taxonID	type	description	language	source
F60287B8811B1C0565BFFF136C07C9D8.taxon	description	(Figs 1 – 3, 5, 13)	en	Hemala, Vladimír, Kocorek, Anna, Lis, Jerzy A. (2020): Megymenum tuberculatum, a new species of Megymenini from Java and a review of distribution of M. brevicorne (Hemiptera: Heteroptera: Dinidoridae). Zootaxa 4808 (3): 439-454, DOI: 10.11646/zootaxa.4808.3.2
F60287B8811B1C0565BFFF136C07C9D8.taxon	materials_examined	Type locality. Indonesia, Java. Type material. Holotype: ♀ (MHNG), INDONESIA: Java: ‘ 418 / 65 Java. / M. Melly. ’ [hw, ivory label] // ‘ ♀ ’ [p, small white label] // ‘ HOLOTYPUS / Megymenum / tuberculatum / sp. nov. / det. V. HEMALA & A. KO- COREK 2017 [p, red label] ’. The holotype is card-mounted, with a pin-hole in pronotum and right hemelytra; right basi- and distiflagellum, left antenna, labial segments III and IV, right and left protarsal segments II and III, right middle leg, and right hind leg all missing.	en	Hemala, Vladimír, Kocorek, Anna, Lis, Jerzy A. (2020): Megymenum tuberculatum, a new species of Megymenini from Java and a review of distribution of M. brevicorne (Hemiptera: Heteroptera: Dinidoridae). Zootaxa 4808 (3): 439-454, DOI: 10.11646/zootaxa.4808.3.2
F60287B8811B1C0565BFFF136C07C9D8.taxon	description	Description of female (holotype). Coloration (Figs 1 – 3). Body dark brown, only labium light brown and membranes of hemelytra yellowish with dark brown spots at the base. Small fields around abdominal trichobothria are also light brown. Punctation. Head, anterior pronotal tubercle, posterior pronotal lobe, scutellum, clavus, connexiva and abdominal laterotergites with small, dense punctures. Punctation of anterior pronotal lobe, ventral sides of pro-, meso- and metathorax and abdominal ventrites with lower density. Pilosity. The vast majority of the body surface without pilosity, only very small areas around coxae and a shallow depression between valvifers VIII and laterotergites IX covered by sparse and very fine light brown colored hairs at their anterior sides. All tibiae apically covered with small semierect setae. Ventral side of tarsi covered by very fine and very dense setae forming a pillow. Structure. Body medium sized, elongate (Fig. 1), widest across mid of abdomen (between apices of lobes on laterotergites IV), only slightly convex dorsally, strongly convex ventrally, thorax trapezoid in cross section (Fig. 5). Head slightly wider than long; clypeus very small, mandibular plates longer than clypeus, together heart-shaped and strongly concave dorsally. Anteocular spines large, triangular but not sharp. Base of head wide (widest across eyes); small compound eyes positioned posterolaterally, nearly in touch with anterior part of pronotum. Antenniferous tubercle very small, short and not visible in dorsal view. Anteocular processes present and visible from dorsal view. Scape (I) very short, shorter than mandibular plate; pedicel (II) nearly flattened, 2.5 times longer than segment I and longer than mandibular plate. Basi- (III) and distiflagellum (IV) mutilated. Bucculae round, very short, but longer than labrum. Labial segment I shorter than segment II, clearly reaching posterior margin of head, segments III and IV mutilated. Thorax. Pronotum (Figs 1, 5) trapezoid in dorsal view, without well differentiated collar-like structure. Anterior margin of pronotum slightly concave; anterolateral margin only very slightly projecting forward to form very small and rounded process, only very slightly concave anteriad from this process and nearly straight