taxonID	type	description	language	source
E8713702D113FFA19266B1EDFE43F801.taxon	description	Fig. 2	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D113FFA19266B1EDFE43F801.taxon	type_taxon	Type species Alluaudensia nigrolineata Lallemand, 1920 by monotypy.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D113FFA19266B1EDFE43F801.taxon	description	Synthesis of Lallemand's description Alluaudensia is an endemic genus from Madagascar known by only one species described from a single female specimen by Lallemand (1920). He augmented his original description in 1949 (Lallemand 1949). A synthesis based on Lallemand's descriptions is presented below: HEAD. Vertex with a light hump carrying the ocelli on its anterior median part. Ocelli closer to each other than to compound eyes. Postclypeus with large furrow. Rostrum reaches mesocoxae. THORAX. Pronotum strongly inclined on anterior part resulting in strongly humped shape; pronotum with dimples on each side and three transverse furrows. Hindlegs with two spines on metatibiae. TEGMINA. Veins prominent. Apical part of tegmina with dense network of veins (Lallemand 1920, 1949).	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D113FFA19266B1EDFE43F801.taxon	distribution	Distribution (Fig. 2 E) Endemic genus of Madagascar.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D113FFA19266B1EDFE43F801.taxon	discussion	Number of species: 1.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D114FFA7924CB462FDA8FD74.taxon	materials_examined	Material examined Holotype MADAGASCAR • 1 ♀; Diego-Suarez; 1893; C. Alluaud leg.; MNHN (EH) 7394.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D114FFA7924CB462FDA8FD74.taxon	description	Description TOTAL LENGTH. 19.0 mm (holotype). HEAD. In dorsal view head 2 times as wide as long. Ocelli closer to each other than to compound eyes and placed on small hump. Postclypeus with large furrow on median part extending over its full height. THORAX. Pronotum very prominent. Scutellum as long as large. COLORATION. Head red with pedicels of antennae black. Anterior third of pronotum red. Posterior part of pronotum and scutellum black. In dorsal view, abdomen with first segment dark and rest red. In ventral view, specimen entirely red. Legs black except hindlegs, femurs brown. Tegmina mainly red-ochre, with a black stripe following the margins. Black stripe large on external and apical margins and thin on internal claval vein. Veins red and black in dark parts of the wing. Hindwings smoked with red veins on anterior part, brown on apical part.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D115FFA49200B7B0FDB9FC93.taxon	description	Fig. 3 H	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D115FFA49200B7B0FDB9FC93.taxon	type_taxon	Type species Amberana elongata Distant, 1908.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D115FFA49200B7B0FDB9FC93.taxon	description	Synthetic description Amberana is an endemic genus from Madagascar described by Distant in 1908. It was revised by Soulier-Perkins & Kunz in 2012. A synthesis based on their descriptions of this genus is presented below: HEAD. Head as wide as long between eyes, anterior margin with carina. Median part of vertex strongly curved, flattened vertically on lower part to form triangle just above delimitation of postclypeus. Ocelli close to each other and separated by small carina. Postclypeus convex, striated laterally with triangular dimple just below frons-postclypeus suture. Middle part of postclypeus can be strongly marked or not by sulciform furrow. Rostrum reaching mesocoxae. THORAX. Anterior margin of pronotum straight or slightly rounded, three small concavities present on each side along anterior margin. Scutellum a bit longer than wide. Hindlegs with one spine on metatibiae. MALE GENITALIA. Male genitalia with sterno-lateral plates and small parameres not exceeding the pygofer margin (Distant 1908; Soulier-Perkins & Kunz 2012).	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D115FFA49200B7B0FDB9FC93.taxon	distribution	Distribution (Fig. 3 H) Endemic genus of Madagascar.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D115FFA49200B7B0FDB9FC93.taxon	discussion	Number of species: 23.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D116FFAA91B2B6DDFEACFD97.taxon	description	Fig. 3	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D116FFAA91B2B6DDFEACFD97.taxon	materials_examined	Material examined MADAGASCAR • 1 ♂; Forêt Tanala; 1901; C. Alluaud leg.; V. Lallemand det.; MNHN (EH) 24113 • 1 ♂; 1898; collection Noualhier; MNHN (EH) 24536 • 1 ♀; Forêt Tanala; 1901; C. Alluaud leg.; MNHN (EH) 24117 • 1 ♀; Forêt Tanala, rég. de Ranomafana Anjorojoro; Mar. 1901; C. Alluaud leg.; V. Lallemand det.; MNHN (EH) 24113 • 1 ♀; 1898; collection Noualhier; MNHN (EH) 24537 • 1 ♀; 1898; collection Noualhier; MNHN (EH) 24538.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D116FFAA91B2B6DDFEACFD97.taxon	description	Description TOTAL LENGTH. Between 6.3 and 6.6 mm. HEAD. In dorsal view (Fig. 3 A, C), median part of vertex with small triangle, indistinct, on lower margin. Ocelli separated by thin carina. Lateral part of vertex curved on upper part and flattened on anterior margin of head. Postclypeus slightly convex forming obtuse angle in lateral view. Postclypeus with triangle dimple below frons-postclypeus suture with clear sulciform mark on middle. THORAX. Anterior part of pronotom with very thin carina. MALE GENITALIA. Subgenital plates long, reaching dorsal third of pygofer, strongly curved; diameter decreasing gently toward tip, which is rounded and covered with short setae (Fig. 3 E). Dorsal margin of parameres possesses two rounded extensions: anterior than other. Sterno-lateral plates wide and exceeding pygofer with triangular shape with dorsal and ventral junction convex and oriented posteriorly (Fig. 3 G). On ventral part of parameres, anterior and posterior margins mainly parallel with junction forming rounded endpoint. Aedeagus (Fig. 3 F) not exceeding height of pygofer. Anterior margin of aedeagus with successively (from base to apex) short bump, small membranous part and with two spined excrescences, lower one being longer. Ventral margin extending in very convex arc. Dorsal part of aedeagus arced in obtuse angle. Apical part of aedeagus bifid with anterior thin spine-like extension and a posterior wider extension with rounded apex; anterior extension one third length of posterior one. COLORATION. Head and anterior third of pronotum uniformly dark brown. Middle of pronotum with continuous transverse yellow stripe (Fig. 3 A – C). Rest of pronotum and scutellum dark to light brown. In ventral view, thorax dark as head (Fig. 3 B). Abdomen and legs reddish brown with tarses dark brown or black (Fig. 3 B – C). Tegmina brown or slightly clearer than rest of body. Tegmina with two distinct yellow bands (Fig. 3 A – B): one extending over clavus, starting from insertion of tegmina to apical part of PCu; other extending on corium between M and Cu. Insertion of tegmina reddish or light brown.