identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
F20687EEFFA8EC302083FD0DF9936AFD.text	F20687EEFFA8EC302083FD0DF9936AFD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kuzicus Gorochov 1993	<div><p>Genus Kuzicus Gorochov, 1993</p><p>Diagnosis. Tenth abdominal tergite of male with large paired apical lobes. Male epiproct small. Male genitalia with large yoke-like epiphallus; apex with projections, either robust, articulated, denticulated or spine-like.</p><p>Discussion. Prior to this review, the genus consists of three subgenera and 14 species in total (Gorochov, 1998; Ingrisch &amp; Shishodia, 1998, 2000; Sänger &amp; Helfert, 2004, 2006a, 2006b; Ingrisch, 2006; Mao et al., 2009; Di et al., 2014). This is an Asian genus with its distribution ranging from Sumatra (Indonesia) north to Japan and China as well as India (Eades et al., 2015).</p><p>The discovery of more species (including those outside of this paper) with intermediate characters led to problems to the subgeneric system. Diagnostic characters previously used for defining subgenus need to be reexamined and a taxonomic revision (using both old and new material) is as such required (A. V. Gorochov, pers. comm.). Thus, the subgenus name for all species are tentatively omitted here.</p><p>Nevertheless, species of subgenus Parakuzicus lacks most important characters of the genus Kuzicus: male epiphallus without apical lobes (either robust unarticulated projections directed anteriorly towards head or denticulated and articulated projection directed posteriorly away from head). It also differs by male tenth abdominal tergite with apical lobe truncated and much shorter, cerci bifurcated from base into a medial (internal) and lateral (external) branch of roughly equal length. Thus, we propose that Parakuzicus should be more accurately considered a separate genus.</p></div>	https://treatment.plazi.org/id/F20687EEFFA8EC302083FD0DF9936AFD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tan, Ming Kai;Dawwrueng, Pattarawich;Artchawakom, Taksin	Tan, Ming Kai, Dawwrueng, Pattarawich, Artchawakom, Taksin (2015): Taxonomic review of Kuzicus Gorochov, 1993 (Tettigoniidae: Meconematinae), with two new species from Thailand and key to species. Zootaxa 3999 (2): 279-290, DOI: 10.11646/zootaxa.3999.2.7
F20687EEFFAEEC372083FF0CFDD96E0C.text	F20687EEFFAEEC372083FF0CFDD96E0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kuzicus	<div><p>Key to species of Kuzicus (for males only)</p><p>1. Tenth abdominal tergite with medial lobes diverging abruptly at the base from one another (Fig. 1I, 2K).......................................................................................... K. koeppeli Sänger &amp; Helfert, 2004</p><p>- Tenth abdominal tergite with medial lobes not diverging at the base from one another; if so, not as abruptly or converged apically (Figs. 1A–1H).................................................................................... 2</p><p>2. Tenth abdominal tergite with one or two pairs of apical lobes (Figs. 1B–1H, 2B–2J). Epiphallus with small ventro-subapical inflation with small denticles and a pair of small semi-articulated, spine-like appendices or lobules directed more or less posteriorly (forward from the apex) or laterally (Fig. 3 B–3J)........................................................ 4</p><p>- Tenth abdominal tergite with two pairs of apical lobes (Figs. 1A, 2A, 6C). Epiphallus without denticles and these appendiceslobules but with a pair of robust unarticulated projections directed anteriorly (backward from the apex) (Figs. 3 A, 6G)..... 3</p><p>3. Tenth abdominal tergite with median lobes with stout ventral teeth apically; with lateral lobes obtuse apically (Figs. 6 C, 6E) Cercus stout and short (Figs. 6 G). Epiphallus with a pair of tongue-shaped lateral processes pointing anteriorly at the apex; with posterior margin producing into a plate in the middle (instead of rounded) (Fig. 6 G).... K. pakthongchai Tan et al. sp. n.