identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
F556879FE750FF900785FA3E28A74B80.text	F556879FE750FF900785FA3E28A74B80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helicopsyche (Feropsyche) bendego Dumas & Nessimian 2019	<div><p>Helicopsyche (Feropsyche) bendego, new species</p><p>(Figures 1 A–1G)</p><p>urn:lsid:zoobank.org:act: 67B653D5-F7CB-4489-B555-4C6E0676444F</p><p>Diagnosis. Helicopsyche bendego sp. nov. resembles H. (F.) cipoensis Johanson &amp; Malm 2006, and H. (F.) muelleri Banks 1920, all from Brazil. All of these species have large, slightly club-like inferior appendages, with prominent basomesal lobes. However, in H. (F.) muelleri the posterior margin of each inferior appendage has a pointed apical projection, absent in the other two species. Moreover, tergum X has linear rows of setae near the mesodorsal borders in all of these species (segment X is destroyed in the lectotype of H. (F.) muelleri according to Johanson (1998), but these rows of setae are present in the illustrations of H. (F.) angelae (Marlier 1964), a junior synonym of the former species). Also, segment X, in dorsal view, is enlarged apically and has a shallow, rectangular mesal cleft in the new species, while in the other species it has almost the same width from its base and is without an apical cleft.</p><p>Description. Adult. Overall color (in alcohol) nearly uniformly golden-brown; antennae, palps, and legs yellowish brown (one maxillary palp missing in the holotype). Cephalic setal warts large, oval; postantennal warts tube-shaped. Maxillary palps 2-segmented, proximal and distal segments about equally long, both covered with many dark-brown setae. Tibial spur formula 2,4,4; each foreleg anterior apical tibial spur about 3x longer than posterior apical spur. Male forewings each 4.4–5.0 mm long (n = 10; holotype male = 4.8 mm), covered with fine, short, brown setae. Forewings with forks I, IV, and V present, discoidal and thyridial cells elongate, discoidal cell slightly shorter; hind wings each with forks I and V present, fork I short, anal veins reduced, hamuli present in continuous row along proximal anterior margin, distal anterior margin abruptly concave and without hamuli. Abdomen with sterna III–IV bearing well-developed and darkly pigmented reticulation; abdominal sternum VI ventral process long, about as long as segment VI, tubular along its length, oriented posteroventrad, larger at base, nearly straight in lateral view, covered with microtrichiae, apex rounded, with lamellae apicoventrally (Figs. 1F, 1G).</p><p>Male genitalia. Segment IX short ventrally; in lateral view with anteromesal margin well-developed, anterodorsal margin almost straight, and anteroventral margin concave; lateral apodeme well-developed, located midlaterally on segment (Fig. 1A); in dorsal view with anterior margin strongly concave (Fig. 1B); in ventral view with anterior margin convex and posterior margin slightly concave with shallow mesal invagination (Fig. 1C). Segment X, in lateral view, triangular, strongly tapering to apex from distal 2/3 (Fig. 1A); in dorsal view broad, slightly surpassing apices of inferior appendages, narrower in basal half, apically rounded with rectangular mesal cleft, mesodorsal borders inverted Y-shaped, stem of Y bordered laterally by pair of nearly linear setal rows each with 6–10 long, stout setae (Fig. 1B). Superior appendages originating dorsolaterally, setose, rounded in lateral view, clavate in dorsal view (Figs. 1A, 1B). Basal plate of inferior appendages triangular in lateral view, not surpassing anteroventral margin of segment IX (Fig. 1A), large and trapezoidal in ventral view (Fig. 1C); primary branches of inferior appendages large, covered by long, stout setae; in lateral view, each nearly clavate, curved dorsad and widening slightly from base to apex, with distal portion about 1.5x broader than its proximal portion, dorsal margin concave, ventral margin convex near base and slightly concave subapically, apicodorsal margin rounded, slightly undulate, with several long, stout setae along margin (Fig. 1A); in dorsal view, 2/3 as wide as base of tergum X, each with inner margin rounded, strongly produced mesad (Fig. 1B); in ventral view partially twisted, ventral surface narrowing to apex, apex rounded, internal portion with two basomesal lobes well-developed, ovate, bearing several spine-like setae apically (10–12 setae, only four visible through appendage in lateral view) (Figs. 1A, 1C), 1–2 spine-like setae distal of these lobes, each on distinct protuberance (Fig. 1C). Phallus tubular, moderately down-curved along its length; phallobase narrow, slightly inflated basodorsally, narrower in middle, apicoventral margin strongly sclerotized, rounded apically; endotheca membranous and partially extending dorsally as lobe over endophallus; phallotremal sclerite not apparent (Figs. 1D, 1E).</p><p>Holotype male: BRAZIL: Rio de Janeiro: Santa Maria Madalena, Parque Estadual do Desengano, Morumbeca dos Marreiros, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.915253&amp;materialsCitation.latitude=-21.876362" title="Search Plazi for locations around (long -41.915253/lat -21.876362)">Ribeirão Macapá</a> (ponte), 21°52’34.9” S, 41°54’54.9” W, 13.iv.2016, 1045 m, LL Dumas, JL Nessimian, CS Portela &amp; JF Barbosa leg. (DZRJ).</p><p>Paratypes: BRAZIL: Rio de Janeiro: same data as holotype, 1 male, 6 females (DZRJ); same data as holotype, 1 male, 1 female (MZSP); same locality, 21.x.2016, LL Dumas, JL Nessimian &amp; JF Barbosa leg., 3 males, 5 females (DZRJ); same locality, 21-23.x.2016, LL Dumas, JL Nessimian &amp; JF Barbosa leg., 2 males (DZRJ); Santa Maria Madalena, Parque Estadual do Desengano, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.918167&amp;materialsCitation.latitude=-21.876528" title="Search Plazi for locations around (long -41.918167/lat -21.876528)">Morumbeca</a> dos <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.918167&amp;materialsCitation.latitude=-21.876528" title="Search Plazi for locations around (long -41.918167/lat -21.876528)">Marreiros</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.918167&amp;materialsCitation.latitude=-21.876528" title="Search Plazi for locations around (long -41.918167/lat -21.876528)">Ribeirão Macapá</a>, 21°52’35.5” S, 41°55’05.4” W, 13.iv.2016, 1039 m, LL Dumas, JL Nessimian, CS Portela &amp; JF Barbosa leg., 1 male, 1 female (DZRJ) ; Santa Maria Madalena, Parque Estadual do Desengano, Morumbeca dos Marreiros, afluente do <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.915363&amp;materialsCitation.latitude=-21.8775" title="Search Plazi for locations around (long -41.915363/lat -21.8775)">Ribeirão Macapá</a>, 21°52’39.0” S, 41°54’55.3” W, 13.iv.2016, 1110 m, LL Dumas, JL Nessimian, CS Portela &amp; JF Barbosa