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D119FFA99248B3A4FD15FCE1.taxon	description	Fig. 4	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D119FFA99248B3A4FD15FCE1.taxon	type_taxon	Type species Bourgoinrana perinetana (Synave, 1957).	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D119FFA99248B3A4FD15FCE1.taxon	materials_examined	Material examined Bourgoinrana perinetana (Synave, 1957) MADAGASCAR • 1 ♂; Marojejy-Ouest; 1600 m a. s. l.; Nov. 1959; P. Soga leg.; A. Soulier-Perkins det.; Institut scientifique de Madagascar; I. G. 22.889 (RIScNBIG) • 1 ♂; Massif du Tsaratanana, versant Sud, Andohanambatoafo; 2030 m a. s. l.; 16 – 18 Dec. 1966; P. Soga leg.; A. Soulier-Perkins det.; MNHN (EH) 4819 • 1 ♀; Massif du Tsaratanana, versant Sud, Andohanambatoafo; 2030 m a. s. l.; 16 – 18 Dec. 1966; P. Soga leg.; A. Soulier-Perkins det.; coll. P. Viette; MNHN (EH) 4816. Bourgoinrana sandrangatensis (Synave, 1957) MADAGASCAR • 1 ♂; A. Soulier-Perkins det.; MNHN (EH) 22380. Bourgoinrana rubescens (Synave, 1957) MADAGASCAR • 1 ♂; Marojezy, Dec. 1972; A. Soulier-Perkins det.; mission CNRS; R. C. P. n ° 225; MNHN (EH) 4833 • 1 ♂; Marojezy; Dec. 1972; A. Soulier-Perkins det.; mission CNRS, R. C. P. n ° 225; MNHN (EH) 4834.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D119FFA99248B3A4FD15FCE1.taxon	description	Synthetic description Bourgoinrana is a genus which had all its species placed in Amberana until its description in 2012. It is an endemic genus of Madagascar, easy to differentiate from the other genera present there, mainly thanks to its flattened shape (Soulier-Perkins & Kunz 2012). A synthesis based on previous taxonomic descriptions and material examined is presented below: HEAD. Vertex as broad as long. Anterior margin of head regularly rounded. Ocelli closer to each other than to compound eyes. Postclypeus with large longitudinal furrow. THORAX. Pronotum as wide as long and dorsoventrally flattened, giving aspect easy to identify to insects from genus. Pronotum with few distinct dimples, not deep. Hindlegs with one spine on metatibiae. MALE GENITALIA. Have sterno-lateral plates and aedeagus with thumb shape on anterior margin (Soulier-Perkins & Kunz 2012).	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D119FFA99248B3A4FD15FCE1.taxon	distribution	Distribution (Fig. 4) Endemic genus of Madagascar.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D119FFA99248B3A4FD15FCE1.taxon	discussion	Number of species: 4.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D11BFFAF92DAB62FFDCEFEAF.taxon	description	Fig. 5	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D11BFFAF92DAB62FFDCEFEAF.taxon	type_taxon	Type species Literna maura (Thunberg, 1822).	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D11BFFAF92DAB62FFDCEFEAF.taxon	description	Synthetic description Literna is an African genus, described long ago in a very brief manner by Stål (1866), making it impossible to currently determine the species with his original publication. Most of its species are present in Madagascar. Lallemand proposed a refined description of this genus in 1912 based on the general body shape and wing venation (Lallemand 1912). He updated his description in 1949 by adding genitalia characters for males and females, but they remained too general to qualify them as characteristic for this genus (Lallemand 1949). The male genitalia have been described for only three species: L. altipeta Lallemand & Synave, 1952; L. muscophila Lallemand & Synave, 1952 and L. pauliani Lallemand & Synave, 1952 (Lallemand & Synave 1952). Hence, the description of the male genitalia for this genus is not presented in this publication. Moreover, some species, such as L. callosipenis (Signoret, 1860), seem to show morphological characters similar to the species now assigned to the new genus Soulierana Bucher & Bouteille gen. nov. Nevertheless, types-specimen (s) were not consulted during this study and as a result, no taxonomic revisions have been made for these taxa. However, the lack of information for the genitalia characters highlights the need for further taxonomic studies. A synthesis based on the older descriptions for the Malagasy species is presented below: HEAD. In dorsal view (Fig. 5 A), head as wide as long. Ocelli closer to each other than to compound eyes and generally separated by carina, barely visible or even absent in some species (Fig. 5 A, C); a carina can be observed on frons. Postclypeus with large furrow which can extend over its entire length, but not necessarily; lateral edges of furrow usually diffused (Fig. 5 C). In some species, such as L. callosipennis (Signoret 1860), L. hovana Lallemand, 1949, L. madegassa Lallemand, 1949 and L. signata Jacobi, 1917, lateral part of postclypeus marked with two carina and deep furrow. Postclypeus striated laterally. THORAX. Anterior margin of pronotum strongly declining; presence of a few dimples on anterior part. Scutellum longer than wide. Hindlegs with one spine on metatibiae. TEGMINA. Tegmina longer than wide (Fig. 5 A) in tapered shape, except for L. ochracea Jacobi, 1917, which possesses oval tegmina strongly rounded on posterior part. One or few bumps on anterior part of tegmina. MALE GENITALIA. Male genitalia with sterno-lateral plates.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D11BFFAF92DAB62FFDCEFEAF.taxon	distribution	Distribution Tropical Africa, South Africa and Madagascar.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D11BFFAF92DAB62FFDCEFEAF.taxon	discussion	Number of species: 25, 16 in Madagascar.