taxonID	type	description	language	source
EA0E87CBFF9D0B05FE81FD3E31680B18.taxon	description	(Fig. 1)	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF9D0B05FE81FD3E31680B18.taxon	materials_examined	Type material. HOLOTYPE:, ‘ CHINA: SW China: Guizhou, Luodian County (ƤÑH), Ban’geng Town (ṞOiş), Ban’geng Village (ṞOi IJ), Guanyin Cave (ḴṘḌ), N 25 ° 31 ′ 6.84 ′′, E 106 ° 38 ′ 54.68 ′′, ca. 540 m, 8. x. 2019, Yun-Chun Li leg. ’ (SNUC). PARATYPE: CHINA: 1 ♀, same label data as holotype (SNUC).	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF9D0B05FE81FD3E31680B18.taxon	diagnosis	Diagnosis. Body length approximately 2.0 mm; with long, sparse pubescence; frons broad, with complete subantennal transverse sulcus; vertexal foveae distinct; male antennomere 10 protuberant at anteromesal corner, antennomere 11 with wide impressed area on mesal surface; median gular carina broad and elevated, with two small gular foveae close to each other; metaventrite with paired elongate lateral carinae in male; legs elongate and simple in both sexes; tergite IV with broad basal sulcus which is densely setose at middle, long discal carinae widely separated and sub-paralleled; aedeagus symmetrical, with relatively stout median lobe, parameres distinctly exceeding apex of median lobe.	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF9D0B05FE81FD3E31680B18.taxon	description	Description. Male (Fig. 1 A). Body reddish-brown, length from anterior margin of clypeus to apex of abdomen 2.02 mm. Head rectangular, length from anterior margin of clypeus to head base 0.44 mm, equals to maximum width across eyes; vertex finely punctate, sparsely pubescent, moderately convex, with pair of small vertexal foveae; antennal tubercles raised; frons flattened posteriorly and raised anteriorly at middle, merging with antennal tubercles to form transverse bridge; subantennal transverse sulcus complete; lacking vertexal sulcus connecting foveae; each eye composed of about 25 facets; antennae relatively elongate, length 0.95 mm, with distinct club (Fig. 1 B) formed by apical two antennomeres; antennomere 1 large, approximately 1.5 times as wide as 2, antennomere 2 cylindrical, antennomeres 3 and 4 successively shorter, narrowed at bases and broadening apically, antennomeres 5 – 9 short and transverse, antennomere 10 approximately twice as wide as antennomere 9, asymmetrical, with its anteromesal corner distinctly protuberant, antennomere 11 largest, curved mesally, with its mesal surface broadly impressed; median gular carina broad and elevated, with two small nude gular foveae close to each other. Pronotum almost glabrous, with sparse pubescence similar to those of head; length along midline 0.43 mm, maximum width 0.44 mm; with nude median and lateral antebasal foveae. Elytra wider than long, length along suture 0.69 mm, maximum width 0.53 mm, dorsal surface with sparse, long setae; each elytron with two punctiform basal foveae in shallow impressions; sutural stria complete, lacking discal longitudinal sulcus. Legs simple, without protuberances. Abdomen longer than wide, length along midline 0.73 mm, maximum width 0.62 mm; slightly constricted near base; tergite IV (first visible tergite) with broad basal impression, densely setose at middle; discal carinae long, widely separated, almost parallel to each other, about 0.4 times of total tergal length; marginal carinae complete and oblique. Length of aedeagus (Fig. 1 C, 1 D) 0.29 mm (including parameres), median lobe relatively stout, with large, oval diaphragm, parameres distinctly exceeding apex of median lobe, each broadened preapically, narrowed at apex, and with two preapical setae; endophallus comprises two elongate sclerites at middle, and numerous weakly sclerotized spine-like structures anteriorly. Female. Similar to male in general appearance; antennomere 10 lacking protuberance, antennomere 11 similarly asymmetrical as male, but mesal surface lacking impression; metaventrite lacking lateral carinae; each eye composed of about 12 facets. Measurements (as for male): body length 2.0 mm; length of head 0.44 mm, equals to width; length of pronotum 0.44 mm; maximum width 0.43 mm; length of elytra 0.50 mm, maximum width 0.67 mm; length of abdomen 0.62 mm, maximum width 0.54 mm. Comparative notes. This is a very distinct species which can be easily separated from all congeners from East Asia by the unique form of the sexually modified antennal club and laterally carinate metaventrite of the male, the simple legs of both sexes (the males of most Batraxis species have modified tibiae or protuberant protrochanters), and the form of the aedeagus.	