</p><p>- Tenth abdominal tergite with median lobes without tooth apically; with lateral lobes tapering apically (Figs. 1A, 2A). Cercus long and slender (Fig. 4 A). Epiphallus with a pair of tongue-shaped lateral processes pointing posteriorly at the apex; with posterior margin rounded (Fig. 3 A)...................................................... K. uvarovi Gorochov, 1993</p><p>4. Tenth abdominal tergite with two pairs of apical lobes (Figs. 1G, 2I)............... K. scorpioides Sänger &amp; Helfert, 2006</p><p>- Tenth abdominal tergite with one pair of apical lobes (Figs. 1B–1H, except 1G, 2B–2J, except 2I); if with two pairs, lateral lobes are small and truncated (Figs. 8 C, 8D)................................................................ 5</p><p>5. Tenth abdominal tergite with medial lobes modified or bifurcate at the apex (Figs. 1B, 1D, 1H, 2B, 2D, 2H, 2J)........... 6</p><p>- Tenth abdominal tergite with medial lobes simple, usually forming a flattened disc at the apex (Figs. 1C, 1E, 1F, 2C, 2E, 2F, 2G)................................................................................................. 9</p><p>6. Tenth abdominal tergite with medial lobes bifurcate at the apex (Figs. 1H, 2J). Epiphallus as shown in Fig. 3 J.................................................................................. K. suzukii (Matsumura &amp; Shiraki, 1908)</p><p>- Tenth abdominal tergite with medial lobes highly modified, forming structure (Figs. 1B, 1D, 2B, 2D, 2H). Epiphallus not as above............................................................................................... 7</p><p>7. Tenth abdominal tergite with apex of medial lobes with minute teeth and warts (Figs. 1B, 2B, 8C, 8E). Cercus with basal process, apex acute (Figs. 1B, 4B, 4C, 8F). Epiphallus at apex with one large tooth and numerous minute denticles on each side (Figs. 3 B, 8E, 8G)..................................................................................... 8</p><p>- Tenth abdominal tergite with apex of median lobes without minute teeth and warts (may have ventral spine-like process); with apex of medial lobes forming a hammer-head structure (in lateral view) (Figs. 1D, 2D). Cercus without basal process, apex with minute teeth (Fig. 4 F). Epiphallus at apex with two pairs of relatively large dentiform processes (Fig. 3 D)....................................................................................... K. compressus Han &amp; Shi, 2014</p><p>8. Tenth abdominal tergite with median lobes denticulated on the dorsal and posterior margins at the apex (Figs. 1B, 2B).......................................................................... K. aspercaudatus Sänger &amp; Helfert, 2006</p><p>- Tenth abdominal tergite with median lobes denticulated on the ventral margin at the apex only (Figs. 8 C, 8F)...................................................................................... K. multidenticulatus Tan et al. sp. n.</p><p>9. Cercus bifurcate at the apex; with ventral process stout, dorsal process slender (Figs. 4 D, 4E). Epiphallus broader and with articulated ventral processes small (Fig. 3 C)....................................... K. cervicercus (Tinkham, 1943)</p><p>- Cercus not bifurcate at the apex (Figs. 4 G–4K). Epiphallus more slender and/ or with articulated ventral processes longer (Figs. 3 E–3H)........................................................................................ 10</p><p>10. Median lobes of tenth abdominal tergite appears distinctively bulbous at the distal end in lateral view (Fig. 2G). Epiphallus with articulated ventral processes short and stout (Fig. 3 G)................ K. megaterminatus Ingrisch &amp; Shishodia, 1998</p><p>- Median lobes of tenth abdominal tergite slightly bulbous or tapering at the distal end in lateral view (Figs. 2E, 2F, 2H). Epiphallus with articulated ventral processes more slender (Figs. 3 E, 3F, 3H)...................................... 11</p><p>11. Epiphallus with dorsal articulation (in addition to ventral articulation) at the apex, but without denticulation (Fig. 3 H).................................................................................. K. multifidous Mao &amp; Shi, 2009</p><p>- Epiphallus without dorsal articulation (only ventral articulation) at the apex, but with denticulation (Figs. 3 E, 3F)........ 12</p><p>12. Epiphallus with ventral articulation long and slender (Fig. 3 E)........................... K. denticulatus (Karny, 1926)</p><p>- Epiphallus with ventral articulation spine-like (Fig. 3 F).................. K. denticuloides Kevan &amp; Jin, 1993 comb. nov.</p></div>	https://treatment.plazi.org/id/F20687EEFFAEEC372083FF0CFDD96E0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tan, Ming Kai;Dawwrueng, Pattarawich;Artchawakom, Taksin	Tan, Ming Kai, Dawwrueng, Pattarawich, Artchawakom, Taksin (2015): Taxonomic review of Kuzicus Gorochov, 1993 (Tettigoniidae: Meconematinae), with two new species from Thailand and key to species. Zootaxa 3999 (2): 279-290, DOI: 10.11646/zootaxa.3999.2.7