leg., 1 male, 1 female (DZRJ) ; São Fidélis, Parque Estadual do Desengano, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.912167&amp;materialsCitation.latitude=-21.876694" title="Search Plazi for locations around (long -41.912167/lat -21.876694)">Morumbeca</a> dos <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.912167&amp;materialsCitation.latitude=-21.876694" title="Search Plazi for locations around (long -41.912167/lat -21.876694)">Marreiros</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.912167&amp;materialsCitation.latitude=-21.876694" title="Search Plazi for locations around (long -41.912167/lat -21.876694)">Ribeirão Macapá</a>, 21°52’36.1” S, 41°54’43.8” W, 13.iv.2016, 1111 m, LL Dumas, JL Nessimian, CS Portela &amp; JF Barbosa leg., 2 males, 1 female (DZRJ) ; Campos dos Goytacazes, Mocotó, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.75675&amp;materialsCitation.latitude=-21.814222" title="Search Plazi for locations around (long -41.75675/lat -21.814222)">Parque Estadual do Desengano</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.75675&amp;materialsCitation.latitude=-21.814222" title="Search Plazi for locations around (long -41.75675/lat -21.814222)">Rio Mocotó</a> (próximo a <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.75675&amp;materialsCitation.latitude=-21.814222" title="Search Plazi for locations around (long -41.75675/lat -21.814222)">Cachoeira do Maracanã</a>), 21°48’51.2” S, 41°45’24.3” W, 17.iv. 2016, 243 m, LL Dumas, JL Nessimian, CS Portela &amp; JF Barbosa leg., 1 male (DZRJ) ; Campos dos Goytacazes, Parque Estadual do Desengano, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.829834&amp;materialsCitation.latitude=-21.832973" title="Search Plazi for locations around (long -41.829834/lat -21.832973)">Itacolomi</a>, afluente do <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.829834&amp;materialsCitation.latitude=-21.832973" title="Search Plazi for locations around (long -41.829834/lat -21.832973)">Córrego da Malhada Branca</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.829834&amp;materialsCitation.latitude=-21.832973" title="Search Plazi for locations around (long -41.829834/lat -21.832973)">Poço Parado</a>, 21°49’58.7” S, 41°49’47.4” W, 24.x. 2016, 919 m, LL Dumas, JL Nessimian &amp; JF Barbosa leg., 1 male, 3 females (DZRJ) ; Itatiaia, Maromba, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.615414&amp;materialsCitation.latitude=-22.330473" title="Search Plazi for locations around (long -44.615414/lat -22.330473)">Rio Preto</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.615414&amp;materialsCitation.latitude=-22.330473" title="Search Plazi for locations around (long -44.615414/lat -22.330473)">Cachoeira do Escorrega</a>, 22°19’49.7” S, 44°36’55.5” W, 26.i.2012, 1381 m, BHL Sampaio &amp; ALH Oliveira leg., 2 males (DZRJ) .</p><p>Distribution. Brazil (Rio de Janeiro).</p><p>Etymology. The specific name “bendego” is in reference to the Bendegó Meteorite, the biggest iron meteorite ever found in Brazilian soil, weighting 5,360 kilograms. It was found in Bahia in 1784 and brought to Museu Nacional in 1888, where it had been on display in museum entrance ever since.</p></div>	https://treatment.plazi.org/id/F556879FE750FF900785FA3E28A74B80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Dumas, Leandro Lourenço;Nessimian, Jorge Luiz	Dumas, Leandro Lourenço, Nessimian, Jorge Luiz (2019): New species of Helicopsyche von Siebold 1856 (Trichoptera: Helicopsychidae) from Brazil, including the redescription of Helicopsyche (Feropsyche) planorboides Machado 1957. Zootaxa 4619 (2): 231-250, DOI: 10.11646/zootaxa.4619.2.2
F556879FE752FF970785F8DD2F064B74.text	F556879FE752FF970785F8DD2F064B74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helicopsyche (Feropsyche) daome Dumas & Nessimian 2019	<div><p>Helicopsyche (Feropsyche) daome, new species</p><p>(Figures 2 A–2F)</p><p>urn:lsid:zoobank.org:act: D377B6D4-2EBD-48C4-86B6-3832141B8EAB</p><p>Diagnosis. Helicopsyche (F.) daome sp. nov. is most similar to H. (F.) cipoensis Johanson &amp; Malm 2006, H. (F.) muelleri, and H. (F.) bendego sp. nov. by the nearly club-like inferior appendages in lateral view. However, it can be easily diagnosed by the presence of a well-developed and strongly rounded protuberance bearing several stout setae subapicomesally on each inferior appendage (absent in the other species). Only H. (F.) planorboides has a protuberance apicoventrally as the new species, but it is less developed and not rounded. Moreover, the overall shape of the inferior appendages and especially of tergum X of these two species are otherwise totally dissimilar. An additional diagnostic character for H. (F.) daome sp. nov. is the subtruncate apex of each inferior appendage in ventral view.</p><p>Description. Adult. Similar to H. (F.) bendego sp. nov. except as follows: Dorsum of head, meso- and metanota dark-brown, warts pale yellow. Each foreleg anterior apical tibial spur about 4x longer than posterior apical spur. Male forewings each 4.4–4.8 mm long (n = 10; holotype male = 4.4 mm). Abdominal sternum VI ventral process long and narrow, longer than segment VI, tubular along its length, oriented posteroventrad, nearly straight in lateral view, with microtrichiae along length, apex tapering, with lamellae apicoventrally (Figs. 2F, 2G).</p><p>Male genitalia. Segment IX short ventrally; in lateral view with anteromesal margin well-developed, anterodorsal margin almost straight, and anteroventral margin concave; lateral apodeme well-developed, located midlaterally on segment (Fig. 2A); in dorsal view horseshoe-like, with anterior margin strongly concave (Fig. 2B); in ventral view with anterior margin almost straight, posterior margin slightly concave (Fig. 2C). Segment X, in lateral view, subtriangular, strongly tapering to apex from distal 1/3 of dorsal margin and distal 2/3 of ventral margin (Fig. 2A); in dorsal view moderately broad, slightly narrower near base, strongly surpassing apices of inferior appendages, mesodorsal borders inverted Y-shaped, stem of Y bordered laterally by pair of nearly linear setal rows each with 7–8 stout setae and two small, stout, apical setae, apex with shallow mesal notch between pair of rounded ends (Fig. 2B). Superior appendages clavate in lateral view, finger-like in dorsal view (Figs. 2A, 2B). Basal plate of inferior appendages ellipsoid in lateral view (Fig. 2A); primary branches of inferior appendages large, covered by large, long setae; in lateral view nearly club-like, slightly widening to apex, with distal portion about 1.5x broader than its proximal portion, dorsal margin concave, ventral margin convex near base and undulate along middle 1/3, with three long, stout setae basally, posterodorsal margin rounded, with several stout setae along apical margins (Fig. 2A); in dorsal view ear-like, with well-developed, nearly rounded protuberance apicoventrally, bearing about 10–12 stout setae (also visible in ventral view) (Fig. 2B); in ventral view, enlarged at base, outer margin convex, with several long, stout setae, inner margin concave, apex subtruncate, with well-developed rounded, setose protuberance (Fig. 2C); basomesal lobes of inferior appendages well-developed, ovate, apically bearing several spine-like setae (12–16 setae) and with a row of 6–8 fine setae at inner margin (Figs. 2A, 2C). Phallus tubular, moderately down-curved along its length; phallobase narrow, inflated basoventrally, with apicoventral margin elongated, strongly sclerotized, rounded apically; endotheca membranous lobe partially extending dorsally over endophallus; phallotremal sclerite small, slightly C-shaped (Figs. 2D, 2E).