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D11DFFAD92D5B4EEFDDDFE59.taxon	description	Fig. 6	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D11DFFAD92D5B4EEFDDDFE59.taxon	type_taxon	Type species Locris rubra (Fabricius, 1794).	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D11DFFAD92D5B4EEFDDDFE59.taxon	materials_examined	Material examined Locris bipunctata (Signoret, 1860) MADAGASCAR • 1 ♂; “ Ambabalame; V. Lallemand det.; Coll. Mus. Congo; 6664 H; Muséum Paris ”; MNHN (EH) 24628 • 1 ♂; “ Lavaudon; Coll. E. de Bergev ”; MNHN (EH) 24115 • 1 ♂; “ Amber-Geb.; Edm. Schmidt det.; Miz Pan Warszawa 12 / 1945 2487 ” • 1 ♂; “ Amber-Geb.; Edm. Schmidt det.; 970; Miz Pan Warszawa 12 / 1945 2488 ” • 1 ♂; “ Amber-Geb.; Edm. Schmidt det.; Miz Pan Warszawa 12 / 1945 2489 ”. Locris nigrolimbata (Lallemand, 1910) KENYA • 1 ♂; “ Afrique orientale Anglaise St. 60; 600 m a. s. l.; Mar. 1911; Alluaud and Jeannel; Coll. Mus. Congo; Coll. V. Lallemand; R det. 6665 ”; MRAC • 1 ♂; “ Afrique orientale Anglaise, WaKikuyu, Fort-Hall, St. 80; 1330 m a. s. l.; Jan. 1912; Alluaud and Jeannel; Coll. Mus. Congo; Coll. V. Lallemand; R det. 6665 ”; MRAC • 1 ♂; “ Nairobi; Apr. 1923; G. Babault leg.; Locris johannae ssn. nigrolimbata ”; MNHN (EH) 24629. TANZANIA • 1 ♂; Mts. Uluguru, Kimboza ofr. Héliophile; 600 m a. s. l.; 24 – 30 Jul. 1971; L. Berger, N. Leleup and J. Debecker leg.; Coll. Mus. Tervuren; H. Synave det.; MNHN (EH) 24635. Locris vicina (Signoret, 1860) MADAGASCAR • 1 ♂; Parc de Zombitse Leobondro, rég. Atsimo-andrefana, bord rivière; 22 ° 40.460 ′ S, 44 ° 51.633 ′ E; 633 m a. s. l.; 20 Mar. 2006; A. Soulier-Perkins leg. and det.; MNHN (EH) 24659 • 1 ♂; Ambodimanga; 1906; S. I. Hammerstem leg.; Edm. Schmidt det.; Miz Pan Warszawa 12 / 1945 2490 • 1 ♂; Ambodimanga; 1906; S. I. Hammerstem leg.; Edm. Schmidt det.; Miz Pan Warszawa 12 / 1945 2491.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D11DFFAD92D5B4EEFDDDFE59.taxon	description	Synthetic description Locris is a large African genus with more than 80 species currently described (Soulier-Perkins 2023). It is known in Madagascar by three species. Stål (1866) initially described Locris in a very brief manner, making it impossible to currently determine the genus with his original publication. Lallemand proposed a refined description of this genus in 1949 on the basis of morphological characters that are not all completely consistent, according to Bouteille et al. (2021). It is true that in addition to a list of general characters supposed to describe the genus, Lallemand (1949) also listed a series of exceptions in order to include more species, probably making the genus paraphyletic. The homogeneity of this genus and its taxonomic unity is therefore questionable. Bouteille et al. (2021) revised the three Malagasy species of Locris and provided descriptions of their genitalia with new characters. Nevertheless, the lack of systematic information for the genitalic characters of the entire genus highlights the need for further taxonomic studies. A synthesis based on previous taxonomic descriptions and material examined is presented below: HEAD. Postclypeus rounded in lateral view with longitudinal carina in median part and transverse ridges. Rostrum short, barely extending to base of median trochanter. Antennae short. Vertex as broad as long with length equal to half length of pronotum. Ocelli closer to each other than to compound eyes. THORAX. Pronotum large with straight posterior margin. Scutellum as long as large with 3 dimples; 2 small dimples above anterior margin and a bigger one on median part. Hindlegs with one spine on metatibiae. TEGMINA. Tegmina 3 times as long as wide. Cu and M linked from base to middle of tegmina. Apical veins form dense network on apex. MALE GENITALIA. No sterno-lateral plates. Subgenital plates wide and sometimes silky at base, forming thin and long filament (Lallemand 1949) curved up or downwards on its apical half. Parameres silky and wide from pygofer’s attachment to its tip and they can form a cup prolonged by one or two structures that can have a little spine curved up- or downwards (Bouteille et al. 2021). Aedeagus can have little bumps on top. Apex with hanging structure (Bouteille et al. 2021) which varies in size and shape, with presence in many species of bifid branch connected in middle. Apex sometimes silky and wider than rest of aedeagus (Bouteille et al. 2021).	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D11DFFAD92D5B4EEFDDDFE59.taxon	distribution	Distribution (Fig. 6) Tropical Africa, South Africa and Madagascar.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D11DFFAD92D5B4EEFDDDFE59.taxon	discussion	Number of species: 87, 3 in Madagascar.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D11FFFAD926EB484FE43FA16.taxon	description	Fig. 7	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D11FFFAD926EB484FE43FA16.taxon	type_taxon	Type species Paramioscarta brunnea (Lallemand, 1920) by monotypy.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D11FFFAD926EB484FE43FA16.taxon	description	Synthetic description Paramioscarta is an endemic monospecific genus of Madagascar. It shares features with the genus Amberana but can be differentiated by its broader general shape and by the large parameres of male genitalia. A synthesis based on Lallemand descriptions (Lallemand 1920, 1949) and material examined is presented below: HEAD. Wider than long. Ocelli closer to each other than to compound eyes and separated by a carina. Median part of vertex strongly domed; lateral margins of dome not parallel, with recess in middle giving this structure a lock shape. Postclypeus rounded in lateral view, striated laterally with triangular dimple just below frons-postclypeus suture as in genus Amberana. Upper part of postclypeus with vertical furrow. THORAX. Three clear dimples just behind anterior margin on each side of pronotum; another one discernable in middle part. Hindlegs with one spine on metatibiae. MALE GENITALIA. Male genitalia with sterno-lateral plates and large parameres clearly exceeding pygofer margin.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D11FFFAD926EB484FE43FA16.taxon	distribution	Distribution (Fig. 7 H) Endemic genus of Madagascar.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D11FFFAD926EB484FE43FA16.taxon	discussion	Number of species: 1.