F20687EEFFAEEC342083F950F9776C77.text	F20687EEFFAEEC342083F950F9776C77.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kuzicus pakthongchai Tan, Dawwrueng, Artchawakom	<div><p>Kuzicus pakthongchai Tan, Dawwrueng, Artchawakom, new species</p><p>Figs. 5, 6</p><p>http://lsid.speciesfile.org/urn:lsid: Orthoptera .speciesfile.org:TaxonName:470490</p><p>Material examined. Holotype (male): Thailand, Nakhon Ratchasima, Sakaerat Environmental Research Station, open area with light trap, N14.50765, E101.92772, 446.7± 9.7 m, dry evergreen forest, attracted to light trap, coll. M. K. Tan, H. Yeo &amp; S. T. Toh, 27 June 2014, 2220 hours (SERS.14.113) (ZRC).</p><p>Paratypes: 2 female: same locality as holotype, headquarter, N14.51039, E101.93057, 445.9± 8.5 m, mix vegetation, attracted to light, coll. M. K. Tan, H. Yeo &amp; S. T. Toh, 29 June 2014, 2330 hours (SERS.14.131); open area with light trap, N14.50763, E101.92760, 424.0± 4.1 m, dry evergreen forest, attracted to light trap, coll. M. K. Tan, H. Yeo &amp; S. T. Toh, 2 July 2014, 2300 hours (SERS.14.164) (all ZRC).</p><p>Diagnosis. This species is similar to the type species Kuzicus uvarovi by its colouration, yellow green with small brown spots on tegmen and one large spot at stridulatory field. This species differs from K. uvarovi by male tenth abdominal tergite with lateral lobes obtuse apically (instead of tapering); male epiphallus with a pair of tongue-shaped lateral (slightly sclerotized) processes pointing anteriorly at the apex (instead of pointing posteriorly); with posterior margin producing into a trilobous (also slightly sclerotized) plate in the middle (instead of rounded).</p><p>Description. Rather large-sized Meconematini but habitus typical of the genus (Figs. 5 A, 5B). Eyes globular and protruding. Fastigium verticis conical, small (about slightly shorter than scapus length), with very weak median sulcus (Fig. 6 A). Last (apical) segment of maxillary palpus equal in length to the fourth (subapical) segment, slightly widened apically. Pronotum covering the base of mirror of tegmen, discus curved into lateral lobe, transverse sulcus in mid length (Fig. 6 B). Anterior margin of pronotal disc slightly convex, posterior margin triangularly rounded (Fig. 6 A). Margin of lateral lobes of pronotum slightly undulated, humeral sinus very weak. Ventral margin of pronotal lateral lobe broadly rounded (Fig. 6 B). Thoracic auditory spiracle (= thoracic foramen) large and oval-shaped (Fig. 6 B). Parapterous, tegmen extending behind the hind knees, hindwings distinctly longer. Procoxal spur short and slender. Both tympana open. Protibiae with 5 outer and 4 inner subapical spines and 1 pair of apical spurs. Mesotibiae with 4 outer and 3 inner subapical spines and 1 pair of apical spurs. Metatibiae ventrally and dorsally with numerous outer and inner spines as well as 2 ventral and 1 dorsal pairs of apical spurs.</p><p>Male. Tenth abdominal tergite with a pair of median lobes and a pair of lateral lobes. Median lobes slender basally with margin between lobes truncated; after basal third, expanded abruptly (about 90°) along external margin to form a plate which bent ventrad laterally; plate tapers apically and rotate from a horizontal to a vertical plate (Fig. 6 C). At the apex, median lobes bent ventrad and turned slightly anteriorly and internally; forming two sclerotized teeth ventrally (Figs. 6 C–6G). Median lobes remain widely spaced throughout, particularly setose along internal margin (Fig. 6 C). Lateral lobes of tenth abdominal tergite flattened, shorter and smaller with apex obtuse (Figs. 6 C, 6E). Epiproct concealed under tenth abdominal tergite. Cerci stout and short; basally slightly swollen, with a short and blunt basal tooth externally; slightly narrower in the middle, slightly swollen and bent internally after middle (Figs. 6 E, 6G). Interiorly, cerci with a