taxonID	type	description	language	source
F542AA7FFFDCFFE000C785233BC9FA16.taxon	materials_examined	Type locality. Kingi Mariout, Cleopatra (Egypt).	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFDCFFE000C785233BC9FA16.taxon	diagnosis	Diagnosis. Length. (female) body 12.2 mm, wing 10.5 mm. Female: Whitish ashy-grey species; frons with sides not parallel, with a trace of upper callus; frontal index 2.64; eyes bare; abdomen without patterns. Local distribution: Coastal strip.	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFDCFFE000C785233BC9FA16.taxon	description	Geographical distribution. Egypt. Specimens examined. Cleopatra (Ramleh), 13. VII. 1924 (1 female “ cotype ”) (PPDD). FIGURES. 1 – 6. Female abdomens. 1. Atylotus aegyptiacus (Kröber, 1926). 2. Atylotus shagrensis sp. nov. 3. Atylotus Agricola (Wiedemann, 1828). 4. Atylotus farinosus (Szilády, 1915). 5. Atylotus agrestis (Wiedemann, 1828). 6. Atylotus pulchellus (Loew, 1858).	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFDFFFE100C787FF3EEDFBA8.taxon	materials_examined	Type locality. Egypt.	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFDFFFE100C787FF3EEDFBA8.taxon	diagnosis	Diagnosis. Length. (male) body 10.8 – 14.5 mm, wing 8.45 – 10.3 mm; (female) body 11.3 – 13.4 mm, wing 9.7 – 10.2 mm. Frons in females greyish tomented, parallel sided, frontal index in female 2.66 – 2.69; eyes bare; post-occipital rim narrow and short in male; wing veins pale yellow, R 4 with short appendix; abdomen whitish grey with 4 greyish brown longitudinal spots, the two median longitudinal spots forming a broad median longitudinal dark brown stripe. Local distribution: Coastal strip, Upper Nile, Lower Nile, Western Desert.	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFDFFFE100C787FF3EEDFBA8.taxon	description	Geographical distribution. China, Egypt, India, Saudi Arabia, Sri Lanka, widespread through Palaearctic and Afrotropical regions. Specimens examined. Sentris, 19. II. 1951 (1 ♀); Kerdasa, 16. III. 1952 (1 ♀); Kom Osheim, 9. V. 1954 (1 ♀); Mansourah, 15. X. 1956 (1 ♀); Khor Galal, 19. IX. 1997 (1 ♀); Kafer El-Sheikh, 10. VI. 1993 (1 ♀); El-Fayoum, 25. X. 1994; El-Fayoum, III. 2008 (1 ♂); Alexandria, Abeis, 1. VIII. 2008 (1 ♂); El-Khanka, 15. VIII. 2 0 0 9 (1 ♀); Bahariya Oasis, VII. 2012 (2 ♂); Tanta, 28. VIII. – 1. IX. 2012. (2 ♀, 3 ♂) (ASUC); El-Marg, 25. V. 1921 (1 ♂); Fayed, 24. IX. 1925 (1 ♂); Gabel-Asfer, 27. X. 1925 (1 ♀); Mirsa Matrouh, III. 1930 (1 ♀); Helwan, 28. III. 1939 (2 ♂, 1 ♀); 30. V. 1945 (1 ♀); El-Fayoum, 5. VIII. 1946 (1 ♀); El-Fayoum, 15. VIII. 1946 (4 ♂, 6 ♀); El-Fayoum, 1. III. 1947 (1 ♀); El-Fayoum, 12. VI. 1948 (3 ♀) (CUC); Gabal-Asfer, 15. I. 1925 (1 ♀); Faraskure, 12. VI. 1925 (1 ♂); Wadi Abu El-Nesour, 23. X. 1925 (1 ♀); Kharga Oasis, 12. VI. 1926 (3 ♀); Kharga Oasis, 20. IX. 1926 (2 ♀) (ESEC); Meadi, 11. V. 1913 (1 ♀); Upper Egypt train, 24. IX. 1914 (1 ♀); Meadi, 20. VII. 1915 (1 ♀); Cairo, 4. VI. 1917 (1 ♀); Alexandria, Nouzha, 15. VII. 1920 (1 male); Ezbet El-Nakhl, 11. V. 1921 (1 ♀); El-Marg, 25. V. 1921 (2 ♂, 1 ♀); Nouzha, 26. VI. 1921 (2 ♀); Nouzha, 5. VII. 1921 (4 ♂ (1 “ cotype ” of Atylotus agrestis), 5 ♀); Nouzha, 18. VII. 1921 (1 ♂, 1 ♀); Cleopatra, 14. VIII. 1921 (1 ♀); Mobar Bez, 3. IX. 1921 (1 ♂); Ezbet El-Nahkl, 20. IV. 1924 (1 ♂, 3 ♀); Ezbet El-Nahkl, 25. IV. 1924 (2 ♂); Faraskure, 7. VII. 1926 (1 ♂); Baharyia Oasis, 1. VI. 1927 (1 ♀); Meadi, 11. VI. 1931 (1 ♂), Meadi, 5. IX. 1931 (1 ♂); El-Fayoum, Kom Osheim, 2. V. 1965 (2 ♀) (PPDD).	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFDFFFE100C787FF3EEDFBA8.taxon	discussion	Remarks. Müller et al. (2012) mentioned this species as a record for the Sinai Peninsula.	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFDEFFE100C786643915F90D.taxon	materials_examined	Type locality: Red Sea (Egypt).	