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D11FFFB39249B34CFB32FA54.taxon	materials_examined	Material examined MADAGASCAR • 1 ♂; Province d’Analalava Maromandia; 1923; R. Decary leg.; V. Lallemand det.; MNHN (EH) 24505 • 1 ♂; Reserve spéciale du Zombitsy, Est de Sakaraha matsabory; 640 m a. s. l.; 13 Feb. 1974; P. Viette and A. Peyrieras leg.; MNHN (EH) 24531 • 1 ♂; Ouest S-P. Antsalova Antsingy, Rés. Nat. 9; Jan. 1975; A. Peyrieras leg.; MNHN (EH) 24532 • 1 ♂; Province de Toliara, massif du Makay; 21 ° 37 ′ 46.1 ″ S, 45 ° 00 ′ 17.7 ″ E; 200 m a. s. l.; 23 Jan. 2011; D. Ouvrard leg.; light trap; MNHN (EH) 24120 • 1 ♂; Sakaraha, Zombitsy; P. Griveaud leg.; MNHN (EH) 24533 • 1 ♂; Sakaraha, Lambomakandro; Mar. 1956; MNHN (EH) 24540 • 1 ♀; Andobo, forêt d’Antsingy, det. Antsalova; 190 m a. s. l.; Feb. 1957; P. Griveaud leg.; MNHN (EH) 24534 • 1 ♀; rég. Atsimo-andrefana, Parc de Zombitse; 22 ° 53.002 ′ S 44 ° 41.956 ′ E; 783 m a. s. l ..; 18 Mar. 2006; A. Soulier-Perkins leg.; light trap; MNHN (EH) 24535 • 1 ♂; Province Toliara, Parc National de Zombitse, 19.8 km, 84 ° E Sakaraha; 22 ° 50 ′ 36 ″ S, 44 ° 42 ′ 36 ″ E; 770 m a. s. l.; 5 – 9 Feb. 2003; Fisher and Griswold leg.; M. Bucher det.; light trap; collection code: BLF 7508; CASENT 3007841 • 1 ♀; Madagascar, Province Toliara, Parc National de Zombitse, 19.8 km, 84 ° E Sakaraha; 22 ° 50 ′ 36 ″ S, 44 ° 42 ′ 36 ″ E; 770 m a. s. l.; 5 – 9 Feb. 2003; Fisher and Griswold leg.; M. Bucher det.; light trap; collection code: BLF 7508; CASENT 3007840 • 1 ♀; Madagascar, Fianarantsoa Province, Forêt d’Analalava, 29.6 km 280 ° W Ranohita; 22 ° 35 ′ 30 ″ S, 45 ° 07 ′ 42 ″ E; 700 m a. s. l.; 1 – 5 Feb. 2003; Fisher and Griswold leg.; M. Bucher det.; collection code: BLF 7387; CASENT 3007856.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D11FFFB39249B34CFB32FA54.taxon	description	Description TOTAL LENGTH. Between 8.5 and 11.5 mm. HEAD. Median part of vertex with very strong carina extending to anterior margin of head. Frons flat but slightly domed in middle. Postclypeus not exceeding anterior margin of head; longitudinal furrow, wide in its lower part. THORAX. Anterior part of pronotum relatively flat and rising up halfway back. Presence of thin and short carina in median part of pronotum, not reaching anterior or posterior margin. TEGMINA. Apical network with concave cells. Two well-marked bumps in anterior part: one between R and bifurcation of M and Cu; second on the bifurcation of R. MALE GENITALIA. Posterior margin of pygofer forming convex arc in apical part and becoming concave on basal margin. Subgenital plates short, wide with narrowing in middle section, slightly exceeding level of end of anal tube. Terminus forming small bulb serrated on apical part. Large sterno-lateral plates. Parameres developed anteriorly with long and thin extension, representing third of total length. On dorsal margin, presence of thumb-like protrusion covered with setae, followed by small tip-shaped extension. Presence of large thorn-like excrescence located internally in posterior part of parameres. Aedeagus developed anteriorly in single pointed protrusion and not exceeding height of pygofer; basal part sclerified but membranous ventrally. Apical part of aedeagus very thin and bifid. Posterior extension with a serrated apex; anterior extension two thirds length of posterior one. COLORATION. Dark brown or light brown with darker legs. In ventral view, abdomen red.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D101FFB19266B096FE43FCEB.taxon	description	Fig. 8	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D101FFB19266B096FE43FCEB.taxon	type_taxon	Type species Pogonorhinella madagascariensis Schmidt, 1910 by monotypy.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D101FFB19266B096FE43FCEB.taxon	description	Synthetic description Pogonorhinella is a monospecific endemic genus of Madagascar which possesses a peculiar shape due to its small head and the narrow anterior margin of the pronotum. A synthesis based on Schmidt, 1910 description and material examined is presented below: HEAD. Small with short vertex. Postclypeus narrow with strong carina and striated on sides, producing obtuse angle in lateral view. Rostrum reaching posterior margin of metacoxae. Antennae long, first segment very large, prominent pedicel. THORAX. Anterior half of pronotum trapezoidal giving impression of neck. Hindlegs with one spine on metatibiae. TEGMINA. Tegmina longer than wide. MALE GENITALIA. Male genitalia without sterno-lateral plates. Aedeagus longer than pygofer height.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D101FFB19266B096FE43FCEB.taxon	distribution	Distribution (Fig. 8 H) Endemic genus of Madagascar.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D101FFB19266B096FE43FCEB.taxon	discussion	Number of species: 1.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D103FFB691AEB643FE66FD97.taxon	materials_examined	Material examined MADAGASCAR • 1 ♂; Province de Toamasina, Andasibe; 18 ° 53.410 ′ S, 48 ° 23.881 ′ E; 1049 m a. s. l.; 5 Nov. 2011; A. Soulier-Perkins leg.; MNHN (EH) 24119 • 1 ♂; Province de Toamasina, Andasibe; 18 ° 57.723 ′ S, 48 ° 25.716 ′ E; 940 m a. s. l.; 8 Nov. 2011; A. Soulier-Perkins leg.; MNHN (EH) 24521 • 1 ♂; forêt d’Amboasary; 1300 m a. s. l.; Jan. 1961; P. Griveaud leg.; MNHN (EH) 24520 • 1 ♀; Province de Toamasina, Andasibe; 18 ° 53.021 ′ S, 48 ° 25.792 ′ E; 951 m a. s. l.; 4 Nov. 2011; A. Soulier-Perkins leg.; MNHN (EH) 24522 • 1 ♀; Region Amoron ï Maia, 35 km d’Ambositra, forêt alt. Ankazomivati; 20 ° 47.002 ′ S, 47 ° 10.964 ′ E; 984 m a. s. l.; 24 Mar. 2006; M Attié leg.; MNHN (EH) 24530 • 1 ♂; Province Diego-Suarez, Sakalava beach, dwarf littoral forest; 12 ° 15 ′ 46 ″ S, 49 ° 23 ′ 51 ″ E; 10 m a. s. l.; 7 – 22 Apr. 2001; R. Harin’Hala leg.; malaise trap; MA- 0104 B- 06; CASENT 3001902 • 1 ♀; Toliara Province, Rés. spéciale d’Ambohijanahary, forêt d’Ankazotsihitafototra, 35.2 km 312 ° NW Ambaravaranala; 18 ° 16 ′ 00 ″ S, 45 ° 24 ′ 24 ″ E; 1050 m a. s. l.; 13 – 17 Jan. 2003; Fisher and Griswold leg.; malaise trap montane rainforest; collection code BLF 7023; CASENT 3007937 • 1 ♀; Province Diego-Suarez, Montaigne français; 12 ° 15 ′ 8 ″ S, 49 ° 38 ′ 51 ″ E; 150 m a. s. l.; 15 Feb. – 6 Mar. 2001; R. Harin’Hala leg.; malaise along forested limestone ridge; MA- 01 - 06 - 06; CASENT 3004723 • 1 ♀; Province d’Antananarivo, 7 km SE of Andasibe National Park headquarters; 18 ° 57.76 ′ S, 48 ° 27.16 ′ E; 1050 m a. s. l.; 9 – 23 Apr. 2001; R. Harin’Hala leg.; malaise trap in tropical rainforest; MA- 01 - 08 A- 06; CASENT 3001997 • 1 ♀; Fianarantsoa province, forêt d’Atsiraamiaty, 7.6 km 285 ° WNW Itremo; 20 ° 35 ′ 36 ″ S, 46 ° 33 ′ 48 ″ E; 1550 m a. s. l.; 22 – 26 Jan. 2003; Fisher and Griswold; malaise trap montane rainforest; collection code BLF 7155; CASENT 3008211.