Habitat in Hispania ” and also cited one synonym “Anonis hispanica frutescens, folio tridentato carnoso” from Magnol (1697), Ray (1704), and Tournefort (1719). The lectotype of this name was designated by F̂rther (in Turland &amp; Jarvis 1997 [August]: 478) from an illustration “Anonis Hispanica frutescens folio tridentato carnoso” published by Magnol (1697: tab. [4] unnumbered). The lectotypification excluded top right-hand plant, which belongs to another species (Fig. 1). This illustration match with the traditional concept and current use of the name as a species with leaflets blade obovate-cuneate, less than six times longer than wider, and with (0–)3–5(–9) teeth in the upper half (see e.g., Losa 1958, Ivimey-Cook 1968, Devesa &amp; López González 1997, Devesa 2000, 2009, López González 2001).</p><p>On the other hand, Devesa &amp; López González (1997 [December]: 254) designated as the “ lectotype ” of the name the same illustration, as “ Lectotypus (Iconotypus): Pierre Magnol, Hortus Regius Monspeliensis, t. [4], 1697, excl. ram. ster. [Designado en este trabajo]”. Although this designation is later than the one published by F̂rther, and therefore ineffective, these authors make an exhaustive exposition about the possible origin of the material used as voucher for Magnol’s illustration (1697: tab. [4] unnumbered). These authors concluded that the origin of the plant cultivated in the Botanical Garden of Montpelier used as voucher could come from seeds collected by Tournefort in Cardona (Catalonia, Spain) during his trip to Spain in 1687 (see Henriques 1898, Devesa &amp; López 1997).</p><p>On the other hand, F̂rther (in Turland &amp; Jarvis 1997: 478) designates an epitype of Ononis tridentata from a specimen preserved at M, as “ Spain, Granada, in salsis et collibus argillosis La Mala prope Granatam, Jul 1837, Boissier (M)”. Unfortunately, this epitypification of O. tridentata published by F̂rther (in Turland &amp; Jarvis 1997: 478) has gone unnoticed (see e.g., Devesa 2000). In the herbarium M there are two specimens of the gathering indicated by F̂rther, barcoded as M0232961 and M0232962 (Fig. 2). Therefore, the epitypification proposed by F̂rther (in Turland &amp; Jarvis 1997) may be further narrowed to a single specimen by a “second-step” epitypification according to Art. 9.17 of the ICN (Turland et al. 2018) (see more details below).</p><p>Consequently, under Art. 9.20, first sentence, and Art. 9.9, the epitype selected by F̂rther (in Turland &amp; Jarvis 1997) must determine the precise application of Ononis tridentata s.str. Although in the epityping it is not mentioned, we assume that F̂rther select an epitype because the lectotype “is demonstrably ambiguous and cannot be critically identified for purposes of the precise application of the name to a taxon” (see ICN Art. 9.9). However, in our opinion, the epitification was not a good choice, since the lectotype of the name, the illustration “Anonis Hispanica frutescens folio tridentato carnoso” in Magnol (1697: tab. [4] unnumbered) can be critically identified for the purposes of its taxonomic identification as was clearly and explicitly mentioned by Devesa &amp; López González (1997).</p><p>Once an epitype is designated in accordance with the provisions of the Shenzhen Code, the identity of the type it supports (in this case the lectotype) is totally irrelevant for the purpose of application of the name. In addition, that although the type being supported by an epitype should be “demonstrably ambiguous and cannot be critically identified for purposes of the precise application of the name to a taxon” (Art. 9.9), even if there is evidence that this is demonstrably not the case (hardly the situation here), there is no provision in the Code for the designated epitype not to continue to determine the application of the name. Therefore, under the current rules O. tridentata must be applied to the species to which F̂rther’s epitype belongs (Art. 9.9 (“An epitype is a specimen or illustration selected to serve as an interpretative type ...”) and Art. 9.20 (“The author who first designates (Art. 7.10, 7.11, and F.5.4) an epitype must be followed.”). Only if the type that an epitype supports is displaced (lost, destroyed, or supersedable under Art. 9.19) does an epitype cease to be the element determining the application of the name.</p><p>The epitype designated by F̂rther (in Turland &amp; Jarvis 1997) “ Spain, Granada, in salsis et collibus argillosis La Mala prope Granatam, Jul 1837, Boissier (M)” is undoubtedly identifiable with the traditional concept and current use to which the name O. tridentata subsp. crassifolia is generally applied (leaflets blade lanceolate–cuneate, less than six times longer than wider, with 3–9 teeth in the upper half, sepals entire, inflorescence peduncle shorter than 10 mm, usually with an inconspicuous arista, inflorescence 1–2-flowered and raceme-like, indumenta of short glandular hairs and sessile glands) (see Devesa &amp; López González 1997, Devesa 2000, 2009, Mota et al. 2011).</p><p>However, the epitypification from the specimen at M generates an abrupt and relevant change in the nomenclature of O. tridentata complex, because the name O. tridentata subsp. tridentata must be applied to populations endemic to the southeastern Iberian Peninsula (with diagnostic characters mentioned above), and another name (in subspecies rank, see below) should be proposed to name the Iberian Peninsula populations traditionally called as O. tridentata subsp. tridentata, with leaflets blade lanceolate–cuneate, less than six times longer than wider, with (0–)3(–5) teeth in the upper half, inflorescence most times 1–2-flowered, axillary; indumenta only of short glandular hairs (see Devesa &amp; López González 1997, Devesa 2000, 2009, López González 2001, Mota et al. 2011, Agulló et al. 2013). Consequently, the names O. barrelieri, O. crassifolia var. intricata, O. tridentata var. canescens, O. tridentata f. intermedia, and O. tridentata var. intermedia f. erecta, traditionally considered as synonyms of O. tridentata subsp. tridentata, should be treated as synonyms of O. tridentata subsp. arbuscula, comb. &amp; stat nov. (see below).</p></div>	https://treatment.plazi.org/id/E4294C700C395C53AEEBC596FBB8812B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferrer-Gallego, P. Pablo	Ferrer-Gallego, P. Pablo (2023): Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae). Phytotaxa 598 (1): 32-58, DOI: 10.11646/phytotaxa.598.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.598.1.3
