taxonID	type	description	language	source
F10F7213FFC9B344FF76EF7CECE5FDC0.taxon	materials_examined	Type: — BRAZIL. Espírito Santo: Itarana, Alto Jatiboca, propriedade de Frederico Seidler, 20 ° 00 ’ 51.0 ” S, 40 ° 54 ’ 42.6 ” W, 816 m, 27 July 2009, A. Salino 14538, M. Megale & A. J. Arruda (holotype BHCB, isotype UC). Plants terrestrial. Rhizomes erect, caudex to 16 cm long x 2.3 – 3.5 cm wide, scales lanceolate, castaneous, 2.2 – 4.3 mm long, with furcate and stellate hairs 0.10 – 0.18 mm long. Fronds 149 – 196 cm long, monomorphic; petioles sulcate adaxially, 70 – 111 cm long, 4.0 – 5.7 mm diam., slightly scaly at base, scales like those of rhizomes and slightly pubescent to glabrescent with a mixture of acicular, furcate, and stellate hairs 0.18 – 0.24 mm long. Laminae (66 –) 81 – 90 cm long, triangular to oblong, papyraceous, verrucose, 1 - pinnate-pinnatifid with a confluent, pinnatifid apex, rarely with apex subconform. Buds usually present in axils of distal pinnae; rachises pubescent to pilose with a mixture of hyaline acicular hairs 0.25 – 0.55 mm and hyaline to castaneous, furcate and stellate hairs 0.09 – 0.18 mm; pinnae 20 – 25 pairs, 14 – 23 x 2.8 – 3.6 cm, perpendicular to ascending, the proximal pair usually deflexed, lanceolate, incised 2 / 3 to 3 / 4 their width, sessile to short-stalked to 2.7 mm, apex acute to short-acuminate, base truncate or oblique in the proximal pinnae; costae, costules, and veins adaxially with scattered acicular hairs 0.27 – 0.45 mm long, laminar surface glabrous or with scattered acicular hairs mainly near the margins; costae, costules, and veins abaxially with a mixture of two sizes of acicular hairs, these 0.08 – 0.20 mm and 0.50 – 0.55 mm, also with furcate and stellate hairs 0.08 – 0.2 mm, laminar surface glabrous; segments 7 – 9 mm wide, entire, acute apically, proximal pinnae with basal acroscopic segments reduced and basal basiscopic segments enlarged; veins 10 – 19 pairs per segment, unbranched or bifurcate, the basal vein pairs from adjacent segments usually connivent at the sinuses, sometimes united just below the sinuses or rarely ending below the sinuses, distal vein of each pair arising from costa. Sori round, medial to inframedial; indusia conspicuous, round or round-reniform, castaneous, entire, margin pilose with acicular and furcate hairs; sporangia each with an acicular hair.	en	Salino, Alexandre, Leroy, Carolina Jesus, Moura, Luiza Costa, Moura, Ingridy Oliveira (2016): Four new species of the fern genus Goniopteris C. Presl (Thelypteridaceae) from Brazilian Atlantic Forest. Phytotaxa 255 (3): 249-258, DOI: 10.11646/phytotaxa.255.3.6, URL: http://dx.doi.org/10.11646/phytotaxa.255.3.6
F10F7213FFC9B344FF76EF7CECE5FDC0.taxon	distribution	Distribution and habitat: — Goniopteris seidleri is known from montane regions of Espírito Santo and Rio de Janeiro states, where it grows in rainforests, usually along the streams, at 660 – 1100 m.	en	Salino, Alexandre, Leroy, Carolina Jesus, Moura, Luiza Costa, Moura, Ingridy Oliveira (2016): Four new species of the fern genus Goniopteris C. Presl (Thelypteridaceae) from Brazilian Atlantic Forest. Phytotaxa 255 (3): 249-258, DOI: 10.11646/phytotaxa.255.3.6, URL: http://dx.doi.org/10.11646/phytotaxa.255.3.6