posteriad from this process. Posterolateral margin slightly rounded, as long as 2 / 3 of anterolateral margin; posterior margin along base of scutellum slightly convex. Pronotal surface tuberculate, with very large rounded tubercle in the middle of its anterior part, positioned at the anterior margin. This tubercle is 1.4 times wider than long. Disc with depressions at anterolateral angles. Scutellum only slightly longer than wide at its base, reaching about middle of abdominal length; regularly narrowing posteriad. Prosternum with tubercles around the coxae, meso- and metasternum nearly flat, without tubercles. External scent efferent system with large ostiole and evaporatorium. Ostiole positioned nearly in ventral 1 / 4 of metapleuron. Peritreme in form of short, narrow spout (about twice as long as ostiole), directed slightly posterolaterad with rounded apex. Evaporatorium wide, placed on the strongly folded cuticle, covering the lateral 1 / 3 of vestibulum, central part of metepisternum anteriorly (in shape of wide semicircle) and narrow strip in central part of mesepimeron posteriorly, surrounding the whole peritreme and nearly the whole metathoracic spiracle. Metathoracic spiracle narrow, straight, placed in the middle of suture between meso- and metapleuron. Hemelytra. Clavus short, slightly surpassing half-length of scutellum. Corium slightly shorter than scutellum, lacking apparent veins; anterodistal (seemingly posterolateral) angle rounded. Membrane large, twice as long as clavus; apically widely rounded, reaching apex of abdomen. Venation on membrane reticulate. Legs. Pro- and mesofemur clavate and shorter than metafemur, metafemur rather cylindrical and longer than pro- and mesofemur. Mesofemur 1.22 times longer than profemur, and metafemur 1.23 times longer than mesofemur. Profemora with two longitudinal series of denticles after four denticles on ventral side apically. Mesofemur and metafemur with two longitudinal series of denticles after three denticles on ventral side apically. Length of dencticles on femora gradually increased towards apex. Femora apically flat between the rows of denticles. Pro- and mesotibia shorter than metatibia, metatibia longer than pro- and mesotibia and bearing the tympanal organ (seen in Dinidoridae females only). Mesotibia 1.13 times longer than protibia, and metatibia 1.44 times longer than mesotibia. Tarsi 3 - segmented; protarsal segments II and III mutilated. Abdomen. Connexiva not covered by hemelytra; posterolateral angles of connexival segments with two distinct small lobes on each one segment, of which anterior one very small, posterior one larger and well developed. Spiracle II exposed, situated slightly more ventrally than spiracles III – VIII. Spiracle VIII clearly smaller than spiracles II – VII, and positioned more laterally on laterotergite VIII. Two trichobothria situated on each laterotergite III – VII, postspiracular, the anterior one positioned more laterally, the posterior one more ventrally; each trichobothrium situated in separate small field. External female genitalia. Valvifers VIII of quarter-circular shape, not fused medially; laterotergites IX trapeziform. Spermatheca (Fig. 13). Apical receptacle (spermathecal bulb) spherical, medium sized. Intermediate part (pumping region) short, well defined, with distinct and large distal and proximal flanges. Spermathecal duct large, strongly dilated (sac-like), with a distinct lateral fold bearing inconspicuous ring sclerite. Spines of spermathecal duct distinct and large, covering only a small