identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
F7118006FF8D7A56FEB1FF27FAB1FB41.text	F7118006FF8D7A56FEB1FF27FAB1FB41.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anatea formicaria Berland 1927	<div><p>Anatea formicaria Berland</p> <p>Figs 1–12, 42</p> <p>Anatea formicaria Berland, 1927: 55, figs 1–7.</p> <p>Not species figured bY Reiskind &amp; Levi, 1967: 21, figs 1–6 (misidentification, we refer this to sp. “J5”, Fig. 41).</p> <p>Diagnosis. Male: From other New Caledonian species shown in Figs 38–41, 43 by pedicel and carapace morphology and careful comparison of palp, particularly embolus length. From Australian species, leg I longer than leg III, pedicel with “node”, cymbium with extended apex, ventral abdomen with ventral plates (but note male of A. elongata sp. nov. is unknown). Female: from undescribed New Caledonian species by general morphology (as male) and details of genitalia; from Australian species by leg I longer than leg III, pedicel with “node”, dorsal scutum present.</p> <p>Type material (not examined). 1♂ tYpe, 1♀ cotYpe, New Caledonia. Found by Berland (dried) in the collection of ants of Ernest André. Berland does not state what he did with the specimens.</p> <p>Other material examined. NEW CALEDONIA: 1♀ QM S34107, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=166.66667&amp;materialsCitation.latitude=-22.083334" title="Search Plazi for locations around (long 166.66667/lat -22.083334)">Rivière Bleu</a>, Parc. 6, 22°05'S 166°40'E, 160 m, 9–22. iv.1987, L. Bonnet de Larbogne, J. Chazeau &amp; A. &amp; S. Tillier, Malaise trap, rainforest sassafras; 1♂ QM S34113, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=166.66667&amp;materialsCitation.latitude=-22.083334" title="Search Plazi for locations around (long 166.66667/lat -22.083334)">Rivière Bleu</a>, Parc. 7, 22°05'S 166°40'E, 170 m, 25.xi–8.xii.1986, L. Bonnet de Larbogne, J. Chazeau &amp; A. &amp; S. Tillier, Malaise trap; 1♂ 1♀ QM S20728, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=166.53334&amp;materialsCitation.latitude=-22.183332" title="Search Plazi for locations around (long 166.53334/lat -22.183332)">Mt Koghis</a>, 22°11'S 166°32'E, 500 m, 23–24.v.1987, R. Raven, rainforest, general and night collection; 1♀, QM S107230, Pic d’Amoa, N slopes, 20°58'S 165°17'E, 500 m, 27.xi.2003 – 30.1.2004, G. Monteith, with Pheidole sp. ant, flight intercept trap, SC11482; 1♂ KS.64090, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=164.76666&amp;materialsCitation.latitude=-20.633333" title="Search Plazi for locations around (long 164.76666/lat -20.633333)">Mt Panie</a>, 20°38'S 164°46'E, 6.vi.1996, M. Moulds; 1♀ WAM T86527, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=165.46666&amp;materialsCitation.latitude=-21.45" title="Search Plazi for locations around (long 165.46666/lat -21.45)">Col de Rossettes</a>, 21°27'S, 165°28'E, 11.ii.1993, Harvey, M.S., Platnick, N. I., Raven, R.J., rainforest litter.</p> <p>Size range. Male (n = 3). Carapace range, 0.75–0.80. Total length 2.05–2.15. Female (n = 3). Carapace range, 0.83–0.92. Total length 2.40–2.55. Berland (1927) gives the total length as 2.2 mm for the species. See Berland (1927) for full male description.</p> <p>Distribution. Recorded from mountainous rainforest sites across the main island of New Caledonia (Fig. 12).</p> <p>Notes. The habitus and genitalia of A. formicaria male and female are figured for comparison with the Australian species. Figure 1 is repeated as Fig. 42 to compare with other New Caledonian species.</p> <p>Whilst Berland (1927) admitted his hypothesis that A. formicaria was a mYrmecomorph could not be verified, he pointed out that it was found in a collection of ants and that the original collector had probably mistaken it for an ant; unfortunately, he did not (apparently) record which species or genera of ants the spiders were placed with in the collection. However, he comments (p. 53) that the spiders would match species of either Pheidole or Monomorium, both genera being well-represented in the New Caledonian ant fauna: En étudiant les Fourmis de Nouvelle-Calédonie, on trouve même sans trop de peine que l’Araignée se rapprocherait beaucoup d’un Pheidole ou d’un Monomorium, genres qui y sont très bien représentés.