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFDEFFE100C786643915F90D.taxon	diagnosis	Diagnosis. Length. (male) body 13.7 – 15 mm, wing 11.2 – 12 mm; (female) body 14.1 – 16 mm, wing 10.5 – 11.9 mm. Frons in females yellowish dusted, with parallel sided; frontal index in female 2.72 – 2.75; subcallus yellowish dusted, lower and median calli distinct; eyes hairy; abdomen reddish yellow with a broad blackish median spots with the longest on 2 nd tergites and the broadest on 3 th and 4 th .. Local distribution: Coastal strip, Western Desert.	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFDEFFE100C786643915F90D.taxon	description	Geographical distribution. Bulgaria, Egypt, Libya, Morocco, Saudi Arabia, Turkey. Specimens examined. Mirsa Matrouh, 7. VIII. 1931 (2 ♀); Hurghada, 4. V. 1939 (1 ♀) (CUC); Abou-Kir, 3. VI. 1924 (1 ♂, 1 ♀, det. by Kröber 1924) (ESEC); Ismailia, 25. VII. 1915 (2 ♀); Egypt, 7. VII. 1916 (1 ♀); Mullah, 22. VII. 1916 (1 ♀); Baharyia Oasis, 1. VI. 1927 (1 ♀) (PPDD).	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFDEFFE600C785013EBBFE6B.taxon	diagnosis	Diagnosis. Length. (male) body 13.5 mm, wing 9.9 mm; (female) body 11.5 mm, wing 8.5 mm. Frons in females with sides not parallel; frontal index in female 2.26; eyes bare; wing with yellow veins; abdominal tergites with 4 greyish brown longitudinal spots. Local distribution: Coastal strip.	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFDEFFE600C785013EBBFE6B.taxon	description	Geographical distribution. Algeria, Egypt, Iraq, Jordan, Palestine, Saudi Arabia, Syria. Specimens examined. Ramleh, 12. V. 1921 (1 ♂, Type of Ochrops farinosus); Dekheila Marioute, IX. 1924 (1 ♀); Dekhela, 24. V. 1925 (1 ♂, Type of Ochrops farinosus) (CUC); Ezbet El-Nakhl, 24. VI. 1924 (1 ♀); Dekheila 15. IX. 1929 (1 ♀) (EESC); Dekheila Mariout, 24. V. 1926 (1 ♀) (PPDD).	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFDEFFE600C785013EBBFE6B.taxon	discussion	Remarks. Müller et al. (2012) mentioned this species as a new record for the Sinai Peninsula.	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFD9FFE600C780213918FBE9.taxon	diagnosis	Diagnosis. Blackish species; lower and middle calli jointed; wings with veins black, whitish only at basal margin; abdomen ferruginous with 3 stripes. Local distribution: Sinai.	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFD9FFE600C780213918FBE9.taxon	description	Geographical distribution. Egypt.	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFD9FFE600C780213918FBE9.taxon	discussion	Remarks. This species is not represented in the Egyptian Reference Collections. Chvála (1988), in his catalogue of Palaearctic Tabanidae, mentioned this as a doubtful species. Müller et al. (2012) also mentioned this species as doubtful and erroneously stated that Efflatoun (1930) placed Tabanus pallescens under “ Tabanidae of doubtful origin ”. While this species was indeed not mentioned at all in the earlier Egyptian works by Kröber (1925 a, b), Efflatoun (1930) and Steyskal & El-Bialy (1967). After examining the paper of original description by Walker (1871) who mentioned the type locality of species as Tor (Sinai) we added this species to the Egyptian fauna depending on this type locality.	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFD9FFE700C786A639C5FA49.taxon	diagnosis	Diagnosis. Length. (male) body 10.5 – 12.1 mm, wing 8.6 – 9 mm; (female) body 10.8 – 12.3 mm, wing 8.9 – 9.5 mm. Frons parallel sided; frontal index in female 3.18 – 3.23; eyes bare; post-occipital rim elongated and puffed out in male; apical part of the wing darker than basal part, R 4 with long appendix; abdomen whitish grey with 4 greyish brown longitudinal spots, the two median spots often forming a broad median longitudinal brown stripe. Local distribution: Coastal strip, Western Desert.	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFD9FFE700C786A639C5FA49.taxon	description	Geographical distribution. Algeria, China, Egypt, Iran, Iraq, Jordan, Palestine, Syria, Turkey. Specimens examined. Bahariya Oasis 19 – 21. V. 2009 (1 ♀); Bahariya Oasis, VII. 2012 (2 m #) (ASUC); Ezbet El-Nakhl, 11. V. 1921 (1 ♂); Siwa Oasis, 24. VIII. 1930 (1 ♀); Kharga Senoh, 15. II. 1937 (1 ♀); El-Fayoum, 30. IV. 1943 (1 ♀); El-Fayoum, Girza Route, 14. IV. 1947 (1 ♀); El-Fayoum, Girza Route, 9. IV. 1950 (4 ♂) (CUC); Kafr-Hakim, 18. VII. 1924 (1 ♀ Kröber det. 1924); Faraskur, 12. VI. 1925 (1 ♀ Kröber det. 1925) (EESC); Kharga Oasis, 9. VII. 1918 (1 ♀); Kharga Oasis, 13. III. 1924 (1 ♀); Ezbet El-Nakhl, 25. IV. 1924 (1 ♂); Bahariya Oasis, 1. VI. 1927 (1 ♀); Kharga Oasis, 13. III. 1934 (1 ♀); Mammura, August 1958 (1 ♀) (PPDD). FIGURES. 