</p><p>Holotype male: BRAZIL: Rio de Janeiro: Petrópolis, Araras, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.259193&amp;materialsCitation.latitude=-22.435194" title="Search Plazi for locations around (long -43.259193/lat -22.435194)">Reserva Biológica de Araras</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.259193&amp;materialsCitation.latitude=-22.435194" title="Search Plazi for locations around (long -43.259193/lat -22.435194)">Trilha da Água</a> (poço), 22°26’06.7” S, 43°15’33.1” W, 13.iv. 2018, 993 m, LL Dumas, JL Nessimian, C Novais &amp; ALD Ferreira leg. (DZRJ).</p><p>Paratypes: BRAZIL: Rio de Janeiro: Petrópolis, Araras, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.257557&amp;materialsCitation.latitude=-22.433277" title="Search Plazi for locations around (long -43.257557/lat -22.433277)">Reserva Biológica de Araras</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.257557&amp;materialsCitation.latitude=-22.433277" title="Search Plazi for locations around (long -43.257557/lat -22.433277)">Rio Araras</a> (bambuzal), 22°25’59.8” S, 43°15’27.2” W, 13.iv–07.v.2018, 1045 m, LL Dumas, JL Nessimian, C Novais &amp; ALD Ferreira leg., 3 males, 1 female (DZRJ) ; Petrópolis, Araras, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.260387&amp;materialsCitation.latitude=-22.437166" title="Search Plazi for locations around (long -43.260387/lat -22.437166)">Reserva Biológica de Araras</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.260387&amp;materialsCitation.latitude=-22.437166" title="Search Plazi for locations around (long -43.260387/lat -22.437166)">Trilha do Caneco</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.260387&amp;materialsCitation.latitude=-22.437166" title="Search Plazi for locations around (long -43.260387/lat -22.437166)">Rio Araras</a> (ponte), 22°26’13.8” S, 43°15’37.4” W, 19.iii.2018, 1086 m, LL Dumas, JL Nessimian, C Novais &amp; ALD Ferreira leg., 2 males, 1 female (MZSP) ; same locality, 20.iii–13.iv.2018, LL Dumas, JL Nessimian, C Novais &amp; ALD Ferreira leg., 47 males (DZRJ); same locality, 13.iv–07.v.2018, LL Dumas, JL Nessimian, C Novais &amp; ALD Ferreira leg., 40 males (DZRJ); Itatiaia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.61222&amp;materialsCitation.latitude=-22.455278" title="Search Plazi for locations around (long -44.61222/lat -22.455278)">Parque Nacional do Itatiaia</a>, Trilha dos Três Picos, 22°27’19.0” S, 44°36’44.0” W, 01.xii. 2012, 822 m, JR Mermudes leg., 1 male (DZRJ) .</p><p>Distribution. Brazil (Rio de Janeiro).</p><p>Etymology. The specific name “daome” is in reference to the Throne of Daomé (now Republic of Benin, West Africa). It had been donated by the ambassadors of King Adandozan (1718–1818) to Prince Regent Dom João VI, in 1811, and probably dates from the 18th to the 19th century. The Daomé Throne was on display in the Museu Nacional public exhibition, being incorporated into the museum’s collection in 1818.</p></div>	https://treatment.plazi.org/id/F556879FE752FF970785F8DD2F064B74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Dumas, Leandro Lourenço;Nessimian, Jorge Luiz	Dumas, Leandro Lourenço, Nessimian, Jorge Luiz (2019): New species of Helicopsyche von Siebold 1856 (Trichoptera: Helicopsychidae) from Brazil, including the redescription of Helicopsyche (Feropsyche) planorboides Machado 1957. Zootaxa 4619 (2): 231-250, DOI: 10.11646/zootaxa.4619.2.2
F556879FE754FF940785F8BD29C04BE6.text	F556879FE754FF940785F8BD29C04BE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helicopsyche (Feropsyche) dinoprata Dumas & Nessimian 2019	<div><p>Helicopsyche (Feropsyche) dinoprata, new species</p><p>(Figures 3 A–3G)</p><p>urn:lsid:zoobank.org:act: 78355E46-538E-44A3-92B7-DB36015CE5F4</p><p>Diagnosis. Helicopsyche (F.) dinoprata sp. nov. is unlike any other species in the subgenus Feropsyche in having broad, elongate inferior appendages pointed apically, each with its basomesal lobe short, forming a small protuberance with a single spine-like seta at its apex. Helicopsyche (F.) piroa Ross 1944, H. (F.) vergelana Ross 1956, and H. (F.) tapadas Denning 1966 also have elongate inferior appendages, however they are obtuse apically, and the basomesal lobes are well-developed. Furthermore, the setation of tergum X is also distinctive for the new species.</p><p>Description. Adult. Similar to H. (F.) bendego sp. nov. except as follows: Each foreleg anterior apical tibial spur about 2x longer than posterior spur. Male forewing length: 4.4–4.9 mm (n = 10; holotype male = 4.8 mm). Abdominal sternum VI ventral process moderately long, shorter than segment VI, tubular along its length, oriented posteroventrad, nearly straight in lateral view, with microtrichiae along length, apex rounded, with lamellae apicoventrally (Figs. 3F, 3G).</p><p>Male genitalia. Segment IX short ventrally; in lateral view with anteromesal margin well-developed, anterodorsal margin elongated, slightly sinuous, and anteroventral margin concave; lateral apodeme well-developed, located midlaterally on segment (Fig. 3A); in dorsal view saddle-like, with anterior margin strongly concave (Fig. 3B); in ventral view with anterior margin straight, anterolateral lobes projecting anterad, posterior margin convex, with mesal portion slightly projected (Fig. 3C). Segment X, in lateral view, elongate, with dorsal and ventral margins subparallel, ventral margin abruptly tapering near apex, apex rounded (Fig. 3A); in dorsal view not surpassing apices of inferior appendages, mesodorsal borders inverted Y-shaped, bearing 2 medially linear rows of 7–8 stout setae along length and a cluster of 6-8 stout setae apically, apex rounded, with slightly mesal notch (Fig. 3B). Superior appendages originate dorsolaterally, setose, rounded in lateral view, nearly club-like in dorsal view (Figs. 3A, 3B). Basal plate of inferior appendages finger-like in lateral view, not surpassing anteroventral margin of segment IX (Fig. 3A), large and rounded at base in ventral view (Fig. 3C); primary branches of inferior appendages each broad, strongly elongate, covered by long, stout setae; in lateral view narrow at base, strongly widening towards midlength, and then tapering apically, apex acute, dorsal margin strongly undulated (Fig. 3A); in dorsal and ventral views curved mesad, tapering apically, with apex subacute, turned mesad (Fig. 3B, 3C); basomesal lobe of each inferior appendage short, not visible in lateral view; in ventral view, slightly protruding from primary branch, bearing single spine-like seta (Fig. 3C). Phallus tubular, strongly down-curved along its length; phallobase narrow, elongate, with apicoventral margin elongate, sclerotized, rounded apically; endotheca membranous lobe partially covering endophallus; phallotremal sclerite small, C-shaped (Figs. 3D, 3E).