F10F7213FFC9B344FF76EF7CECE5FDC0.taxon	etymology	Etymology: — The epithet refers to the surname of the owner of some preserved forests at the type locality of this species. The Seidler family have protected forests in a mountain region in the state of Espírito Santo, and they are also very receptive to the botanists interested in the regional flora. Additional specimens examined (paratypes): — BRAZIL: Espírito Santo: Itarana, Alto Jatiboca, Fazenda do Frederico Seidler, Floresta Estacional Semidecidual, 665 m, 20 o 00 ’ 49 ” S, 40 o 54 ’ 27 ”, 26 January 2003, A. Salino 8333, F. A. Carvalho, J. A. Lombardi & R. C. Mota (BHCB); ibidem, 683 m, 20 ° 00 ’ 47.6 ” S, 40 ° 54 ’ 23.3 ” W, 19 November 2015, A. Salino 16080, I. O. Moura & L. C. Moura. Rio de Janeiro: Santa Maria Madalena, Parque Estadual do Desengano, Serra do Rifa, 25 November 1977, J. P. P. Carauta 2804 & B. C. de Souza (BHCB, RB). Notes: — Goniopteris seidleri is most similar to Goniopteris lugubris (mainly in southern, southeastern, and central Brazil) and Goniopteris montana (endemic to southeastern Brazil) but differs from both by having erect rhizomes forming a caudex to 16 cm tall, no costal scales, pinnae with reduced basal acroscopic segments and enlarged basal basiscopic segments usually overlapping the rachises, and abaxially glabrous laminar surface between the veins. Goniopteris lugubris and G. montana have short-to long-creeping rhizomes, costal scales present, basal basiscopic segments of pinnae not overlapping the rachises, and abaxial laminar surface between the veins pilose with simple (G. lugubris) or stellate (G. montana) Hairs. Sometimes, G. montana has rhizomes decumbent to erect, but never forming a caudex.	en	Salino, Alexandre, Leroy, Carolina Jesus, Moura, Luiza Costa, Moura, Ingridy Oliveira (2016): Four new species of the fern genus Goniopteris C. Presl (Thelypteridaceae) from Brazilian Atlantic Forest. Phytotaxa 255 (3): 249-258, DOI: 10.11646/phytotaxa.255.3.6, URL: http://dx.doi.org/10.11646/phytotaxa.255.3.6
F10F7213FFCBB342FF76EB7AECD8FE54.taxon	materials_examined	Type: — BRAZIL. Bahia. Jussari, RPPN Serra do Teimoso, 15 º 09 ’ 15.8 ” S, 39 º 31 ’ 24.2 ” W. 260 – 400 m, 02 November 2002, A. Salino & J. G. Jardim 8160 (holotype BHCB!, isotype CEPEC!). Plants terrestrial; rhizomes short-creeping bearing lanceolate, brown scales, 3.2 – 4.3 mm, with furcate and stellate trichomes. Leaves 59 – 104 cm long, monomorphic. Petioles sulcate adaxially, 25 – 55 cm long, 1.6 – 5.1 mm, with few scales at bases, hirsute with furcate and stellate trichomes 0.04 to 0.1 mm long. Laminae 33 – 50 cm long, obovate, chartaceous, not verrucose, 1 - pinnate-pinnatifid, the apex abruptly reduced and conform with distal pair of pinnae adnate to apical pinna. Buds present at bases of distal pinnae. Rachises hirsute mainly along the grooves with furcate and stellate trichomes 0.04 – 0.07 mm long, also with scattered short linear-lanceolate to linear scales 0.25 – 1.25 mm long. Pinnae 4 – 9 pairs per leaf, alternate, 16.5 – 23.5 cm × 3.4 – 4.5 cm, with the basal pair shorter than median pinnae, ascending, narrowly elliptic, sessile to short-stipitate to 0.4 mm, the apices acuminate and bases cuneate; pinnae incised 2 / 3 their width; abaxial side lacking hairs or only with some simple hairs at the sinuses, with linear to linear-lanceolate costal scales 0.25 – 1.25 mm evident at costal bases, lacking scales on costules and veins; adaxial side glabrous, except for some simple and furcate hairs to 0.07 mm and