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF9D0B05FE81FD3E31680B18.taxon	etymology	Etymology. The new species is named after Yun-Chun Li, who collected the type material of the new species.	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF9D0B05FE81FD3E31680B18.taxon	distribution	Distribution. Southwestern China: Guizhou.	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF9D0B05FE81FD3E31680B18.taxon	discussion	Comments. This is the first Batraxis species known to inhabit a cave environment. Both specimens were found walking on the ground in the dark zone of the cave (Y. - C. Li, pers. comm.). However, B. liyunchuni sp. nov. exhibits no particular morphological adaptations (e. g., distinctly elongate appendages, reduction of eyes) as usually found in other obligate troglobitic pselaphines (e. g., Zopherobatrus Yin & Li, 2015). With this regard, further field work both in and outside the cave is needed to determinate the biology of this species.	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF9E0B04FC07FD40379202F8.taxon	description	(Fig. 2)	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF9E0B04FC07FD40379202F8.taxon	materials_examined	Type material. HOLOTYPE:, ‘ CHINA: SW China: Guizhou, Pingtang County (ỸṚH), Wanzi Village (ḾTIJ), Ba-mao-chong (ƂĿ Ṗ), Ganxiao Cave (Ȓ ḊḌ), N 27 ° 46 ′ 17.18 ′′, E 107 ° 17 ′ 39.26 ′′, ca. 1,200 m, 6. x. 2019, Yun-Chun Li leg. ’ (SNUC). PARATYPES: CHINA: 3 ♀♀, same label data as holotype (SNUC).	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF9E0B04FC07FD40379202F8.taxon	diagnosis	Diagnosis. Body length 1.65 – 1.81 mm; with dense, moderately long pubescence on head, pronotum, and elytra; male with strongly modified vertex and frons comprising complicated structures, vertex laterally with large setose area; female with head unmodified, frons roughly punctate instead; aedeagus symmetrical, apices of parameres pointed, endophallus mainly composed of weakly to strongly sclerotized spine-like structures.	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF9E0B04FC07FD40379202F8.taxon	description	Description. Male (Fig. 2 A). Body reddish-brown, length (from anterior margin of clypeus to apex of abdomen) 1.65 mm. Head (Fig. 2 B) sub-rectangular and strongly transverse, length from anterior margin of clypeus to head base 0.35 mm, maximum width across eyes 0.43 mm; vertex finely punctate, densely pubescent, strongly concave at middle, with pair of markedly raised projections each densely setose apically, anterior area deeply and transversely sulcate, lateral portions covered by dense, short setae; antennal tubercles only faintly raised; frons abruptly elevated at middle, forming vertical plane with clypeus; each eye composed of about 45 facets; antennae relatively elongate, length 0.68 mm, with distinct club formed by apical three antennomeres; antennomere 1 and 2 distinctly larger than antennomeres 3 – 8, antennomere 3 elongate, antennomeres 4 – 8 each moniliform, antennomere 9 approximately twice size of 8, subconical, antennomere 10 subconical and strongly transverse, antennomere 11 largest, broadest near middle and narrowing apically. Pronotum with moderately dense pubescence; slightly wider than long, length along midline 0.33 mm, maximum width 0.37 mm; with complete transverse antebasal sulcus connecting foveae, with nine short longitudinal ridges at base. Elytra much wider than long, length along suture 0.51 mm, maximum width 0.69 mm, dorsal surface with dense, long setae; each