L-shaped area postero-lateraly of the ring sclerite. Measurements (of holotype, in mm; n = 1). Body length (from apex of clypeus to apex of abdomen) 13.18; head: length (from apex of clypeus to anterior margin of pronotum) 2.01, width (maximum width across eyes) 2.56, height (height across compound eye) 1.62, interocular width (between inner margins of compound eyes) 1.78; lengths of antennal segments: scape (I) — 0.85, pedicel (II) — 1.71, basiflagellum (III) —?, distiflagellum (IV) —?; length of labial segments: I — 1.32, II — 1.40, III —?, IV —?; pronotum: length (medially) 3.72, anterior width (between anterolateral angles) 4.65, length of anterior tubercle (medially) 1.24, width of anterior tubercle (in middle) 1.74; posterior width (maximum width between humeral angles) 5.97; scutellum: length (medially from base to apex) 3.87, width (maximum width at base) 3.41; corium: length 4.03, width 1.86; abdomen width (maximum width between apices of lobes on laterotergites IV) 7.32; length of femora: profemur 2.79, mesofemur 3.41, metafemur 4.18; length of tibiae: protibia 2.48, mesotibia 2.79, metatibia 4.03; length of tarsi: protarsus?, mesotarsus 1.24, metatarsus 1.24. Male. Unknown.	en	Hemala, Vladimír, Kocorek, Anna, Lis, Jerzy A. (2020): Megymenum tuberculatum, a new species of Megymenini from Java and a review of distribution of M. brevicorne (Hemiptera: Heteroptera: Dinidoridae). Zootaxa 4808 (3): 439-454, DOI: 10.11646/zootaxa.4808.3.2
F60287B8811B1C0565BFFF136C07C9D8.taxon	diagnosis	Differential diagnosis. The habitus of M. tuberculatum sp. nov. is very similar to that of M. brevicorne (Fabricius, 1787) (see Figs 1, 4), but the new species differs in the characters on head, antennae, pronotum, prosternum, connexiva and spermatheca. The crucial character for separating the new species is distinctly larger anterodorsal tubercle on pronotum, but in contrast to M. brevicorne, the new species has also: i) anteocular spines on head present (only sporadically found in M. brevicorne; see Figs 1, 4), ii) pedicel more wider and more flattened, iii) pronotum with not well differentiated collar-like structure due to the low concavity of its anterolateral margin (see Figs 5, 6), iv) prosternum with tubercles around the coxae (not present in M. brevicorne), v) connexiva with smaller and not so sharp lobes as in M. brevicorne (see Figs 1, 2, 4), vi) intermediate part (pumping region) of spermatheca clearly shorter than in M. brevicorne (see Figs 13, 14), vii) spermathecal duct slightly shorter and more dilated lateraly than in M. brevicorne (see Figs 13, 14), viii) slightly more distinct lateral fold of spermathecal duct, but inconspicuous ring sclerite (slightly less distinct lateral fold and more distinct ring sclerite in M. brevicorne; see Figs 13, 14) and ix) L-shaped spine area of spermathecal duct (only longitudinal in M. brevicorne; see Figs 13, 14). Anterodorsal tubercle on pronotum of the new species is the largest among all known species of the genus Megymenum which possess this tubercle (Figs 5 – 12).	en	Hemala, Vladimír, Kocorek, Anna, Lis, Jerzy A. (2020): Megymenum tuberculatum, a new species of Megymenini from Java and a review of distribution of M. brevicorne (Hemiptera: Heteroptera: Dinidoridae). Zootaxa 4808 (3): 439-454, DOI: 10.11646/zootaxa.4808.3.2
