taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
4E574931E2725135B5A689A4E7DD70E0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/347596	https://doi.org/10.3897/zookeys.881.38026.figure1	Figure 1. Dorsal drawing of a male Onthophagus humboldti sp. nov.	Figure 1. Dorsal drawing of a male Onthophagus humboldti sp. nov.	2019-12-31	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.		Zenodo	biologists	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.			
4E574931E2725135B5A689A4E7DD70E0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/347597	https://doi.org/10.3897/zookeys.881.38026.figure2	Figure 2. Drawings of the clypeal horn of a O. humboldti sp. nov. and b O. micropterus; pronotum of c O. humboldti sp. nov. and d O. micropterus; head of e O. humboldti sp. nov. and f O. micropterus; and elytral apex of g U. bonplandi sp. nov. and h U. dybasi. Scale bars: 1 mm.	Figure 2. Drawings of the clypeal horn of a O. humboldti sp. nov. and b O. micropterus; pronotum of c O. humboldti sp. nov. and d O. micropterus; head of e O. humboldti sp. nov. and f O. micropterus; and elytral apex of g U. bonplandi sp. nov. and h U. dybasi. Scale bars: 1 mm.	2019-12-31	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.		Zenodo	biologists	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.			
4E574931E2725135B5A689A4E7DD70E0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/347598	https://doi.org/10.3897/zookeys.881.38026.figure3	Figure 3. Aedeagui of a O. humboldti sp. nov. b U. bonplandi sp. nov. c brachypterous wing of O. humboldti sp. nov.	Figure 3. Aedeagui of a O. humboldti sp. nov. b U. bonplandi sp. nov. c brachypterous wing of O. humboldti sp. nov.	2019-12-31	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.		Zenodo	biologists	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.			
4E574931E2725135B5A689A4E7DD70E0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/347600	https://doi.org/10.3897/zookeys.881.38026.figure5	Figure 5. Known distribution of O. humboldti sp. nov. (orange circle) and U. bonplandi sp. nov. (red triangle). The distribution of the proposed sister species of these new taxa is also depicted, O. micropterus (blue rhombus) and U. dybasi (black square).	Figure 5. Known distribution of O. humboldti sp. nov. (orange circle) and U. bonplandi sp. nov. (red triangle). The distribution of the proposed sister species of these new taxa is also depicted, O. micropterus (blue rhombus) and U. dybasi (black square).	2019-12-31	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.		Zenodo	biologists	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.			
4E574931E2725135B5A689A4E7DD70E0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/347601	https://doi.org/10.3897/zookeys.881.38026.figure6	Figure 6. Known distribution of brachypterous Scarabaeinae in Costa Rica.	Figure 6. Known distribution of brachypterous Scarabaeinae in Costa Rica.	2019-12-31	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.		Zenodo	biologists	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.			
4E574931E2725135B5A689A4E7DD70E0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/347602	https://doi.org/10.3897/zookeys.881.38026.figure7	Figure 7. Present day distribution of Onthophagus humboldti sp. nov. (blue dot) and O. micropterus (red dot) and lines indicating proposed localities (rhombi) depressed by 1500 m (14 km in straight line) generated by the last glacial maximum, ~ 25 - 23 ka, in the Cordillera de Talamanca. All mountain systems are 150 m lower than present day height and an estimated sea level descent of 120 m is depicted. Dotted black lines represent present-day sea levels.	Figure 7. Present day distribution of Onthophagus humboldti sp. nov. (blue dot) and O. micropterus (red dot) and lines indicating proposed localities (rhombi) depressed by 1500 m (14 km in straight line) generated by the last glacial maximum, ~ 25 - 23 ka, in the Cordillera de Talamanca. All mountain systems are 150 m lower than present day height and an estimated sea level descent of 120 m is depicted. Dotted black lines represent present-day sea levels.	2019-12-31	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.		Zenodo	biologists	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.			
4E574931E2725135B5A689A4E7DD70E0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/347604	https://doi.org/10.3897/zookeys.881.38026.figure9	Figure 9. A cytochrome c-oxidase I (COI- 5 P) mitochondrial DNA sequence-based BOLD taxon ID tree of the nearest species of O. humboldti sp. nov.	Figure 9. A cytochrome c-oxidase I (COI- 5 P) mitochondrial DNA sequence-based BOLD taxon ID tree of the nearest species of O. humboldti sp. nov.	2019-12-31	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.		Zenodo	biologists	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.			
CD2B9D8B870A5AA787E67D6936B3E89F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/347597	https://doi.org/10.3897/zookeys.881.38026.figure2	Figure 2. Drawings of the clypeal horn of a O. humboldti sp. nov. and b O. micropterus; pronotum of c O. humboldti sp. nov. and d O. micropterus; head of e O. humboldti sp. nov. and f O. micropterus; and elytral apex of g U. bonplandi sp. nov. and h U. dybasi. Scale bars: 1 mm.	Figure 2. Drawings of the clypeal horn of a O. humboldti sp. nov. and b O. micropterus; pronotum of c O. humboldti sp. nov. and d O. micropterus; head of e O. humboldti sp. nov. and f O. micropterus; and elytral apex of g U. bonplandi sp. nov. and h U. dybasi. Scale bars: 1 mm.	2019-12-31	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.		Zenodo	biologists	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.			
CD2B9D8B870A5AA787E67D6936B3E89F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/347598	https://doi.org/10.3897/zookeys.881.38026.figure3	Figure 3. Aedeagui of a O. humboldti sp. nov. b U. bonplandi sp. nov. c brachypterous wing of O. humboldti sp. nov.	Figure 3. Aedeagui of a O. humboldti sp. nov. b U. bonplandi sp. nov. c brachypterous wing of O. humboldti sp. nov.	2019-12-31	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.		Zenodo	biologists	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.			
CD2B9D8B870A5AA787E67D6936B3E89F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/347599	https://doi.org/10.3897/zookeys.881.38026.figure4	Figure 4. Dorsal drawing of a male Uroxys bonplandi sp. nov.	Figure 4. Dorsal drawing of a male Uroxys bonplandi sp. nov.	2019-12-31	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.		Zenodo	biologists	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.			
CD2B9D8B870A5AA787E67D6936B3E89F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/347600	https://doi.org/10.3897/zookeys.881.38026.figure5	Figure 5. Known distribution of O. humboldti sp. nov. (orange circle) and U. bonplandi sp. nov. (red triangle). The distribution of the proposed sister species of these new taxa is also depicted, O. micropterus (blue rhombus) and U. dybasi (black square).	Figure 5. Known distribution of O. humboldti sp. nov. (orange circle) and U. bonplandi sp. nov. (red triangle). The distribution of the proposed sister species of these new taxa is also depicted, O. micropterus (blue rhombus) and U. dybasi (black square).	2019-12-31	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.		Zenodo	biologists	Kohlmann, Bert;Solis, Angel;Alvarado, Guillermo E.			