sclerotized process in the middle: longer and flattened, bladelike and bent basally (Fig. 6 G). At the apex, cerci with a short acute sclerotized tooth along dorso-inner margin (Figs. 6 G). Yoke-like epiphallus typical of the genus, but with distal appendages only (Figs. 6 F, 6G). Epiphallus transverse at the base, with a short shaft that widens posteriorly; at the apex with a pair of tongue-shaped lateral (slightly sclerotized) processes pointing anteriorly, with posterior margin producing into a trilobous (also slightly sclerotized) plate in the middle (Fig. 6 G). Subgenital plate with styli relatively long and slender, gently curved inwards (Fig. 6 F).</p><p>Female. Tenth abdominal tergite with posterior margin distinctly concave in the middle, truncated laterally (Fig. 6 H). Epiproct small and triangular (Fig. 6 H). Cerci slender and short, slightly swollen in the middle before tapering to an acute apex (Fig. 6 H). Subgenital plate transverse, much wider than long; with posterior margin feebly notched in the middle (Fig. 6 I). Ovipositor long with base slightly swollen; feebly sinuated (Fig. 6 J). Valves smooth with acute apex; ventral valve forming a small hook at the apex (Fig. 6 J). At the base, with two small sclerotized tooth ventrally (Fig. 6 I).</p><p>Colouration. Green when alive; white to ochre in alcohol (Figs. 5 A, 5B). Fastigium darken yellow brown with margin along vertex black; median sulcus distinctly pale (Fig. 6 A). Scapus with inner half yellow brown; pedicel with anterior and posterior margin black (Fig. 6 A). Antennae yellow brown with some segments black (Fig. 6 A). Dorsal disc of pronotum darkened yellow in the middle, darker in the anterior half than posterior half of disc; anterior of transverse sulcus with two elongated black spot diverging anteriorly; disc with anterior and posterior margin black in the middle (Fig. 6 A). Tegmen with one large spot at stridulatory field; with some small rounded infumated brown spots throughout, randomly and widely distributed; with cells near posterior margin infumated dark.</p><p>Measurements. See Table 1.</p><p>Etymology. This species is named after Pak Thong Chai District in which Sakaerat Environmental Research Station spans.</p></div>	https://treatment.plazi.org/id/F20687EEFFAEEC342083F950F9776C77	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tan, Ming Kai;Dawwrueng, Pattarawich;Artchawakom, Taksin	Tan, Ming Kai, Dawwrueng, Pattarawich, Artchawakom, Taksin (2015): Taxonomic review of Kuzicus Gorochov, 1993 (Tettigoniidae: Meconematinae), with two new species from Thailand and key to species. Zootaxa 3999 (2): 279-290, DOI: 10.11646/zootaxa.3999.2.7
F20687EEFFADEC392083FA69F9E76A60.text	F20687EEFFADEC392083FA69F9E76A60.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kuzicus multidenticulatus Tan, Dawwrueng, Artchawakom	<div><p>Kuzicus multidenticulatus Tan, Dawwrueng, Artchawakom, new species</p><p>Figs. 7, 8</p><p>http://lsid.speciesfile.org/urn:lsid: Orthoptera .speciesfile.org:TaxonName:470491</p><p>Material examined. Holotype (male): Thailand, Nakhon Ratchasima, Sakaerat Environmental Research Station, open area with light trap, dry evergreen, attracted to light trap, 400-450m, 13 September 2014, coll. P. Dawwrueng, N. Makbun &amp; T. Dowwiangkan (THNHM).</p><p>Paratype: 1 female, same locality and information as holotype (THNHM).</p><p>Diagnosis. This species is very similar to Kuzicus aspercaudatus Sänger &amp; Helfert, 2006 by median lobes of male tenth abdominal tergite long and slender, bent ventrad to form two lobes; and male epiphallus with apex minutely denticulated but differ by male tenth abdominal tergite with small lateral apical lobes (absent in K. aspercaudatus); median lobe of male tenth abdominal tergite with a row of smaller teeth along the ventral margin at the apex (instead of denticulated on the dorsal and posterior margins in K. aspercaudatus).