</p><p>Holotype male: BRAZIL: Rio de Janeiro: Santa Maria Madalena, Parque Estadual do Desengano, Morumbeca dos Marreiros, afluente do <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.915363&amp;materialsCitation.latitude=-21.8775" title="Search Plazi for locations around (long -41.915363/lat -21.8775)">Ribeirão Macapá</a>, 21°52’39.0” S, 41°54’55.3” W, 21.x.2016, 1110 m, LL Dumas, JL Nessimian &amp; JF Barbosa leg. (DZRJ).</p><p>Paratypes: BRAZIL: Rio de Janeiro: same data as holotype, 1 male, 2 females (DZRJ); same data as holotype, 1 male, 1 female (MZSP); Santa Maria Madalena, Parque Estadual do Desengano, Morumbeca dos Marreiros, Trilha da Mina, afluente do Ribeirão Macapá, 21°52’3935.6” S, 41°54’55.0” W, 22.x.2016, 1011 m, LL Dumas, JL Nessimian &amp; JF Barbosa leg., 2 males, 3 females (DZRJ) ; São Fidélis, Parque Estadual do Desengano, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.915028&amp;materialsCitation.latitude=-21.875305" title="Search Plazi for locations around (long -41.915028/lat -21.875305)">Morumbeca</a> dos <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.915028&amp;materialsCitation.latitude=-21.875305" title="Search Plazi for locations around (long -41.915028/lat -21.875305)">Marreiros</a>, afluente do <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.915028&amp;materialsCitation.latitude=-21.875305" title="Search Plazi for locations around (long -41.915028/lat -21.875305)">Ribeirão Macapá</a>, 21°52’31.1” S, 41°54’54.1” W, 21.x.2016, 1058 m, LL Dumas, JL Nessimian &amp; JF Barbosa leg., 5 males, 2 females (DZRJ) ; Itatiaia, Parque Nacional do Itatiaia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.61222&amp;materialsCitation.latitude=-22.455278" title="Search Plazi for locations around (long -44.61222/lat -22.455278)">Trilha</a> dos <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.61222&amp;materialsCitation.latitude=-22.455278" title="Search Plazi for locations around (long -44.61222/lat -22.455278)">Três Picos</a>, 22°27’19.0” S, 44°36’44.0” W, 01.xii. 2012, 822 m, JR Mermudes leg., 2 males, 2 females (DZRJ) ; Itatiaia, Maromba, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.615414&amp;materialsCitation.latitude=-22.330473" title="Search Plazi for locations around (long -44.615414/lat -22.330473)">Rio Preto</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.615414&amp;materialsCitation.latitude=-22.330473" title="Search Plazi for locations around (long -44.615414/lat -22.330473)">Cachoeira do Escorrega</a>, 22°19’49.7” S, 44°36’55.5” W, 26.i.2012, 1381 m, BHL Sampaio &amp; ALH Oliveira leg., 1 male (DZRJ) ; Mangaratiba, Parque Estadual do Cunhambebe, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.98761&amp;materialsCitation.latitude=-22.89939" title="Search Plazi for locations around (long -43.98761/lat -22.89939)">Sahy</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.98761&amp;materialsCitation.latitude=-22.89939" title="Search Plazi for locations around (long -43.98761/lat -22.89939)">Trilha Sahy-Rubião</a>, 22°53’57.8” S, 43°59’15.4” W, vii–viii. 2016, 400 m, LS Barbosa leg., 9 males, 7 females (DZRJ) ; Paraty, Sertão do Taquari, afluente do <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.69589&amp;materialsCitation.latitude=-23.041777" title="Search Plazi for locations around (long -44.69589/lat -23.041777)">Rio Taquari</a>, 23°02’30.4” S, 44°41’45.2” W, 18.viii.2012, 50 m, PM Souto leg., 1 male, 1 female (DZRJ) ; Cachoeiras de Macacu, Guapiaçu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-42.738945&amp;materialsCitation.latitude=-22.42086" title="Search Plazi for locations around (long -42.738945/lat -22.42086)">Reserva Ecológica de Guapiaçu</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-42.738945&amp;materialsCitation.latitude=-22.42086" title="Search Plazi for locations around (long -42.738945/lat -22.42086)">Rio Manoel Alexandre</a>, 22°25’15.1” S, 42°44’20.2” W, 23.viii. 2018, 135 m, LL Dumas &amp; JL Nessimian leg., 2 males (DZRJ) .</p><p>Distribution. Brazil (Rio de Janeiro).</p><p>Etymology. This species is named for Maxakalissaurus topai Kellner et al. 2006, an aeolosaurid dinosaur affectionally known as “Dinoprata”. Maxakalissaurus topai is the biggest dinosaur described in Brazil, measuring about 13 meters and nine tons, and lived about 80 million years ago. It was found in the city of Prata, Minas Gerais, Brazil, in 1998. A reconstructed Maxakalisaurus topai skeleton was on display at the Museu Nacional, being one of the main attractions of the public exhibition.</p></div>	https://treatment.plazi.org/id/F556879FE754FF940785F8BD29C04BE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Dumas, Leandro Lourenço;Nessimian, Jorge Luiz	Dumas, Leandro Lourenço, Nessimian, Jorge Luiz (2019): New species of Helicopsyche von Siebold 1856 (Trichoptera: Helicopsychidae) from Brazil, including the redescription of Helicopsyche (Feropsyche) planorboides Machado 1957. Zootaxa 4619 (2): 231-250, DOI: 10.11646/zootaxa.4619.2.2