some sparse reduced linear scales on costae. Segments 5.5 – 7.8 mm wide, slightly arcuate, entire, rounded or obtuse at tips; veins 10 – 15 pairs per segment, unbranched, ending in clavate tips adaxially, the basal vein pair (sometimes two pairs) from adjacent segments united below the sinuses and forming an excurrent vein, the distal one of each pair arising from the costa, rarely from the costule. Sori inframedial, indusia reniform or round-reniform, sometimes spathulate, entire, the surface with simple and furcate hairs; sporangia glabrous.	en	Salino, Alexandre, Leroy, Carolina Jesus, Moura, Luiza Costa, Moura, Ingridy Oliveira (2016): Four new species of the fern genus Goniopteris C. Presl (Thelypteridaceae) from Brazilian Atlantic Forest. Phytotaxa 255 (3): 249-258, DOI: 10.11646/phytotaxa.255.3.6, URL: http://dx.doi.org/10.11646/phytotaxa.255.3.6
F10F7213FFCBB342FF76EB7AECD8FE54.taxon	distribution	Distribution and habitat: — Goniopteris smithii is endemic to the southern Bahia state, where it grows in Atlantic rainforests usually on hillsides at 250 – 400 m.	en	Salino, Alexandre, Leroy, Carolina Jesus, Moura, Luiza Costa, Moura, Ingridy Oliveira (2016): Four new species of the fern genus Goniopteris C. Presl (Thelypteridaceae) from Brazilian Atlantic Forest. Phytotaxa 255 (3): 249-258, DOI: 10.11646/phytotaxa.255.3.6, URL: http://dx.doi.org/10.11646/phytotaxa.255.3.6
F10F7213FFCBB342FF76EB7AECD8FE54.taxon	etymology	Etymology: — The epithet honors Dr. Alan R. Smith from the University of California at Berkeley (UC), an authority on Thelypteridaceae. Additional specimens examined (paratypes): — BRAZIL. Bahia: Jussari, estrada de Jussari para Palmira, RPPN Serra do Teimoso, 15 º 09 ’ 37 ” S, 39 º 32 ’ 10 ” W, 14 February 2004, P. Fiaschi 1949 et al. (CEPEC, NY), ibidem, RPPN Serra do Teimoso, 15 º 09 ’ 29 ” S, 39 º 31 ’ 43 ” W, 09 April 1998, A. M. Amorim 2366 et al. (CEPEC, NY, UC), ibidem, 18 June 2003, A. M. Amorim 3715 et al. (CEPEC); Camacan, RPPN Serra Bonita, Fazenda Paris, Interior da Cabruca, 15 º 30 ’ 52 ” S, 39 º 40 ’ 27 ” W, 250 m, 17 June 2009, F. B. Matos 1796 (CEPEC); Camacan, estrada Camacan-Jacareci, Serra Bonita, 15 º 22´54 ” S, 39 º 32´57 ” W, 265 m, 12 December 2014, A. Salino 16032 et al. (BHCB, UC). Notes: — Goniopteris smithii is most similar to G. nicaraguensis (Mexico to Panama), G. oroniensis (endemic to Costa Rica), and G. tristis (Costa Rica, Panama, Guianas, Colombia, Ecuador, Peru, Bolivia, and northern Brazil) by the laminae with a conform apex and proximal pinnae with long-cuneate bases, but differs by the basal vein pair from adjacent segments united below the sinuses and scales abaxially on rachises and costae. Goniopteris. nicaraguensis also differs in having proximal pinnae with short-cuneate bases, in having exindusiate sori and setulose sporangia (vs. proximal pinnae with long-cuneate bases, indusiate sori, and glabrous sporangia in G. smithii). In addition, G. nicaraguensis has dense furcate trichomes 0.1 mm long on the rachises and along costae adaxially, and rachises adaxially with stalked reddish trichomes 0.05 mm long, as well as simple or furcate trichomes 0.06 – 0.08 mm long on the costae and veins abaxially. Goniopteris oroniensis differs from G. smithii by the erect to suberect rhizomes, flexuose rachises, basal pinnae long-petiolulate, exindusiate sori, and setose sporangia. Goniopteris