elytron with two nude foveae; sutural sulcus complete, lacking discal longitudinal sulcus. Legs simple, without protuberances. Abdomen wider than long, length along midline 0.64 mm, maximum width 0.42 mm; tergite IV (first visible tergite) with short discal carinae widely separated; both inner and outer marginal carinae complete and parallel; tergite VIII (Fig. 2 C) strongly transverse, with two lateral and one middle projection along anterior margin; sternite IX (Fig. 2 D) composed of two weakly-sclerotized, elongate plates. Length of aedeagus (Figs 2 E, 2 F) 0.24 mm (including parameres), median lobe and parameres symmetrical; each paramere roundly transverse and with pointed apex on posterior margin; endophallus with numerous weakly-sclerotized spine-like structure at middle portion, and with more elongate and more strongly sclerotized spines, and one elongate sclerite at basal portion. Female. Similar to male in general appearance; head lacking modification, anterior portion of frons roughly punctate; each eye composed of about 8 facets. Measurements (as for male): body length 1.77 – 1.81 mm; length of head 0.34 – 0.37 mm, width 0.37 – 0.41 mm; length of antennae 0.67 – 0.70 mm; length of pronotum 0.32 – 0.33 mm; width 0.34 – 0.35 mm; length of elytra 0.45 mm, width 0.65 – 0.67 mm; length of abdomen 0.64 mm, width 0.43 – 0.48 mm. Comparative notes. This is a very distinct species which can be easily separated from all congeners by the unique form of the sexually modified head of the male, and the configuration of the aedeagus.	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF9E0B04FC07FD40379202F8.taxon	etymology	Etymology. The new specific epithet alienoceps is an adjective combined from Latin ‘ aliēnus (exotic, foreign) and ‘ - ceps (- headed) ’, referring to the strongly modified head of the male.	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF9E0B04FC07FD40379202F8.taxon	distribution	Distribution. Southwestern China: Guizhou.	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF9E0B04FC07FD40379202F8.taxon	discussion	Comments. Two Japanese species, i. e., N. latifrons Jeannel, 1958, and N. plagiatus Jeannel, 1958, have been reported to inhabit cave environments (JEANNEL 1958; TA- NOKUCHI 1985). In Guizhou all individuals of N. alienoceps sp. nov. were collected under a few decomposing sticks on wet ground in a completely dark environment deep within the cave (Y. - C. Li, pers. comm.).	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF9F0B03FC36FDB6324D0138.taxon	diagnosis	Revised diagnosis. General body form slender, with extremely elongate appendages. Head narrowed apically; median longitudinal carina on vertex faintly present; lacking frontal rostrum; with indistinct antennal tubercles; eyes strongly reduced to lacking. Pronotal disc moderately convex; median longitudinal sulcus present or absent, lateral longitudinal sulci usually present; lacking antebasal spines; lacking spines on lateral margins. Elytra strongly constricted at base, each elytron with two basal foveae; discal stria lacking to distinct; disc moderately to strongly convex. Tergite IV (first visible tergite) longest, with single marginal carina.	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF980B02FEB8FDBE32D106B8.taxon	description	(Fig. 3)	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF980B02FEB8FDBE32D106B8.taxon	materials_examined	Type material. HOLOTYPE:, ‘ CHINA: SW China: Chongqing, Fengjie County (Ŧ ṂH), Xinglong Town (ĽD ẘ), Da-ban-qiao Stockaded Village (L ṞŘḂ), Taoyuan Village (ḢḎIJ), Zhangjiagou (K Ẏ NJ), Chuan Cave (Ŷ ḌT), N 30 ° 46 ′ 60 ′′, E 109 ° 28 ′ 10.2 ′′, ca. 905 m, 22. iii. 2016, Yun-Chun Li leg. ’ (SNUC). PARATYPE: CHINA: 1 ♀, same label data except ‘ 14 th Commune (14 "), anonymous cave (XZ Ḍ), N 30 ° 50 ′ 42 ′′, E 109 ° 34 ′ 21 ′′, ca. 905 m’ (SNUC).	