identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
95D7CEE5AB59532DA7A47CDE469569A7.text	95D7CEE5AB59532DA7A47CDE469569A7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Candolleomyces conicus Haqnawaz, Niazi & Khalid 2025	<div><p>Candolleomyces conicus Haqnawaz, Niazi &amp; Khalid sp. nov.</p><p>Figs 3, 4</p><p>Etymology.</p><p>The species name “ conicus ” (Latin) refers to the conical pileus.</p><p>Holotype.</p><p>Pakistan • Punjab Province: Lahore District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=74.30167&amp;materialsCitation.latitude=31.494167" title="Search Plazi for locations around (long 74.30167/lat 31.494167)">University of the Punjab</a>, (31°29'39"N, 74°18'06"E, 227 m a. s. l.), on loamy soil, rich in organic matter, under Dalbergia sissoo, 10 August 2024, Muhammad Haqnawaz, PU–08 (LAH 38582). GenBank: PV 265173 [ITS], PV 265184 [LSU], PV 357393 [tef -1α].</p><p>Diagnosis.</p><p>Candolleomyces conicus is different from the closest species, C. sindhudeltae, by its relatively larger (20–90 mm diam.), conical, plane, reddish gray to light gray pileus with uplifted margins; brownish gray to dull reddish-brown lamellae, a large stipe (50–130 × 4–8 mm), a sheathing annulus with a wavy fibrillous margin, lecythiform, fusiform, tibiform cheilocystidia, and the pileipellis, a transition between hymeniderm and epithelium.</p><p>Description.</p><p>Pileus 20–90 mm diam., parabolic to campanulate with undulating margin in young stages, conical to campanulate than plane with uplifted and undulating margin when old; with a conical umbo, appressed scales, cracked at center, presence of veil elements; dull orange (7.5 YR 7 / 3) with grayish brown (7.5 YR 6 / 2) center when young, reddish gray (5 R 6 / 1) to light gray (10 YR 8 / 2) with dull orange (5 YR 6 / 3) center when old, with light gray (5 YR 8 / 1) squamules. Lamellae adnate, narrow to broad, even, with 3–7 tiers of lamellulae, light brownish (5 YR 7 / 1) when-young, brownish gray (5 YR 6 / 1) to dull reddish brown (2.5 YR 5 / 3) when mature. Stipe 50–130 × 4–8 mm, central, flexuous, tomentose, broad at base and narrow at apex, fibrillose, grayish white (N 8 / 0) to light gray (7.5 Y 7 / 2). Annulus sheathing, wavy and fibrillous at the margin.</p><p>Basidiospores [200 / 4 / 2], (6 –) 6.5–8.5 (– 9) × (4 –) 4.5–5 (– 6) µm, Q = 1.3–1.8, Qav = 1.65, ellipsoid to amygdaliform, thick – walled, smooth, guttulate, central germ pore present, red in water. Basidia (12 –) 13–21 (– 22) × (6 –) 7–9 (– 10) µm, clavate, hyaline in water, thick – walled, smooth, with 2–4 sterigmata. Cheilocystidia (11 –) 12–49 (– 50) × (10 –) 11–44 (– 45) µm, utriform, narrowly clavate to broadly clavate, ovoid, globose to subglobose, capitate to lecythiform, cylindrical, oblong, lageniform, flexuose, fusiform, tibiform, hyaline to olive yellow in water, thick-walled. Pleurocystidia absent. Pileipellis a transition between hymeniderm and epithelium, one-layered, thin-walled, 36–60 × 29–50 µm, globose to subglobose and ellipsoid to broadly ellipsoid, clavate to broadly clavate, cylindrical cells, hyaline to olive yellow in water. Stipitipellis a cutis, hyphae subregular, branched 6–17 µm in diameter, thin-walled, septate, and hyaline to yellow in water. Caulocystidia (18 –) 19–39 (– 40) × (9 –) 10–15 (– 16) µm, utriform, clavate to broadly clavate, ellipsoid, oblong, hyaline to olive yellow in water. Clamp connections present in all tissues.