identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
F74687C8FFBBFFB1E69AFCEDFD878E4A.text	F74687C8FFBBFFB1E69AFCEDFD878E4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius (Raphirus) umbrinus Erichson 1839	<div><p>Quedius (Raphirus) umbrinus ERICHSON, 1839</p><p>Quedius (Raphirus) kuboni ŠTOURAČ, 1998: 15 ff.; nov.syn.</p><p>C o m m e n t: Quedius umbrinus previously already had ten junior synonyms, more than any other representative of the subgenus Raphirus STEPHENS, 1829 in the Palaearctic region. The species is widespread and at the same time subject to enormous intraspecific variation (ASSING 2018).</p><p>The original description of Q. kuboni is based on a male holotype and eleven paratypes collected in three mountain ranges (Rodopi, Rila, Pirin) in Bulgaria. Based on the illustrations of the aedeagus provided by ŠTOURAČ (1998), the type material is undoubtedly conspecific with Q. umbrinus .</p></div>	https://treatment.plazi.org/id/F74687C8FFBBFFB1E69AFCEDFD878E4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker	Assing, Volker (2019): On the taxonomy oI some West Palaearctic Quedius species, with descriptions oI new species and new synonymies (Coleoptera, Staphylinidae, Staphylininae). Linzer biologische Beiträge 51 (1): 189-201, DOI: 10.5281/zenodo.3762449
F74687C8FFBBFFB4E69AFB47FEDA8B45.text	F74687C8FFBBFFB4E69AFB47FEDA8B45.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius (Raphirus) iridicolor QUEDENFELDT 1882	<div><p>Quedius (Raphirus) iridicolor QUEDENFELDT, 1882 (Figs 1-7, Map 1)</p><p>Quedius iridicolor QUEDENFELDT, 1882: 181 . Quedius duplex FAUVEL, 1886: 68 f. Quedius (Sauridus) iridicolor: COIFFAIT (1978). Quedius (Sauridus) iridicolor: HERMAN (2001). Quedius (Microsaurus) iridicolor: SMETANA (2004). Quedius (Microsaurus) iridicolor: SCHÜLKE &amp; SMETANA (2015). T y p e m a t e r i a l e x a m i n e d: Holotype ♀: ̎ Tetuan / iridicolor / Type / 67191 / Quedius iridicolor Quedf. / Hist .- Coll . ( Coleoptera), Nr. 67191, Quedius iridicolor Quedf . *, Tetuan, Coll. Quedenfeldt, Coll. Richter, A. Bau V., Zool. Mus. Berlin / HOLOTYPE Quedius iridicolor Quedenfeldt, 1882 , labelled by MFNB 2019 / Quedius iridolor Quedenfeldt , det. V. Assing 2019̎ (MNB). A d d i t i o n a l m a t e r i a l e x a m i n e d: Morocco: 1♁, 2♀♀, Ar Rif, road</p><p>Chefchaouen - Ksal el Kebr, 35°05'N, 5°25'W, 680 m, 3-7.VI.2007, leg. Hlaváč (cAss); 1♁,</p><p>Chefchaouen env., 35°12'N, 5°19'W, 270 m, 3.VI.2007, leg. Hlaváč (cAss); 1 ex. [apex of</p><p>abdomen missing], W Tetuan, El Fendek, 350 m, pitfall trap, XI-XII.1985, leg. Heinz (MNB).</p><p>Spain: 1♁, Andalucía, Jaén, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.3166666&amp;materialsCitation.latitude=35.2" title="Search Plazi for locations around (long -5.3166666/lat 35.2)">Hornos</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.3166666&amp;materialsCitation.latitude=35.2" title="Search Plazi for locations around (long -5.3166666/lat 35.2)">Sima del Campamento</a>, 11.VIII.2012, leg. G.E.V. (cAss); 1♁, Andalucía, Jaén, Hornos, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.45&amp;materialsCitation.latitude=36.75" title="Search Plazi for locations around (long -5.45/lat 36.75)">Sima de la Tuberia</a>, 23.X.2011, leg. G.E. V. (cAss). 1♁, 1♀, Andalucía, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.45&amp;materialsCitation.latitude=36.75" title="Search Plazi for locations around (long -5.45/lat 36.75)">Cádiz</a>, 15 km NE Ubrique, 36°45'N, 5°27'W, 770 m, calcareous oak forest, 28.XII.2009, leg. Assing (cAss) ; 1♀, Andalucía, Cádiz, Ronda env., Sierra de Ubrique, 1000 m, 25.III.1994, leg. Assing (cAss) .