F60287B8811B1C0565BFFF136C07C9D8.taxon	etymology	Etymology. The species name is the Latin adjective tuberculatum (= tuberculate), referring to the characteristic large anterodorsal tubercle of pronotum, which is distinctly larger than such tubercle in all other known species of the genus.	en	Hemala, Vladimír, Kocorek, Anna, Lis, Jerzy A. (2020): Megymenum tuberculatum, a new species of Megymenini from Java and a review of distribution of M. brevicorne (Hemiptera: Heteroptera: Dinidoridae). Zootaxa 4808 (3): 439-454, DOI: 10.11646/zootaxa.4808.3.2
F60287B8811B1C0565BFFF136C07C9D8.taxon	biology_ecology	Biology. Unknown.	en	Hemala, Vladimír, Kocorek, Anna, Lis, Jerzy A. (2020): Megymenum tuberculatum, a new species of Megymenini from Java and a review of distribution of M. brevicorne (Hemiptera: Heteroptera: Dinidoridae). Zootaxa 4808 (3): 439-454, DOI: 10.11646/zootaxa.4808.3.2
F60287B8811B1C0565BFFF136C07C9D8.taxon	distribution	Distribution. Indonesia: Java (without exact locality). Note on collector name. The collector name “ M. Melly ” written on the first ivory label probably refers to André Melly (1802 – 1851) who gathered a large entomological collection in MHNG (over 22 000 species from around the world) consisting especially of beetles (Coleoptera), but also of other insect orders (viz Schaum 1852).	en	Hemala, Vladimír, Kocorek, Anna, Lis, Jerzy A. (2020): Megymenum tuberculatum, a new species of Megymenini from Java and a review of distribution of M. brevicorne (Hemiptera: Heteroptera: Dinidoridae). Zootaxa 4808 (3): 439-454, DOI: 10.11646/zootaxa.4808.3.2
F60287B8811F1C0E65BFFBF96A6DC9F3.taxon	description	(Figs 4, 6, 14)	en	Hemala, Vladimír, Kocorek, Anna, Lis, Jerzy A. (2020): Megymenum tuberculatum, a new species of Megymenini from Java and a review of distribution of M. brevicorne (Hemiptera: Heteroptera: Dinidoridae). Zootaxa 4808 (3): 439-454, DOI: 10.11646/zootaxa.4808.3.2
F60287B8811F1C0E65BFFBF96A6DC9F3.taxon	materials_examined	Material examined. CHINA: unspecified: no exact data, 1 ♀, Nonfried lgt., A. Kocorek det. (DBOU). Guangdong: Canton [= Guangzhou], 12. x. 1911, 1 ♀, S. V. Mell lgt., J. A. Lis 1988 det. (DBOU). Hainan: Qiongshan, vi. 1961, 1 ♂ [lost pygophore], from winter melon [= Benincasa hispida], X. S. Tan lgt., A. Kocorek det. (DBOU; translated from Chinese by D. Rédei); Bawangling Mts., Baotie env., 19 ° 05.3 ′ N 109 ° 07.4 ′ E, 415 – 800 m a. s. l., 7. – 8. v. 2011, 1 ♂, at light, M. Fikáček, V. Kubeček & L. Li lgt., V. Hemala det. (NMPC). Yunnan: no exact locality, no date, 1 ♂, no collector, R. Kumar 1973 det. (DBOU). — NEPAL: Central Nepal, Chitwan (Roy. Nat. Park), Sauraha village, 27.35 N 84.30 E, 166 m a. s. l., 1 ♀, D. Král lgt., V. Hemala det. (NMPC). — THAILAND: Chiang-Mai prov., Chiang-Mai surroundings, 1. vii. 1995, 4 ♂♂ 1 ♀ 12 L, W. G. Ullrich lgt., A. Kocorek det. (WULG); North Thailand, Muang Nan, viii. 1995, 1 ♂, W. G. Ullrich lgt., A. Kocorek det. (WULG); Chiang Mai-Chiang Dao, 5. – 10. vii. 1997, 1 ♀, M. Klícha lgt., V. Hemala det. (NMPC); ca. 5 km S of Mae Hong Son, Pai River, high water, flooding detri- tus / vegetation, 19 ° 14 ′ 12.2 ″ N 97 ° 53 ′ 41.1 ″ E, 23. vii. 1997, 1 ♂ 4 ♀♀, W. G. Ullrich lgt., A Kocorek det. (WULG); Chiang Dao, 27. v. – 2. vi. 2002, 1 ♀, B. Makovský lgt., V. Hemala det. (NMPC); Trat prov., Ko Chang island, 12 ° 05 ′ N 102 ° 21 ′ E, 0 – 200 m a. s. l., 6. – 13. vii. 2002, 1 ♂ 1 ♀, R. + H. Fouquè lgt., V. Hemala & A. Kocorek det. (NMPC); Phuket, White Buddha, 7 ° 51 ′ 39 ″ N 98 ° 19 ′ 30 ″ E, 50 m a. s. l., 21. vii. 2010, 1 ♀, L. Juříčková lgt., V. Hemala det. (NMPC). — LAOS: no exact data, 1 spec. [lost abdomen], no collector, J. A. Lis 1988 det. (DBOU). — VIETNAM: Annam, no date, 2 ♂♂, no collector, R. Kumar 1973 det. (DBOU); Mai lam, NE of Hanoi, 12. – 16. iv. 1966, 1 ♂, Gy. Topál lgt., A. Kocorek det. (HNHM); Tonkin, no date, 1 ♂, K. U. Tyduna lgt., J. A. Lis 1988 det. (DBOU). — MA- LAYSIA: Perak: Talping, x. 1977, 1 ♂, P. Pfanner lgt., V. Hemala & A. Kocorek det. (MHNG). Federal Territory of Kuala Lumpur: Kuala Lumpur, 16. viii. 1983, 1 ♀, S. Zabanski lgt., B. Lis det. (DBOU). — INDONESIA: Sumatra: Medan, no date, 1 ♂ 1 ♀, Mjöberg lgt., V. Hemala det. (NHRS); Tjinta Radja, no date, 1 ♀, Mjöberg lgt., V. Hemala det. (NHRS).	