E4294C700C3C5C51AEEBC34DFB888177.text	E4294C700C3C5C51AEEBC34DFB888177.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ononis crassifolia Dufour ex Boissier, P 1840	<div><p>Ononis crassifolia Dufour ex Boissier (1840: 155)</p><p>Boissier’s protologue (1840: 155) of Ononis crassifolia, numbered “441” in the Voyage botanique dans le midi de l’Espagne, consists of a name “ Ononis crassifolia Dufour. ” followed by three synonyms: 1) “ Ononis tridentata β canescens. DC. Prod.” cited from Candolle (1825: 161), 2) “Anonis Hispanica frutescens foliis crithmi chrysanthemi Tourn. Herb.” cited from Tournefort (1719: 408), and 3) “Cicer montanum fruticans angustifolium serratum minus Barrel. Ic, tab. 419” cited from Barrelier (1714: t. 419). The protologue also includes the provenance “In collibus argillosis et salsis regionis montanae inferioris, circa Cacin et la Mala prov. Granatensis. Alt. 2500’-3000’. Fl. Jun. Jul. / Hab. in Hispaniâ praecipuè interior, regno Granatensi, Valentino (Cav.), Castellâ nova prope Cuenca (Quer), Aragoniâ (Asso), Catalauniâ (Quer)”, and followed by a complete description in French “Je ne sais si cette espèce est ou n’est pas une variété de l’ O. tridentata L.-Ic. Cav. tab. 152, que je n’ai jamais vue et qui se distingue par des feuilles linéaires entières ou tridentées à l’extrémité; mais ces plantes, dissent-elles être réunies, ce qui n’est point prouvé, le nom de Linné devrait être rejeté comme ne pouvant s’appliquer à la forme que nous décrivons et qui est de beaucoup la plus commue. L’ O. crassifolia forme des arbrisseaux de deux à quatre pieds de haut couverts d’un duvet blanchâtre, les feuilles sont épaisses, charnues, caduques, cassantes et glauques comme celles de l’ H. squamatum, de certains Atriplex et d’autres plantes particulières aux terrains argileux à sources salées, ells sont obovées et munies de six à huit petites dents ou crénelures peu prononcées et toutes égales entre elles; les fleurs sont roses avec l’extrémité de la carène plus forncée, les legumes sont velus, ovales, atténués par les deux estrémités, et deux fois plus longs que le calice”.</p><p>This name was not typified by Burdet et al. (1988). On the other hand, Devesa &amp; López González (1997: 258) mentioned “Material original: G., herb. Boissier [n.v.]”. However, this statement cannot be considered as an effective typification according to Art. 7.11 of the ICN because the citation does not include the term “ type ” (typus) or an equivalent.</p><p>Among the original elements used by Boissier to describe his Ononis crassifolia, the references of Barrelier (1714: t. 419) provided an illustration that can be considered an original element. In addition, there is a gathering of Boissier with material collected “in salsis et collibus argillosis La Mala” [La Malahá] in July 1837, with several specimens preserved at M, MEL, and P (see below). These all specimens are undoubtedly identifiable with the species to which the name Ononis crassifolia is generally applied (leaflets blade obovate-cuneate, less than six times longer than wider, with 3–9 teeth in the upper half, sepals entire, inflorescence peduncle shorter than 10 mm, usually with an inconspicuous arista, inflorescence 1–2-flowered and raceme-like). I designate as the lectotype of the name Ononis crassifolia a specimen of this gathering preserved at P, barcoded P03447750 (Fig. 3). As mentioned above, a specimen of this gathering was selected as the epitype of O. tridentata by F̂rther (in Turland &amp; Jarvis 1997: 478). Therefore, O. crassifolia is a heterotypic synonym of O. tridentata .</p><p>Ononis tridentata f. latifolia Lange (1865: 153)</p><p>The protologue of Ononis tridentata f. latifolia published by Lange (1865: 153) includes the epithet “3. latifolia ” followed by the synonyms “ O. crassifolia Duf. ined., Willk. l. c. ex p.; Barr. ic. 420; O. Barrelieri Duf. ex p.), a bief diagnosis “foliis late obovatis, 5–7-dentatis”, and two gatherings “Vidi specimina lecta ad la Mala pr. Granada (Boiss.)! la Armilla fl. granat. (P. del Campo)!”. The protologue includes the gathering of Boissier from “La Mala” (Granada, Spain). A specimen of this gathering was designated as the epitype of the name Ononis crassifolia (M [and in this work as a second-step epitypification with barcode M0232962]), the gathering bears also several duplicates or isoepitypes (see e.g., M0232961, MEL2471775, P03447783 [stem mounted on the right half of the sheet], P03447784 [two stems mounted on the upper left half of the sheet], P03447754, P03447785 [three stems mounted on the left half of the sheet], P03447669, P03447750). On the other hand, there are other relevant syntypes of the second gathering cited in the protologue from Armilla and collected by Pedro del Campo (e.g., COI00053682, LY0175028, MPU1207861, P03447766, P03447752, P03490744, P03079357, P02993329, VTA011733). These sheets bear a printed label annotated as: “Pedro del Campo, Pl. de la Prov. de Grenade, 1852. / Distr. par E. Bourgeau. / 33. Ononis crassifolia, L. Duf. / (Coss.) / Granada, plaine salée de Armilla. / 6 Juin. Among all this material, I designate the specimen with barcode P02993329 (Fig. 4) as the lectotype of the name Ononis tridentata f. latifolia, the specimens at COI (barcode COI00053682), LY (barcode LY0175028), MPU (barcode MPU1207861), P (barcodes P03447766, P03447752, P03079357, and P03490744), and VTA (barcode VTA011733) are therefore isolectotypes.</p></div>	https://treatment.plazi.org/id/E4294C700C3C5C51AEEBC34DFB888177	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferrer-Gallego, P. Pablo	Ferrer-Gallego, P. Pablo (2023): Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae). Phytotaxa 598 (1): 32-58, DOI: 10.11646/phytotaxa.598.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.598.1.3
E4294C700C3E5C51AEEBC368FBF587AC.text	E4294C700C3E5C51AEEBC368FBF587AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ononis tridentata Linnaeus 1753	<div><p>Ononis tridentata Linnaeus (1753: 718) tridentata</p><p>Lectotype (designated by F̂rther in Turland &amp; Jarvis (1997: 478)): [illustration] “Anonis Hispanica frutescens folio tridentato carnoso” in Magnol (1697: tab. [4] unnumbered) [excluding top right-hand plant] (Fig. 1)</p><p>Epitype (first-step designated by F̂rther in Turland &amp; Jarvis (1997: 478)):— [SPAIN], Granada, in salsis et collibus argillosis La Mala [ La Malahá] prope Granatam, Jul 1837, Boissier (M); (second-step designated here): M (barcode M0232962!) (Fig. 2). Isoepitypes: M (barcodes M0232961!), MEL (barcode MEL2471775!), P (barcodes P03447783! [stem mounted on the right half of the sheet], P03447784! [two stems mounted on the upper left half of the sheet], P03447754!, P03447785! [three stems mounted on the left half of the sheet], P03447669!, P03447750!) .</p><p>= O. crassifolia Dufour ex Boissier (1840: 155)</p><p>≡ O. tridentata subsp. crassifolia (Dufour ex Boiss.) Nyman (1878: 160)</p><p>Lectotype (designated here):—[SPAIN], Granada, “in salsis et collibus argillosis La Mala propé Granatam, Jul 1837, E. Boissier s.n., P (barcode P03447750!) (Fig. 3). Isolectotypes: M (barcodes M0232961! and M0232962!), MEL (barcode MEL2471775!), P (barcodes P03447783! [stem mounted on the right half of the sheet], P03447784! [two stems mounted on the upper left half of the sheet], P03447754!, P03447785! [three stems mounted on the left half of the sheet], P03447669!).</p><p>= O. tridentata f. latifolia Lange (1865: 153)</p><p>Lectotype (designated here):—[SPAIN], Granada, “plaine salée de Armilla”, 6 Jun 1852, Pedro del Campo 33, P (barcode P02993329) (Fig. 4). Isolectotypes: COI (barcode COI00053682!), LY (barcode LY0175028!), MPU (barcode MPU1207861!), P (barcodes P03447766!, P03447752!, P03079357! and P03490744!), VTA (barcode VTA011733!).</p></div>	https://treatment.plazi.org/id/E4294C700C3E5C51AEEBC368FBF587AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferrer-Gallego, P. Pablo	Ferrer-Gallego, P. Pablo (2023): Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae). Phytotaxa 598 (1): 32-58, DOI: 10.11646/phytotaxa.598.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.598.1.3