</p> <p>Reiskind &amp; Levi (1967) suggest different candidates as a model for the species they identify as Anatea formicaria (which we have re-identified as a closelY related species, “J5”, see Figs 41, 42), they write: “ Anatea formicaria is an accurate mimic of the small myrmicine ant Chelaner croceiventre (Emery), 2.6 mm long, which has been collected at the same locality as the spider. The color pattern (dark brown anterior and light, yellow-brown posterior) of both is quite rare in ants and is also found in specimens of Xiphomyrma tenuierius Emery, 2.9 mm. long, and a species of Lordomyrma, 4.8 mm long, two myrmicine ants found in the same area of rain forest (E. O. Wilson, pers. comm.).” The colour description given above, does not match the specimens we have seen of either A. formicaria or “J5”, in which the cephalothorax is only slightly darker than the yellow-amber ground colour of the abdominal scute (see Figs 41, 42). This difference maY be due to time since preservation or could represent colour polymorphism within the species “J5”, as is recorded in some other myrmecomorphs (see examples in Cushing, 1997). Berland’s original description of A. formicaria (1927) gives colouration of cephalothorax, pedicel and abdominal scuta as “light reddish fawn” (“fauve rougeâtre clair”) but this is of minimal help as these were dried specimens of uncertain age.</p> <p>Three of the Anatea samples we have examined each contain a single ant, which we deduce may have been included by collector or sorter of the specimens as a putative model. All three ants are of similar appearance (but may not be conspecific) and are from the genus Pheidole. The Pheidole specimens are of pale golden colour, similar to A. formicaria or our species “J5” (and possibly similar to juveniles of species “J1”). All these three Anatea species occur together: QM S10730 and QM S10732 that now contain A. fomicaria and species “J5”, respectively, were separated from QM S88025, which now contains species “J1”; QM S88026 contains adults and a juvenile of species “J1” and a subadult A.? formicaria. Of note is the size of the ants, which at c. 1.6 mm total length are considerably smaller than adult Anatea of either similarly coloured species, and slightly smaller than the sub adult male Anatea spp. included in two of the samples.</p> </div>	https://treatment.plazi.org/id/F7118006FF8D7A56FEB1FF27FAB1FB41	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Smith, Helen M.;Harvey, Mark S.;Agnarsson, Ingi;Anderson, Gregory J.	Smith, Helen M., Harvey, Mark S., Agnarsson, Ingi, Anderson, Gregory J. (2017): Notes on the Ant-mimic Genus Anatea Berland (Araneae: Theridiidae) and Two New Species from Tropical Australia. Records of the Australian Museum (Rec. Aust. Mus.) 69 (1): 1-13, DOI: 10.3853/j.2201-4349.69.2017.1672, URL: http://dx.doi.org/10.3853/j.2201-4349.69.2017.1672
F7118006FF8C7A55FF06FF27FE79F9F4.text	F7118006FF8C7A55FF06FF27FE79F9F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anatea monteithi Smith & Harvey & Agnarsson & Anderson 2017	<div><p>Anatea monteithi Smith sp. nov.</p> <p>Figures 13–24, 31, 35–37</p> <p>Holotype ♀ QM S25842, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.61667&amp;materialsCitation.latitude=-16.716667" title="Search Plazi for locations around (long 145.61667/lat -16.716667)">Mt Formartine South</a>, 10 km N Kuranda, [16°43'S 145°37'E, Queensland, Australia] 700 m, 23.xi.1990, G. Monteith, G. Thompson, pyrethrum trees &amp; logs. Paratypes. 