en	Hemala, Vladimír, Kocorek, Anna, Lis, Jerzy A. (2020): Megymenum tuberculatum, a new species of Megymenini from Java and a review of distribution of M. brevicorne (Hemiptera: Heteroptera: Dinidoridae). Zootaxa 4808 (3): 439-454, DOI: 10.11646/zootaxa.4808.3.2
F60287B8811F1C0E65BFFBF96A6DC9F3.taxon	description	Redescription of selected structures. Head slightly wider than long; clypeus very small, shorter than mandibular plates, together heart-shaped and strongly concave dorsally (Fig. 4). Anteocular spines present only sporadically (in most specimens absent), instead small rounded anteocular tubercles present (Fig. 4). Base of head wide (widest across eyes); small compound eyes positioned posterolaterally, nearly touching anterior part of pronotum. Antenniferous tubercle very small, short and not visible in dorsal view. Anteocular tubercles visible from dorsal view. Scape (I) very short, shorter than mandibular plate. Pedicel (II) is longest antennal segment, only slightly flattened, more circular in cross-section, about 2.2 times longer than segment I and longer than mandibular plate. Basiflagellum (III) more circular in cross-section, very slightly curved apically and 1.65 – 1.69 times longer than segment I. Distiflagellum (IV) apically narrowed, relatively short, only 1.46 – 1.59 times longer than scape. Bucculae round, very short, but longer than labrum. Labial segment I shorter than segment II, clearly reaching posterior margin of head, segment III and IV very short (shortest segments of labium; 2.0 – 2.26 times shorter than segment I) with nearly of same length or segment IV slightly shorter. Labial segment IV apically narrowed. Pronotum (Figs 4, 6) trapezoid in dorsal view, with well differentiated collar-like structure due to the distinct concavity of its anterolateral margin. Anterolateral margin apparently or strongly projecting forward to form distinct process, strongly concave anteriad of this process and only slightly concave posteriad of this process. Posterolateral margin slightly rounded, as long as 2 / 3 of anterolateral margin. Posterior margin along base of scutellum slightly convex. Pronotal surface tuberculate, with distinct medium sized rounded tubercle in middle of its anterior part, narrowly positioned near the anterior margin; this tubercle only 1.12 – 1.18 times wider than long. Disc with strong depressions at anteriolateral angles. Spermatheca (Fig. 14) with medium sized and spherical apical receptacle (spermathecal bulb), short and distinct intermediate part (pumping region) with distinct and large distal and proximal flanges, large and strongly dilated (sac-like) spermathecal duct with a distinct lateral fold bearing distinct ring sclerite. Spines of spermathecal duct distinct and large, covering only a small longitudinal area posteriorly of the ring sclerite. Measurements [median (minimum – maximum); in mm]. Females (n = 17; for pedicel: n = 16; for basiflagellum: n = 9; for distiflagellum IV: n = 7; for labial segments: n = 11; for profemora and protibiae: n = 14; for mesofemora and mesotibiae: n = 13; for