E4294C700C3E5C5FAEEBC5D4FECC8433.text	E4294C700C3E5C5FAEEBC5D4FECC8433.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ononis tridentata subsp. arbuscula (Desvaux) P. P. Ferrer	<div><p>Ononis tridentata subsp. arbuscula (Desvaux) P.P. Ferrer</p><p>As mentioned above, due to the epitype designation for the name Ononis tridentata published by F̂rther (in Turland &amp; Jarvis 1997: 478), a new name in subspecies rank should be proposed for the Iberian populations that until now were known with the traditional concept (pre-epitypification) of O. tridentata subsp. tridentata .</p><p>In this sense, there is a relevant early name published by Desvaux (1814: 76), Ononis arbuscula (see Losa 1958, Devesa &amp; López González 1997). Nicaise Auguste Desvaux (1784-1856) was a French botanist at Angers (France). He was appointed director of the Angers botanic garden in 1817. Desvaux did not personally collect botanical specimens beyond the region of Angers. He received specimens from many correspondents, but the collector is not noted and the area stated is rarely specific (Howard et al. 1981). In his Journal de Botanique Appliquée Desvaux proposed a large number of species. It would not be true to say that his work was neglected; but it was difficult to follow. His descriptions, though accurate as far as they go, are brief and, like so many older ones, omit details later deemed essential. His data of locality are frequently very general and by no means always correct. He never cited collectors; authentic material, where it exists, has been, therefore, hard to identify as such. Until 1896, when it was presented to the Museum d’Histoire Naturelle at Paris, his personal herbarium remained in the possession of his family, presumably more or less inaccesible (Weatherby 1936). Following Desvaux’s death in 1856, his general herbarium was sold by his son to Alphonse Lavallée (Bureau 1896, Bonnet 1907). Lavallée moved the collection to Segrez (Seine-et-Oise). It was given to the Museum of Natural History, Paris, by his widow in 1896 (Bureau 1896). The collection has been subsequently integrated into the general herbarium (Howard et al. 1981). Several types are preserved at P, P-JU, and PC, other material is kept at B (Willdenow), FI, G, and M (Stafleu &amp; Cowan 1976).</p><p>An unusually large proportion of Desvaux’s species have been misunderstood or left as frankly doubtful. One of these species could be Ononis arbuscula . The identity of this name has long been controversial, it could not be assigned with confidence to any known member of the group due to the ambiguity of the name and the ensuing nomenclatural confusion and uncertainty among users (see Devesa &amp; López González 1997). One option would have been to reject this troublesome name altogether. However, a precise typification would allow the use of the name Ononis arbuscula .</p><p>The protologue of Ononis arbuscula Desvaux (1814: 76) includes a description in Latin “Caule fruticoso racemis infra tuberculosis, apicè tomentosis; foliis crassis glabris subspathulatis apicè dentatis; calicibus sub-glabris; leguminibus solitariis pedunculatis hirsutis” followed by the geographical provenance “Habitât in Europeâ australis”.</p><p>In the Willdenow herbarium at B there is a specimen of Ononis tridentata . The sheet, with barcode B -W 13318 -00 0, bears six poorly preserved plant fragments plant of this species, and is annotated “ On. tridentata / 1.” at the top of the sheet, and “Roestel. W.” at the base of the sheet. The sheet also bears an original label handwritten by Willdenow “Diadelphia Decandria / Ononis tridentata fruticosa, / foliis ternatis cyanosis sub-/ linearibus tridentatis, pedun-/ culis bifloris Lin. Sp. Pl. ed R3. p. 431. / Habitat in Hispania ” (image available at https://herbarium.bgbm.org/ object/BW13318000). However, due to the absence of original annotations by Desvaux explicitly linked to his Ononis arbuscula in the sheet at B and according to Art. 9.4 (a) of the Shenzhen Code (Turland et al. 2018), this specimen cannot be considered original material.</p><p>Fortunately, there is a herbarium sheet at P, with barcode P03447770 that bear original material of Desvaux of Ononis arbuscula . This sheet bears three stems, with leaves, flowers and fruits, and five labels. The stem mounted on the right half of the herbarium sheet is labeled as “On[onis] tridentara / Ononis tridentata / Cav. / en Hispania / prope Granatam. This specimen can be identified as belonging to O. tridentata subsp. crassifolia . The stem mounted on the center of the sheet is labeled as “ Ononis arbuscula Desv. ”. The stem mounted on the left half of the sheet is accompanied by two handwritten labels by Desvaux, annotated as: 1) Ononis tridentata L. Cav. ic. / 2. t. 152 / - arbuscula Desv. J. Bot. 1814 / 1 p. 76 / Mag. hort. t. 16 / hab. in Hispania ”, and 2) “Spec. nova / hab. Hisp.”. The sheet also bears a printed label “Herb. Mus. Paris / Herbier de A. N. Desvaux / Donné par Mme Vve Lavallée en 1896” (Fig. 5). Among these three stems, I designate as the lectotype of the name Ononis arbuscula the stem mounted on the left half of the sheet, which bears two original labels handwritten by Desvaux. This specimen shows important diagnostic features (leaflets blade lanceolate-cuneate, less than six times longer than wider, with (0–)3(–5) teeth in the upper half, inflorescence most times 1–2-flowered, axillary; indumenta only of short glandular hairs) and matches both the historical and current concepts of this taxon (until now named under O. tridentata subsp. tridentata) (see e.g., see Devesa &amp; López González 1997, Devesa 2000, 2009, López González 2001, Mota et al. 2011, Agulló et al. 2013).</p><p>Ononis tridentata s.l. includes several names validly published as nomenclatural and taxonomic synonyms, and the typification of the most relevant names is necessary for their correct application in this taxonomically difficult group. In this sense, the names O. barrelieri, O. crassifolia var. intricata, O. tridentata var. canescens, O. tridentata f. intermedia, and O. tridentata var. intermedia f. erecta should be treated as synonyms of O. tridentata subsp. arbuscula, comb. &amp; stat nov., and not as synonyms of O. tridentata Linnaeus s.str. due to the epitype of this name designated by F̂rther in 1997.</p></div>	https://treatment.plazi.org/id/E4294C700C3E5C5FAEEBC5D4FECC8433	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferrer-Gallego, P. Pablo	Ferrer-Gallego, P. Pablo (2023): Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae). Phytotaxa 598 (1): 32-58, DOI: 10.11646/phytotaxa.598.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.598.1.3
