taxonID	type	description	language	source
95D7CEE5AB59532DA7A47CDE469569A7.taxon	description	Figs 3, 4	en	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
95D7CEE5AB59532DA7A47CDE469569A7.taxon	etymology	Etymology. The species name “ conicus ” (Latin) refers to the conical pileus.	en	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
95D7CEE5AB59532DA7A47CDE469569A7.taxon	diagnosis	Diagnosis. Candolleomyces conicus is different from the closest species, C. sindhudeltae, by its relatively larger (20 – 90 mm diam.), conical, plane, reddish gray to light gray pileus with uplifted margins; brownish gray to dull reddish-brown lamellae, a large stipe (50 – 130 × 4 – 8 mm), a sheathing annulus with a wavy fibrillous margin, lecythiform, fusiform, tibiform cheilocystidia, and the pileipellis, a transition between hymeniderm and epithelium.	en	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
95D7CEE5AB59532DA7A47CDE469569A7.taxon	description	Description. Pileus 20 – 90 mm diam., parabolic to campanulate with undulating margin in young stages, conical to campanulate than plane with uplifted and undulating margin when old; with a conical umbo, appressed scales, cracked at center, presence of veil elements; dull orange (7.5 YR 7 / 3) with grayish brown (7.5 YR 6 / 2) center when young, reddish gray (5 R 6 / 1) to light gray (10 YR 8 / 2) with dull orange (5 YR 6 / 3) center when old, with light gray (5 YR 8 / 1) squamules. Lamellae adnate, narrow to broad, even, with 3 – 7 tiers of lamellulae, light brownish (5 YR 7 / 1) when-young, brownish gray (5 YR 6 / 1) to dull reddish brown (2.5 YR 5 / 3) when mature. Stipe 50 – 130 × 4 – 8 mm, central, flexuous, tomentose, broad at base and narrow at apex, fibrillose, grayish white (N 8 / 0) to light gray (7.5 Y 7 / 2). Annulus sheathing, wavy and fibrillous at the margin. Basidiospores [200 / 4 / 2], (6 –) 6.5 – 8.5 (– 9) × (4 –) 4.5 – 5 (– 6) µm, Q = 1.3 – 1.8, Qav = 1.65, ellipsoid to amygdaliform, thick – walled, smooth, guttulate, central germ pore present, red in water. Basidia (12 –) 13 – 21 (– 22) × (6 –) 7 – 9 (– 10) µm, clavate, hyaline in water, thick – walled, smooth, with 2 – 4 sterigmata. Cheilocystidia (11 –) 12 – 49 (– 50) × (10 –) 11 – 44 (– 45) µm, utriform, narrowly clavate to broadly clavate, ovoid, globose to subglobose, capitate to lecythiform, cylindrical, oblong, lageniform, flexuose, fusiform, tibiform, hyaline to olive yellow in water, thick-walled. Pleurocystidia absent. Pileipellis a transition between hymeniderm and epithelium, one-layered, thin-walled, 36 – 60 × 29 – 50 µm, globose to subglobose and ellipsoid to broadly ellipsoid, clavate to broadly clavate, cylindrical cells, hyaline to olive yellow in water. Stipitipellis a cutis, hyphae subregular, branched 6 – 17 µm in diameter, thin-walled, septate, and hyaline to yellow in water. Caulocystidia (18 –) 19 – 39 (– 40) × (9 –) 10 – 15 (– 16) µm, utriform, clavate to broadly clavate, ellipsoid, oblong, hyaline to olive yellow in water. Clamp connections present in all tissues.	en	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
A8A3BE215D7C5A369EC6CDEF0D6F6CF7.taxon	description	Figs 5, 6	en	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
A8A3BE215D7C5A369EC6CDEF0D6F6CF7.taxon	etymology	Etymology. The species name “ denticulatus ” (Latin) refers to the toothed cap margins.	en	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
A8A3BE215D7C5A369EC6CDEF0D6F6CF7.taxon	diagnosis	Diagnosis. Candolleomyces denticulatus differs from C. rubrobruneus in having toothed cap margins, abundant squamules when young, a grayish-white pileus with a pale orange center, grayish-white edges of the lamellae, and cheilocystidia that are globose, ellipsoid, pedunculate, globose, oblong, or lecythiform; at the same time, caulocystidia are tibiiform, lecythiform, oblong, fusiform, and moniliform.	en	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