F556879FE759FF980785FF562A984C88.text	F556879FE759FF980785FF562A984C88.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helicopsyche (Feropsyche) luziae Dumas & Nessimian 2019	<div><p>Helicopsyche (Feropsyche) luziae, new species</p><p>(Figures 4 A–4G)</p><p>urn:lsid:zoobank.org:act: A23278E4-1E62-4CF9-8629-0BD1DAE20496</p><p>Diagnosis. The new species closely resembles Helicopsyche (F.) planorboides by the overall shape of tergum X, the racket-like inferior appendages as viewed laterally and the finger-like elongated basomesal lobes. However, H. (F.) luziae sp. nov. can be distinguished from H. (F.) planorboides by the absence of an apicoventral projection on each inferior appendage and by the tab-like basal plate, which is rounded in the second species. Also, the new species can be distinguished by the short midlength projection on tergum X (almost hidden in lateral view), which is robust and spine-like as viewed dorsally but tab-like in lateral view in H. (F.) planorboides .</p><p>Description. Adult. Similar to H. (F.) bendego sp. nov. except as follows: Overall color (in alcohol) nearly uniformly dark-brown. Cephalic setal warts large, drop-like; postantennal warts apparently absent. Maxillary palps with proximal and distal segments covered with several light-brown setae. Each foreleg anterior apical tibial spur about 4x longer than posterior spur. Male forewing length: 3.6–4.4 mm (n = 3; holotype male = 4.4 mm). Hind wings each with forks I, III, and V present. Abdominal sternum VI ventral process much shorter than segment VI, slightly wider at base, nearly straight in lateral view, with microtrichiae along length, its apex oblique in lateral view, subtruncate in ventral view, with few lamellae apicoventrally (Figs. 4F, 4G).</p><p>Male genitalia. Segment IX short ventrally; in lateral view with anteromesal margin well-developed, anterodorsal margin slightly concave, and anteroventral margin almost straight; lateral apodeme well-developed, located midlaterally on segment (Fig. 4A); in dorsal view with anterior margin strongly concave, inverted V-shaped, lateral margins sinuous (Fig. 4B); in ventral view with anterior margin almost straight, bearing acute anterolateral lobes projected anterad, dorsal margin slightly concave, bearing shallow recess mesally (Fig. 4C). Segment X, in lateral view, finger-like, wider at base, with two small projections, one spine-like midlaterally and one tab-like apicolaterally (Fig. 4A, 4C); in dorsal view subrectangular, slightly surpassing the apex of inferior appendages, mesodorsal borders straight, parallel, long, forming mesal crest, bearing scattered stout setae subapically and two apical clusters of 8–10 stout setae, small midlateral projections directed anterad, apex with deep mesal V-shaped notch, each apical portion with acute and elevated apex (Fig. 4B). Superior appendages originate dorsolaterally, wide, setose, round in lateral view, slightly elongate in dorsal view (Figs. 4A, 4C). Basal plate of inferior appendages sickle-like in lateral view, not surpassing anteroventral margin of segment IX (Fig. 4A), broad and trapezoidal in ventral view, nearly rounded anteriorly (Fig. 4C); primary branches of inferior appendages covered by stout, long setae; in lateral view each racket-like, with distal portion rounded and about 3x as broad as its proximal portion, dorsal margin strongly concave, ventral margin slightly concave and undulating, apicodorsal margin slightly undulating, with long, stout setae along apical margin (Fig. 4A); in dorsal view oblong, with inner margin strongly projecting mesad, with many long, stout setae along apical margin, apex rounded, bearing 3–4 long, stout setae (Fig. 4B); in ventral view, with three mesal longitudinal ridges, bearing many stout setae, external portion sickle-like, with 4–6 stout setae apically, apex subacute, first subventral mesal longitudinal ridge with 2–3 stout setae sub-basally, middle mesal longitudinal ridge short, forming small protuberance subapically, without setae, subdorsal mesal longitudinal ridge extending from midlength to apex, with 3–4 short, stout setae (two more-apical ones on distinct protuberances), dorsomesal margin with 4–6 stout setae (two more-apical ones on distinct protuberances) (Fig. 4C); basomesal lobes of inferior appendages finger-like, very long, strongly protruding beyond primary branches in lateral view (Fig. 4A), in ventral view wide at base, with several subapical and many apical short, stout setae (Fig. 4C). Phallus tubular, strongly down-curved along its length; phallobase narrow, slightly inflated basodorsally and mesoventrally, with apicoventral margin elongate, semi-membranous apically in lateral view, in ventral view with two sclerotized lobes curved ventroanterad (lobes tab-like in lateral view); endotheca membranous lobe extending dorsally over endophallus; phallotremal sclerite small, slightly C-shaped (Figs. 4D, 4E).</p><p>Holotype male: BRAZIL: Rio de Janeiro: São Fidélis, Itacolomi, afluente de 1ª ordem do <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.85975&amp;materialsCitation.latitude=-21.830473" title="Search Plazi for locations around (long -41.85975/lat -21.830473)">Rio do Colégio</a>, 21°49’49.7” S, 41°51’35.1” W, 23.x. 2016, 421 m, LL Dumas, JL Nessimian &amp; JF Barbosa leg. (DZRJ).</p><p>Paratypes: BRAZIL: Rio de Janeiro: same data as holotype, 1 female (DZRJ); same data as holotype, 1 male (MZSP); Petrópolis, Araras, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.28086&amp;materialsCitation.latitude=-22.42739" title="Search Plazi for locations around (long -43.28086/lat -22.42739)">Ponte Funda</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.28086&amp;materialsCitation.latitude=-22.42739" title="Search Plazi for locations around (long -43.28086/lat -22.42739)">Cachoeira da Ponte Funda</a>, 22°25’38.6” S, 43°16’51.1” W, 18.iii.2018, 1072 m, LL Dumas, JL Nessimian, C Novais &amp; ALD Ferreira leg., 1 male (DZRJ) .</p><p>Distribution. Brazil (Rio de Janeiro).</p><p>Etymology. This species is named for Luzia Woman, an upper paleolithic period skeleton that was found in a cave in Brazil in 1975 by the French archaeologist Annette Laming-Emperaire. At about 11,500 years old, she was believed to be the oldest human skeleton found in the Americas. The Luzia skull was on display in the public exhibition of the Museu Nacional.</p></div>	https://treatment.plazi.org/id/F556879FE759FF980785FF562A984C88	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Dumas, Leandro Lourenço;Nessimian, Jorge Luiz	Dumas, Leandro Lourenço, Nessimian, Jorge Luiz (2019): New species of Helicopsyche von Siebold 1856 (Trichoptera: Helicopsychidae) from Brazil, including the redescription of Helicopsyche (Feropsyche) planorboides Machado 1957. Zootaxa 4619 (2): 231-250, DOI: 10.11646/zootaxa.4619.2.2