E4294C700C305C5DAEEBC6A5FEE68092.text	E4294C700C305C5DAEEBC6A5FEE68092.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ononis tridentata var. canescens Candolle 1825	<div><p>Ononis tridentata var. canescens Lamarck ex Candolle (1825: 161)</p><p>Ononis tridentata var. canescens was described by Candolle (1825: 161) with the accompanying diagnosis “ramis cano-tomentosis, foliis pallidis cuneiformibus quinquedentatis”, and followed by the provenance “in Arragoniâ” and the references “Asso fl. arr. n. 671. var. 2” from Asso (1779: 95 [2. Varietas]) and “Barr. icon. t. 416 [see 419]” from Barrelier (1714: ic. 419 [not 416]). The protologue also includes a reference to Lamarck’s Encyclopédie méthodique (1785: 508), as “ β. canescens (Lam! dict. I. p. 508)”. The indication in the protologue as “(v. s.)” must be treated as the acronym of “ vidit sicco ” (i.e., seen in a herbarium sheet).</p><p>Lamarck (1785: 508) mentioned as a variety into Ononis tridentata or his “Bugrane à trois dents”: “ β. Eadem? foliis cuneiformibus quinquedentatis, pedunculis unifloris. Flor. Aragon. p. 95. nº. 671. var. 2. Cicer montanum fruticans, incanum, angustifolium, serratum minus. Barrel. Ic. 419” and “La plante β forme pareillement un sous-arbrisseau trèsrameux &amp; haut d’un pied ou d’un pied &amp; demi; mais elle en diffère par plusieurs caractères assez remaquables. Ses tiges sont blanches &amp; beaucoup plus cotonneuses; ses feuilles sont plus courtes, cuneiforms, blanchâtres, &amp; la plupart à cinq petites dents à leur sommet. Les pédoncules sont courts, latéraux, plus souvent uniflores, &amp; soutiennent des fleurs purpurines dont le calice est moins velu. Ces plantes croissant en Espagne, &amp; nous ont été communiquées par M. l’Abbé Pourret (v. s.)”.</p><p>The reference of Barrelier (1714: t. 419) provided an illustration that can be considered original material used by Candolle to describe O. tridentata var. canescens . On the other hand, in the Lamarck’ herbarium at P, there is a sheet, with barcode P00296101, which bears four stems, with leaves, flowers, and fruits, and an original label handwritten by Pourret and Lamarck, annotated as “ Ononis tridentata / varietas pedunculis, floris / calycibus glabris / ad Balbastr. / in Arrag. / [handwritten by Pourret?] Barrel. ic. 419 [handwritten by Lamarck]” (image available at https://mediaphoto. mnhn.fr/media/1665502967319P49a40tO4W7dyGh7). This specimen matches the description published by Lamarck (1785: 508) and Candolle (1825: 161). In addition, there is a sheet in the Candolle herbarium at G-DC, with barcode G00477895 (Fig. 6), this specimen can be treated as a duplicate of the sheet at P (P00296101). The sheet preserved at G-DC bears two stems, with leaves and fruits, and two original labels handwritten by Candolle, annotated as “ Arragon ” and “ Ononis tridentate β. Lam.” (Fig. 6).</p><p>In conclusion, among the original elements, the illustration published by Barrelier (1714: ic. 419 [not 416 as was cited in the protologue]) and the specimen at G-DC, I designate as the lectotype of the name the specimen G00477895 .</p></div>	https://treatment.plazi.org/id/E4294C700C305C5DAEEBC6A5FEE68092	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferrer-Gallego, P. Pablo	Ferrer-Gallego, P. Pablo (2023): Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae). Phytotaxa 598 (1): 32-58, DOI: 10.11646/phytotaxa.598.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.598.1.3
E4294C700C325C5CAEEBC774FB078396.text	E4294C700C325C5CAEEBC774FB078396.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ononis crassifolia var. intricata Willkomm 1852	<div><p>Ononis crassifolia var. intricata Willkomm (1852: 114)</p><p>Willkomm (1852: 114) indicates in the protologue of Ononis crassifolia var. intricata: “β. intricata. Caules humiles, saepe prostrati inde a basi ramosissimi, ramulis intricatis.Foliola floresque dimidio minores, calycis laciniae angustiores. Hab. in argillosis salsuginosis Aragoniae inferioris (v. c. inter Magallón et Borja, Zaragoza et Muel copiose) atque in Castella nova cum varietate primaria et genuina. Flor. Julio.”.</p><p>I have found several specimens that are part of the original material cited in the protologue. The specimen with barcode COI00053668 bears two stems poorly preserved, and two labels, a label handwritten by Willkomm is annotated as “ Ononis tridentata L. / b. intermedia Lge. / β. intricata.”, and the second one is a printed label “Willkomm. it. hisp. secund. / 397. / Ononis crassifolia Duf. β. intricata Wk. / In gypsaceis Arragoniae inferioris copiose, v. c. / inter Magallon et Borja. / Julio 1850 ” (Fig. 7). There are duplicates of this specimen at BM and P, with barcodes BM000751116, P03447783 [stem mounted on the left half of the sheet], and P03447652, with the same printed label of the Willkomm herbarium. These three specimens are syntypes. On the other hand, the specimen with barcode COI00053669 bears two stems, with leaves and flowers, and a label annotated as “Joh. Lange, plantae ex Hispania 1851 -52. / Ononis crassifolia L. Duf. / β. intricata Willk. / In collibus gypsaicis ad Valladolid / 1 Jul.”. There are duplicates of this gathering preserved at JE and P, with barcodes JE00010313 and P03447749 (QR code PL01757118) .</p></div>	https://treatment.plazi.org/id/E4294C700C325C5CAEEBC774FB078396	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferrer-Gallego, P. Pablo	Ferrer-Gallego, P. Pablo (2023): Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae). Phytotaxa 598 (1): 32-58, DOI: 10.11646/phytotaxa.598.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.598.1.3
E4294C700C335C5CAEEBC209FAED855B.text	E4294C700C335C5CAEEBC209FAED855B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ononis barrelieri Dufour, BORD 1860	<div><p>Ononis barrelieri Dufour (1860: 324)</p><p>The protologue of the name Ononis barrelieri Dufour (1860: 324) includes three names “ O. tridentata var. 2ª Asso Syn. p. 95.” cited from Asso (1779: 95 [2. Varietas: foliis cuneiformibus, quinquedentatis, pedunculis unifloris. Hujus synonymum est / Cicer montanum, fruticans, incanum, angustifolium, serratum minus Barrel. Ic. 419. Provenit in collibus Casarauguste, circa Epila, Calatayud, Pozuelo, Hijar, Belchite, ubi gypsum adest]), “Cicer montanum fruticans incanum angustifolium serratum minus Barr. ic. 419 (et 299?)” cited from Barrelier (1714: ic. 419), and “C. latifolium, etc. (haud floridum) Barr. ic. 420” cited from Barrelier (1714: ic. 420), followed by a brief description in Latin. The protologue also includes some greographical provenances “Hab. in montibus aridis Navarrae (Tudela, Peralta), Aragoniae (Fuentes, Sanper, etc.), and a relevenat description in French “L’espèce décrite et figurée par