</p><p>Ecology and habitat.</p><p>Terrestrial, caespitose, on loamy soil rich in organic matter, under the dead trunk of Dalbergia sissoo .</p><p>Additional material examined.</p><p>Pakistan • Punjab Province: Kot Addu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=70.844444&amp;materialsCitation.latitude=30.546667" title="Search Plazi for locations around (long 70.844444/lat 30.546667)">Pirhar Gherbi</a> (30°32'48"N, 70°50'40"E, 136 m a. s. l.), on loamy soil, 11 Aug. 2023, Muhammad Haqnawaz, HQ–210 (LAH 38583). GenBank: PV 265174 [ITS], PV 265185 [LSU], PV 357394 [tef -1α] .</p><p>Notes.</p><p>According to phylogenetic analyses, the new species Candolleomyces conicus forms a separate clade from its sister species, such as C. sindhudeltae and C. undulatus . Morpho-anatomically, Candolleomyces conicus is different from C. sindhudeltae by having a plane, reddish gray pileus with uplifted margins, dull reddish-brown lamellae, a long stipe (50–130 mm length), lecythiform, and tibiform cheilocystidia. Candolleomyces sindhudeltae has an umbonate, grayish white, scalloped to cracked pileus with a decurved margin, small stipe (20–35 mm length), ovoid-pedunculate cheilocystidia (Haqnawaz et al. 2023 b). Candolleomyces undulatus has small (10–27 mm), straight shape, light purplish gray pileus with light brownish gray center, 1–3 tiers of lamellulae, light reddish gray lamellae, small stipe (15–46 × 2–4 mm), cylindrical, spherical with broad peduncle of basidia, ovoid, spheropedunculate, and globose caulocystidia (Li et al. 2025).</p></div>	https://treatment.plazi.org/id/95D7CEE5AB59532DA7A47CDE469569A7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Haqnawaz, Muhammad;Khalid, Muhammadah;Li, Qirui;Lu, Pingzhu;Bibi, Aysha;Naseer, Arooj;Afshan, Najam ul Sehar;Rehman Niazi, Abdul;Khalid, Abdul Nasir	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
A8A3BE215D7C5A369EC6CDEF0D6F6CF7.text	A8A3BE215D7C5A369EC6CDEF0D6F6CF7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Candolleomyces denticulatus M. Khalid, Haqnawaz & Afshan 2025	<div><p>Candolleomyces denticulatus M. Khalid, Haqnawaz &amp; Afshan sp. nov.</p><p>Figs 5, 6</p><p>Etymology.</p><p>The species name “ denticulatus ” (Latin) refers to the toothed cap margins.</p><p>Holotype.</p><p>Pakistan • Punjab Province: Lahore District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=74.30167&amp;materialsCitation.latitude=31.494167" title="Search Plazi for locations around (long 74.30167/lat 31.494167)">University of the Punjab</a>, (31°29'39"N, 74°18'06"E, 227 m a. s. l.), on loamy soil, rich in organic matter, under Punica granatum Linn, 15 Aug. 2024, Muhammadah Khalid, Abdul Nasir Khalid, and Muhammad Haqnawaz, PU–10 (LAH 38584). GenBank: PV 265175 [ITS], PV 265186 [LSU], PV 357395 [tef -1α].</p><p>Diagnosis.</p><p>Candolleomyces denticulatus differs from C. rubrobruneus in having toothed cap margins, abundant squamules when young, a grayish-white pileus with a pale orange center, grayish-white edges of the lamellae, and cheilocystidia that are globose, ellipsoid, pedunculate, globose, oblong, or lecythiform; at the same time, caulocystidia are tibiiform, lecythiform, oblong, fusiform, and moniliform.</p><p>Description.