A8A3BE215D7C5A369EC6CDEF0D6F6CF7.taxon	description	Description. Pileus 20 – 40 mm diam., parabolic in young stages, convex to plane when old, veil remnants present, dense squamules present in young stages, mostly removed when old, margin straight and split in symmetry to toothed, cracked surface, light gray (10 YR 8 / 1) to grayish white (N 8 / 0) with dull reddish brown (10 YR 5 / 4) to pale orange (5 YR 8 / 3) center, grayish white (N 8 / 0) squamules. Lamellae adnate, narrow, even to rarely eroded, with 3 – 7 tiers of lamellulae, forked, grayish brown (5 YR 6 / 2) to light brownish gray (5 YR 7 / 2), edge grayish white (N 8 / 0). Stipe 20 – 30 mm, central, equal, flexuous, surface scabrous, dull reddish brown (10 YR 5 / 4) to grayish white (N 8 / 0). Annulus absent. Basidiospores [150 / 3 / 2], (6.0 –) 6.5 – 11.0 (– 11.6) × (4.2 –) 4.5 – 6.0 (– 6.3) µm, Q = 1.2 – 1.7, Qav = 1.56, amygdaliform in side view, ellipsoid in frontal view, smooth, guttulate, thick-walled, light red in water. Basidia (11 –) 11.9 – 21.1 (– 21.5) × (7.5 –) 7.9 – 9.1 (– 9.2) µm, clavate, hyaline to light gray (5 Y 7 / 1) in 5 % KOH, thin-walled, smooth, with 2 – 4 sterigmata, basal clamp connections present. Cheilocystidia 25 – 42 × 8 – 20 µm, narrowly utriform to broadly utriform, pedunculate, ovoid to conical, globose to subglobose, lecythiform, ellipsoid – pedunculate, narrowly lageniform, thick-walled, hyaline to yellowish gray (2.5 Y 5 / 1) in 5 % KOH. Pleurocystidia absent. Pileipellis an epitheloid hymeniderm, thick – walled, cells abundantly globose to subglobose, clavate to broadly clavate, ellipsoid to broadly ellipsoid, rarely utriform, oblong, lecythiform, cylindrical, 20 – 55 × 15 – 45 µm, hyaline to light gray (5 Y 7 / 1) in 5 % KOH. Stipitipellis made of 4.3 – 5.4 μm diam., subregular hyphae, rarely branched, hyaline in 5 % KOH, thin-walled septate, clamp connections abundant. Caulocystidia 27 – 64 × 8 – 18 µm, utriform to utriform pedunculate, slightly cylindrical, flexuous, ellipsoid, tibiiform, lecythiform, oblong, capitate, fusiform, moniliform, hyaline to light gray (5 Y 7 / 1) in 5 % KOH, thick-walled.	en	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
4A084D8F3EA752A09C280DE6BC3CECEC.taxon	description	Figs 7, 8	en	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
4A084D8F3EA752A09C280DE6BC3CECEC.taxon	etymology	Etymology. The species name “ kanhattiensis ” (Latin) refers to the type locality of the species, ‘ Kanhatti Garden’, Khushab, Pakistan.	en	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
4A084D8F3EA752A09C280DE6BC3CECEC.taxon	diagnosis	Diagnosis. This species is different from its closest species, Candolleomyces rubrobrunneus, by having sub-globose to campanulate, dull orange pileus with decurved margins, shiny stipe with clavate base, light brownish gray basidiospores, and cylindrical, ellipsoid, and globose cheilocystidia.	en	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
4A084D8F3EA752A09C280DE6BC3CECEC.taxon	description	Description. Pileus 15 – 25 mm in diam., sub-globose to campanulate, convex, slightly umbonate, decurved with even to undulating margins, smooth surface, orange (5 YR 7 / 6) to dull orange (7.5 YR 7 / 4), with distinct dark orange (7.5 YR 8 / 6) center. Lamellae adnate, narrow, even, with 3 – 7 tiers of lamellulae, dull orange (5 YR 7 / 3), entire edge. Stipe 30 – 65 × 2 – 5 mm, cylindrical, central, equal, grayish white (N 8 / 0), shiny and smooth upwards, clavate base, narrow toward pileus. Annulus absent. Basidiospores [150 / 3 / 2], (5.5 –) 6 – 6.5 (– 