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D103FFB691AEB643FE66FD97.taxon	description	Description TOTAL LENGTH. Between 6.5 and 7.6 mm. HEAD. Small head and short vertex. From dorsal view, head 1.5 times wider than long in midline between eyes. Antennae long with first segment broad and large pedicel. Postclypeus very slim. TEGMINA. Tegmina long and narrow. MALE GENITALIA. Subgenital plates bit longer than pygofer height. Subgenital plates with large base becoming thinner until last third; dorsal and ventral margin parallel and curved dorsally, ending in flat junction. Dorsal and ventral margin of parameres mainly parallel with large spine on dorsal margin. External extremity with two hooks shape, one curving dorsally, other one ventrally joining two margins. Aedeagus really long and slender, measuring around three times as long as pygofer height and curving strongly dorsally. Apical part of aedeagus bifid with ventral thin spine and spoon-shaped dorsal one covering ventral spine. COLORATION. Dorsal body all black or dark blue from the anterior extremity of vertex to end of tegmina. Ventral part of head and coxa orange or ochre. Thorax and abdomen black. Legs black except for first half of each femur.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D104FFB69240B7CCFDA8FB19.taxon	type_taxon	Type species Rhinaulax analis (Fabricius, 1794).	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D104FFB69240B7CCFDA8FB19.taxon	description	Synthetic description Rhinaulax is a small genus with only three species described. Among them, Rhinaulax limbata, described by Signoret in 1860 is from Madagascar. Since its description, Jacobi (1917) and Lallemand (1949) mentioned this species briefly in their works as present in Madagascar based on the original description. Signoret mentioned neither specific material nor the locality of the type specimens, which indeed seems to be unfindable, as suggested by Jacobi.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D104FFB69240B7CCFDA8FB19.taxon	discussion	Number of species: 3, 1 in Madagascar.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D104FFB791A0B045FA8AF9A4.taxon	description	urn: lsid: zoobank. org: act: 25 F 2 E 561 - 3864 - 4 E 4 D- 9085 - BBB 74 D 41641 E Figs 9 – 13	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D104FFB791A0B045FA8AF9A4.taxon	type_taxon	Type species Literna rugosa (Fallou, 1890).	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D104FFB791A0B045FA8AF9A4.taxon	diagnosis	Diagnosis Postclypeus strongly compressed laterally with the presence of a deep and narrow furrow on the median part; medial margin of postclypeus composed of two very thick carinae. Very pronounced tegmina veins forming numerous cavities and bumps on the general surface of the tegmina. Presence of a strongly pronounced lobe on the dorso-posterior margin of the pygofer below the anal tube. This very typical character is only observed in Soulieriana gen. nov. and separates it from all other genera within Cercopidae.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D104FFB791A0B045FA8AF9A4.taxon	etymology	Etymology This new genus is named after the last name of our hemipterist colleague Adeline Soulier-Perkins with the postfix ‘ ana’ in reference to Amberana and Bourgoinrana. The gender is feminine.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D104FFB791A0B045FA8AF9A4.taxon	description	Description HEAD. Head wider than long. Ocelli closer to each other than to compound eyes and separated by strong carina extending over the frons to anterior margin of head. Postclypeus compressed laterally; in frontal view, presence of deep and narrow median furrow bordered by edges and forming two clearly bulges striated laterally. Rostrum reaching mesocoxae. THORAX. Hindlegs with one spine on metatibiae. TEGMINA. Tegmina longer than wide with large number of bumps, making them strongly embossed; cells of apical network very concave. MALE GENITALIA. With sterno-lateral plates. Pygofer not straight in its dorsoposterior part; presence of a lobe clearly identifiable just below anal tube. Anal tube with sparsely sclerified area on anterior part.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D104FFB791A0B045FA8AF9A4.taxon	distribution	Distribution (Figs 9 H, 10 H, 11 H, 12 I, 13 I) Endemic genus of Madagascar.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D104FFB791A0B045FA8AF9A4.taxon	discussion	Number of species: 5.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D105FFB5925EB3E0FB45F9CB.taxon	description	Fig. 9	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D105FFB5925EB3E0FB45F9CB.taxon	materials_examined	Material examined MADAGASCAR • 1 ♂; Province de Toliara, massif du Makay, forêt sèche “ humide ”; 21 ° 37 ′ 41.6 ″ S, 45 ° 00 ′ 13.7 ″ E; 203 m a. s. l.; 23 Jan. 2011; D. Ouvrard leg.; MNHN (EH) 24704 • 1 ♂; Province de Toliara, massif du Makay, forêt sèche; 21 ° 36.236 ″ S, 45 ° 06.464 ″ E; 243 m a. s. l.; 12 Jan. 2011; D. Ouvrard leg.; MNHN (EH) 24541 • 1 ♂; Province de Toliara, massif du Makay, forêt sèche; 21 ° 36 ′ 236 ″ S, 45 ° 06 ′ 464 ″ E; 243 m a. s. l.; 12 Jan. 2011; D. Ouvrard leg.; MNHN (EH) 24700 • 1 ♀; Province de Toliara, massif du Makay forêt sèche; 21 ° 36 ′ 236 ″ S, 45 ° 06 ′ 464 ″ E; 243 m a. s. l.; 12 Jan. 2011; D. Ouvrard leg.; MNHN (EH) 24696 • 1 ♀; Province de Toliara, massif du Makay forêt sèche; 21 ° 36 ′ 236 ″ S, 45 ° 06 ′ 464 ″ E; 243 m a. s. l.; 12 Jan. 2011; D. Ouvrard leg.; MNHN (EH) 24697 • 1 ♂; Province d’Antsiranana, Forêt d’Anabohazo, 21.6 km 247 ° WSW Macromandia; 14 ° 18 ′ 32 ″ S, 47 ° 54 ′ 52 ″ E; 120 m a. s. l.; 11 – 16 Mar. 2001; Fisher and Griswold leg.; malaise trap in tropical dry forest; California Academy of Sciences; collecting code: BLF 336; CASENT 3001856 • 1 ♂; Province d’Antsiranana, Ampasindava, Forêt d’Ambilanivy, 3.9 km 181 ° S Ambaliha; 13 ° 47 ′ 55 ″ S, 48 ° 9 ′ 42 ″ E; 600 m a. s. l.; 4 – 9 Mar. 2001; Fischer and Griswold leg.; malaise trap in rainforest; California Academy of Sciences; collection code: BLF 3251; CASENT 3001283.