taxonID	type	description	language	source
E43B726B0471FF9F224C59F2FCB6754C.taxon	description	(Figs 1 – 5)	en	Short, Andrew E. Z., Mcintosh Iv, Charles E. (2015): Hydrophilus harpe sp. nov., a remarkable new species of giant water scavenger beetle from Brazil (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 55 (2): 665-671, DOI: 10.5281/zenodo.5372500
E43B726B0471FF9F224C59F2FCB6754C.taxon	materials_examined	Type material. HOLOTYPE: J, BRAZIL: RIO GRANDE DO NORTE: “ BRAZIL, R. G. N. / Ceara-Mirim / VII- 6 & 7 - 1969 / P. & P. Spangler ”, “ USNMENT / [Matrix Barcode] / 00776385 ” (USNM). PARATYPES (3 spec.): BRAZIL: BAHÍA: 15 km E. Itabuna, 3. vii. 1969, leg. P. & P. Spangler, USNMENT 00776384 (1 ♀, USNM). MINAS GERAIS: Aguas Vermelhas, xii. 1983, leg. M. Alvarenga (1 J, CMNH). RIO GRANDE DO NORTE: Same data as holotype, USNMENT 00776386 (1 J, SEMC).	en	Short, Andrew E. Z., Mcintosh Iv, Charles E. (2015): Hydrophilus harpe sp. nov., a remarkable new species of giant water scavenger beetle from Brazil (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 55 (2): 665-671, DOI: 10.5281/zenodo.5372500
E43B726B0471FF9F224C59F2FCB6754C.taxon	description	Description. Size and form. Body length = 42 – 47 mm. Body broadly oval, strongly convex. Color. Dorsum of head, pronotum, and elytra charcoal black, usually with a metallic green iridescent sheen which is most easily visible under direct light. Ventral surface of head, thorax, and abdomen uniformly black; protarsi slightly paler. Antennomeres 1 – 6 pale yellow, antennomeres 7 – 9 brown. Maxillary palps reddish-brown. Head. Antennae made up of nine articles, including a three-segmented pubescent club. Length of scape subequal to cardo, pedicel and following three segments combined subequal to length of scape. Cupule large, ca. two-thirds length of preceding six antennomeres combined or ten times length of preceding antennomeres. Maxillary palps with four palpomeres; in male, second palpomere ca. five times as long as first palpomere, third palpomere ca. half as long as second palpomere, fourth palpomere ca. half as long as third palpomere; third palpomere greatly inflated, ca. three times as large as the apical palpomere. Maxillary palps unmodified in female. Combined length of palps subequal to width of head. Labial palps with three palpomeres; in male, second palpomere ca. five times as long as palpomere one and ca. two times as long as apical palpomere; second palpomere very swollen, ca. three times as wide as apical palpomere; lateral outer margin with dense row of long setae; inner lateral margin with a small patch of setae at apex; in female, second segment only slightly wider than apical palpomere. Combined length of labial palps subequal to combined length of the first two maxillary palpomeres. Mentum triangular; glabrous except for a few short, fine setae along lateral margins. Thorax. Prosternum divided into two lobes which are conical and tapered to a point with tufts of setae on the anterior margin. The sternal keel formed by the fusion of the meso- and metaventrite extending slightly past posterior margin of ventrite 2. Anterior tenth of keel with a small depression lined with transverse grooves. Keel glabrous except for few, short setae along anterior margin of depression. Apical protibial spurs of male dimorphic, one rounded and shield-like, the other slightly curved and pointed at apex (Fig. 3). Single row of long setae on inner margin of protibiae. Male protasal claws asymmetrical, with outer claw extremely long and recurved, the apex explanate and forming a sickle-shaped hook (Fig. 3) and with inner claw shorter and not expanded or flattened apically. In males, ventral face of meso- and metatarsi set with rows of thick, dense setae appearing as a brush; in females, they are set with only 2 – 3 regular rows. Abdomen with five