7 – 13. Female Frons. 7 – 11. 7. Atylotus shagrensis sp. nov. 8. Atylotus Agricola (Wiedemann, 1828). 9. Atylotus farinosus (Szilády, 1915). 10. Atylotus agrestis (Wiedemann, 1828). 11. Atylotus pulchellus (Loew, 1858). Wings. 12 – 13. 12. Atylotus agrestis (Wiedemann, 1828). 13. Atylotus pulchellus (Loew, 1858).	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFD8FFE400C784413F2BFC32.taxon	materials_examined	Type material. Holotype female, Egypt: Wadi Shagra, near Mersa Alam, 24. IV. 1933 (PPDD).	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFD8FFE400C784413F2BFC32.taxon	diagnosis	Diagnosis. Eyes with short yellow pubescence, frontal index 2.6, calli completely absent, R 4 with long appendix, abdominal tergites 1 – 3 dark reddish brown with black spots, tergites 4 – 7 blackish. Length: (female) body 12.5 mm, wing 11.6 mm.	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFD8FFE400C784413F2BFC32.taxon	description	Description. Head: dark reddish-brown. Eyes with short yellow pubescence, without bands; frons parallel-sided, blackish with golden-yellow hairs, frontal index 2.6, calli completely absent, with whitish ocellar tubercle; face and genae polished brown tomentose; palpi yellowish with mixed white and a few black hairs; antennae pale reddish-yellow becoming darker towards apex, basal two segments with golden-yellow hairs, basal flagellomere broad. Thorax: blackish-brown with long and dense whitish and golden-yellow hairs; mesonotum including scutellum with dense long yellowish hairs, notopleural lobes reddish-brown with yellow hairs; legs reddish-yellow, fore tibia only blackish; wings hyaline with brown veins and a long appendix to R 4; halteres pale brown with whitish knob. Abdomen: with the first three tergites dark reddish-brown, the 1 st with black spots around the base of the scutellum, 2 nd and 3 rd with large black spots on the middle of each segment and a small black triangular spot on the sides of the posterior margin of each segment, tergites 4 – 7 blackish, all tergites covered with long golden-yellow and whitish hairs, venter reddish-brown and covered with long golden-yellow and white hairs. Male unknown. Local distribution: W. Shagra (Eastern Desert). Geographical distribution. Egypt. Specimens examined. Wadi Shagra, 24. IV. 1933 (1 ♀, holotype) (PPDD).	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFD8FFE400C784413F2BFC32.taxon	etymology	Etymology: This new species is named after its type locality. Affinities: Atylotus shagrensis sp. nov. is closely related to the Egyptian species A. agricola (Wiedemann, 1828) and the European species A. loewianus (Villeneuve), 1920 because of the parallel-sided frons and the abdominal pattern. The new species can be easily separated from A. agricola by the absence of frontal calli, blackish colour of frons, polished brown colour of subcallus and reddish-brown colour of the abdomen. Atylotus shagrensis sp. nov. is distinguished from A. loewianus by the hairy eyes, absence of frontal calli, colour of subcallus and frons, and reddish-brown colour of the abdomen.	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
F542AA7FFFD8FFE400C784413F2BFC32.taxon	discussion	Remarks. Four female specimens of Atylotus loewianus Villeneuve, which were collected in Croatia (8. VIII. 1996), were sent to us for examination by Prof. Dr. Stjepan Krčmar from his personal collection (SKC).	en	Abu El-Hassan, Gawhara M. M., Badrawy, Haitham B. M., Fadl, Hassan H., Mohammad, Salwa K. (2019): Review of the horse flies of the genus Atylotus Osten Sacken, 1876 (Diptera: Tabanidae) from Egypt, with description of a new species. Zootaxa 4551 (1): 79-86, DOI: 10.11646/zootaxa.4551.1.5
