taxonID	type	description	language	source
F00587B70379FFD3341FFC9006CE01B5.taxon	type_taxon	Type species: Belgrandiella andalucensis Boeters, 1983 by original designation Redefinition (modified from Arconada et al., 2007): Shell narrow, long, cylindrical to slightly conical. Surface smooth or with microsculpture of fine, irregularly shaped pits, best seen on the protoconch. The edge of the aperture is sharp; only its columellar border may be slightly broadened. Horny operculum of yellowish colour. The animal lacks pigmentation, has no eyes, gill with 7 – 15 leaflets, rectum forms two z-like loops, penis simple (only G. arconadae has a small lateral appendix), female genital tract with bursa and single receptaculum. The genus comprises the following eight species: G. algarvensis, G. andalucensis, G. arconadae, G. ballesterosi, G. kolendaensis sp. nov.; G. murita (Boeters, 2003) comb. nov., G. pilelongata, and G. ramosae. All are stygobionts from the Iberian Peninsula.	en	Quiñonero-Salgado, Sergio, López-Soriano, Joaquín (2024): New data on the genus Guadiella Boeters, 2003 (Gastropoda: Hydrobiidae) from the northern half of the Iberian Peninsula, with the description of a new species. Nemus 14: 102-111, DOI: 10.5281/zenodo.17031682
F00587B70379FFD4341FFA3504C1061C.taxon	materials_examined	Material: Holotype RMNH. MOL. 350854 (Naturalis) (Fig. 2); Paratypes: 3 s. RMNH. MOL. 350855 (Naturalis). Other material examined: 40 shells in SQS collection, from the type locality. Type locality: Fuente de Morales, Calanda (Teruel province), Spain. [30 TYL 370357]. 381 m a. s. l. (Fig. 6 A). This is a spring with abundant and permanent water flow, which rises in a small cavity and is channelized with a rubber pipe. It is hard to localize and to access, due to its abandoned state, and the amount of vegetation around. It is located in a slope close to the Guadalope River, a tributary of Ebro River. This is also the type locality of Hadziella forneri Quiñonero-Salgado, López-Soriano & Rolán, 2023.	en	Quiñonero-Salgado, Sergio, López-Soriano, Joaquín (2024): New data on the genus Guadiella Boeters, 2003 (Gastropoda: Hydrobiidae) from the northern half of the Iberian Peninsula, with the description of a new species. Nemus 14: 102-111, DOI: 10.5281/zenodo.17031682
F00587B70379FFD4341FFA3504C1061C.taxon	etymology	Etymology: The specific name derives from the Celtiberian village of Kolenda, origin of the present village of Calanda.	en	Quiñonero-Salgado, Sergio, López-Soriano, Joaquín (2024): New data on the genus Guadiella Boeters, 2003 (Gastropoda: Hydrobiidae) from the northern half of the Iberian Peninsula, with the description of a new species. Nemus 14: 102-111, DOI: 10.5281/zenodo.17031682
F00587B70379FFD4341FFA3504C1061C.taxon	description	Description: The shell is elongate, slender, with inflated whorls and a deep, well-defined suture (Fig. 1 A). The last whorl is much higher and therewith flatter. Protoconch of 1 – 1.5 whorls with a marked microsculpture of fine, irregularly shaped pits (most pits have grooves in the shape of a 6 - pointed star in the centre and a more or less hexagonal outline, but the number of points varies from 5 to 8) (Fig. 1 D). Teleoconch of 3 whorls, surface with poorly defined, low, irregular radial ribs with microsculpture of irregular, very fine and shallow pits (Fig. 1 B). Aperture with slanted ovoid outline, slightly wider at base, edge reflected. A narrow ridge inside the aperture, close to the edge, supports the operculum. The final part of the last whorl is very slightly decollated, forming a narrow umbilical groove between aperture and penultimate whorl (Figs. 1, 2, 4 A). As only empty shells were found, no operculum has been seen. It is assumed to be horny, as in other species of the genus. Dimensions: Shell height 0.87 – 1.16 mm, and width 0.34 – 0.50 mm. The aperture / last whorl has dimensions of 0.26 – 0.34 mm. See Table 1 and Fig. 5. Habitat: Stygobitic. Shells were washed out from their habitat, likely after sudden rises of the subterranean water levels. Shells had a mainly fresh appearance, given their transparent colour.	en	Quiñonero-Salgado, Sergio, López-Soriano, Joaquín (2024): New data on the genus Guadiella Boeters, 2003 (Gastropoda: Hydrobiidae) from the northern half of the Iberian Peninsula, with the description of a new species. Nemus 14: 102-111, DOI: 10.5281/zenodo.17031682