</p><p>Pileus 20–40 mm diam., parabolic in young stages, convex to plane when old, veil remnants present, dense squamules present in young stages, mostly removed when old, margin straight and split in symmetry to toothed, cracked surface, light gray (10 YR 8 / 1) to grayish white (N 8 / 0) with dull reddish brown (10 YR 5 / 4) to pale orange (5 YR 8 / 3) center, grayish white (N 8 / 0) squamules. Lamellae adnate, narrow, even to rarely eroded, with 3–7 tiers of lamellulae, forked, grayish brown (5 YR 6 / 2) to light brownish gray (5 YR 7 / 2), edge grayish white (N 8 / 0). Stipe 20–30 mm, central, equal, flexuous, surface scabrous, dull reddish brown (10 YR 5 / 4) to grayish white (N 8 / 0). Annulus absent.</p><p>Basidiospores [150 / 3 / 2], (6.0 –) 6.5–11.0 (– 11.6) × (4.2 –) 4.5–6.0 (– 6.3) µm, Q = 1.2–1.7, Qav = 1.56, amygdaliform in side view, ellipsoid in frontal view, smooth, guttulate, thick-walled, light red in water. Basidia (11 –) 11.9–21.1 (– 21.5) × (7.5 –) 7.9–9.1 (– 9.2) µm, clavate, hyaline to light gray (5 Y 7 / 1) in 5 % KOH, thin-walled, smooth, with 2–4 sterigmata, basal clamp connections present. Cheilocystidia 25–42 × 8–20 µm, narrowly utriform to broadly utriform, pedunculate, ovoid to conical, globose to subglobose, lecythiform, ellipsoid – pedunculate, narrowly lageniform, thick-walled, hyaline to yellowish gray (2.5 Y 5 / 1) in 5 % KOH. Pleurocystidia absent. Pileipellis an epitheloid hymeniderm, thick – walled, cells abundantly globose to subglobose, clavate to broadly clavate, ellipsoid to broadly ellipsoid, rarely utriform, oblong, lecythiform, cylindrical, 20–55 × 15–45 µm, hyaline to light gray (5 Y 7 / 1) in 5 % KOH. Stipitipellis made of 4.3–5.4 μm diam., subregular hyphae, rarely branched, hyaline in 5 % KOH, thin-walled septate, clamp connections abundant. Caulocystidia 27–64 × 8–18 µm, utriform to utriform pedunculate, slightly cylindrical, flexuous, ellipsoid, tibiiform, lecythiform, oblong, capitate, fusiform, moniliform, hyaline to light gray (5 Y 7 / 1) in 5 % KOH, thick-walled.</p><p>Ecology and habitat.</p><p>Gregarious and caespitose, terrestrial, on alluvial soil rich in organic matter.</p><p>Additional material examined.</p><p>Pakistan • Punjab Province: Kot Addu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=70.844444&amp;materialsCitation.latitude=30.551943" title="Search Plazi for locations around (long 70.844444/lat 30.551943)">Pirhar Gherbi</a> (30°33'07"N, 70°50'40"E, 137 m a. s. l.), on loamy soil, 21 Aug. 2024, Muhammadah Khalid, Abdul Nasir Khalid, and Muhammad Haqnawaz, KA–103 (LAH 38585). GenBank: PV 265176 [ITS], PV 265187 [LSU], PV 357396 [tef -1α] .</p><p>Notes.</p><p>Phylogenetically, Candolleomyces rubrobrunneus, C. pabbiensis, and C. kanhattiensis nov. sp. formed a separate branch from the new taxon C. denticulatus . Morpho-anatomically, Candolleomyces denticulatus is different from C. rubrobrunneus by having a toothed pileus margin, abundant squamules at the immature stage, grayish white pileus with pale orange center, grayish white edges of lamellae, yellowish gray basidiospores, ellipsoid pedunculate, utriform, oblong, lecythiform, cylindrical cheilocystidia; tibiiform, lecytheform, fusiform, moniliform caulocystidia. In contrast, C. rubrobrunneus has a straight margin, a dark brown pileus with a brownish, pale, fibrillose to floccose veil, grayish-orange to brown edges of lamellae, olive-brown basidiospores, broadly fusiform capitate cheilocystidia, and