tristis differs in having simple, furcate, and stellate hairs along costae abaxially and, simple hairs on the costules and veins abaxially; it shares with G. smithii the usual presence of buds on adaxial side of the distal pinnae. In southeastern Brazil there are two other endemic species with cuneate pinna bases: Goniopteris cuneata (Christensen 1913: 253) Brade (1972: 215) and G. monosora (Presl 1851: 36) Brade (1972: 216). However, these species have basal veins from adjacent segments meeting the sinuses, and G. cuneata usually has 7 – 15 pinna pairs, simple and furcate hairs along costae, costules, and veins abaxially, and simple hairs on abaxial surfaces between the veins, furcate or simple hairs on sporangial stalks, and costal scales are absent. Goniopteris monosora usually has 10 – 23 pinna pairs, simple and furcate hairs along costae, costules, and veins abaxially, and simple hairs on abaxial surfaces between the veins (such hairs sometimes absent), indusia with only simple hairs, and simple hairs on sporangial stalks.	en	Salino, Alexandre, Leroy, Carolina Jesus, Moura, Luiza Costa, Moura, Ingridy Oliveira (2016): Four new species of the fern genus Goniopteris C. Presl (Thelypteridaceae) from Brazilian Atlantic Forest. Phytotaxa 255 (3): 249-258, DOI: 10.11646/phytotaxa.255.3.6, URL: http://dx.doi.org/10.11646/phytotaxa.255.3.6
F10F7213FFCDB342FF76E8CEE8CFF898.taxon	description	Goniopteris subdimorpha differs from all Brazilian Goniopteris species by the dense, simple pluricellular tortuous hairs 1.1 – 2.7 mm along the abaxial costae, costules, veins, and laminar surface between veins, and these hairs are always arched over the sori. Type: – BRAZIL. Minas Gerais: Sobrália, fazenda Santa Luzia, Floresta Estacional Semidecidual Submontana, 145 – 150 m, 19 ° 12 ’ 34.2 ” S, 42 ° 11 ’ 54.3 ” W, 07 September 2004, A. Salino 9695 (holotype BHCB!, isotype UC!). Plants terrestrial. Rhizomes erect, caudices up to 10 cm tall, scales lanceolate, castaneous, with furcate and stellate hairs. Fronds clustered, 36 – 80 cm long, monomorphic to subdimorphic (fertile fronds with longer petioles); petioles sulcate adaxially, 20 – 61 cm, long, 3.0 – 4.8 mm diam., scaly at bases, scales like those of rhizomes, pilose with simple and stalked stellate and furcate hairs, these denser in adaxial grooves. Laminae 22.5 – 61.0 cm long, lanceate, papyraceous, not verrucose, 1 - pinnate-pinnatifid, apex gradually reduced, confluent and pinnatifid. Buds or plantlets absent on rachises; rachises pilose with simple and stalked stellate and furcate hairs, these denser in adaxial grooves; pinnae 15 – 16 pairs, 8 – 19 × 1.8 – 2.7 cm, perpendicular to ascending, the proximal pair rarely deflexed, linear-elliptic, incised 1 / 3 – 1 / 2 their width, sessile, apices acuminate, bases obtuse; costae, costules, veins, and laminar surfaces adaxially pilose with simple hairs 0.1 mm long, costae also with stellate and furcate stipitate hairs 0.1 mm long; costae abaxially with a mixture of dense, simple, pluricellular, tortuous long hairs 1.1 – 2.7 mm, simple short hairs 0.24 – 0.85 mm long, stalked furcate hairs 0.1 – 0.3 mm long, and reduced linear scales, rarely also with stalked stellate hairs 0.10 – 0.16 mm long, costules, veins and laminar surfaces with simple long and short hairs like those of costae; segments 3.9 – 4.8 mm wide, entire, rounded apically; veins 7 – 11 pairs per segment, unbranched, the basal vein pairs from adjacent segments connivent at the sinuses. Sori rounded, medial or sometimes inframedial, indusia round to reniform with simple hairs; sporangia glabrous.	