</p><p>C o m m e n t: The original description of Q. iridicolor is based on ̎unum specimen̎ from ̎ Marocco (Tetuan)̎ (QUEDENFELDT, 1882), that of Q. duplex on an unspecified number of syntypes from ̎O. Daya̎ and ̎A. Teniet-el-Had̎ in Algeria (FAUVEL 1886). FAUVEL (1902) provided additional records of Q. duplex from North Africa and did not rule out synonymy of this name with Q. iridicolor . The synonymy of Q. duplex with Q. iridicolor was eventually established by COIFFAIT (1978). Despite the previous assignment to the subgenus Sauridus MULSANT &amp; REY, 1876 (COIFFAIT 1978, HERMAN 2001), currently a junior synonym of Raphirus STEPHENS, 1829, SMETANA (2004) and SCHÜLKE &amp; SMETANA (2015) erroneously attribute the species to Microsaurus DEJEAN, 1833. Quedius iridicolor undoubtedly belongs to Raphirus and is most likely the adelphotaxon of Q. nigriceps.</p><p>D i a g n o s i s: According to COIFFAIT (1978), Q. iridicolor is distinguished from the similar Q. nigriceps KRAATZ, 1857 by shorter, more slender, and less convex elytra, by a shorter head with more convex eyes, by coarser and sparser punctation of the anterior tergites, denser punctation of the posterior tergites, an apically dilated median lobe of the aedeagus, and a more slender paramere with parallel lateral margins. An examination of the holotype and the additional material listed above revealed, however, that these characters are partly incorrect or unsuitable for a reliable identification of Q. iridicolor and separation from Q. nigriceps. This applies to the punctation of the abdomen, the shape and length of the elytra (see ratio EL/PL below), and some aedeagal characters mentioned. Moreover, the coloration of both species is highly variable. The only external differences found are that the head tends to be relatively more transverse (see ratio HW/HL), the pronotum narrower in relation to head (ratio PW/HW; but note the considerable overlap) with the lateral margins of the pronotum mostly less convex in dorsal view, and the pair of posterior punctures on the vertex (the two punctures close to the posterior constriction of the head on either side) are arranged differently (pvp in Fig. 1). The anterior one of these punctures is situated more or less directly in front of the posterior one, whereas in Q. nigriceps it is situated obliquely antero-mediad or both punctures are arranged in a transverse line (i.e., at the same distance from the posterior constriction).</p><p>Measurements (in mm) and ratios (range): Quedius iridicolor: HL: 0.97-1.09; HW: 1.16- 1.32; PW: 1.35-1.53; EL: 0.88-1.02; HW/HL: 1.14-1.21; PW/HW: 1.12-1.24; EL/PL: 0.62-0.69.</p><p>Quedius nigriceps (exclusively material from the south of the Iberian Peninsula): HL: 0.95-1.11; HW: 1.04-1.23; PW: 1.25-1.55; PL: 1.25-1.58; EL: 0.86-1.11; HW/HL: 1.10- 1.14; PW/HW: 1.20-1.27; EL/PL: 0.67-0.73.</p><p>The aedeagus (Figs 2-7) of Q. iridicolor is distinguished from that of Q. nigriceps as follows: the apex of the median lobe is more acute in ventral view and the subapical tooth is connected to the apex by a distinct keel; the paramere is usually more slender and characterized by shorter, more irregular, and closer series of peg-setae. For illustrations of the aedeagus of Q. nigriceps see Figs 8-10.</p><p>D i s t r i b u t i o n: Quedius iridicolor evidently represents a Mauretanian faunal element and is currently known only from Northwest Africa (Morocco, Algeria) and South Spain. COIFFAIT (1978) erroneously states that Q. iridicolor replaces Q. nigriceps in the southern Iberian Peninsula. I have examined Q. nigriceps from South Portugal across Andalucía to Alicante. In one cave in Jaén, both species were even found syntopically.</p></div>	https://treatment.plazi.org/id/F74687C8FFBBFFB4E69AFB47FEDA8B45	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker	Assing, Volker (2019): On the taxonomy oI some West Palaearctic Quedius species, with descriptions oI new species and new synonymies (Coleoptera, Staphylinidae, Staphylininae). Linzer biologische Beiträge 51 (1): 189-201, DOI: 10.5281/zenodo.3762449