1♂, WAM T99420, Daintree NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.43611&amp;materialsCitation.latitude=-16.237501" title="Search Plazi for locations around (long 145.43611/lat -16.237501)">Alexandra Range</a> lookout point, 16°14'15"S 145°26'10"E, 6.iv.2009, K. Edward, J. Waldock, sieved litter (WT 11); 1♂, AM KS.126452, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.60583&amp;materialsCitation.latitude=-16.751112" title="Search Plazi for locations around (long 145.60583/lat -16.751112)">Black Mountain Rd</a> , start of track to Mt Formartine, 16°45'04"S 145°36'21"E, 425 m, 6.xii.2008, G. Milledge, H. Smith, beat, sweep; [right palp on SEM stub s/1049; right leg I on s/1050]; both Queensland, Australia.</p> <p>Other material examined (Queensland, Australia): 1♀ AM KS.116502, Lake Barrine, E of Yungaburra, 17°15'S 145°38'E, 31.viii.2011, M. Zabka, B. Patoleta; [abdomen on SEM stub s/1048, legs I, IV on s/1047; epi cleared]. 1 juv. AM KS.7647, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.36667&amp;materialsCitation.latitude=-16.166666" title="Search Plazi for locations around (long 145.36667/lat -16.166666)">Thornton Peak</a>, N of Daintree, 16°10'S 145°22'E, Nov. 1975, M. Gray, leaf litter sample (RF survey site 40). QM 1 juv. (unregistered), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.85&amp;materialsCitation.latitude=-17.266666" title="Search Plazi for locations around (long 145.85/lat -17.266666)">Bellenden Ker Ra</a>, 0.5 km S Cable Tower 7, 17°16'S 145°51'E, 500 m, 25–31.x.1981, Earthwatch / Qld Museum, rainforest, pyrethrum logs, stones, tree trunks.</p> <p>Etymology. The species is named for Geoff Monteith, in recognition of his enormous contributions to the study of Australian invertebrates through his collecting activities for the Queensland Museum.</p> <p>Diagnosis. From New Caledonian species by leg III longer than legs I and II; pedicel without “node”; male without extended tip to cymbium; from A. elongata sp. nov. by pedicel shorter than carapace; female insemination ducts with short conjoined section, entering spermathecae posteriorly (Fig. 22).</p> <p>Description</p> <p>Colour (in alcohol) (Figs 13–19). Cephalothorax, mouthparts, sternum, pedicel and male scutum dark chestnut brown, glossy, except pedicel rugose, female abdomen and parts of male abdomen not covered by scutum dark charcoal grey, slightly paler ventrally, cuticle glossy between sparse but prominent setal bases; dorsal and ventral abdominal apodemes dark brown, obvious. Legs: (female) coxae and trochanters white on legs III, IV, suffused with brownish black on legs I, II; femora pale brown laterally, dark dorsally and ventrally (legs I, II) or vis. v. (legs III, IV), femora legs III, IV with distal white band extending onto proximal patella, traces of same legs I, II; tibiae-tarsi mid-brown proximally to amber brown distally, darkest coloration ventral on anterior legs to retrolateral on leg IV; male legs similar but darker, stronger coloration.</p> <p>Carapace (Figs 13–15, 18) longer than wide, widest at coxae II, smooth transition to caput; fovea absent; in profile gently domed, highest at coxae I (male WAM T99420 rather flatter than female holotYpe, other male similar to female); eye group 3/5 carapace width; AME largest (Figs 14, 15) and prominent above slightly concave clypeus, clypeus&gt; 2 × AME diameter in male, slightly lower in female. Labium bluntly triangular (Fig. 16), cheliceral bases shorter than maxillae, flexible with slender, curved fangs. Female palpal claw weakly palmate with three or four large teeth. Legs: 4312 (holotype female missing both fourth legs but confirmed in female from other material). Legs with sparse dorsal macrosetae on patellae and tibiae (a few visible in Figs 13, 14), hadrotarsine flat-topped setae (see Agnarsson, 2004) present on distal tarsus I, few on II. Pedicel sclerotized, open “S” in lateral view, slightly expanded anterior to apex and constriction at abdominal articulation (Fig. 18). Abdomen egg-shaped, widest anterior to mid-point (Figs 17, 19), constricted by sclerotized band around spinnerets; male with dorsal scutum covering central