en	Salino, Alexandre, Leroy, Carolina Jesus, Moura, Luiza Costa, Moura, Ingridy Oliveira (2016): Four new species of the fern genus Goniopteris C. Presl (Thelypteridaceae) from Brazilian Atlantic Forest. Phytotaxa 255 (3): 249-258, DOI: 10.11646/phytotaxa.255.3.6, URL: http://dx.doi.org/10.11646/phytotaxa.255.3.6
F10F7213FFCDB342FF76E8CEE8CFF898.taxon	distribution	Distribution and habitat: — Goniopteris subdimorpha is known only from the type locality in Minas Gerais state, where it grows in semideciduous forests at 140 – 150 m.	en	Salino, Alexandre, Leroy, Carolina Jesus, Moura, Luiza Costa, Moura, Ingridy Oliveira (2016): Four new species of the fern genus Goniopteris C. Presl (Thelypteridaceae) from Brazilian Atlantic Forest. Phytotaxa 255 (3): 249-258, DOI: 10.11646/phytotaxa.255.3.6, URL: http://dx.doi.org/10.11646/phytotaxa.255.3.6
F10F7213FFCDB342FF76E8CEE8CFF898.taxon	etymology	Etymology: — The epithet refers to the subdimorphic fronds (fertile fronds with much longer petioles than sterile fronds). Additional specimens examined (paratypes): — BRAZIL. Minas Gerais: Sobrália, fazenda Santa Luzia, floresta estacional semidecidual submontana, 145 – 150 m, 19 ° 12 ’ 34.2 ” S, 42 ° 11 ’ 54.3 ” W, 07 September 2004, A. Salino 9694 (BHCB, UC). Notes: — Goniopteris subdimorpha differs from all Brazilian Goniopteris species by the dense, simple pluricellular tortuous hairs 1.1 – 2.7 mm along the abaxial side of costae, costules, veins, and laminar surfaces between veins, and these hairs are always arched over the sori. Goniopteris subdimorpha is similar to G. littoralis (Salino 2002: 332) Salino & Almeida (2015: 40), endemic to southeastern Brazil, and G. windischii Salino, endemic to the southern Bahia, Brazil, by having slightly dimorphic fronds (i. e., fertile fronds with longer petioles) and often inframedial sori. However, G. windischii has short to medium-creeping rhizomes, and has rachises with a mixture of short (to 0.1 mm) and long (0.3 – 0.6 mm) hyaline hairs, costae and costules abaxially with a dense mixture of short simple, furcate or stellate hairs ca. 0.1 mm and long stout simple hairs 0.35 – 0.45 mm, and abaxially the laminar surfaces between veins with only short simple hairs 0.1 – 0.15 mm. Goniopteris subdimorpha has abaxial laminar surfaces between veins with only very long simple hairs 1.1 – 2.7 mm. Goniopteris litoralis has long hairs on abaxial side of costae, but these hairs are maximally 1 mm long and are usually unicellular and erect; also the rachises are densely covered with furcate and stellate red hairs to 0.1 mm long, and only few simple hairs to 0.7 mm long in the adaxial grooves.	en	Salino, Alexandre, Leroy, Carolina Jesus, Moura, Luiza Costa, Moura, Ingridy Oliveira (2016): Four new species of the fern genus Goniopteris C. Presl (Thelypteridaceae) from Brazilian Atlantic Forest. Phytotaxa 255 (3): 249-258, DOI: 10.11646/phytotaxa.255.3.6, URL: http://dx.doi.org/10.11646/phytotaxa.255.3.6