F74687C8FFBEFFB6E69AFC36FE7289E7.text	F74687C8FFBEFFB6E69AFC36FE7289E7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius (Raphirus) cohaesus Eppelsheim 1888	<div><p>Quedius (Raphirus) cohaesus EPPELSHEIM, 1888</p><p>Quedius cohaesus EPPELSHEIM, 1888: 60 f.</p><p>Quedius pseudonigriceps REITTER, 1909: 113; nov.syn.</p><p>C o m m e n t: The original description of Q. cohaesus is based on three syntypes from ̎Turcmenien̎ (EPPELSHEIM 1888), of which SOLODOVNIKOV (2004) designated the lectotype, that of Q. pseudonigriceps on an unspecified number of syntypes from ̎ Herzegowina, Türkei, Kleinasien̎ (REITTER 1909). Quedius cohaesus has three junior synonyms, Q. pseudonigriceps five (SCHÜLKE &amp; SMETANA 2015, SALNITSKA &amp; SOLODOVNIKOV 2018). According to the latest edition of the Palaearctic catalogue, the distribution of Q. cohaesus ranges from Bulgaria across Turkey and the Caucasus region to Iran and Turkmenistan, whereas SALNITSKA &amp; SOLODOVNIKOV (2018) state that the distribution of this species includes Iran, Turkmenistan, Tajikistan, and Afghanistan. There is no reference to the records from other regions in the latter study, probably because it focuses on Middle Asia. The distributions given for Q. pseudonigriceps are Southeast Europe northwards to Austria, Turkey, and the Caucasus region (Georgia) (SCHÜLKE &amp; SMETANA 2015), and South Europe, West Asia eastwards to Middle Asia (Kyrgyzstan, Kazakhstan) (SALNITSKA &amp; SOLODOVNIKOV 2018). SALNITSKA &amp; SOLODOVNIKOV (2018) regard Q. cohaesus and Q. pseudonigriceps as distinct species, provide redescriptions and illustrations of the aedeagi based on material from Middle Asia, and state that Q. cohaesus is most similar to Q. pseudonigriceps, but ̎easily</p><p>distinguished by the presence of an apical seam of palisade fringe VII and normally developed elytra, as well as by the characters of the aedeagus̎ (p. 143).</p><p>Although I have not studied material from Middle Asia personally, the argumentation and conclusions laid out by SALNITSKA &amp; SOLODOVNIKOV (2018) are difficult to follow and Q. pseudonigriceps is considered a junior synonym of Q. cohaesus for two main reasons:</p><p>1. Based on personal observations and abundant material, the species previously identified as Quedius pseudonigriceps is one of the most widespread and most common representatives of the genus in the Balkans, Turkey, and the Caucasus region (Georgia, Armenia). In this region, the species always has normally developed elytra and a palisade fringe at the posterior margin of tergite VII. This would mean that, according to SALNITSKA &amp; SOLODOVNIKOV (2018), there is an abrupt character displacement in the range of the species somewhere in North Iran. True, there are several examples of wing di- or polymorphic Quedius species, sometimes even with clinal variation. It does not seem plausible, however, that there should be another, highly similar macropterous species ( Q. cohaesus) replacing the macropterous morph of Q. pseudonigriceps in this region.</p><p>2. The aedeagus of Quedius species is subject to - sometimes considerable - interspecific variation. This particularly applies to widespread species (ASSING 2018). SALNITSKA &amp; SOLODOVNIKOV (2018) may state that there are differences between Q. cohaesus and Q. pseudonigriceps in aedeagal characters, but no such discrete differences are visible in the illustrations they provide. Instead, they illustrate a continuum of aedeagal variation.</p><p>In consequence, the available evidence suggests that a) Q. cohaesus is distributed on both sides of the Caspian Sea and that b) this species is wing-dimorphic in Middle Asia and monomorphic elsewhere.