abdomen but leaving bare small surround of cuticle in dorsal view (Fig. 13), sclerotized around pedicel and genital plate (Fig. 17); female without scutum (Fig. 18) and only small sclerotized band around pedicel insertion (Fig. 19). One pair of dorsal apodemes, prominent in female, embedded in scutum of male; three circular ventral plates/apodemes in both sexes (Figs 17, 19). Genitalia. Male: Cy blunt with two short, broad based tooth-like modified spines (Fig. 21). Palp with Co, TTA, MA (Fig. 35); TR extends ventrally to wrap over the ES, Co arises from apicodorsal T and apicolateral tegular rim (Figs 35, 37). TTA large, arising basally, extending beyond tip of Cy. ES tip resting between TTA and Co. MA base and basal E conjoined proventrally (Figs 35, 36). Female: externallY with ridges and grooves in lateral profile (Fig. 23), projecting lip on posterior margin of ovoid fossa. Internally (Fig. 22), ID exit fossa posteriorly and almost immediately turn anterolaterally, entering spermathecae Sp at posterior end. Fertilisation ducts FD exit adjacent to ID, with terminal nodule (possibly degenerate second pair of Sp) before terminal part of FD.</p> <p>Measurements. Male. Carapace range, 0.81 to 0.88 (n = 2). WAM T99420. Total length, 2.35; carapace length, 0.88; width, 0.62; height, 0.37; abdomen length, 1.25; width, 0.91; pedicel length, 0.50; clypeus height, 0.22. Eyes: AME, 0.10; PME, 0.05; ALE, 0.06; PLE, 0.06; Limbs (femur + (patellatibia) + metatarsus + tarsus = total): leg I, 0.49 + 0.51 + 0.28 + 0.24 = 1.51; leg II, 0.44 + 0.54 + 0.25 + 0.29 = 1.51; leg III, 0.50 + 0.56 + 0.24 + 0.33 = 1.63; leg IV, 0.55 + 0.84 + 0.33 + 0.36 = 2.08.</p> <p>Female. Carapace range, 0.86 (n = 2). Holotype QM S25842. Total length, 2.88; carapace length, 0.86; width, 0.69; height, 0.33; abdomen length, 1.68; width, 1.32; pedicel length, 0.56; clypeus height, 0.16. Eyes: AME, 0.10; PME, 0.06; ALE, 0.07; PLE, 0.06; Limbs (femur + (patella-tibia) + metatarsus + tarsus = total): leg I, 0.49 + 0.60 + 0.31 + 0.26 = 1.66; leg II, 0.48 + 0.58 + 0.26 + 0.30 = 1.61; leg III, 0.50 + 0.61 + 0.26 + 0.34 = 1.71; leg IV, missing (but longest in female AM KS.116502).</p> <p>Distribution. Northeast Queensland. Recorded from rainforests, mainly at higher altitudes, between Thornton Peak and Bellenden Ker, and on the Atherton Tableland at Yungaburra (Fig. 31).</p> <p>Notes. Without observations of behaviour, it is unclear exactly if, or how, ant mimicry is achieved in the Australian species, but as in New Caledonian species, the long pedicel and overall appearance are ant-like. Relative leg length, with both III and IV longer than I or II is unusual and may indicate behavioural modifications.</p> <p>The tropical rainforests of northeastern Queensland occupied by A. monteithi hold an extremely high diversity of ants, with over 66 genera recorded in these areas (Antwiki, 2017). If A. monteithi is indeed an ant mimic, candidate models are found in several myrmecine genera. Examples of genera with species in an appropriate size range for adult spiders include Monomorium, Orectognathus, Pheidole, Pristomyrmex and Vombisidris.</p> </div>	https://treatment.plazi.org/id/F7118006FF8C7A55FF06FF27FE79F9F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Smith, Helen M.;Harvey, Mark S.;Agnarsson, Ingi;Anderson, Gregory J.	Smith, Helen M., Harvey, Mark S., Agnarsson, Ingi, Anderson, Gregory J. (2017): Notes on the Ant-mimic Genus Anatea Berland (Araneae: Theridiidae) and Two New Species from Tropical Australia. Records of the Australian Museum (Rec. Aust. Mus.) 69 (1): 1-13, DOI: 10.3853/j.2201-4349.69.2017.1672, URL: http://dx.doi.org/10.3853/j.2201-4349.69.2017.1672