metafemora and metatibae: n = 12; for tarsi: n = 11). Body length (from apex of clypeus to apex of abdomen) 13.72 (12.79 – 14.88); head: length (from apex of clypeus to anterior margin of pronotum) 1.86 (1.71 – 2.17), width (maximum width across eyes) 2.56 (2.40 – 2.87), height (height across compound eye) 1.55 (1.55 – 1.86), interocular width (between inner margins of compound eyes) 1.86 (1.71 – 2.02); lengths of antennal segments: scape (I) — 0.85 (0.70 – 0.93), pedicel (II) — 1.86 (1.71 – 2.09), basiflagellum (III) — 1.40 (1.40 – 1.75), distiflagellum (IV) — 1.24 (1.24 – 1.32); length of labial segments: I — 1.40 (1.24 – 1.55), II — 1.55 (1.32 – 2.16), III — 0.62 (0.47 – 0.78), IV — 0.62 (0.47 – 0.70); pronotum: length (medially in most exposed view) 3.57 (3.41 – 3.95), anterior width (between anterolateral angles) 4.88 (4.57 – 5.43), length of anterior tubercle (medially) 1.05 (0.93 – 1.16), width of anterior tubercle (transversally) 1.24 (1.09 – 1.50); posterior width (maximum width between humeral angles) 6.36 (6.05 – 7.29); scutellum: length (medially from base to apex) 4.00 (3.72 – 4.31), width (maximum width at base) 3.95 (3.64 – 4.31); corium: length 4.50 (4.19 – 4.96), width 2.09 (1.86 – 2.33); abdomen width (maximum width between apexes of lobes on laterotergites IV) 7.36 (6.67 – 7.91); length of femora: profemur 2.95 (2.71 – 3.10), mesofemur 3.26 (3.10 – 3.64), metafemur 4.19 (3.57 – 4.26); length of tibiae: protibia 2.56 (2.33 – 2.95), mesotibia 2.95 (2.64 – 3.57), metatibia 4.19 (3.80 – 4.65); length of tarsi: protarsus 1.55 (1.55 – 1.55), mesotarsus 1.71 (1.55 – 1.71), metatarsus 1.71 (1.55 – 1.71). Males (n = 12; for pedicel: n = 11; for basiflagellum: n = 7; for distiflagellum: n = 6; for labial segments, femora, protibia and metatibia: n = 8; for mesotibia and protarsus: n = 7; for mesotarsus: n = 6; for metatarsus: n = 3). Body length (from apex of clypeus to apex of abdomen) 12.12 (11.44 – 13.02); head: length (from apex of clypeus to anterior margin of pronotum) 1.71 (1.55 – 1.94), width (maximum width across eyes) 2.48 (2.17 – 2.64), height (height across compound eye) 1.55 (1.47 – 1.71), interocular width (between inner margins of compound eyes) 1.71 (1.71 – 1.86); lengths of antennal segments: scape (I) — 0.78 (0.62 – 0.81), pedicel (II) — 1.71 (1.47 – 1.78), basiflagellum (III) — 1.32 (1.24 – 1.52), distiflagellum (IV) — 1.24 (1.16 – 1.40); length of labial segments: I — 1.32 (1.24 – 1.32), II — 1.32 (1.24 – 1.39), III — 0.65 (0.45 – 0.78), IV — 0.55 (0.45 – 0.62); pronotum: length (medially in most exposed view) 3.30 (3.10 – 3.64), anterior width (between anterolateral angles) 4.54 (4.19 – 5.19), length of anterior tubercle (medially) 0.97 (0.85 – 1.01), width of anterior tubercle (transversally) 1.09 (1.01 – 1.47); posterior width (maximum width between humeral angles) 6.05 (5.74 – 6.82); scutellum: length (medially from base to apex) 3.57 (3.49 – 4.03), width (maximum width at base) 3.57 (3.26 – 4.03); corium: length 4.23 (3.95 – 4.65), width 1.86 (1.75 – 2.09); abdomen width (maximum width between apexes of lobes on laterotergites IV) 6.57 (6.12 – 7.70); length of femora: profemur 2.64 (2.48 – 2.95), mesofemur 3.17 (2.87 – 3.38), metafemur 3.76 (3.64 – 3.95); length of tibiae: protibia 2.48 (2.33 – 2.64), mesotibia 2.95 (2.64 – 3.10), metatibia 3.95 (3.72 – 4.34); length of tarsi: protarsus 1.55 (1.40 – 1.55), mesotarsus 1.55 (1.40 – 1.71), metatarsus 1.55 (1.47 – 1.71).	en	Hemala, Vladimír, Kocorek, Anna, Lis, Jerzy A. (2020): Megymenum tuberculatum, a new species of Megymenini from Java and a review of distribution of M. brevicorne (Hemiptera: Heteroptera: Dinidoridae). Zootaxa 4808 (3): 439-454, DOI: 10.11646/zootaxa.4808.3.2