7) × (3.9 –) 4 – 4.2 (– 4.5) μm, Q = 1.541 – 1.55, Qav = 1.53, ellipsoid to ovoid, thick – walled, dull reddish in water, smooth, apiculate, germ-pore absent, inner contents visible. Basidia (18 –) 19 – 22 (– 24) × (6 –) 7 – 8.5 (– 9) μm, sub-clavate to clavate, cylindrical, hyaline in KOH, thin-walled, smooth, with 2 – 4 small sterigmata. Cheilocystidia (18 –) 19 – 45 (– 46) × (8.5 –) 9 – 16 (– 17) μm, avl × avw = 34 × 14 μm, ovoid, clavate, lageniform, broadly cylindrical, ellipsoid to broadly ellipsoid, utriform, globose to subglobose, thick – walled. Pleurocystidia absent. Pileipellis an irregular epithelium, 15 – 25 × 8 – 20 μm, made up of oblong, subglobose, ellipsoid to broadly ellipsoid, clavate hyaline to yellowish in water, thin-walled, smooth, up to 39 μm broad. Stipitipellis a cutis made up of thin-walled hyphae, 4 – 12 μm diam., smooth, hyaline in KOH, branched, clamp connections present.	en	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
DDB3C8BBB6D95AF79F5BB4EBF9CB4AD9.taxon	description	Figs 9, 10	en	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
DDB3C8BBB6D95AF79F5BB4EBF9CB4AD9.taxon	etymology	Etymology. The species name “ swaticus ” (Latin) refers to the type locality, Swat.	en	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
DDB3C8BBB6D95AF79F5BB4EBF9CB4AD9.taxon	diagnosis	Diagnosis. Candolleomyces swaticus is different from Candolleomyces candolleanus by its large (20 – 60 mm), light gray with yellow-orange center, umbonate pileus, dull reddish-brown lamellae with 3 – 7 tiers, scales on stipe surface, bright brown basidiospores, capitate, lageniform, flexuose, tibiform, moniliform cheilocystidia.	en	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
DDB3C8BBB6D95AF79F5BB4EBF9CB4AD9.taxon	description	Description. Pileus 20 – 60 mm diam., campanulate at young stage, convex to plane when old, broadly umbonate, margin straight, split and irregularly wavy, presence of veil elements, smooth dull surface; light gray (10 YR 8 / 1) with dull yellow orange (10 YR 7 / 3) center. Lamellae adnexed, narrow to broad, average, with 3 – 7 tiers of lamellulae, dull reddish brown (2.5 YR 5 / 4). Stipe 25 – 55 × 4 – 8 mm, central, flexuous, tomentose, equal in width, scaly, grayish white (N 8 / 0). Annulus absent. Basidiospores [150 / 3 / 3], (6 –) 6.5 – 7.5 (– 8) × (4 –) 4.2 – 4.8 (– 5) µm, Q = 1.3 – 1.8, Qav = 1.65, amygdaliform in side view, ovoid in frontal view, thick – walled, smooth, guttulate, central germ pore present, bright brown in water. Basidia 15 – 20 × 6.3 – 10.2 µm, clavate to broadly clavate, hyaline in water, thick – walled, smooth, with 3 – 4 sterigmata. Cheilocystidia (30 –) 31 – 49 (– 50) × (10 –) 11 – 19 (– 20) µm, utriform to slightly lecythiform, capitate, cylindrical to cylindrical with median constriction at upper side, narrowly lageniform to broadly lageniform, flexuose, broadly fusiform, tibiform, moniliform, hyaline to slightly yellowish in water, thick – walled. Pleurocystidia absent. Pileipellis an irregular epithelium, one layer, of thick – walled, 25 – 75 × 18 – 70 µm, globose to subglobose and ellipsoid to broadly ellipsoid, clavate to broadly clavate, cylindrical cells, hyaline to olive yellow in water. Stipitipellis a cutis, made of hyphae subregular, branched, 6 – 17 µm in diameter, thin-walled, septate, and hyaline to yellow in water. Caulocystidia not recorded. Clamp connections present in all tissues.	en	Haqnawaz, Muhammad, Khalid, Muhammadah, Li, Qirui, Lu, Pingzhu, Bibi, Aysha, Naseer, Arooj, Afshan, Najam ul Sehar, Rehman Niazi, Abdul, Khalid, Abdul Nasir (2025): Morphology and multi-locus phylogeny reveal four new Candolleomyces (Psathyrellaceae) species from Pakistan. MycoKeys 121: 143-166, DOI: 10.3897/mycokeys.121.157282