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF980B02FEB8FDBE32D106B8.taxon	diagnosis	Diagnosis. Body length 3.3 – 3.5 mm; vertex lacking sulcus connecting foveae, mediobasal ridge faintly present; eyes strongly reduced, with only a few ommatidia; pronotum lacking median longitudinal sulcus, with thin lateral longitudinal sulci; elytra moderately constricted at base, with distinct discal sulcus; mesotibia with long apical spine in male; tergite IV with sides nearly parallel, narrowed at middle; median lobe of aedeagus with ventral lobe at right side, dorsal lobe weakly sclerotized.	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF980B02FEB8FDBE32D106B8.taxon	description	Description. Male (Fig. 3 A). length from anterior margin of clypeus to apex of abdomen 3.46 mm. Body reddish brown, antennae, mouthparts and tarsi lighter in color; most part of dorsal surface covered with long pubescence. Head (Fig. 3 B) lengthily oval, length from anterior margin of clypeus to head base 0.76 mm, maximum width across eyes 0.52 mm; surface finely punctate; vertex slightly convex, foveae below level of posterior margins of eyes, lacking U-shaped sulcus connecting vertexal foveae, faint median carina extending from head base to anterior margin of eyes; eyes strongly reduced, indicated by three small ommatidia; postocular margins rather long, narrowing from eyes towards base; length of antennae 2.30 mm, antennomeres each elongate; maxillary palpi with minute palpomere 1 and elongate palpomeres 2 – 4; palpomere 4 lacking obvious thickened setae at anterolateral margins, with small apical palpal cone. Pronotum (Fig. 3 C) wider than long, with round lateral margins, length along midline 0.62 mm, maximum width 0.77 mm, disc finely punctate; lacking antebasal sulcus. Elytra approximately as long as wide, length along suture 1.08 mm, maximum width 1.0 mm; each elytron with two small basal foveae; disc finely punctate, covered with long setae. All legs elongate, mesotibiae (Fig. 3 D) with long, slightly curved apical spine. Abdomen slightly longer than wide, length along midline 1.0 mm, maximum width 0.85 mm. Tergite IV (first visible tergite) longest, lacking modification; tergites V – VI combined much shorter than tergite IV; tergite VII as long as V and VI combined; tergite VIII semicircular. Sternite VIII semicircular, emarginate at middle of posterior margin. Aedeagus (Figs 3 F – G) asymmetrical, length 0.42 mm; basal capsule with triangular foramen; with moderately long basoventral projection; ventral lobe broadened at apical part, split to two major parts at apex; broad articulated dorsal lobe elongate, strongly sclerotized at base and weakly so apically; parameres fused to flattened, lamellate ventral membrane. Female. Similar to male in general appearance; mesotibiae lacking apical spine; each eye composed of five ommatidia. Measurements (as for male): body length 3.38 mm; length of head 0.79 mm, width across 0.53 mm; length of antennae 2.25 mm; length of pronotum 0.63 mm; maximum width 0.74 mm; length of elytra 1.09 mm, maximum width 1.0 mm; length of abdomen 1.03 mm, maximum width 0.87 mm. Comparative notes. The new species can be readily separated from Z. tianmingyii by the weakly developed vertexal sulcus and mediobasal ridge of the head, the presence of eyes in both sexes, the lack of median longitudinal sulcus of the pronotum, a relatively much border elytral base, a much more elongate apical spine of the male mesotibiae, and a totally different structure of the aedeagus.	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF980B02FEB8FDBE32D106B8.taxon	etymology	Etymology. The Latin lusciosus (- a, - um) is an adjective meaning ‘ purblind, dim-sighted’, referring to the strongly reduced eyes of the new species.	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF980B02FEB8FDBE32D106B8.taxon	distribution	Distribution. Southwestern China: Chongqing.	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
EA0E87CBFF980B02FEB8FDBE32D106B8.taxon	discussion	Comments. Both individuals were found under a rock in relatively moist environments of the dark zones in the two caves (Y. - C. Li, pers. comm.).	en	Yin, Zi-Wei (2020): New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 60 (1): 163-168, DOI: 10.37520/aemnp.2020.009