F00587B70379FFD4341FFA3504C1061C.taxon	distribution	Distribution: Only known from the type locality. Differentiating characters: Guadiella kolendaensis sp. nov. is similar to G. arconadae regarding microsculpture, but the latter has a wider apex, more inflated whorls, and the narrow ridge inside the aperture is much closer to the edge. The apex of Guadiella murita comb. nov. is slenderer, resulting in an inverted conical shape, whereas the microsculpture of the protoconch differs in having a dense pattern of irregularly arranged micropunctures. Guadiella pilelongata also has a slender apex, deep sutures with a narrow subsutural ramp and a protoconch with a fine microsculpture of variable shaped polygonal micropits. Like G. kolendaensis sp. nov., G. ballesterosi has a suboval and inclined aperture, but the sutures are deeper and it has very fine polygonal depressions on the protoconch and irregular spiral lines on the teleoconch, in addition to be larger in size. Guadiella andalucensis has lower upper whorls and a much broader final whorl, resulting in a much plumper shell. Guadiella ramosae is very similar in outline to G. kolendaensis sp. nov., but the whorls are more compressed whereas the apex is wider. Guadiella algarvensis has a substantially larger shell, with one additional whorl, is relatively narrower, and it narrows towards the apex; the microsculpture of the protoconch is much finer, formed by very dense and irregularly displayed micropunctures.	en	Quiñonero-Salgado, Sergio, López-Soriano, Joaquín (2024): New data on the genus Guadiella Boeters, 2003 (Gastropoda: Hydrobiidae) from the northern half of the Iberian Peninsula, with the description of a new species. Nemus 14: 102-111, DOI: 10.5281/zenodo.17031682
F00587B7037EFFD437B9FD9304E606BB.taxon	description	(Fig. 3 A-C, 4 B)	en	Quiñonero-Salgado, Sergio, López-Soriano, Joaquín (2024): New data on the genus Guadiella Boeters, 2003 (Gastropoda: Hydrobiidae) from the northern half of the Iberian Peninsula, with the description of a new species. Nemus 14: 102-111, DOI: 10.5281/zenodo.17031682
F00587B7037FFFDA35EEFF6F04FE04DF.taxon	description	(Figs. 3 D-F, 4 C)	en	Quiñonero-Salgado, Sergio, López-Soriano, Joaquín (2024): New data on the genus Guadiella Boeters, 2003 (Gastropoda: Hydrobiidae) from the northern half of the Iberian Peninsula, with the description of a new species. Nemus 14: 102-111, DOI: 10.5281/zenodo.17031682
F00587B7037FFFDA35EEFF6F04FE04DF.taxon	materials_examined	Type locality: Merindad de Río Ubierna, spring at north side of San Martín de Ubierna, Burgos province, 900 m a. s. l. (Fig. 6 C). Material examined: 43 shells in SQS collection from the type locality, collected in 2023 (SQS leg.)	en	Quiñonero-Salgado, Sergio, López-Soriano, Joaquín (2024): New data on the genus Guadiella Boeters, 2003 (Gastropoda: Hydrobiidae) from the northern half of the Iberian Peninsula, with the description of a new species. Nemus 14: 102-111, DOI: 10.5281/zenodo.17031682
F00587B7037FFFDA35EEFF6F04FE04DF.taxon	discussion	Remarks: Based on anatomical traits and the stygobitic habitat, in the original description the species was provisionally assigned to Guadiella. A single SEM image of the shell was provided, but no data related to its microsculpture. The type locality was visited by the first author of this note, which allowed the gathering of fresh, new material. New SEM images reveal a protoconch microsculpture formed by small undulations or excavations (Figs. 3 D-E), irregularly displayed, and a rough surface of the teleoconch, with few growth lines (Fig. 3 F). A number of shells from this new material were measured, allowing description of shell variability (Figs. 3 D-F, 4 C, 5 and Table I).	en	Quiñonero-Salgado, Sergio, López-Soriano, Joaquín (2024): New data on the genus Guadiella Boeters, 2003 (Gastropoda: Hydrobiidae) from the northern half of the Iberian Peninsula, with the description of a new species. Nemus 14: 102-111, DOI: 10.5281/zenodo.17031682