F556879FE75BFF9E0785FEEC2F3F4DF0.text	F556879FE75BFF9E0785FEEC2F3F4DF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helicopsyche (Feropsyche) petri Dumas & Nessimian 2019	<div><p>Helicopsyche (Feropsyche) petri, new species</p><p>(Figures 5 A–5G)</p><p>urn:lsid:zoobank.org:act: 6B01EAB2-6C74-4BEB-A0CF-DAF3F480179F</p><p>Diagnosis. The new species is similar to Helicopsyche (F.) monda Flint 1983 and H. (F.) lambda Flint 1983 by the overall shape of the inferior appendages and their basomesal lobes in lateral view. It differs from both in that the inferior appendages are wider dorsally, each with its inner margin strongly projected mesad, forming a rounded large lobe, and the basomesal lobes are less trianguloid, bearing spine-like setae. Furthermore, H. (F.) petri sp. nov. has tergum X without a mesal notch and is truncate apically, while in H. (F.) monda and H. (F.) lambda it has a shallow mesal notch and is more rounded apically.</p><p>Description. Adult. Similar to H. (F.) bendego sp. nov. except as follows: Overall color (in alcohol) uniformly yellowish-brown; antennae, palps and legs pale yellow. Each foreleg anterior apical tibial spur about 3x longer than posterior spur. Male forewings each 3.2–3.9 mm long (n = 10; holotype male = 3.4 mm). Abdominal sternum VI ventral process long, about as long as segment VI, tubular along its length, oriented posteroventrad, slightly wider at base, curved slightly upward in lateral view, with microtrichiae along length, apex rounded, with lamellae apicoventrally (Figs. 5F, 5G).</p><p>Male genitalia. Segment IX short ventrally; in lateral view with anteromesal margin well-developed, anterodorsal margin almost straight, and anteroventral margin concave; lateral apodeme well-developed, located midlaterally on segment (Fig. 5A); in dorsal view horseshoe-like, with anterior margin strongly concave (Fig. 5B); in ventral view with anteroventral and posteroventral sclerotized bars, lateral portions well-developed, anterolateral lobes projected anterad (Fig. 5C). Segment X, in lateral view, short, triangular, tapering to apex from base (Fig. 5A); in dorsal view subtrapezoidal, not surpassing apices of inferior appendages, mesodorsal borders well-marked, inverted Y-shaped, bearing sparse stout setae near middle of lateral margins and pair of apicolateral clusters of 4 stout setae, apex truncate, without mesal notch (Fig. 5B). Superior appendages ovate in lateral view, clavate in dorsal view (Figs. 5A, 5B). Basal plate of inferior appendages subtriangular in lateral and ventral views, almost reaching the anteroventral margin of segment IX as viewed laterally (Figs. 5A, 5C); primary branches of inferior appendages each in lateral view nearly drop-like, broad, distal portion about 2x broader than its proximal portion, dorsal margin slightly convex, ventral margin sinuous, apicodorsal margin concave, undulating, with long, stout setae along apical margin, apex pointed (Fig. 5A); in dorsal and ventral views broad, curved mesad, tapering apically (Figs. 5B, 5C); in dorsal view with dorsal portion rounded, strongly projecting mesad (visible also in ventral view), with about 12–15 long, stout setae at inner margin, inner margin undulate, apex narrow, subacute, turned mesad, with an apical short, stout seta (Fig. 5B); in ventral view with inner margin undulating, bearing several long, stout setae (Fig. 5C); basomesal lobes of inferior appendages each short, slightly protruding beyond primary branch in lateral view (Fig. 5A); in ventral view with right-angled median corner, bearing 5-6 short, dark spine-like setae (Fig. 5C). Phallus tubular, moderately down-curved along its length; phallobase narrow, short, slightly inflated mesodorsally, its apicoventral margin elongate, rounded apically, apex semi-membranous; endotheca membranous lobe partially covering endophallus; phallotremal sclerite small, strongly C-shaped (Figs. 5D, 5E).</p><p>Holotype male: BRAZIL: Rio de Janeiro: Rio das Ostras, Reserva Biológica União, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-42.039223&amp;materialsCitation.latitude=-22.424778" title="Search Plazi for locations around (long -42.039223/lat -22.424778)">Trilha Interpretativa do Pilão</a>, riacho, 22°25’29.2” S, 42°02’21.2” W, 07.iii.2014, 42 m, FC Avelino &amp; LS Barbosa leg. (DZRJ).</p><p>Paratypes: BRAZIL: Rio de Janeiro: same data as holotype, 8 males (DZRJ); same data as holotype, 2 males (MZSP); Nova Iguaçu, Tinguá, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.437336&amp;materialsCitation.latitude=-22.585667" title="Search Plazi for locations around (long -43.437336/lat -22.585667)">Reserva Biológica do Tinguá</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.437336&amp;materialsCitation.latitude=-22.585667" title="Search Plazi for locations around (long -43.437336/lat -22.585667)">Rio Tinguá</a> (atrás da brigada), 22°35’08.4” S, 43°26’14.4” W, 13.ix. 2016, 159 m, LL Dumas, JL Nessimian &amp; JF Barbosa leg., 1 male (DZRJ) ; Nova Iguaçu, Tinguá, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.441086&amp;materialsCitation.latitude=-22.58875" title="Search Plazi for locations around (long -43.441086/lat -22.58875)">Reserva Biológica do Tinguá</a>, estrada de captação de água do <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.441086&amp;materialsCitation.latitude=-22.58875" title="Search Plazi for locations around (long -43.441086/lat -22.58875)">Macuco</a>, afluente do <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.441086&amp;materialsCitation.latitude=-22.58875" title="Search Plazi for locations around (long -43.441086/lat -22.58875)">Rio Tinguá</a>, 22°35’19.5” S, 43°26’27.9” W, 15.ix. 2016, 211 m, LL Dumas, JL Nessimian &amp; JF Barbosa leg., 1 male (DZRJ) ; Miguel Pereira, Vera Cruz, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.431667&amp;materialsCitation.latitude=-22.52136" title="Search Plazi for locations around (long -43.431667/lat -22.52136)">Reserva Biológica do Tinguá</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.431667&amp;materialsCitation.latitude=-22.52136" title="Search Plazi for locations around (long -43.431667/lat -22.52136)">Córrego Fazenda</a> das <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.431667&amp;materialsCitation.latitude=-22.52136" title="Search Plazi for locations around (long -43.431667/lat -22.52136)">Pedras</a>, 22°31’16.9” S, 43°25’54.0” W, 10.xi. 2016, 754 m, LL Dumas, JL Nessimian, B Clarkson &amp; JF Barbosa leg., 1 male (DZRJ) .</p><p>Distribution. Brazil (Rio de Janeiro).</p><p>Etymology. This species is dedicated to the second and last Brazilian emperor Dom Pedro II ( Petrus means Pedro in Latin), who was born in Rio de Janeiro in 1825 and died in 1891. The Imperial Palace (Paço de São Cristóvão) housed the Brazilian Imperial Family between 1822 and 1889, and also hosted the Republican Constituent Assembly from 1889 to 1891, before being assigned to the use of the National Museum in 1892. Dom Pedro II was considered a patron of arts and science and was an avid amateur scientist, playing a key role in advances in science, technology and education in Brazil. By the end of the 19th century, reflecting the personal preferences of Dom Pedro II, the National Museum started to invest in several scientific areas. The Emperor himself contributed with several of the collections artifacts, which he acquired during many of his trips abroad. In 1946, the Imperial Palace was incorporated into the Universidade Federal do Rio de Janeiro, hosting several researchers and six post-graduations courses, as Zoology, Botany and Anthropology. Unfortunately, the palace, which housed a large part of the 20 million objects in the museum’s collection, was engulfed by a fire on 2 September 2018.</p></div>	https://treatment.plazi.org/id/F556879FE75BFF9E0785FEEC2F3F4DF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Dumas, Leandro Lourenço;Nessimian, Jorge Luiz	Dumas, Leandro Lourenço, Nessimian, Jorge Luiz (2019): New species of Helicopsyche von Siebold 1856 (Trichoptera: Helicopsychidae) from Brazil, including the redescription of Helicopsyche (Feropsyche) planorboides Machado 1957. Zootaxa 4619 (2): 231-250, DOI: 10.11646/zootaxa.4619.2.2