Cavanilles (tab. 152) sous le nom d’O. tridentata L. diffère de celle-ci par ses feuilles linéaires, la plupart entières. – Stipule embrassant la moitié de la tige et adherente, s’avançant de chaque côté en un lobe lancéolé-aigu entier, par conséquent bifide. Le pétiole s’unit par une articulation si faible à la feuille que celle-ci se détache très facilement. Pédoncule tomenteux; calice velu, à dents aiguës; corolle de moyenne grandeur. / La figure 420 de Barrelier représente un individu qui n’est ni fleuri ni à même de fleuir. Les folioles y sont larges, arrondies. L’épithète d’ angustifolium (fig. 419) est évidemment comparative avec la fig. 420; elle se rapporte à un individu non-seulement en fleur, mais très avancé dans la fructification; et, malgré l’épithète, les folioles son ovalaires. J’ai rencontré ces modifications dans les mêmes montagnes, et Barrelier dit aussi les avoir trouvées ensemble”. The references of Barrelier (1714: t. 419 and 420) provided illustrations that can be considered original material used by Dufour to describe O. barrelieri, and are identifiable with the species to which this name is generally applied.</p><p>Jean-Marie Léon Dufour (1780-1865) was a French physician, botanist, and entomologist, born in Saint-Seversur-Adour, Landes (France). This author described a large number of species for the Spanish flora. The herbarium of this author is preserved at NTM, and further material is preserved at BORD, BR, CN, FI, K, LD, LY, P, P-JU (Stafleu 2000, Boone 2003). We have found a relevant original material of this species in the herbarium of Léon Dufour at BORD. The sheet Herbier Léon Dufour 5.1.061 (BORD) bears five stems, with leaves, flowers and fruits, and two labels: 1) “298. hisp. / Ononis Barrelieri. Duf. / O. tridentata var. Asso / Cicer montanum fruticans / incanum Barrel. ic. 420 / in montib. arid. / Peralta, / Fuentes”; 2) “ Ononis tridenta- / ta. Lin. / Benamaurel” (Fig. 8). Therefore, this material belonging to two different gatherings. The stem mounted on the lower left corner of the sheet can be identified as belonging to O. tridentata subsp. tridentata and could be linked to the label annotated as “Benamaurel” a locality of Granada province (Spain). The other four stems can also be identified as belonging to O. tridentata subsp. tridentata, and could be linked to the label annotated as “Peralta”, a locality cited in the protologue (Navarra province, Spain), and “Fuentes” another locality cited in the protologue and located in Aragón (Spain).</p><p>In conclusion, among the original elements mentioned, i.e., the illustrations of Barrelier (1714: t. 419 &amp; 420), and the specimens at BORD, I prefer to designate the specimen on the sheet Herbier Léon Dufour 5.1.061 (BORD) [except the stem mounted on the lower left corner of the sheet] as the lectotype of Ononis barrelieri. This specimen corresponds with the traditional concept and current use of the name Ononis tridentata subsp. tridentata .</p></div>	https://treatment.plazi.org/id/E4294C700C335C5CAEEBC209FAED855B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferrer-Gallego, P. Pablo	Ferrer-Gallego, P. Pablo (2023): Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae). Phytotaxa 598 (1): 32-58, DOI: 10.11646/phytotaxa.598.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.598.1.3
E4294C700C365C59AEEBC0BDFD14804A.text	E4294C700C365C59AEEBC0BDFD14804A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ononis tridentata fm. intermedia Lange 1865	<div><p>Ononis tridentata f. intermedia Lange (1865: 153)</p><p>The protologue of Ononis tridentata f. intermedia published by Lange (1865: 153) includes the epithet “2. intermedia ” followed by the synonyms “ O. crassifolia Willk. l. c. ex p., Barr. ic. 419; O. Barrelieri Duf. ex p.), a bief diagnosis “foliis cuneatis v. anguste obovatis, 3–5-dentatis”, and several gatherings “Hujus specimina vidi ex Hispania (Barnad. in herb. Vahl)! in collibus argillosis ad Murciam urbem (Guirao)! Beniajan, Fuensanta! inter Baza et Cullar de Baza (Willk.)! Barbastro Aragoniae (Vahl.)! Valladolid (β)!”. There is a syntype in the herbarium COI, with barcode COI00053670. The specimen, two stems with leaves and flowers, is accompanied by a handwritten label by Willkomm, annotated as “1205. Ononis tridentata L. = crassifolia Duf. / in collib. gypsaceis inter Baza et Cullar de Baza / 10-VII-1845 / Willkomm / forma videtus media inter O. tridentata et crassifolia / Duf.” (Fig. 9). This specimen is designated as the lectotype of the name Ononis tridentata f. intermedia .</p></div>	https://treatment.plazi.org/id/E4294C700C365C59AEEBC0BDFD14804A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferrer-Gallego, P. Pablo	Ferrer-Gallego, P. Pablo (2023): Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae). Phytotaxa 598 (1): 32-58, DOI: 10.11646/phytotaxa.598.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.598.1.3
E4294C700C365C59AEEBC26DFF2687CE.text	E4294C700C365C59AEEBC26DFF2687CE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ononis tridentata var. intermedia fm. erecta Willkomm	<div><p>Ononis tridentata var. intermedia f. erecta Willkomm</p><p>Willkomm (in Willkomm &amp; Lange 1877: 413) mentioned in the protologue of the name O. tridentata b [var.] intermedia α [f.] erecta “pedalis et ultra, ramis erectis, flexuosis simplicibus aut patule ramulosis, pedunculis brevibus, laciniis calyc. tubo duplo longioribus, vexilo is triente longiore ( O. tridentata Coss. ap. Bourg. pl. exs. n. 1122! Cicer sylv. fruticans incanum angustifolium serratum fl. purp. Hispan. Barr. Ic. 299!)” and “in Catal. c. Cardona, Manresa, Vich, Csta.! Aragon. australi en el Barranco del Maduro, Pardo! in regno Murc. pr. Beniajam, Guir.! Lge.! Granat. pr. Vera, Bourg.! inter Baza et Cullar de Baza. Wk.”. The specimen with barcode COI00053676 match with the gathering cited in the protologue as “Catal. c. Cardona, Manresa, Vich, Csta.!”. This material is a branch, with leaves and flowers, accompanied by a handwritten label, annotated as: “193 / A. [Antonio] C. [Cipriano] Costa / Ononis tridentata L. / Cardona, Manresa. Vich. / Junio – Legit.” 193 / A. [Antonio] C. [Cipriano] Costa / Ononis tridentata L. / Cardona, Manresa. Vich. / June – Legit.” (Fig. 10). There is at P, with barcode P03447663, another specimem collected in 1856 by Costa in “Cardona, Vich” which can probably be treated as a duplicate of the specimen at COI.</p><p>On the other hand, there are three sheets at P, with barcodes P03447772, P03447659 and P03447662 that contain specimens belonging to the gathering mentioned in the protologue as “Granat. pr. Vera, Bourg.!”. These three specimens are accompanied by a printed label “ E. Bourgeau, P. d’Espagne, 1851 / 1122. Ononis tridentata, L. / (Coss.) / Bords de la rivière à Vera (province de Grenade). / 15 Avril.” .</p><p>Finally, there is a specimen at COI, barcode COI00053672 which is part of the gathering cited in the protologue as “in regno Murc. pr. Beniajam, Guir.!”. This material, two stems, with leaves and flowers, is accompanied by a label handwritten by Willkomm “ Ononis tridentata L. / b. intermedia Lge. / α. erecta .” and an original label of A. Guirao, annotated as “Herbarium A. Guirao / Ononis crassifolia Duf. ined. / O. tridentata β. canescens / Ipse legi sponte cresc. in collibus argi- / lloso-gypsaceis ad Beniajan pr. / Murcia et alibi. / Leg. Maj. 1851”. There is a duplicate of this specimen at P (barcode P03447782). These all specimens are syntypes of O. tridentata b [var.] intermedia α [f.] erecta .</p></div>	https://treatment.plazi.org/id/E4294C700C365C59AEEBC26DFF2687CE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferrer-Gallego, P. Pablo	Ferrer-Gallego, P. Pablo (2023): Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae). Phytotaxa 598 (1): 32-58, DOI: 10.11646/phytotaxa.598.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.598.1.3