F60287B8811F1C0E65BFFBF96A6DC9F3.taxon	biology_ecology	Biology. Megymenum brevicorne is trophically bound especially on plants from the families Cucurbitaceae and Passifloraceae sucking on their stems and fruits (Miller 1929, Hoffmann 1932 a, Schaefer & Ahmad 1987). The species was reported most often as a minor pest of Passiflora quadrangularis, Cucurbita maxima and Trichosan- thes cucumerina, on which the larval development was observed (Miller 1929, Hoffmann 1932 a). It was recorded also on other Cucurbitaceae (e. g. Cucumis sativus, Lagenaria siceraria), on some Fabaceae (Vigna unguiculata sesquipedalis, Phaseolus lunatus) and even on some other plant families — Celosia cristata (Amaranthaceae), Im- patiens balsamina (Balsaminaceae), Manihot esculenta (Euphorbiaceae), Quercus spp. (Fagaceae), Cinnamomum camphora (Lauraceae), Eucalyptus spp. (Myrtaceae), on which the development is supposed, but not confirmed yet (Hoffmann 1931, 1932 a; Corbett 1933; Cheo 1935; Chen & Gu 2000; Hua 2000). Sen (1965) stated the species as a “ major pest ” of Millettia pinnata (Fabaceae) without giving any further details and therefore this record also needs a confirmation. Feeding of M. brevicorne is probable also on Benincasa hispida (Cucurbitaceae) on which the species was recorded in Qiongshan (Hainan, China) in 1961 (sitting record) (this paper). Megymenum brevicorne was listed also in the list of pests of various leguminous trees (Fabaceae) in India without indicating the taxon on which it was observed (Vir & Verma 1996). Larvae of M. brevicorne were successfully reared on Mormodica charantia (Cucurbitaceae), Basella alba (Basellaceae) and Citrus aurantiifolia (Rutaceae) (Durai 1993). Larval development was not fully described in nature, but only in laboratory conditions (Miller 1929, Durai 1993). Number of generations per year is not known, but length of life from the egg hatching to adult death was 74 – 103 days in laboratory conditions (Durai 1993).	en	Hemala, Vladimír, Kocorek, Anna, Lis, Jerzy A. (2020): Megymenum tuberculatum, a new species of Megymenini from Java and a review of distribution of M. brevicorne (Hemiptera: Heteroptera: Dinidoridae). Zootaxa 4808 (3): 439-454, DOI: 10.11646/zootaxa.4808.3.2
F60287B8811F1C0E65BFFBF96A6DC9F3.taxon	distribution	Distribution. CHINA: unspecified (Fabricius 1787, 1794, 1803; Gmelin 1790, as Sina; Burmeister 1835; Westwood 1837; Dallas 1851; Dohrn 1859; Walker 1868; Stål 1870; Distant 1902; Kirkaldy 1909; Miller 1929; Hoffmann 1935; Durai 1987; Hua 1989; Lis 1990, 1992; Zhang 1995; Waterhouse 1998; Chen & Gu 2000; Lin et al. 2000; Li 2011; this paper), Beijing (Hua 2000), Chongqing (Lin et al. 1992), Fujian (Wu 1932; Hoffmann 1935 [both previous as Fukien]; Tang 1935; Yang 1940 as Fokien; Lin et al. 1999; Hua 2000), Guangdong (Burmeister 1834 a; Kirkaldy 1909; Oshanin 1911; Hoffmann 1931, 1935, 1948; Wu 1933; Cheo 1935; Tang 1935; Yang 1940; Lis 1990; Lis & Kocorek 1996; Hua 2000; this paper), Guangxi (Zhang et al. 1994; Hua 2000; Xie et al. 2000, 2011; Yang et al. 2005; Liang et al. 2014), Guizhou (Yang 1940 as Kweichow; Chen 1987; Lin et al. 1992; Hua 2000; Liu 2006), Hainan (Hoffmann 1932 c, 1935; Tang 1935; Yang 1940; Hua 2000; this paper), Hebei (Hoffmann 1948 as Hopei; Wang et al. 2000), Hong Kong (Mayr 1868; Hoffmann 1935; Durai 1987), Hubei (Lei & Zhou 1998; Hua 2000), Hunan (Lin et al. 1992), Jiangxi (Chang [= Zhang] 1974, as Kiangsi; Zhang & Lin 1986, 1992; Zhang 1994; Hua 2000; Li et al. 2008), Macao (Kirkaldy 1910; Hoffmann 1935; Easton & Pun 1997), Shaanxi (He et al. 2007), Sichuan (Hua 2000), Tibet [= Xizang] (Zhang 1986; Zhang & Lin 1987, 1988; Hu 1990; Zheng & Jin 1990; Wany et al. 1992; Hua 2000), Yunnan (Hoffmann 1948; Jiang 1985; Zhang & Lin 1990; Hua 2000; Chen et al. 2009; He et al. 2011; this paper; record in Zhang 2011 is misidentification of M. spinosum), Zhejiang (Yang 1940 as Chekiang; Chen 1990; Hua 2000; Wang & Tong 2014). TAIWAN (Lis 1992 as Formosa; Zheng & Lin 2013). INDIA: unspecified (Lethierry & Severin 1893; Kirkaldy 1909; Hoffmann 1935; Tang 1935; Durai 1987; Lis 1990), “ east part ” (Dohrn 1859 as M. inerme), “ North India ” (Atkinson 1882), Andhra Pradesh (Biswas & Bal 2007), Assam (Atkinson 1882, 1889; Distant 1902; Kirkaldy 1909; Schouteden 1913; Hoffmann 1935; Yang 1940; Durai 1987), Jharkhand (Sen 1965), Meghalaya (Distant 1902; Miller 1929), Mizoram (Chakraborty & Bal 2007), Nicobar Islands (Distant 1902; Kirkaldy 1909; Miller 1929; Hoffmann 1935; Tang 1935; Yang 1940; Durai 1987; Lis 1990), Rajasthan (Lethierry 1891 from Tetara), Sikkim (Yang 1940), West Bengal (Distant 1902; Miller 1929; Banerjee 1958; Chakraborty et al. 1994;? Herrich-Schaeffer 1840;? Hoffmann 1935,? Tang 1935 [previous three ambiguously as Bengal]).? BANGLADESH (? Herrich-Schaeffer 1840;? Hoffmann 1935;? Tang 1935 [all ambiguously as Bengal]; Ahmad & Khan 1979 without any exact locality). NEPAL (new record; the record in Jyoti & Rajbhandari 2015 is misidentification of an unidentified species of the genus Coridius Illiger, 1807). BHOUTAN (Yang 1940). SRI LANKA (Longstaff 1912; Hoffmann 1935 as Ceylon; Durai 1987; Lis 1990). MYANMAR: unspecified (Kirkaldy 1909; Hoffmann 1935; Tang 1935; Durai 1987; Lis 1992 [all previous as Burma]; Yang 1940 as Birmaine), Yangon Region (Distant 1901, 1902; Miller 1929; Tang 1935 [all as Rangoon]). THAILAND (Hoffmann 1935 as Siam; Durai 1987; Lis 1990; Lis & Kocorek 1996; Lis et al. 2011; Rakowiecka & Lis 2012; this paper). LAOS (Lis 1990; this paper). CAMBODIA (Yang 1940, as Cambodge). VIETNAM (Hoffmann 1948; Sienkiewicz 1964 [both as M. inerme]; Lis 1992; this paper). MALAYSIA: unspecified (? Corbett 1933 as Malaya [= ‘ British Malaya’, the term describing a set of states on the Malay Peninsula and the island of Singapore that were under the British control in 1874 – 1957 (e. g. Andaya & Andaya 1982)]; Durai 1986, 1987; listed in Lis 1990), Perak (this paper), Federal Territory of Kuala Lumpur (this paper).? SINGAPORE (? Corbett 1933 as Malaya [= ‘ British Malaya’, the term describing a set of states on the Malay Peninsula and the island of Singapore that were under the British control in 1874 – 1957 (e. g. Andaya & Andaya 1982)]). INDONESIA:? Java (Yang 1940: country is only listed; cited only in Hoffmann 1948), Sumatra (Breddin 1900; Kirkaldy 1909; Schouteden 1933; Hoffmann 1935; Yang 1940; Durai 1987; Lis 1990, 1992; this paper). Erroneous records. Japan: Hua’s (2000) reference to the distribution of M. brevicorne in Japan seems to be erroneous, because there is reliable documentation of only one species of the genus Megymenum Guérin-Méneville, 1831 — M. gracilicorne Dallas, 1851 (Hayashi et al. 2016).	en	Hemala, Vladimír, Kocorek, Anna, Lis, Jerzy A. (2020): Megymenum tuberculatum, a new species of Megymenini from Java and a review of distribution of M. brevicorne (Hemiptera: Heteroptera: Dinidoridae). Zootaxa 4808 (3): 439-454, DOI: 10.11646/zootaxa.4808.3.2