F556879FE75DFF9C0785FDCE2E0B4CA4.text	F556879FE75DFF9C0785FDCE2E0B4CA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helicopsyche (Feropsyche) shaamunensu Dumas & Nessimian 2019	<div><p>Helicopsyche (Feropsyche) shaamunensu, new species</p><p>(Figures 6 A-G)</p><p>urn:lsid:zoobank.org:act: D30B23D9-C044-4B3C-A913-649AB8FF2CA8</p><p>Diagnosis. Helicopsyche (F.) shaamunensu sp. nov. resembles Helicopsyche (F.) lazzariae Holzenthal, Bahnik &amp; Calor 2016 by the overall deltoid shape of the primary branches of inferior appendages. The new species can be easily distinguished from H. (F.) lazzariae by the presence of basomesal lobes of inferior appendages, which are absent in the second species, and by the shape and setation of tergum X in dorsal view.</p><p>Description. Adult. Similar to H. (F.) bendego sp. nov. except as follows: Overall color (in alcohol) uniformly golden-brown; antennae, palps, and legs pale yellow. Maxillary palps with proximal segment longer than distal one. Each foreleg anterior apical tibial spur about 2x longer than posterior spur. Male forewings each 3.2 mm long (holotype male). Abdominal sternum VI ventral process long, about as long as segment VI, slender, tubular along its length, oriented posteroventrad, slightly wider at apex, nearly straight in lateral view with apex subacute, with lamellae apically (Figs. 6F, 6G).</p><p>Male genitalia. Segment IX short ventrally; in lateral view with anteromesal margin well-developed, anterodorsal margin strongly convex near dorsum, and anteroventral margin slightly convex; lateral apodeme well-developed, located midlaterally in the segment (Fig. 6A); in dorsal view with anterior margin strongly concave, lateral margins concave (Fig. 6B); in ventral view subrectangular, with anterior and posterior margins slightly concave (Fig. 6C). Segment X, in lateral view, subtriangular, strongly tapering to apex from apical 2/3 (Fig. 6A); in dorsal view slightly narrowed near base, about as long as inferior appendages, mesodorsal borders inverted V-shaped, bearing 1–3 short, stout setae subapically and pair of clusters of short, stout setae apically, apex rounded on either side of slight mesal notch (Fig. 6B). Superior appendages originate dorsolaterally, setose, rounded in lateral view, club-like and laterally directed in dorsal view (Figs. 6A, 6B). Basal plate of inferior appendages triangular in lateral and ventral views, tapering to apex, slightly surpassing anteroventral margin of segment IX (Figs. 6A, 6C); primary branches of inferior appendages large, covered by long, stout setae; in lateral view each deltoid, with distal portion about 2.5x broader than its proximal portion, dorsal margin slightly convex, ventral margin slightly convex near base, with 3 stout setae, and strongly convex near apex, with 5–6 stout setae, posterodorsal margin concave, slightly undulated, with sparse long, stout setae along apical margin, apex pointed (Fig. 6A); in dorsal view curved mesad, tapering apically, dorsal margin projecting at midlength, bearing several stout setae along apical margin, apex subacute, with single short, stout seta (Fig. 6B); in ventral view, with inner margin sinuous, with 3 stout setae basally and row of 4–5 long, stout setae subapically (Fig. 6C); basomesal lobes of inferior appendages well-developed, narrow basally, widening apically, bifid at apex and protruding beyond primary branches in lateral view (Fig. 6A); in ventral view each of these lobes bearing 3–4 stout setae on upper apical surface and with small lateral protuberance bearing 2–3 short, stout setae at inner margin (Fig. 6C). Phallus tubular, strongly down-curved along its length; phallobase narrow, narrowest in middle, inflated basodorsally, with apicoventral margin elongate, strongly sclerotized, rounded apically, semi-membranous apically in lateral view; endotheca large, long, membranous lobe extending dorsally over endophallus; endophallic membranes not inflated; phallotremal sclerite of moderate size, slightly C-shaped (Figs. 6D, 6E).</p><p>Holotype male: BRAZIL: Rio de Janeiro: Nova Iguaçu, Tinguá, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.44542&amp;materialsCitation.latitude=-22.583918" title="Search Plazi for locations around (long -43.44542/lat -22.583918)">Reserva Biológica do Tinguá</a>, afluente do <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.44542&amp;materialsCitation.latitude=-22.583918" title="Search Plazi for locations around (long -43.44542/lat -22.583918)">Rio Tinguá</a>, 22°35’02.1” S, 43°26’43.5” W, 13.ix. 2016, 233 m, LL Dumas, JL Nessimian &amp; JF Barbosa leg. (DZRJ).</p><p>Distribution. Brazil (Rio de Janeiro).</p><p>Etymology. This species is named for Sha-Amun-En-Su, an Egyptian priestess and singer who lived in Thebes during the first half of the 8th century B.C., responsible for ceremonial duties at the Temple of Karnak, dedicated to the god Amun. The wonderful wooden sarcophagus and its mummy were given as presents to the Brazilian emperor Dom Pedro II during his second travel to Egypt, in 1876, by the Khedive Ismail Pasha. In 1889 they became part of the great Egyptian artifacts collection of the Museu Nacional, one of the specialties of the museum.</p></div>	https://treatment.plazi.org/id/F556879FE75DFF9C0785FDCE2E0B4CA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Dumas, Leandro Lourenço;Nessimian, Jorge Luiz	Dumas, Leandro Lourenço, Nessimian, Jorge Luiz (2019): New species of Helicopsyche von Siebold 1856 (Trichoptera: Helicopsychidae) from Brazil, including the redescription of Helicopsyche (Feropsyche) planorboides Machado 1957. Zootaxa 4619 (2): 231-250, DOI: 10.11646/zootaxa.4619.2.2