E4294C700C365C46AEEBC5F0FCEA83BF.text	E4294C700C365C46AEEBC5F0FCEA83BF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ononis tridentata subsp. arbuscula (Desvaux) P. P. Ferrer	<div><p>Ononis tridentata subsp. arbuscula (Desvaux) P.P. Ferrer, comb. &amp; stat. nov.</p><p>≡ O. arbuscula Desvaux (1814: 76)</p><p>Lectotype (designated here):—[SPAIN] “Habitat in Hispania”, s.d., A.N. Desvaux 25, P (barcode P03447770 [the lectotype is the stem mounted on the left half of the sheet, accompanied by two labels handwritten by Desvaux]) (Fig. 5).</p><p>= O. tridentata β [var.] canescens Lamarck ex Candolle (1825: 161)</p><p>Lectotype (designated here):—[SPAIN], “ Arragon ” [ Aragón], s.d., s.n., G (barcode G00477895!) (Fig. 6). Isolectotype: P (barcode P00296101!)</p><p>= O. crassifolia β [var.] intricata Willkomm (1852: 114)</p><p>Lectotype (designated here):—[SPAIN], Zaragoza, “Arragoniae inferioris copiose inter Magallon et Borja”, Julio 1850, H.M. Willkomm 397, COI (barcode COI00053668!) (Fig. 7). Isolectotypes: BM (barcode BM000751116!), P (barcodes P03447783 [stem mounted on the left half of the sheet]!, P03447652!).</p><p>= O. barrelieri Dufour (1860: 324)</p><p>Lectotype (designated here):—[SPAIN], Peralta ( Navarra), “ Fuentes ” ( Aragón), s.d., L. Dufour 298, BORD (Herbier Léon Dufour 5.1.061!) [except the stem mounted on the lower left corner of the sheet] (Fig. 8).</p><p>= O. tridentata f. intermedia Lange (1865: 153)</p><p>≡ O. tridentata var. intermedia (Lange) Nyman (1889: 86)</p><p>Lectotype (designated here):—[SPAIN], Granada, “inter Baza et Cullar de Baza, 10 Jul 1845, H.M. Willkomm 1205, COI (barcode COI00053670!) (Fig. 9).</p><p>= O. tridentata var. intermedia f. erecta Willkomm (in Willkomm &amp; Lange 1877: 413)</p><p>Lectotype (designated here):—[SPAIN], Cataluña, “Catal. c. Cardona, Manresa,Vich, June, A.C. Costa 193, COI (barcode COI00053676!) (Fig. 10). Probable isolectotype: Jul-Aug. 1856, A.C. Costa s.n., P (barcode P03447663!).</p><p>= O. rentonarensis M.B. Crespo &amp; Serra (1993: 38)</p><p>Holotype:—[SPAIN], Alicante, La Torre de les Maçanes, Sierra del Rentonar, 1 Oct 1989, L. Serra s.n., VAL (number 142414!; ex VAB 90/0452). Isotypes: VAL 23831!, VAL 118502! (ex VF 18627, ex VAB 90/0456), BM000751131!, ABH3256! M0274811!, G00370381!, MA517204!, BC, MUB, JACA, LSH0275, RNG</p></div>	https://treatment.plazi.org/id/E4294C700C365C46AEEBC5F0FCEA83BF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferrer-Gallego, P. Pablo	Ferrer-Gallego, P. Pablo (2023): Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae). Phytotaxa 598 (1): 32-58, DOI: 10.11646/phytotaxa.598.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.598.1.3
E4294C700C295C44AEEBC221FF0B82B7.text	E4294C700C295C44AEEBC221FF0B82B7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ononis tridentata subsp. angustifolia (Lange) Devesa & G. Lopez 1997	<div><p>Ononis tridentata subsp. angustifolia (Lange) Devesa &amp; G. López (1997: 258)</p><p>The protologue of the name Ononis tridentata f. angustifolia included a brief diagnosis “foliis lineari lanceolatis v. lineari-cuneatis, integerrimis v. 2-5-dentatis” preceded by the reference to two works: “ O. tridentata Cav. ic. tab. 152, Barr. ic. 300; Willk. Strand- und Steppengew. p. 115” (Lange 1865: 152).</p><p>The references cited in the protologue (Barrelier 1714: ic. 300, Cavanilles 1793: Tab. 152) included illustrations of this species that can be considered as original elements used by Lange to describe Ononis tridentata f. angustifolia . The Cavanilles’s drawing illustrates a stem with leaves, flowers, and fruits, and several details of a flower, fruit, and seed (figures a [Calix], b [Petala expansa, ubi 1 vexillum, 2 alae, 3 carina], c [Genitalia], d [Legumen], and e [Semen]) (Cavanilles 1793: 42, Tab. 152). The Barrelier’s drawing “Cicer sylvestre, fruticans, angustifolium, non serratum, Hispanicum” illustrates a stem with flowers and fruits, and several details of the leaves, a separate flower and a separate fruit (Barrelier 1714: ic. 300). These two illustrations certainly matche with the current concept of the name and could be chosen as lectotype.</p><p>Cavanilles (1793: 42) cites the following provenances as localities of Ononis tridentata “Habitat copiose prope Llanera in montibus vulgo Port de Carcer: crescit etiam inter Domeño et Chelva; Xixona et Tibi; Ialance et Cofrentes; Eldam et Noveldam”. In the Cavanilles herbarium at MA there is a herbarium sheet, with code MA 61824, that bears material of this species collected by Cavanilles himself in “prope Eldam” (Alicante province) in May 1791. On the other hand, there is a specimen collected by Cavanilles and preserved at BM, with barcode BM001122985. This material is a stem with leaves and flower, and is annotated “ Ononis tridentata L. Cav. ! Ic.” handwritten by Cavanilles, and “Cavanilles, 1803” handwritten by J.E. Smith. This material (specimens at MA and BM) could have been used by Cavanilles as voucher to illustrate the species in his drawing number 152 (see Cavanilles 1793), but it cannot be treated as original material of O. tridentata subsp. angustifolia .</p><p>In the protologue are cited several gatherings, as: “Specimina hujus formae vidi ex Aragonia: 2 mill. a Zaragoza (herb. Vahl)! Barranco del maduro pr. Castellote (Pardo)! Serrania de Cuenca inter Mira et Villora (Willk. Nr. 530)! Valenc. inter Alicante et Alcoy (Boiss. et Reut.)! Sierra Castalla supra Alicante (Leresche)! Rio de Aguas (prov. Almeria) 1 Decbr. 1852 c. fl. et fr.!” (Lange 1865: 152).</p><p>Devesa &amp; López González (1997: 258) indicate that they have not seen original material for this name. On the other hand, Bolòs &amp; Vigo (1984: 531) mentioned that they could not see the type of Lange’s name, and many of the localities cited by Lange in the protologue correspond to the distribution area of O. tridentata var. tridentata . In addition, Bolòs &amp; Vigo (1984: 531) recognize “ var. edentula Webb in Bourg.” within O. tridentata and include the geographic provenance “Xàtiva” (taken from the locality indicated in the exsiccatum of Bourgeau “S. Felipe de Xativa”) as the classic locality. However, the Philip Barker Webb’s name ( Ononis tridentata α [var.] edentula Webb ex Willk. in Willk. &amp; Lange) was not effectively published in Bourgeau’s exsiccatum but in Willkomm’s publication (in Willkomm &amp; Lange 1877: 413) (see below).