</p><p>Quedius (Microsaurus) invreae GRIDELLI, 1924 and Q. (M.) puncticollis (THOMSON, 1867)</p><p>C o m m e n t: According to the latest edition of the Palaearctic catalogue (SCHÜLKE &amp; SMETANA 2015), the distribution of Q. invreae ranges from France, Great Britain and Scandinavia across Central Europe southeastwards to the Russian South European territory and that of Q. puncticollis from France, the British Isles (including Ireland), and Scandinavia across Central Europe southwards to Italy and eastwards to the Caucasus region and West Siberia. Recently, Q. puncticollis was recorded also from Greece and Middle Asia (ASSING 2017, SALNITSKA &amp; SOLODOVNIKOV 2018). In his key to the Central European Quediina, SOLODOVNIKOV (2012) doubted the taxonomic status of Q. invreae and suspected synonymy of these names.</p><p>Inspired by records of specimens preliminarily identified as Q. invreae from Armenia, material of Q. puncticollis and Q. invreae in the authoŕs collection was revised. This study revealed that both names undoubtedly represent distinct taxa and that both species are easily distinguished. The aedeagi of both species may be rather similar, but the paramere of Q. invreae is broader and usually weakly concave apically, more rarely truncate or weakly convex, whereas in Q. puncticollis it is more slender and apically distinctly convex. The best characters for the separation of both species, however, are the colour, shape, and especially the chaetotaxy of sternite VIII, especially in the male, as already stated by LOHSE (1964). In Q. invreae, sternite VIII is darker, often completely black or at least black in anterior three-fourths with the posterior fourth sometimes slightly paler, whereas in Q. puncticollis tergite VIII is paler with at least the anterior and posterior portions pale-brownish and the median portion usually blackish-brown. More importantly, however, the posterior margin of the male sternite VIII is broadly concave and furnished with extremely long black setae (especially laterally), the longest of them longer than antennomere I in Q. invreae . In Q. puncticollis, on the other hand, the posterior margin is shallowly concave only in the middle and furnished with short brown setae, the longest of them barely half as long as antennomere I. The posterior margin of the female sternite VIII is furnished with relatively short setae in both species, but these setae are black in Q. invreae and brown in Q. puncticollis . The illustrations of the aedeagus of Q. puncticollis provided by SOLODOVNIKOV (2012: figures 191t-v) clearly refer to Q. invreae .</p><p>According to LOHSE (1964), Q. puncticollis is the most common representative of the genus in mole nests. This may be true of the environs of Hamburg, G.A. Lohsés hometown, but certainly not of regions farther south. Based on personal observations (studies of mole nests, extensive pitfall trap studies), Q. puncticollis is rare, much more so than Q. invreae, and the southernmost record examined is from South Germany (München). While most of the few specimens of Q. puncticollis in cAss were extracted from mole nests (some also collected with pitfall traps), material of Quedius invreae was exclusively collected with pitfall and flight interception traps (especially in warmer habitats; never from mole nests). A re-examination of a specimen from Greece, which the record of Q. puncticollis in ASSING (2017) is based on, revealed that it was misidentified and in fact belongs to Q. invreae (first record from Greece). The same most likely applies to records of Q. puncticollis from Middle Asia (SALNITSKA &amp; SOLODOVNIKOV 2018).</p></div>	https://treatment.plazi.org/id/F74687C8FFBEFFB6E69AFC36FE7289E7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker	Assing, Volker (2019): On the taxonomy oI some West Palaearctic Quedius species, with descriptions oI new species and new synonymies (Coleoptera, Staphylinidae, Staphylininae). Linzer biologische Beiträge 51 (1): 189-201, DOI: 10.5281/zenodo.3762449