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D105FFB5925EB3E0FB45F9CB.taxon	description	Description This species was first described by Stål (1866) from a single female specimen. Here, we propose its integration into the genus Soulierana Bucher & Bouteille gen. nov. with the description of the male genitalia for the first time. TOTAL LENGTH. Between 5.2 and 5.5 mm. HEAD. Median part of vertex relatively flat, forming very obtuse angle with postclypeus. Median furrow on postclypeus wide, deep, beginning below front margin of head and straight at upper extremity. Lateral bulges very thin and comparable to two carinae, parallel and keeping same width over entire length; in dorsal view, postclypeus slightly exceeding anterior margin of head; in lateral view, postclypeus convex and forming obtuse angle slightly pronounced. THORAX. Scutellum relatively flat. TEGMINA. Three bumps clearly distinguishable on each tegmen. Apical network not dense with fewer than 15 concave cells per tegmen. MALE GENITALIA. Subgenital plates longer than length of pygofer, margin wider on first third and curved dorsally on apical third; on apical margin, flattened internal part covered with setae forming rounded protuberance. Sterno-lateral plates greatly exceeding posterior margin of pygofer. Parameres inserted almost entirely in pygofer and not very visible in lateral view, their dorsal margin bearing dorsal extension, posterior tip forming spine hook-shaped clearly visible in dorsal view. Aedeagus developed anteriorly in one-pointed protrusions; ventral margin forming convex arc and going straight up linearly towards dorsal part. Aedeagus bifid on its apical part; posterior extension with thin serrated apex, on last third; anterior dorsal extension with rounded apex representing one third length of posterior one. COLORATION. Specimens entirely black except for thorax ochre-yellow in ventral view.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D107FFBB9252B30AFD06F827.taxon	description	Fig. 10	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D107FFBB9252B30AFD06F827.taxon	materials_examined	Material examined Lectotype MADAGASCAR • ♀; collection Fallou; MNHN (EH) 24695. Paralectotype MADAGASCAR • 1 ♀; collection Fallou; MNHN (EH) 24694. Other material MADAGASCAR • 1 ♂; Province de Toamasina Andasibe, forêt humide, s / Hisbiscus; 18 ° 58.871 ′ S, 48 ° 25.202 ′ E; 944 m a. s. l.; 9 Nov. 2011; A. Soulier-Perkins leg.; MNHN (EH) 24118 • 1 ♂; Province de Toamasina Andasibe, forêt humide, s / Hisbiscus; 18 ° 58.871 ′ S, 48 ° 25.202 ′ E; 944 m a. s. l.; 9 Nov. 2011; A. Soulier-Perkins leg.; MNHN (EH) 24634 • 1 ♂; Province de Toamasina Andasibe, forêt humide, s / Hisbiscus; 18 ° 58.871 ′ S, E 48 ° 25.202 ′ E; 944 m a. s. l.; 9 Nov. 2011; A. Soulier-Perkins leg.; MNHN (EH) 24698 • 1 ♂; Province de Toamasina Andasibe, forêt humide, s / Hisbiscus; 18 ° 58.871 ′ S, 48 ° 25.202 ′ E; 944 m a. s. l.; 9 Nov. 2011; A. Soulier-Perkins leg.; MNHN (EH) 24699 • 1 ♂; Province de Toamasina Andasibe, forêt humide, s / Hisbiscus; 18 ° 58.871 ′ S, 48 ° 25.202 ′ E; 944 m a. s. l.; 9 Nov. 2011; A. Soulier-Perkins leg.; MNHN (EH) 24703.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D107FFBB9252B30AFD06F827.taxon	description	Description TOTAL LENGTH. Between 9.0 and 10.0 mm. HEAD. Median part of vertex strongly domed forming an obtuse angle with postclypeus. Frons concave in its middle. Postclypeus not exceeding anterior margin of head. Furrow on postclypeus wide, rounded at its upper extremity. Lateral bulges parallel, narrower than width of furrow and keeping same width over their entire length. In lateral view, postclypeus convex, forming an obtuse angle slightly pronounced. Rostrum reaching mesocoxae. THORAX. Scutellum thick and main dimple deep; tip of scutellum slightly raised. TEGMINA. Tegmina 2.5 times as long as wide with many tubers all over surface; anal vein on clavus strongly wave-shaped. Apical network dense with more than 20 concave cells per tegmina. MALE GENITALIA. Under anal lobe, posterior margin of pygofer forming convex arc; margin becoming concave on ventral part. Subgenital plates shorter than length of pygofer; in lateral view, dorsal margin forming lobe on anterior part. Apical extremity of subgenital plates folded inwards with flattened internal part covered with setae. Sterno-lateral plates extend from pygofer in tapered shape; slightly dorsally oriented with convex apex. Dorsal margin of parameres bearing thin dorsal extension and posterior area excavated in its middle forming two separate bulbs. Aedeagus developed anteriorly in one-pointed protrusion at end of convex arc. Aedeagus bifid on apical part; posterior extension with serrated apex on last quarter. Anterior dorsal extension with a rounded apex representing one quarter length of posterior one. COLORATION. Head red with end of rostrum and pedicels black. Pronotum and tegmina red; apical network sometimes with black colouration. In ventral view, thorax and abdomen fully red. Red legs with tibias fore and mid-leg black; all tarsi black.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D107FFBB9252B30AFD06F827.taxon	discussion	Nomenclatural note The type specimens designated by Fallou in the monograph of Monecphora rugosa were conserved in his personal collection located in the MNHN. The number of type specimens was not mentioned in his original publication. Nevertheless, in Fallou’s collection, the pinholes at the bottom of the collection boxes correspond to the number of specimens referenced. Only two female specimens were placed under the pinned label of Monecphora rugosa, with no supernumerary pinholes, suggesting that only these two specimens were considered to be the types. We choose to designate the better-shaped specimen as lectotype and the second one as paralectotype.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D10AFFBE9242B462FE0DFCE2.taxon	description	urn: lsid: zoobank. org: act: 27 ACD 9 DF- 474 C- 408 C- 990 E- 71 D 6 AFEE 05 A 8 Fig. 11	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D10AFFBE9242B462FE0DFCE2.taxon	diagnosis	Diagnosis In general shape, Soulierana bigidea Bucher gen. et sp. nov. is most similar to S. rugosa. It is distinguished by the combination of the following characters: anterior part of the frons protruding and raised upward; median furrow on the postclypeus very tight and lateral bulges wide; colouring ochre in dorsal view; colouring green and pale pink in ventral view.