exposed ventrites; ventrites 2 – 5 subequal in length. Ventrites densely covered with uniform short pubescence; ventrites 3 – 5 with a medial glabrous patch of varying size: glabrous patches on ventrites 4 and 5 subequal in length, glabrous patch on ventrite 3 ca. one-third length of other patches. Glabrous patches on ventrites 3 and 4 subequal in width, ca. one-fourth width of patch on ventrite 5. Posterior margin of ventrites 3 and 4 slightly overhangs onto following segment; overhang of fourth segment margin far more pronounced. Glabrous patch on ventrite 5 creased medially due to elevated ridge down abdominal midline. Aedeagus as in Fig. 4.	en	Short, Andrew E. Z., Mcintosh Iv, Charles E. (2015): Hydrophilus harpe sp. nov., a remarkable new species of giant water scavenger beetle from Brazil (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 55 (2): 665-671, DOI: 10.5281/zenodo.5372500
E43B726B0475FF9A22205AA3FE7A7293.taxon	materials_examined	Type material examined. LECTOTYPE: J (here designated): “ MUSEUM PARIS / Rosario / Claine 1898 ” (MNHN: Régimbart Collection). PARALECTOTYPE: Same data as Lectotype (1 ♀, MNHN: Régimbart Collection). Additional material examined: (2 spec.): “ Rosario / de S. Fe ”, “ Dr. Stem. / pelmann. ”, “ 37. ” (1 J, MNHN: Régimbart Collection), “ Republ. / Argentina ” (1 ♀, MNHN: Régimbart Collection),	en	Short, Andrew E. Z., Mcintosh Iv, Charles E. (2015): Hydrophilus harpe sp. nov., a remarkable new species of giant water scavenger beetle from Brazil (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 55 (2): 665-671, DOI: 10.5281/zenodo.5372500
E43B726B0475FF9A22205AA3FE7A7293.taxon	diagnosis	Differential diagnosis. See Differential diagnosis for H. harpe.	en	Short, Andrew E. Z., Mcintosh Iv, Charles E. (2015): Hydrophilus harpe sp. nov., a remarkable new species of giant water scavenger beetle from Brazil (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 55 (2): 665-671, DOI: 10.5281/zenodo.5372500
E43B726B0475FF9A22205AA3FE7A7293.taxon	distribution	Distribution. Argentina, Uruguay (HANSEN 1999).	en	Short, Andrew E. Z., Mcintosh Iv, Charles E. (2015): Hydrophilus harpe sp. nov., a remarkable new species of giant water scavenger beetle from Brazil (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 55 (2): 665-671, DOI: 10.5281/zenodo.5372500
E43B726B0475FF9A22205AA3FE7A7293.taxon	discussion	Remarks. Of the specimens of H. masculinus he examined, RÉGIMBART (1901) remarked as follows: ‘ République Argentine: Rosario (Claine, 1898), collection du Muséum de Paris et la mienne’. It is clear from the description he saw both males and females though he does not indicate the number of specimens he examined. In the collection of Régimbart, which is maintained separately at MNHN, the Hydrophilus specimens are arranged as he had them in the same order as presented in the 1901 publication. There are four specimens situated Figs 6 – 9. Hydrophilus (D.) masculinus (Régimbart, 1901), male lectotype: 6 – mesotarsus, 7 – metatarsus, 8 – protarsus, 9 – label. Arrows in Figs 6 – 7 note the dense tuft of ventrally-facing setae. by the name ‘ masculinus ’, including a male and female pair with identical information to those listed above. The male is here designated as the lectotype for H. masculinus to fix the identity of this species. The other two specimens (also from Argentina) do not bear Rosario labels and it is unclear if they were part of the original type series or not. Consequently, we have opted not to designate these two specimens as paralectotypes as it is unclear if they were part of the original type series.	en	Short, Andrew E. Z., Mcintosh Iv, Charles E. (2015): Hydrophilus harpe sp. nov., a remarkable new species of giant water scavenger beetle from Brazil (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 55 (2): 665-671, DOI: 10.5281/zenodo.5372500