F74687C8FFBCFFB6E69AFBD3FC8A8F73.text	F74687C8FFBCFFB6E69AFBD3FC8A8F73.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius (Microsaurus) cruentus	<div><p>Quedius (Microsaurus) cruentus (OLIVIER, 1795)</p><p>Quedius xanthurus IABLOKOFF- KHNZORIAN, 1961: 147 f.; nov.syn.</p><p>T y p e m a t e r i a l e x a m i n e d: Holotype ♀ [antennae missing]: ̎Kafan, N. Gand, ASSR. 27.7.52 / Holotypus Quedius cruentus Iablokoff-Khnzorian, rev. V. Assing 2019 / Quedius cruentus (Olivier), det. V. Assing 2019̎ (cKhn).</p><p>C o m m e n t: The original description of Q. xanthurus is based on a unique holotype from ̎Nerkin Hand (Armenie orientale)̎ collected ̎dans un vieux creux, 27.7.1952 ̎ (IABLOKOFF- KHNZORIAN 1961). The species was attributed to the subgenus Sauridus MULSANT &amp; REY, 1876, now a junior synonym of Raphirus STEPHENS, 1829, by COIFFAIT (1978). An examination of the holotype, however, revealed that it is conspecific with Q. cruentus . Hence the synonymy proposed above.</p></div>	https://treatment.plazi.org/id/F74687C8FFBCFFB6E69AFBD3FC8A8F73	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker	Assing, Volker (2019): On the taxonomy oI some West Palaearctic Quedius species, with descriptions oI new species and new synonymies (Coleoptera, Staphylinidae, Staphylininae). Linzer biologische Beiträge 51 (1): 189-201, DOI: 10.5281/zenodo.3762449
F74687C8FFBCFFB7E69AFAAFFE378818.text	F74687C8FFBCFFB7E69AFAAFFE378818.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius (Microsaurus) transcaucasicus	<div><p>Quedius (Microsaurus) transcaucasicus IABLOKOFF- KHNZORIAN, 1961</p><p>Quedius transcaucasicus IABLOKOFF- KHNZORIAN, 1961: 146 f.</p><p>Quedius iablokofi COIFFAIT, 1967: 396; unnecessary replacent name.</p><p>M a t e r i a l e x a m i n e d: 1♀: ̎Idhsewan, Kiranu, ASSR - 17-5-51̎ (cKhn).</p><p>C o m m e n t: The original description of Q. transcaucasicus is based on a unique holotype from ̎Schischkert (Arménie orientale) dans un chẻne creux ả la lisiẻre supérieure de la forêt (env. 1900 m), 31.7.1959 ̎ (IABLOKOFF- KHNZORIAN 1961). The examined female is the sole specimen found in the Khnzorian collection, but apparently not the holotype, as the locality and date indicated on the label do not agree with those given in the description. Nevertheless, the specimen is in good agreement with the characters mentioned by IABLOKOFF- KHNZORIAN (1961). Quedius transcaucasicus is characterized particularly by its conspicuous coloration (head and pronotum black; elytra and abdomen reddish).</p><p>Quedius transcaucasicus was subsequently replaced with the nomen novum Q. iablokofi by COIFFAIT (1967), who believed the name was preoccupied by Q. transcaucasicus GEMMINGER &amp; HAROLD, 1968. The latter, however, is a nomen nudum and consequently unavailable (HERMAN 2001).</p></div>	https://treatment.plazi.org/id/F74687C8FFBCFFB7E69AFAAFFE378818	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker	Assing, Volker (2019): On the taxonomy oI some West Palaearctic Quedius species, with descriptions oI new species and new synonymies (Coleoptera, Staphylinidae, Staphylininae). Linzer biologische Beiträge 51 (1): 189-201, DOI: 10.5281/zenodo.3762449