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D10AFFBE9242B462FE0DFCE2.taxon	etymology	Etymology This specific epithet is derived from the name of the band The Big Idea, which was regularly listened to during the writing of this article and as a tribute to Manon Bucher's childhood friends.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D10AFFBE9242B462FE0DFCE2.taxon	materials_examined	Type material Holotype MADAGASCAR • 1 ♂; Province Antsiranana, Marojejy Nat’l Park, 5 km W of Manantenina village, Camp Mantella; 14 ° 26.29 ′ S, 49 ° 46.44 ′ E; 490 m a. s. l.; 18 – 26 Sep. 2005; M. Irwin and R. Harin’Hala leg.; malaise trap, low altitude, rainforest; California Academy of Sciences; MA 31 - 31; CASENT 8107935. Paratypes MADAGASCAR • 1 ♂; same collection data as for holotype; 26 Sept. – 4 Oct. 2005; MA- 31 - 32; CASENT 8107937; MNHN (EH) 24707 • 1 ♂; same collection data as for holotype; 26 Sept. – 4 Oct. 2005; MA 31 - 32; CASENT 8107938; MNHN (EH) 24706 • 1 ♂; same collection data as for holotype; 26 Sept. – 4 Oct. 2005; California Academy of Sciences; MA 31 - 32; CASENT 8107943; MNHN (EH) 24705 • 1 ♀; same collection data as for holotype.; 11 – 18 Feb. 2005; MA- 31 - 11; CASENT 8107906; MNHN (EH) 24701 • 1 ♂; same collection data as for holotype; 18 – 30 May 2005; MA 31 - 22; CASENT 8107921 • 1 ♂; same collection data as for holotype; 28 Aug. – 11 Sep. 2005; MA- 31 - 29; CASENT 8107924 • 1 ♂; same collection data as for holotype; 14 – 22 Oct. 2005; MA- 31 - 34; CASENT 8107949 • 1 ♀; same collection data as for holotype; 18 – 25 Mar. 2005; MA- 31 - 16; CASENT 8107914.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D10AFFBE9242B462FE0DFCE2.taxon	description	Description TOTAL LENGTH. Between 7.0 and 7.6 mm (holotype is 7.6 mm). HEAD. Head in dorsal view 1.3 times wider eye to eye than long in midline. Median part of vertex moderately domed, flattening slightly on lower part. Frons concave, bearing very marked median carina in middle part; anterior margin of frons exceeding anterior margin of vertex giving impression that superior part of postclypeus is raised up. Postclypeus not exceeding anterior margin of head. Furrow on postclypeus very narrow, thinner than lateral bulges which are much wider than in other species of this genus and narrowing ventrally. In lateral view, postclypeus barely convex and almost straight down to anteclypeus to form very pronounced obtuse angle. THORAX. Pronotum in dorsal view 1.9 times as wide as long in midline. Anterior pronotum with small median carina, sometimes not distinguishable, on posterior part. Scutellum wide and main dimple deep. TEGMINA. Veins clearly distinct and with few sharp bumps on anterior part. Apical network not dense with less than 15 concave cells per tegmen. MALE GENITALIA. Pygofer much wider in its dorsal than its ventral part; its basal apex forms protruding tip oriented downward. Subgenital plates long; in lateral view they are wider in first third and dorsal margin forms convex arc; apex protruding into sprouting extension on which flattened internal part is partially covered with setae. Sterno-lateral plates exceed pygofer; ventral margin shrinking and joining dorsal margin to form strong convex extremity. Dorsal margin of parameres bearing thin dorsal extension; posterior area excavated to form three separate parts, middle one being small thorn-like excrescence. Aedeagus developed anteriorly in one-pointed protrusion which is wide and rounded at apex; ventral margin declining in convex arc to form basal protuberance and going up towards apical part in three discernable waves. Aedeagus bifid on apical part; posterior extension with serrated apical two-thirds, serrations wide and spaced. Anterior dorsal extension with rounded apex, representing one third length of posterior one. COLORATION. In dorsal view head, thorax and tegmina ochre with exception of insertion point of tegmina green. In ventral view, thorax green, abdomen pale pink orange spotted with green. Legs ochre apart from hindlegs light green.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D10CFFBC925FB622FD43FC45.taxon	description	urn: lsid: zoobank. org: act: E 8 C 7 DD 9 F- 1 E 93 - 4105 - B 061 - ED 7900 BA 2 BC 8 Fig. 12	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D10CFFBC925FB622FD43FC45.taxon	diagnosis	Diagnosis In general shape, Soulierana claudinae Bouteille gen. et sp. nov. is most similar to S. bigidea Bucher gen. et sp. nov. This new species is distinguished by the combination of the following characters: the colouring is golden except for the apex and the middle dot of the tegmina which are brown, lateral bulges on the postclypeus are wide and spaced by a large furrow.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D10CFFBC925FB622FD43FC45.taxon	etymology	Etymology This specific epithet is derived from the name of E. Bouteille’s twin sister Claudine Bouteille.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D10CFFBC925FB622FD43FC45.taxon	materials_examined	Type material Holotype MADAGASCAR • 1 ♂; Province Diego-Suarez, Parc National, Montagne d’Ambre; 12 ° 31 ′ 13 ″ S, 49 ° 10 ′ 45 ″ E; 1125 m a. s. l.; 14 – 30 May 2001; R. Harin’Hala leg.; malaise trap; MA- 0101 D- 11; CASENT 3001255. Paratypes MADAGASCAR • 1 ♂; Province Diego-Suarez, Sakalava Beach, dwarf littoral forest; 12 ° 15 ′ 46 ″ S, 49 ° 23 ′ 51 ″ E; 10 m a. s. l.; 16 – 31 May 2001; R. Harin’Hala leg.; malaise trap, across sandy trail; MA- 01 - 04 B- 10; CASENT 3001220; MNHN (EH) 25189 • 1 ♂; Province Diego-Suarez, Sakalava Beach, dwarf littoral forest; 12 ° 15 ′ 46 ″ S, 49 ° 23 ′ 51 ″ E; 10 m a. s. l.; 16 – 31 May 2001; R. Harin’Hala leg.; malaise trap, across sandy trail; MA- 01 - 04 B- 10; CASENT 3001222.