F10F7213FFCFB34EFF76E98AE8A7FEE0.taxon	description	Goniopteris windischii is most similar to G. litoralis and G. subdimorpha by the slightly dimorphic fronds (fertile with longer petioles), but differs from those species mainly by the short- to medium-creeping rhizomes, rachises with a mixture of short (to 0.1 mm) and long (0.3 – 0.6 mm) hyaline hairs, rounded to obtuse segment apices, and medial to inframedial sori.	en	Salino, Alexandre, Leroy, Carolina Jesus, Moura, Luiza Costa, Moura, Ingridy Oliveira (2016): Four new species of the fern genus Goniopteris C. Presl (Thelypteridaceae) from Brazilian Atlantic Forest. Phytotaxa 255 (3): 249-258, DOI: 10.11646/phytotaxa.255.3.6, URL: http://dx.doi.org/10.11646/phytotaxa.255.3.6
F10F7213FFCFB34EFF76E98AE8A7FEE0.taxon	materials_examined	Type: — BRAZIL. Bahia. Ilhéus, Distrito de Rio do Braço, Fazenda Estrela da Manhã, na estrada para Banco do Pedro, 09 December 2014, A. Salino 15990, L. A. A. Goes Neto, R. S. Viveros, J. M. Costa & J. F. Souza (Holotype, BHCB, Isotypes, CEPEC, RB, UC). Plants terrestrial. Rhizomes short- to medium-creeping, scales lanceolate with filiform apices, castaneous, 3.6 – 5.5 mm long, with acicular and furcate hairs 0.07 – 0.10 mm long. Fronds 32 – 72 cm long, monomorphic to subdimorphic with fertile fronds with longer petioles and contracted pinnae; petioles stramineous, sulcate adaxially, 16 – 41 cm long, 1 – 3 mm diam., sparsely scaly, scales like those of rhizomes, and pubescent with a mixture of stipitate, furcate, and stellate hairs (0.08 – 0.12 mm long), rarely some simple hairs. Laminae 16 – 34 cm long, triangular to oblong, chartaceous, verrucose, 1 - pinnate-pinnatifid with a confluent, pinnatifid apex. Buds absent; rachises pilose in adaxial grooves with furcate and stellate hairs 0.05 – 0.20 mm, abaxially hirsute with a mixture of simple hairs 0.3 – 0.6 mm and furcate and stellate hairs 0.06 – 0.15 mm; pinnae 7 – 13 pairs, 5.5 – 10.5 × 1.7 – 2.2 cm, slightly to strongly ascending, the proximal pair perpendicular or ascending, linear to linear-lanceolate, incised 1 / 2 – 2 / 3 their width, sessile to short-stalked to 0.1 mm, apices acute to short-acuminate, bases truncate or oblique in the proximal pinnae; costae and costules abaxially hirsute with a mixture of simple hairs 0.3 – 0.5 mm long and furcate and stellate hairs 0.06 – 0.1 mm long, veins and laminar surfaces between veins with only short simple hairs 0.15 – 0.25 mm long, adaxially on costae with only moderate to dense simple hairs 0.1 – 0.45 mm; segments 2.5 – 5 mm wide, entire, rounded to obtuse apically, very rarely acute; veins 5 – 11 pairs per segment, unbranched, the basal vein pairs from adjacent segments connivent at sinuses or just above the sinus, distal vein of pair arising from costule. Sori round, inframedial; indusia round or round-reniform, margins with acicular hairs 0.19 – 0.30 mm; sporangia glabrous.	en	Salino, Alexandre, Leroy, Carolina Jesus, Moura, Luiza Costa, Moura, Ingridy Oliveira (2016): Four new species of the fern genus Goniopteris C. Presl (Thelypteridaceae) from Brazilian Atlantic Forest. Phytotaxa 255 (3): 249-258, DOI: 10.11646/phytotaxa.255.3.6, URL: http://dx.doi.org/10.11646/phytotaxa.255.3.6
F10F7213FFCFB34EFF76E98AE8A7FEE0.taxon	distribution	Distribution and habitat: — Goniopteris windischii is endemic to the Atlantic Forest of southern Bahia, Brazil, where it grows inside or in the edges of the semideciduous and evergreen forests, sometimes along streams, at 665 – 1000 m.	en	Salino, Alexandre, Leroy, Carolina Jesus, Moura, Luiza Costa, Moura, Ingridy Oliveira (2016): Four new species of the fern genus Goniopteris C. Presl (Thelypteridaceae) from Brazilian Atlantic Forest. Phytotaxa 255 (3): 249-258, DOI: 10.11646/phytotaxa.255.3.6, URL: http://dx.doi.org/10.11646/phytotaxa.255.3.6