F74687C8FFBDFFBAE69AFD96FC78886D.text	F74687C8FFBDFFBAE69AFD96FC78886D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius (Microsaurus) morulus Assing 2019	<div><p>Quedius (Microsaurus) morulus nov.sp. (Figs 11-12, 16-18)</p><p>T y p e m a t e r i a l: Holotype ♁: ̎ IRAN - Mazandaran, 36.297°N, 53.453°E, 1140 m, window trap, 5.IX.2017, Barimani / Holotypus ♁ Quedius morulus sp. n., det. V. Assing 2019̎ (cAss) . Paratypes: 1♁ [aedeagus missing]: ̎ IRAN - Mazandaran, 36.296°N, 53.447°E, 1230 m, window trap, 29.VII.2017, Barimani ̎ (cAss); 1♀: ̎ IRAN - Mazandaran, 36.292°N, 53.437°E, 1285 m, window trap, 22. V.2017, Barimani ̎ (cAss); 1♁: ̎ IRAN - Mazandaran, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=53.41&amp;materialsCitation.latitude=36.14" title="Search Plazi for locations around (long 53.41/lat 36.14)">6 km W Part Kola</a>, 2050 m, 36.14°N, 53.41°E, Fagus orientalis trap, 3, V.2015, leg. Barimani ̎ (cAss) .</p><p>E t y m o l o g y: The specific epithet (Latin, adjective: black) alludes to the uniformly blackish coloration of the body.</p><p>D e s c r i p t i o n: Body length 11.5-13.5 mm; length of forebody 6.0-7.0 mm. Habitus as in Fig. 12. Coloration: body (including appendages) black, except for the reddish-brown tarsomeres V.</p><p>Head (Fig. 12) 1.16-1.28 times as broad as long, on average larger and more transverse in male than in female; punctation fine, shallow, and rather dense; one macropuncture near middle of dorsal margin of eye, one near posterior margin of eye, one approximately halfway between posterior margin of eye and posterior constriction of head, and two near posterior constriction of head; interstices with fine, very shallow to distinct transverse microreticulation; frons with shallow impression, this impression with vortexlike arrangement of microsculpture. Eyes approximately as long as postocular region. Antenna with antennomeres IV-V approximately as long as broad and VI-X very weakly transverse.</p><p>Pronotum (Fig. 12) 1.08-1.11 times as broad as long and 1.22-1.31 times as broad as head; dorsal series each composed of 1+2 macropunctures; usually with 1-3 smaller sublateral punctures behind level of posterior pair of dorsal punctures; disc with very fine and shallow microsculpture composed of transverse or oblique striae.</p><p>Elytra (Fig. 12) 0.67-0.71 times as long as pronotum; punctation dense and distinct; interstices on average slightly broader than diameter of punctures, without microreticulation. Scutellum with fine transverse microreticulation and fine non-setiferous micropunctation. Hind wings fully developed.</p><p>Abdomen with dense and distinct punctation, that of posterior tergites less dense than that of anterior tergites; pubescence long and suberect, greyish to blackish; interstices with extremely fine transverse microsculpture visible only at high magnification (100 x); posterior margin of tergite VII with palisade fringe.</p><p>♁: sternite VIII with shallowly concave posterior excision; aedeagus 1.5-1.6 mm long and shaped as in Figs 16-18.</p><p>♀: sternite VIII with weakly convex posterior margin, pubescence unmodified.</p><p>C o m p a r a t i v e n o t e s: Based on external (habitus, size, punctation pattern, etc.), Q. morulus is similar to the widespread Q. truncicola FAIRMAIRE &amp; LABOULBÊNE, 1856. It is distinguished from this species by the black abdomen and by the morphology of the aedeagus (median lobe and paramere longer; median lobe apically of different morphology; paramere with subparallel lateral margins). For illustrations of the aedeagus of Q. truncicola see, e.g., COIFFAIT (1978) and SOLODOVNIKOV (2012).</p><p>D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The type specimens were collected in four geographically close localities in Mazandaran province, North Iran, all of them with window traps in forest habitats at altitudes of 1140-2050 m.</p></div>	https://treatment.plazi.org/id/F74687C8FFBDFFBAE69AFD96FC78886D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker	Assing, Volker (2019): On the taxonomy oI some West Palaearctic Quedius species, with descriptions oI new species and new synonymies (Coleoptera, Staphylinidae, Staphylininae). Linzer biologische Beiträge 51 (1): 189-201, DOI: 10.5281/zenodo.3762449