subcapitate apex of caulocystidia (Nayana and Pradeep 2024). However, C. pabbiensis is distinguished from our new taxon by conical to broadly parabolic pileus with straight margins, grayish white lamellae, light brown basidiospores, and plump cheilocystidia with an obtuse to subobtuse apex (Haqnawaz et al. 2024 b). Candolleomyces kanhattiensis differs from the new taxon C. denticulatus by having a sub-globose to campanulate pileus with decurved, undulating margins, a shiny stipe with a clavate base, greenish gray basidiospores, and cylindrical, ellipsoid, and globose cheilocystidia (in this study).</p></div>	https://treatment.plazi.org/id/A8A3BE215D7C5A369EC6CDEF0D6F6CF7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Haqnawaz, Muhammad;Khalid, Muhammadah;Li, Qirui;Lu, Pingzhu;Bibi, Aysha;Naseer, Arooj;Afshan, Najam ul Sehar;Rehman Niazi, Abdul;Khalid, Abdul Nasir	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
4A084D8F3EA752A09C280DE6BC3CECEC.text	4A084D8F3EA752A09C280DE6BC3CECEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Candolleomyces kanhattiensis Bibi, Haqnawaz, Afshan & Khalid 2025	<div><p>Candolleomyces kanhattiensis Bibi, Haqnawaz, Afshan &amp; Khalid sp. nov.</p><p>Figs 7, 8</p><p>Etymology.</p><p>The species name “ kanhattiensis ” (Latin) refers to the type locality of the species, ‘ Kanhatti Garden’, Khushab, Pakistan.</p><p>Holotype.</p><p>Pakistan • Punjab Province: Khushab District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=72.24972&amp;materialsCitation.latitude=32.42278" title="Search Plazi for locations around (long 72.24972/lat 32.42278)">Kanhatti Garden</a> (32°25'22"N, 72°14'59"E, 555 m a. s. l.), on loamy soil, rich in organic matter, under Phoenix dactylifera, 15 Aug. 2022, Ayesha Bibi, KUN–03 (LAH 38586). GenBank: PV 265177 [ITS], PV 265189 [LSU], PV 357397 [tef -1α].</p><p>Diagnosis.</p><p>This species is different from its closest species, Candolleomyces rubrobrunneus, by having sub-globose to campanulate, dull orange pileus with decurved margins, shiny stipe with clavate base, light brownish gray basidiospores, and cylindrical, ellipsoid, and globose cheilocystidia.</p><p>Description.</p><p>Pileus 15–25 mm in diam., sub-globose to campanulate, convex, slightly umbonate, decurved with even to undulating margins, smooth surface, orange (5 YR 7 / 6) to dull orange (7.5 YR 7 / 4), with distinct dark orange (7.5 YR 8 / 6) center. Lamellae adnate, narrow, even, with 3–7 tiers of lamellulae, dull orange (5 YR 7 / 3), entire edge. Stipe 30–65 × 2–5 mm, cylindrical, central, equal, grayish white (N 8 / 0), shiny and smooth upwards, clavate base, narrow toward pileus. Annulus absent.</p><p>Basidiospores [150 / 3 / 2], (5.5 –) 6–6.5 (– 7) × (3.9 –) 4–4.2 (– 4.5) μm, Q = 1.541–1.55, Qav = 1.53, ellipsoid to ovoid, thick – walled, dull reddish in water, smooth, apiculate, germ-pore absent, inner contents visible. Basidia (18 –) 19–22 (– 24) × (6 –) 7–8.5 (– 9) μm, sub-clavate to clavate, cylindrical, hyaline in KOH, thin-walled, smooth, with 2–4 small sterigmata. Cheilocystidia (18 –) 19–45 (– 46) × (8.5 –) 9–16 (– 17) μm, avl × avw = 34 × 14 μm, ovoid, clavate, lageniform, broadly cylindrical, ellipsoid to broadly ellipsoid, utriform, globose to subglobose, thick – walled. Pleurocystidia absent. Pileipellis an irregular epithelium, 15–25 × 8–20 μm, made up of oblong, subglobose, ellipsoid to broadly ellipsoid, clavate hyaline to yellowish in water, thin-walled, smooth, up to 39 μm broad. Stipitipellis a cutis made up of thin-walled hyphae, 4–12 μm diam., smooth, hyaline in KOH, branched, clamp connections present.