F10F7213FFCFB34EFF76E98AE8A7FEE0.taxon	etymology	Etymology: — Epithet is in honor of the Brazilian pteridologist Dr. Paulo Günter Windisch, who encouraged me to study Thelypteridaceae. Additional specimens examined (paratypes): — BRAZIL. Bahia: Almadina. Fazenda São José, estrada Coaraci- Almadina, Serra do Corcovado, 14 ° 43 ’ 12 ” S, 39 ° 36 ’ 11 ” W, 320 m, 10 December 2014, A. Salino et al. 16000 (BHCB, UC); Caatiba, rodovia BA- 265, trecho Caatiba / Barra do Choça, a 6 km a W de Caatiba, ca. 1000 m, 15 March 1979, S. A. Mori & T. S. dos Santos 11565 (CEPEC); Camacã, Assentamento Conjunto Recordação, 05 September 2001, D. L. Santana 757 et al. (CEPEC); Camacã, estrada Camacan-Jacaraci, Serra Bonita, 15 ° 24 ’ 15 ” S, 39 ° 32 ’ 06 ” W, 190 m, 12 December 2014, A. Salino et al. 16031 (BHCB); Ibicaraí, 3 km W da cidade, 14 ° 52`S, 39 ° 38´W, 16 February 1988, J. R. Pirani 2328 et al. (BHCB, SPF); Ilhéus, BR- 415 Km 22, 14 October 1986, J. C. Hage 2128 & E. B. dos Santos (CEPEC, SJRP), ibidem, Centro de Pesquisas do Cacau, 24 May 1965, R. P Belém & M. Magalhães 1106 (CEPEC, UB, US), ibidem, 05 December 1968, J. Almeida & T. S. Santos 288 (CEPEC, NY), ibidem, 15 May 1965, R. P. Belém & M. Magalhâes 987 (US), ibidem, 1821, Riedel s. n. (P 00167488), ibidem, 28 October 1978, S. A. Mori 10978 (CEPEC, NY); Itacaré, ramal à esquerda da Rod. BR- 101, com entrada no Km 11 do trecho, 12 June 1979, L. A. Mattos Silva et al. 437 (CEPEC); Jussari, RPPN Serra do Teimoso, 15 ° 09 ’ 15.8 ” S, 39 ° 31 ’ 24.2 ” W, 02 November 2002, A. Salino 8163 & J. Jardim (BHCB), ibidem, 432 m, 03 November 2013, L. L. Giacomin et al. 1978 (BHCB); Santo Amaro, saída de Santo Amaro ao entroncamento de Valença, Rod. BR- 101, 07 May 1969, J. S. de Jesus 351 & T. S. dos Santos 400 (NY). Notes: — Goniopteris windischii is most similar to G. littoralis and G. subdimorpha by having slightly dimorphic fronds (fertile with longer petioles), but differs from those species mainly by the short- to medium-creeping rhizomes, rachises with a mixture of short (to 0.1 mm) and long (0.3 – 0.6 mm) hyaline hairs, rounded to obtuse segment apices, and sori medial to inframedial. Goniopteris littoralis has erect rhizomes, rachises with short, furcate and stellate red hairs 0.1 – 0.3 mm, rarely with some simple hairs, acute (rarely rounded) segment apices, and sori always inframedial. In addition, G. windischii has pinnae usually linear to linear-lanceolate, and the proximal pinna pair perpendicular to the rachis, while G. littoralis has pinnae usually lanceolate, sometimes linear-lanceolate, and the proximal pair strongly deflexed. Goniopteris subdimorpha has erect rhizomes and dense, simple pluricellular tortuous hairs 1.1 – 2.7 mm along the abaxial costae, costules, veins, and laminar surfaces between veins, and these hairs are always arched, covering the sori.	en	Salino, Alexandre, Leroy, Carolina Jesus, Moura, Luiza Costa, Moura, Ingridy Oliveira (2016): Four new species of the fern genus Goniopteris C. Presl (Thelypteridaceae) from Brazilian Atlantic Forest. Phytotaxa 255 (3): 249-258, DOI: 10.11646/phytotaxa.255.3.6, URL: http://dx.doi.org/10.11646/phytotaxa.255.3.6