</p><p>Description. Habitus typical of the genus (Fig. 7 A, 7B). Description similar to Kuzicus pakthongchai sp. n. by rather large size, eyes globular and protruding. Fastigium verticis conical but with inconspicuous median sulcus (Fig. 8 A). Last (apical) segment of maxillary palpus slightly shorter than the fourth (subapical) segment, also slightly widened apically (Figs. 8 A, 8B). Pronotal disc with anterior and posterior margin similar to Kuzicus pakthongchai sp. n. and covering base of mirror tegmen (Fig. 8 A). Margin of lateral lobes of pronotum slightly undulated (Fig. 8 B). Thoracic auditory spiracle (= thoracic foramen) large and inverted pyriform (Fig. 8 B). Parapterous, tegmen extending behind the hind knees, hindwings distinctly longer. Procoxal spur short. Meso- and metatibiae with 5 outer and 4 inner subapical spines.</p><p>Male. Tenth abdominal tergite with a pair of median lobes and lateral lobes. Median lobes long and slender; converge slightly towards one another in the middle before diverging apically, never in contact with one another; each broaden apically into a bulbous apex that contacts one another (Figs. 8 C, 8D). At the apex, median lobes bent ventrad to form two lobes: basal lobe smaller, produced with a small and slender external tooth; apical lobe larger, produced into a larger internal tooth with a row of smaller teeth along the ventral margin (Figs. 8 D, 8E). Lateral lobes of tenth abdominal tergite long and slender, reaching slightly before middle of median lobes, with apex obtuse (Figs. 8 C, 8D). Epiproct concealed under tenth abdominal tergite. Cerci with basal half swollen and robust, internally with a sclerotize spine-like process; then curve internally and taper into an acute and sclerotized apex (Figs. 8 C, 8F). Yoke-like epiphallus typical of the genus, but with distal appendages only (Figs. 8 E, 8G). Epiphallus transverse at the base, with a very short and broad shaft that widens posteriorly; at the apex with a pair of tooth-like sclerotized processes pointing posteriorly; and with fairly thick bristles. Posterior margin producing into a trilobous plate in the middle; with lateral lobe denticulated and heavily sclerotized (Figs. 8 E, 8G). Subgenital plate with styli relatively short and robust, gently curved outwards (Fig. 8 E).</p><p>Female. Tenth abdominal tergite with posterior margin distinctly concave in the middle, truncated laterally; more deeply excised than Kuzicus pakthongchai sp. n. in the middle (Fig. 8 H). Epiproct and cerci similar to Kuzicus pakthongchai sp. n. (Fig. 8 H). Subgenital plate trapezoid (Fig. 8 I). Ovipositor straight and very long, reaching apex of tegmen; with base slightly swollen (Fig. 8 J). Valves smooth with acute apex; ventral valve forming a small hook at the apex (Fig. 8 J). At the base, with two small sclerotized tooth ventrally (Fig. 8 J).</p><p>Colouration. Unicolorous green when alive; white to ochre when preserved (Figs. 7 A, 7B). Tegmen also with some small rounded infumated brown spots throughout, randomly and widely distributed; with cells near posterior margin infumated dark.</p><p>Measurements. See Table 2.</p><p>Etymology. The species name refers to the numerous ventral teeth male at the apex of the median lobes of tenth abdominal tergite.</p></div>	https://treatment.plazi.org/id/F20687EEFFADEC392083FA69F9E76A60	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tan, Ming Kai;Dawwrueng, Pattarawich;Artchawakom, Taksin	Tan, Ming Kai, Dawwrueng, Pattarawich, Artchawakom, Taksin (2015): Taxonomic review of Kuzicus Gorochov, 1993 (Tettigoniidae: Meconematinae), with two new species from Thailand and key to species. Zootaxa 3999 (2): 279-290, DOI: 10.11646/zootaxa.3999.2.7
F20687EEFFA0EC392083FC02FBF36C1A.text	F20687EEFFA0EC392083FC02FBF36C1A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kuzicus denticuloides (Kevan 1993) Kevan 1993	<div><p>Kuzicus denticuloides (Kevan, 1993), new combination</p><p>(Figs. 2F, 3F, 4I)</p><p>http://lsid.speciesfile.org/urn:lsid: Orthoptera .speciesfile.org:TaxonName:470492</p><p>Remarks. Kevan described this species as Xiphidiopsis in (Kevan &amp; Jin, 1993). Ingrisch &amp; Shishodia (1998) mentioned that this species should belong to the genus Kuzicus instead.</p></div>	https://treatment.plazi.org/id/F20687EEFFA0EC392083FC02FBF36C1A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tan, Ming Kai;Dawwrueng, Pattarawich;Artchawakom, Taksin	Tan, Ming Kai, Dawwrueng, Pattarawich, Artchawakom, Taksin (2015): Taxonomic review of Kuzicus Gorochov, 1993 (Tettigoniidae: Meconematinae), with two new species from Thailand and key to species. Zootaxa 3999 (2): 279-290, DOI: 10.11646/zootaxa.3999.2.7