</p><p>Ecology and habitat.</p><p>Gregarious, rich clayey soils in damp places, some in rotting leaves, compost piles, or dead small sticks of wood.</p><p>Additional specimens examined.</p><p>Pakistan • Punjab Province, District Khushab, in groups on muddy nutrient-rich soil under Dalbergia sissoo, sometimes on woody debris, 05 Aug. 2023, Ayesha Bibi, KHU–33 (LAH 38587). GenBank: PV 265178 [ITS], PV 265188 [LSU], PV 357398 [tef -1α] .</p><p>Notes.</p><p>Phylogenetically and morpho-anatomically, Candolleomyces kanhattiensis is closely related to C. rubrobrunneus, C. denticulatus nov. sp., and C. pabbiensis . It can be distinguished from C. rubrobrunneus by its sub-globose to campanulate, dull orange with dark orange center, decurved, undulating margin of pileus, shiny stipe with clavate base, greenish gray to light brownish gray basidiospores, and cylindrical, ellipsoid, and globose cheilocystidia. In contrast, C. rubrobrunneus has a conical, reddish brown to dark brown pileus with straight margins and a fibrillose to floccose surface, a pruinose apex of the stipe, broadly fusiform, capitate, flexuose cheilocystidia, and brownish yellow basidiospores (Nayana and Pradeep 2024). Candolleomyces pabbiensis exhibits a conical to broadly parabolic pileus with straight margins, grayish white lamellae, light brown basidiospores, and plump cheilocystidia with an obtuse to subobtuse apex (Haqnawaz et al. 2024 b). Candolleomyces denticulatus has toothed margins, abundant squamules when young, a grayish white pileus, and reddish gray to yellowish gray basidiospores. Furthermore, the cheilocystidia are conical, and polymorphic caulocystidia are also present (in this study).</p></div>	https://treatment.plazi.org/id/4A084D8F3EA752A09C280DE6BC3CECEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Haqnawaz, Muhammad;Khalid, Muhammadah;Li, Qirui;Lu, Pingzhu;Bibi, Aysha;Naseer, Arooj;Afshan, Najam ul Sehar;Rehman Niazi, Abdul;Khalid, Abdul Nasir	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
DDB3C8BBB6D95AF79F5BB4EBF9CB4AD9.text	DDB3C8BBB6D95AF79F5BB4EBF9CB4AD9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Candolleomyces swaticus Naseer, Haqnawaz & Khalid 2025	<div><p>Candolleomyces swaticus Naseer, Haqnawaz &amp; Khalid sp. nov.</p><p>Figs 9, 10</p><p>Etymology.</p><p>The species name “ swaticus ” (Latin) refers to the type locality, Swat.</p><p>Holotype.</p><p>Pakistan • Khyber Pakhtunkhwa Province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=72.409164&amp;materialsCitation.latitude=35.076942" title="Search Plazi for locations around (long 72.409164/lat 35.076942)">Swat</a> (35°04'37"N, 72°24'33"E, 1480 m a. s. l.), on loamy soil, 11 Aug. 2016, Arooj Naseer, A. N. Khalid, SK–23 (LAH 37132). GenBank: PV 265182 [ITS], PV 265191 [LSU].</p><p>Diagnosis.</p><p>Candolleomyces swaticus is different from Candolleomyces candolleanus by its large (20–60 mm), light gray with yellow-orange center, umbonate pileus, dull reddish-brown lamellae with 3–7 tiers, scales on stipe surface, bright brown basidiospores, capitate, lageniform, flexuose, tibiform, moniliform cheilocystidia.</p><p>Description.