F74687C8FFB0FFBBE69AFD58FE7D8871.text	F74687C8FFB0FFBBE69AFD58FE7D8871.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius (Microsaurus) limans Assing 2019	<div><p>Quedius (Microsaurus) limans nov.sp. (Figs 13-15, 19-23)</p><p>T y p e m a t e r i a l: Holotype ♁: ̎ ISRAEL - Har Hermon, 2000 m, doline, pitfall, 33°18'17''N, 35°47'14''E, 3.VII.2010, C. Drees / Holotypus ♁ Quedius limans sp. n., det. V. Assing 2019̎ (cAss).</p><p>E t y m o l o g y: The specific epithet is the present participle of the Latin verb limare (to rasp) and alludes to the rasper- or grater-shaped apex of the median lobe of the aedeagus.</p><p>D e s c r i p t i o n: Large species; body length 14.5 mm; length of forebody 8.2 mm. Habitus as in Fig. 13. Coloration: body (including appendages) black with bright-reddish elytra and brown tarsomeres V.</p><p>Head (Fig. 14) large (sexual dimorphism?), 1.21 times as broad as long, broadest behind eyes; one macropuncture near middle of dorsal margin of eye, one approximately halfway between posterior margin of eye and posterior constriction of head, and one near posterior constriction of head; postero-lateral portion of head with dense macropunctation; dorsal surface with dense fine and distinct micropunctation; interstices with fine, but distinct transverse microsculpture. Eyes flat, approximately 0.7 times as long as postocular region in dorsal view. Antenna short, 3.5 mm long; antennomeres IV- X all distinctly transverse.</p><p>Pronotum (Fig. 14) 1.11 times as broad as long and 1.18 times as broad as head; dorsal series composed of 1+2 macropunctures; one smaller sublateral puncture situated behind level of posterior pair of dorsal punctures; disc with fine non-setiferous micropunctation and with fine microsculpture predominantly composed of striae directed antero-posteriad in lateral portion, oblique in median portion, and transverse in posterior portion.</p><p>Elytra (Fig. 14) 0.67 times as long as pronotum; punctation very dense and fine; pubescence blackish and depressed; interstices without microreticulation. Scutellum with fine transverse microreticulation and fine non-setiferous micropunctation. Hind wings fully developed.</p><p>Abdomen with dense and fine punctation, that of tergite VII as dense as that of tergite III; interstices with fine transverse microsculpture; pubescence depressed, greyish; posterior margin of tergite VII with palisade fringe.</p><p>♁: sternite VIII posteriorly with deep concavity and long brown marginal setae; aedeagus 1.6 mm long and shaped as in Figs 19-23.</p><p>♀: unknown.</p><p>C o m p a r a t i v e n o t e s: In habitus, coloration, and the short antennae, Q. limans somewhat resembles Q. brevicornis THOMSON, 1860. It is distinguished from this species by even larger size, the pronotal punctation pattern, much finer and denser punctation of the elytra and the abdomen, a deeper posterior excision of the male sternite VIII, and by the different shape of the aedeagus. For illustrations of the aedeagus of Q. brevicornis see, e.g., COIFFAIT (1978) and SOLODOVNIKOV (2012).</p><p>D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The type locality is situated in North Israel. The slightly teneral holotype was collected with a pitfall trap in a doline at an altitude of 2000 m.</p></div>	https://treatment.plazi.org/id/F74687C8FFB0FFBBE69AFD58FE7D8871	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Assing, Volker	Assing, Volker (2019): On the taxonomy oI some West Palaearctic Quedius species, with descriptions oI new species and new synonymies (Coleoptera, Staphylinidae, Staphylininae). Linzer biologische Beiträge 51 (1): 189-201, DOI: 10.5281/zenodo.3762449