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D10CFFBC925FB622FD43FC45.taxon	description	Description TOTAL LENGTH. Between 7.0 and 7.2 mm (holotype is 7.0 mm). HEAD. Head in dorsal view 1.4 times wider eye to eye than long in midline. Head triangular, covered with golden silk. Central part of vertex salient and wide towards frons. Frons with median ridge. Ridges of postclypeus very wide. Space between two longitudinal ridges three times width of one ridge. Parallel ridges extending horizontally from furrows towards sides of postclypeus. THORAX. Pronotum in dorsal view 1.9 times as wide as long in midline, covered with golden silk. Dimples slightly darker than pronotal ground colour. Posterior margin bilobed. Scutellum with 2 small dimples along anterior margin and large central dimple. TEGMINA. Veins distinct forming few sharp bumps on anterior part; single brown dot present in middle, apex of tegmina brown. MALE GENITALIA. Pygofer much wider in dorsal than ventral part. Subgenital plates long; in lateral view, plates wider in first two thirds. Intern margin of apex covered with setae; basal apex forming tip oriented downward. Dorsal margin of parameres bearing two excrescences: rounded extension and tiny thin dorsal curved; another rounded excrescence in middle of parameres. Aedeagus bifid on apical part; posterior extension serrated on dorsal margin of last two-thirds, serrations fine and spaced, wider from base on first third and thinner on last two thirds. Anterior dorsal extension with protruding tip oriented ventrally on apex and one quarter length of posterior one. As in Soulierana bigidea Bucher gen. et sp. nov., ventral margin of aedeagus declining in convex arc, forming rounded protuberance, much longer than Soulierana bigidea and going up straight towards apical part with a single wave. Dorsal margin declining in concave arc to form pointed protuberance as long as ventral protuberance. COLORATION. Body, tegmina and legs gilded. Apex and middle dot of tegmina brown black. In dorsal view, two closed tegmina form smiley face.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D10EFF82925CB680FDB7FA73.taxon	description	urn: lsid: zoobank. org: act: A 8 AC 3 F 00 - 92 E 9 - 4 BFC- 92 E 1 - 5 C 16 D 0 C 398 F 7 Fig. 13	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D10EFF82925CB680FDB7FA73.taxon	diagnosis	Diagnosis This new species is easily distinguishable from all the other Soulierana Bucher & Bouteille gen. nov. by its small size (<7.0 mm) and its typical pale red colouration and the colouration of its legs, but also by the length of its long rostrum reaching the anterior margin of the abdomen, a pronotum 2.1 times as wide as long and short tegmina embracing the abdomen.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D10EFF82925CB680FDB7FA73.taxon	etymology	Etymology ‘ Kely ’ means ‘ small’ and ‘ mena ’ ‘ red’ in Malagasy, ‘ kelymena ’ therefore refers to the small size and colour of the species.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D10EFF82925CB680FDB7FA73.taxon	materials_examined	Type material Holotype MADAGASCAR • 1 ♂; Province Diego-Suarez, Montaigne Français; 12 ° 18 ′ 8 ″ S, 49 ° 38 ′ 51 ″ E; 150 m a. s. l.; 15 Feb. – 6 Mar. 2001; R. Harin’Hala leg.; malaise trap, along forested limestone ridge; MA- 01 - 06 - 06; CASENT 3004738. Paratypes MADAGASCAR • 1 ♂; Antsiranan Réserve Spéciale de l’Ankarana 22.9 km SW of Anivorano Nord; 12 ° 54 ′ 32 ″ S, 49 ° 6 ′ 35 ″ E; 80 m a. s. l.; 10 – 16 Feb. 2001; B. Fischer and C. Griswoid leg.; BLF 2857; CASENT 8080999; MNHN (EH) 4890 • 1 ♂; Province Diego-Suarez, Montagne d’Ambre; 12 ° 31 ′ 13 ″ S, 49 ° 10 ′ 45 ″ E; 1125 m a. s. l.; 11 Feb. – 4 Mar. 2001; R. Harin’Hala leg.; malaise trap; MA- 01 - 01 D- 04; CASENT 3001751; MNHN (EH) 4894 • 1 ♀; Province Diego-Suarez, Parc National Montagne d’Ambre; 12 ° 31 ′ S, 49 ° 11 ′ E; 975 m a. s. l.; 19 Mar. – 5 Apr. 2001; M. Irwin, E. I. Schlinger and R. Harin’Hala leg.; malaise trap; MA- 01 - 01 B- 07; CASENT 3001827; MNHN (EH) 4893 • 1 ♂; Province Diego-Suarez, Montaigne Français; 12 ° 18 ′ 8 ″ S, 49 ° 38 ′ 51 ″ E; 150 m a. s. l.; 15 Feb. – 6 Mar. 2001; R. Harin’Hala leg.; malaise trap, along forested limestone ridge; MA- 01 - 06 - 06; CASENT 3004724 • 1 ♀; Province Diego-Suarez, Parc National Montagne d’Ambre; 12 ° 30 ′ 52 ″ S, 49 ° 10 ′ 53 ″ E; 960 m a. s. l.; 12 Feb. – 4 Mar. 2001; R. Harin’Hala leg.; malaise trap; MA- 01 - 01 A- 07; CASENT 3001228 • 1 ♀; Province Diego-Suarez, Montaigne Français; 12 ° 18 ′ 8 ″ S, 49 ° 38 ′ 51 ″ E; 150 m a. s. l.; 15 Feb. – 6 Mar. 2001; R. Harin’Hala leg.; malaise trap, along forested limestone ridge; MA- 01 - 06 - 06; CASENT 3004733.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
E8713702D10EFF82925CB680FDB7FA73.taxon	description	Description TOTAL LENGTH. Between 6.3 and 6.7 mm (holotype is 6.7 mm). HEAD. Head in dorsal view 2.8 times wider eye to eye than long in midline, vertex short with anterior margin rounded. Ocelli closer to each other than to eyes. Antenna with large pedicel bearing long flagellum. Postclypeus with narrow central furrow strongly marked by keel on each side. Long rostrum reaching ventral anterior margin of abdomen. THORAX. Pronotum in dorsal view 2.1 times as wide as long in midline. Anterior margin rounded, with two central small dimples framed on each side by bigger one. Posterior margin slightly wave-shaped. Scutellum triangular with large anterior base and sharp posterior point. TEGMINA. Short tegmina, embracing the abdomen. Veins clearly distinct and few bumps on apical part where network is not dense with only few cells. MALE GENITALIA. Subgenital plates elongated and curved laterally to sharp apex wearing setae. Sterno-lateral plates slightly rounded. Aedeagus developed anteriorly in one-pointed protrusion; ventral margin declining in convex arc to form protuberance and going straight up to apical part. Parameres bearing rounded lobe on dorsal margin and with hook-shaped apical apex. Aedeagus bifid, posterior margin ending in acute apex bearing teeth on each side in its last third. Anterior margin with slim extension, long as one third of posterior one, ending in rounding apex. Anterior protrusion pointing into rounded tooth. COLORATION. Body pale red. Tegmina of same colouration with dark red veins and some of apical cells dark brown. Tibia and tarsus of first pair of legs dark brown. Midlegs with apical half of tibia and tarsus dark brown. Hindlegs with single black tibial spine; distal extremities of tibia and tarsus dark brown, rest of each pair of legs pinkish ochre.	en	Bucher, Manon, Bouteille, Emilien, Cesne, Maxime Le (2024): Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar. European Journal of Taxonomy 935: 166-202, DOI: 10.5852/ejt.2024.935.2555, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547