</p><p>Among the gathrings mentioned by Lange in the protologe. I have located several relevant specimens that can be treated as syntypes preserved in the Martin Vahl herbarium at C and Willkomm herbarium at COI. I found a specimen at C that has belonged to Vahl’s herbarium and partly fits to the description included in the protologue. This specimen, without code, belongs to the gathering cited by Lange in the protologue as “Specimina hujus formae vidi ex Aragonia: 2 mill. a Zaragoza (herb. Vahl)!”, and bears a stem with leaves, and is annotated (on back of the sheet): “ Ononis / tridentata / varietas glandulosa” and “Legi copiose in Arragonia / mens, Aug: 1787. / 2 mill. à Sarragoza” (Fig. 11). This specimen is undoubtedly original material of the name Ononis tridentata f. angustifolia, and is not in direct conflict with the protologue. However, the specimen is in direct conflict with the current usage of the name and can be ascribed to Ononis tridentata subsp. tridentata (with leaflets blade obovate-cuneate, less than six times longer than wider, with 3–9 teeth in the upper half). A lectotypification of O. tridentata f. angustifolia using this specimen would be technically sound, but it would be undesirable in terms of nomenclatural stability since it does not represent the current concept and use of the name O. tridentata subsp. angustifolia .</p><p>The specimen with barcode COI00053677 belonging to the collection mentioned in the protologue as “ Aragonia […] Barranco del maduro pr. Castellote (Pardo)!”. This specimen consists of three stems, with leaves and flowers, accompanied by an original handwritten label from the Willkomm herbarium, annotated as: “Herbarium Willkommii. / Ononis tridentata L. / Abundat in Aragoniae australis collibus aridis. / Legit. Loscos Salvador Pardo. / 18”. On the other hand, the specimen with barcode COI00053673 are three stems, with leaves, flowers, and fruits. This material was collected in Cuenca, between Mira and Villora, it is well-preserved and is accompanied by an original label from the Willkomm herbarium, annotated as: “Willkomm. it. hisp. secund. / 530. / Ononis tridentata L. / angustifolia Lge. / Serrania de Cuenca in argillosis pinetorum inter / Mira et Villora. / Aug. 1850. / α β edentula Wbb.” (Fig. 12). There is a duplicate specimen of this material preserved at P, with barcode P03447781 [the three stems are mounted on the right half of the herbarium sheet]. This material is accompanied by the same printed label that the sheet at COI.</p><p>On the same sheet at P, with barcode P03447781 (QR-code PL01757605), is mounted (on the left half of the sheet) a specimen (two stems) that can be treated as a syntype mentioned by Lange in the protologue as “Valenc. inter Alicante et Alcoy (Boiss. et Reut.)!”. This material is accompanied by a label annotated as “ Ononis tridentata Cav. / Montes Regni Valentini. / Boissier et Reuter. in collibus gypsosis / inter Alicante et Alcoi. / Julio 1858.”.</p><p>In all cases the material cited above matches with the traditional concept and current use of the name O. tridentata subsp. angustifolia (see e.g., Devesa &amp; López González 1997, Devesa 2000, López González 2001). In conclusion, I designate as the lectotype of the name the specimen COI00053673. This specimen is a well preserved and very complete material, and show relevant diagnostic characters (e.g., leaflets linear, more than six times longer than wider, glabrous).</p></div>	https://treatment.plazi.org/id/E4294C700C295C44AEEBC221FF0B82B7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferrer-Gallego, P. Pablo	Ferrer-Gallego, P. Pablo (2023): Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae). Phytotaxa 598 (1): 32-58, DOI: 10.11646/phytotaxa.598.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.598.1.3
E4294C700C2C5C43AEEBC3A9FF11870B.text	E4294C700C2C5C43AEEBC3A9FF11870B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ononis tridentata var. dentata Willkomm 1877	<div><p>Ononis tridentata var. dentata Willkomm (1877: 413)</p><p>The protologue of Ononis tridentata var. dentata Willkomm (1877: 413) includes a brief diagnosis “foliis lineariscuneatis, apice 2-3-5-dentatis” followed by two synonyms “( O. tridentata L. ex descr., Cav. Ic. II, t. 152! Anonis Hispanica frutescens, folio tridentato carnoso Tourn. Inst. 408!)”. The protologue also includes several provenances and gatherings “(omnium formarum rarissima!) pr. urbem Murcia, Guir.! in regno Valentino. inter Villena, Elda, Novelda et alibi, Cav.! in Baet pr. Puerto de Sta. Maria, Clem.! in hb. Bout.”.</p><p>Among the original elements used by Willkomm to describe his Ononis tridentata var. dentata, the reference of Cavanilles (1793: t. 152) provided an illustration that can be considered an original element. On the other hand, there is a relevant specimen at COI that bears original material of this name. The specimen, with barcode COI00053679 is labeled “Herbarium Willkommii. / Ononis tridentata L. / a. angustifolia Lge. α. genuine / β. dentata / ( Ononis tridentata L. / Cav. Ic. T. 152!) / Murcia. /Lefit. A. Guiraro / 18” (Fig. 14). This material, four fragments plant with leaves, flowers and fruits, is complete and well-preserved, and it is designate as the lectotype of the name Ononis tridentata var. dentata .</p></div>	https://treatment.plazi.org/id/E4294C700C2C5C43AEEBC3A9FF11870B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferrer-Gallego, P. Pablo	Ferrer-Gallego, P. Pablo (2023): Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae). Phytotaxa 598 (1): 32-58, DOI: 10.11646/phytotaxa.598.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.598.1.3
E4294C700C2C5C43AEEBC0BDFF04810F.text	E4294C700C2C5C43AEEBC0BDFF04810F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ononis tridentata var. integrata Amo, COI 1873	<div><p>Ononis tridentata var. integrata Amo (1873: 447)</p><p>Ononis tridentata var. integrata was described by Amo (1873: 447) with the accompanying diagnosis “Folia saepiùs integerrima, rarò bi-tridentata”, and followed by a complete description in Spanish and the provenance “Hab. en la Alcarria, Serranía de Cuenca y reino de Valencia (Quer, Pal., Cav.), entre Jaen y Campillo de Arenas (Wk.)”.</p><p>The herbarium of Amo y Mora is preserved at GDA (Vizoso &amp; Quesada 2018). This collection contains the material collected by himself and the contributions from Spanish botanists as well as foreign collaborators, such as Guirao, Cutanda, Willkomm, Lange, and Reuter (Negrillo Galindo &amp; Marín Calderón 1985 a, 1985b). In this herbarium there is a folder that contains two specimens of Ononis tridentata . On the folder is annotated “1730 / Ononis tridentata ”. A sheet (without number or barcode) bears a specimen of this species, and two labels: “31. Ononis tridentata L. / Murcia”, and “Instituto de Granada / Gabinete de Historia Natural / Ononis / tridentata / Lin, / N.