F7118006FF8E7A5BFF7EF91AFC11FE45.text	F7118006FF8E7A5BFF7EF91AFC11FE45.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anatea elongata Smith & Harvey & Agnarsson & Anderson 2017	<div><p>Anatea elongata Smith sp. nov.</p> <p>Figures 25–31</p> <p>Holotype ♀ ANIC 42 002257, 12 km SSE <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.63333&amp;materialsCitation.latitude=-11.85" title="Search Plazi for locations around (long 142.63333/lat -11.85)">Heathlands</a>, 11°51'S 142°38'E [Queensland, Australia], 25.iv–7.vi.1992, T. McLeod, FIT#2, F.I.T., ANIC 1248, closed forest.</p> <p>Etymology. The species is named in reference to the pedicel, the longest compared to carapace length so far observed in genus Anatea.</p> <p>Diagnosis. From New Caledonian species by leg III longer than legs I and II; pedicel longer than carapace, without “node”; from A. monteithi sp. nov. by pedicel longer than carapace; insemination ducts with long conjoined section, crossing ventral spermathecae to entry point towards anterior (Fig. 30).</p> <p>Description</p> <p>Colour (in alcohol). Probably similar to A. monteithi but specimen extremely faded, pedicel appears rather reddishbrown; coxae brown, trochanters white on legs III and IV; otherwise apparently similar to A. monteithi except extent of abdominal colour cannot be established as only 1 patch remains.</p> <p>Carapace (Figs 25, 26, 29) generally similar to female of A. monteithi; eye group slightly over half carapace width (Fig. 29); eyes and mouthparts similar to A. monteithi. Female palpal claw strongly palmate with at least 8 teeth. Legs: 43(12). Pedicel sclerotized, almost straight ventrally in anterior half then arching posteriorly, narrowest anteriorly and thickest, slightly ridged at mid-point, much longer than carapace (Figs 25–27). Abdomen as in A. monteithi. Legs missing any visible setae but setal bases suggest macrosetae probably present as in A. monteithi.</p> <p>Genitalia. ExternallY with fine edge curving anterior to ovoid fossa, but otherwise apparentlY smooth in profile. Internal genitalia not dissected but visible through cuticle (Fig. 30), ID exit fossa to posterior, running together for 2/3 of distance to genital groove, then turning away from each other and simultaneously turning dorsally, crossing ventral Sp and entering anteroventrally. Sp lying to either side of fossa in ventral view. FD not visible.</p> <p>Measurements. Female.Total length, 3.04; carapace length, 0.93; width, 0.72; height, 0.38; abdomen length, 1.32; width, 0.90; pedicel length, 1.06; clypeus height, 0.17. Eyes: AME, 0.09; PME, 0.06; ALE, 0.06; PLE, 0.06; Limbs (femur + (patella-tibia) + metatarsus + tarsus = total): leg I, 0.52 + 0.60 + 0.30 + 0.26 = 1.68; leg II, 0.50 + 0.60 + 0.28 + 0.30 = 1.68; leg III, 0.58 + 0.68 + 0.32 + 0.38 = 1.96; leg IV, 0.70 + 0.98 + 0.42 + 0.42 = 2.52.</p> <p>Male. Unknown</p> <p>Distribution. Northeast Queensland. Recorded only from the type locality on the Cape York Peninsula (Fig. 31).</p> <p>Notes. The single female has lost most of its colour and may not be robust enough to excise the genitalia without severely</p> <p>damaging the specimen. Due to the lack of pigment, the external view is sufficient to see the route of the ducts and this is clearlY different from A. monteithi.</p> <p>The habitat around the type locality is generally savannah woodland, or “closed forest” on the label, a contrast to the rainforest habitat of A. monteithi. The condition of the specimen is not good enough to determine gloss (except carapace) or be sure of colour. However, if the species is an ant mimic the extreme extension of the pedicel, should facilitate the identification of the model in ant samples taken from the area.</p> </div>	https://treatment.plazi.org/id/F7118006FF8E7A5BFF7EF91AFC11FE45	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Smith, Helen M.;Harvey, Mark S.;Agnarsson, Ingi;Anderson, Gregory J.	Smith, Helen M., Harvey, Mark S., Agnarsson, Ingi, Anderson, Gregory J. (2017): Notes on the Ant-mimic Genus Anatea Berland (Araneae: Theridiidae) and Two New Species from Tropical Australia. Records of the Australian Museum (Rec. Aust. Mus.) 69 (1): 1-13, DOI: 10.3853/j.2201-4349.69.2017.1672, URL: http://dx.doi.org/10.3853/j.2201-4349.69.2017.1672