</p><p>Pileus 20–60 mm diam., campanulate at young stage, convex to plane when old, broadly umbonate, margin straight, split and irregularly wavy, presence of veil elements, smooth dull surface; light gray (10 YR 8 / 1) with dull yellow orange (10 YR 7 / 3) center. Lamellae adnexed, narrow to broad, average, with 3–7 tiers of lamellulae, dull reddish brown (2.5 YR 5 / 4). Stipe 25–55 × 4–8 mm, central, flexuous, tomentose, equal in width, scaly, grayish white (N 8 / 0). Annulus absent.</p><p>Basidiospores [150 / 3 / 3], (6 –) 6.5–7.5 (– 8) × (4 –) 4.2–4.8 (– 5) µm, Q = 1.3–1.8, Qav = 1.65, amygdaliform in side view, ovoid in frontal view, thick – walled, smooth, guttulate, central germ pore present, bright brown in water. Basidia 15–20 × 6.3–10.2 µm, clavate to broadly clavate, hyaline in water, thick – walled, smooth, with 3–4 sterigmata. Cheilocystidia (30 –) 31–49 (– 50) × (10 –) 11–19 (– 20) µm, utriform to slightly lecythiform, capitate, cylindrical to cylindrical with median constriction at upper side, narrowly lageniform to broadly lageniform, flexuose, broadly fusiform, tibiform, moniliform, hyaline to slightly yellowish in water, thick – walled. Pleurocystidia absent. Pileipellis an irregular epithelium, one layer, of thick – walled, 25–75 × 18–70 µm, globose to subglobose and ellipsoid to broadly ellipsoid, clavate to broadly clavate, cylindrical cells, hyaline to olive yellow in water. Stipitipellis a cutis, made of hyphae subregular, branched, 6–17 µm in diameter, thin-walled, septate, and hyaline to yellow in water. Caulocystidia not recorded. Clamp connections present in all tissues.</p><p>Ecology and habitat.</p><p>Terrestrial, gregarious, under a coniferous tree, on alluvium soil, during the rainy season.</p><p>Additional material examined.</p><p>Pakistan • Khyber Pakhtunkhwa Province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=72.409164&amp;materialsCitation.latitude=35.076942" title="Search Plazi for locations around (long 72.409164/lat 35.076942)">Swat</a> (35°04'37"N, 72°24'33"E, 1480 m a. s. l.), on loamy soil, 15 Aug. 2019, Arooj Naseer, A. N. Khalid, SK–01 (LAH 37131). GenBank: PV 265183 [ITS], PV 265190 [LSU] .</p><p>Notes.</p><p>According to phylogenetic analyses, Candolleomyces swaticus is closely related to C. candolleanus and C. badhyzensis . Morpho-anatomically, the former is different from the latter by having a campanulate pileus when young, being light gray with a dull yellow-orange center, having absence of hairy material (vs. C. candolleanus, which is covered with hairy material), having the presence of a broad umbo, and having irregular, wavy cap margins. Candolleomyces candolleanus has a parabolic pileus at a young stage, a bright brown pileus, and straight to even cap margins (Wächter and Melzer 2020). The next closely related taxon is Candolleomyces badhyzensis, which differs from the new species by a broadly bell-shaped, yellowish-brown pileus and cylindrical to ellipsoid to phaseoliform, pale brown basidiospores (vs. amygdaliform to ovoid, grayish red) (Kalamees 1981).</p></div>	https://treatment.plazi.org/id/DDB3C8BBB6D95AF79F5BB4EBF9CB4AD9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Haqnawaz, Muhammad;Khalid, Muhammadah;Li, Qirui;Lu, Pingzhu;Bibi, Aysha;Naseer, Arooj;Afshan, Najam ul Sehar;Rehman Niazi, Abdul;Khalid, Abdul Nasir	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