º 1730”. A second one sheet (without number or barcode) bears a specimen of this species and a label annotated as “31. Ononis tridentata . Pert. / Murcia”. Unfortunately, due to the absence of original annotations by Amo y Mora explicitly linked to Ononis tridentata var. integrata or to the protologue of the name in these two sheets at GDA, these specimens cannot be considered original material.</p><p>On the other hand, there is a revelant specimen in the Willkomm herbarium at COI. The specimen barcoded COI00053681 was collected by Willkomm in “Inter Campillos de Arenas et urbem Jaen” in 1848 (Fig. 13). This material match with the gathering cited by Amo in the protologue, as “entre Jaen y Campillo de Arenas (Wk.)”, and therefore it can be treated as a syntype. The specimen (two stems with leaves, flowers, and fruits) is well-preserved and matches the protologue of Ononis tridentata var. integrata . Therefore, this specimen is designated as the lectotype of the name.</p></div>	https://treatment.plazi.org/id/E4294C700C2C5C43AEEBC0BDFF04810F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferrer-Gallego, P. Pablo	Ferrer-Gallego, P. Pablo (2023): Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae). Phytotaxa 598 (1): 32-58, DOI: 10.11646/phytotaxa.598.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.598.1.3
E4294C700C2C5C40AEEBC5A4FF23804B.text	E4294C700C2C5C40AEEBC5A4FF23804B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ononis tridentata var. edentula Webb ex Willkomm, COI 1877	<div><p>Ononis tridentata var. edentula Webb ex Willkomm (1877: 413)</p><p>Willkomm’s protologue of Ononis tridentata α [var.] edentula includes a brief diagnosis “foliis lineari-lanceolatis linearibusve canaliculatis, interdum subteretibus, integerrimis mucronatis”, preceded by the epithet “α. edentula Wbb.” and followed by these two names and indications: “Cicer silv. fruticans, angustifolium non serratum, Hispanicum Barr. Ic. 300! O. tridentata Willk. Sert. p. 41 et pl. exs. 1850, n. 530, Bourg. pl. exs. n. 1724!. In the protologue also was mentioned several gatherings: “[…] in Catalaun. int. Hortalets et Tost atque in Conca de Tremp, CSTA.!, in regno Valent. pr. Játiva, BOURG.! in Serranía de Cuenca int. Mira et Villora, WK., int. urbem Jaen et Campillos de Arenas, WK. […]” (see Willkomm &amp; Lange 1877: 413).</p><p>I have found several syntypes at COI and P herbaria. Willkomm cited in the protologue material collected in Cuenca: “in Serranía de Cuenca int. Mira et Villora, WK”. A specimen belonging to this gathering is kept at COI, with barcode COI00053673. This material has been selected as the lectotype of the name O. tridentata f. angustifolia (see above) and is also a syntype of the name O. tridentata var. edentula .</p><p>On the other hand, there is a specimen at COI, with barcode COI00053680 that is part of the Bourgeau’s exsiccatum cited in the protologue as “Bourg. pl. exs. n. 1724” and also as “in regno Valent. pr. Játiva, BOURG.!”. This specimen are three well-preserved stems, with leaves and flowers, and an original printed label of the Bourgeau’s exsiccatum, annotated as: “E. Bourgeau, Pl. d’Espagne, 1852 / 1724. Ononis tridentata, Ass. / ver. edentula Webb. / (Webb.) / Collines arides près S. Felipe de Xativa. / 17 Juin.” (Fig. 13). In the herbarium P there are several duplicates of Bourgeau 1724, with barcodes P03600614, P03447655, P03447761, P03447763, P03447771. These specimens are syntypes.</p><p>In the same sheet that the specimen COI00053680 is preserved the specimen barcoded COI00053681. This specimen consists of two stems, with leaves and flowers, and is accompanied by an original handwritten label by Willkomm (Fig. 13). The material was collected on August 22, 1845 from the locality cited in the protologue as “int. urbem Jaen et Campillos de Arenas, WK.”, this material is therefore also a syntype.</p><p>In conclusion, among the original material, I designate as the lectotype of Ononis tridentata var. edentula the specimen COI00053680 from the Bourgeau’s exsiccatum. The lectotype selected is a complete and well-preserved specimen that shows relevant diagnostic features (e.g., leaflets blade linear to linear-ensiform, more than six times longer than wider, glabrous, with 0–1(2) apical teeth), and it clearly represents the traditional concept of this name (Willkomm &amp; Lange 1877) and reflects the current application of Ononis tridentata var. edentula as a synonym of Ononis tridentata subsp. angustifolia (see e.g., Devesa &amp; López González 1997, Devesa 2000, López González 2001).</p></div>	https://treatment.plazi.org/id/E4294C700C2C5C40AEEBC5A4FF23804B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferrer-Gallego, P. Pablo	Ferrer-Gallego, P. Pablo (2023): Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae). Phytotaxa 598 (1): 32-58, DOI: 10.11646/phytotaxa.598.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.598.1.3
E4294C700C2F5C40AEEBC26CFB3D86EF.text	E4294C700C2F5C40AEEBC26CFB3D86EF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ononis tridentata subsp. angustifolia (Lange) Devesa & G. Lopez 1997	<div><p>Ononis tridentata subsp. angustifolia (Lange) Devesa &amp; G. López (1997: 258)]</p><p>≡ O. tridentata f. angustifolia Lange (1865: 152) [basionym]</p><p>≡ O. tridentata var. angustifolia (Lange) Nyman (1889: 86)</p><p>Lectotype (designated here):—[SPAIN], Cuenca, Serrania de Cuenca in argillosis pinetorum inter Mira et Villora, August 1850, H.M. Willkomm 530, COI (barcode COI00053673!) (Fig. 12). Isolectotype: P (barcode P03447781! [the three stems are mounted on the right half of the herbarium sheet]).</p><p>= O. tridentata var. integrata Amo (1873: 447)</p><p>Lectotype (designated here):—[SPAIN], “Inter Campillos de Arenas et urbem Jaen”, 22 Aug 1848, H.M. Willkomm 1205, COI (barcode COI00053681) (Fig. 13)</p><p>= O. tridentata β [var.] dentata Willkomm in Willkomm &amp; Lange (1877: 413)</p><p>Lectotype (designated here):—[SPAIN], Murcia, s.d., Guirao s.n., COI (barcode COI00053679!) (Fig. 14).</p><p>= O. tridentata α [var.] edentula Webb ex Willkomm in Willkomm &amp; Lange (1877: 413)</p><p>Lectotype (designated here):—[SPAIN], Valencia, “Collines arides près S. Felipe de Xativa”, 17 Jun 1852, E. Bourgeau 1724, COI (barcode COI00053680!) (Fig. 13). Isolectotypes: P (barcodes P03600614!, P03447655!, P03447761!, P03447763!, P03447771!).</p><p>= O. tridentata subvar. linearifolia Font Quer ex Bolòs &amp; Vigo (1974: 72)</p><p>Holotype:—[SPAIN], Lleida, La Pobla de Segur, ad 550 m, 6 Jun 1918, P. Font i Quer s.n., BC (number 87936!).</p></div>	https://treatment.plazi.org/id/E4294C700C2F5C40AEEBC26CFB3D86EF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ferrer-Gallego, P. Pablo	Ferrer-Gallego, P. Pablo (2023): Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae). Phytotaxa 598 (1): 32-58, DOI: 10.11646/phytotaxa.598.1.3, URL: http://dx.doi.org/10.11646/phytotaxa.598.1.3