F556879FE75FFF9D0785FE822F9D4ADB.text	F556879FE75FFF9D0785FE822F9D4ADB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helicopsyche (Feropsyche) planorboides Machado 1957	<div><p>Helicopsyche (Feropsyche) planorboides Machado</p><p>(Figures 7 A–7B, 8A–8G)</p><p>Helicopsyche (Feropsyche) planorboides Machado 1957: 193 [Type locality: Brazil, Minas Gerais, Tarumirim, Rio Doce valley; type depository not designated, now in DZRJ; ♂; ♀; larva; pupa; case].— Johanson 1995:115 [catalog].— Johanson 2002: 135 [distribution].— Paprocki et al. 2004: 6 [checklist].— Paprocki and França 2014: 18 [checklist].</p><p>The types of Helicospyche (F.) planorboides were donated by Dr. Angelo Machado to Coleção Entomológica Professor José Alfredo Pinheiro Dutra, Universidade Federal do Rio de Janeiro (DZRJ). Specimens were stored in vials with alcohol; however, they are greatly damaged, stiffened, and distorted due to the lower concentration of alcohol in which they were stored. The male holotype is slightly better preserved, but with broken wings and several missing parts: antennae, palps, all legs, portions of thorax, and some genital structures (inferior appendages and phallus). Thus, this redescription of H. (F.) planorboides was also based on additional specimens which have the same pattern when compared with the male holotype, especially the distinctive tergum X, avoiding the complete damage of the type specimens in this way.</p><p>Diagnosis. Helicopsyche (F.) planorboides is easily distinguished by the unusual tergum X, which has a pair of large tab-like midlength projections, a distal thorn-like process in lateral view, and a longitudinally cleft apically. Only H. (F.) luziae sp. nov. has a similar pattern of tergum X among American species of the subgenus Feropsyche, but in the new species the tergum X projections are much less developed. Furthermore, both species can be also distinguished by the general structure of the inferior appendages, as discussed in the H. (F.) luziae sp. nov. diagnosis. Also, H. (F.) planorboides is recorded for the first time for Rio de Janeiro and Espírito Santo states.</p><p>Description. Adult. Similar to H. (F.) bendego sp. nov. except as follows: Antennae golden-brown, palps and legs yellowish-brown. Cephalic setal warts narrow, drop-like; postantennal warts apparently absent. Unpaired eversible membranous structure in dorsal portion of head, located close to coronal suture, concealed by pair of movable sclerites in male holotype and some of additional specimens included here; membranous structure tubular, directed anteriorly between antennal scapes, extremely developed, about as long as antennal scapes (Figs. 7A, 7B). Maxillary palps covered with several yellow setae. Each foreleg anterior apical tibial spur about 3x longer than posterior spur. Male forewings each 4.8 mm (n = 2). Forewing discoidal cell about half as long as thyridial cell; hindwings each with forks I, III, and V present. Abdominal sternum VI ventral process much shorter than segment VI, lanceolate, slightly narrower at base in ventral view, nearly straight in lateral view, apex acute in lateral and ventral views, with microtrichiae along length but without lamellae apicoventrally (Figs. 8F, 8G).</p><p>Male genitalia. Segment IX short ventrally; in lateral view with anteromesal margin well-developed, anterodorsal margin slightly sinuous, and anteroventral margin almost straight; lateral apodeme well-developed, located midlaterally in the segment (Fig. 8A); in dorsal view with anterior margin strongly concave, inverted V-shaped, lateral margins slightly convex (Fig. 8B); in ventral view with anterior margin very strongly concave, dorsal margin concave in middle (Fig. 8C). Segment X, in lateral view, taller at base, with ventral and dorsal margins parallel from midlength, rounded apically, bearing two large projections: midlateral projection tab-like, apical projection forming thorn-like, both upturned (Fig. 8A); in dorsal view surpassing apices of inferior appendages, dorsally with two marginal rows of 5–6 setae at apical fourth, apex with longitudinal V-shaped notch at apical fourth, midlateral projections acute, apical projections forming pair of auriculate rims with crenulate margins (Fig. 8B). Superior appendages originate dorsolaterally, large, setose, rounded in lateral view, club-like in dorsal view (Figs. 8A, 8B). Basal plate of inferior appendages tapering anterad in lateral and ventral views, surpassing anteroventral margin of segment IX (Figs. 8A, 8C); primary branches of inferior appendages covered by stout, long setae; in lateral view each racket-like, with distal portion rounded, about 3x as broad as its proximal portion, basodorsal margin strongly concave, ventral margin slightly concave, apicodorsal margin strongly convex, slightly undulating, with long, stout setae along apical margin (Fig. 8A); in dorsal view curved mesad, subrectangular, with small protuberance subapically on inner margin, bearing several small spine-like setae (Fig. 8B); in ventral view, with inner margin concave and with rounded protuberance near base bearing 6 long stout setae, and subapically projecting strongly mesad and bearing several apical spines (Fig. 8C); basomesal lobes of inferior appendages thumb-like, very long, strongly protruding beyond primary branches in lateral view (Fig. 8A), in ventral view finger-like, slightly broader at base, with row of 4 marginal short, stout, subapicolateral setae on distal 1/3 (Fig. 8C). Phallus tubular, strongly down-curved along its length; phallobase narrow, slightly inflated basally and mesoventrally, with apicoventral margin elongate, sclerotized apically, in lateral view with lateral sclerotized lobes curved anteroventrad (not visible in ventral view); endotheca membranous lobe extending dorsally over endophallus; phallotremal sclerite small, C-shaped, apically with two semi-membranous structures (Figs. 8D, 8E).</p><p>Material examined: BRAZIL: Minas Gerais: Tarumirim, Rio Doce valley, v.1956, male holotype (DZRJ) ; same data as holotype, 5 female paratypes (DZRJ); Espírito Santo: Santa Teresa, Reserva Biológica Santa Lúcia, 09-10.i.2003, AL Carvalho leg., 3 males (DZRJ) ; Rio de Janeiro: Campos dos Goytacazes, Mocotó, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.75514&amp;materialsCitation.latitude=-21.814888" title="Search Plazi for locations around (long -41.75514/lat -21.814888)">Parque Estadual do Desengano</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.75514&amp;materialsCitation.latitude=-21.814888" title="Search Plazi for locations around (long -41.75514/lat -21.814888)">Rio Mocotó</a> (poço), 21°48’53.6” S, 41°45’18.5” W, 17.iv. 2016, 237 m, LL Dumas, JL Nessimian, CS Portela &amp; JF Barbosa leg., 1 male, 1 female (DZRJ) — male used for illustrations of genital structures here; São Fidélis, Itacolomi, afluente de 1ª ordem do <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.85975&amp;materialsCitation.latitude=-21.830473" title="Search Plazi for locations around (long -41.85975/lat -21.830473)">Rio do Colégio</a>, 21°49’49.7” S, 41°51’35.1” W, 23.x. 2016, 421 m, LL Dumas, JL Nessimian &amp; JF Barbosa leg., 1 male (DZRJ) .</p><p>Distribution. Brazil (Espírito Santo, Minas Gerais, and Rio de Janeiro).</p><p>Notes on membranous head structure. Glandular structures on the male head are unusual among Trichoptera, being common only in some microcaddisflies ( Hydroptilidae), like many Hydroptila Dalman 1819, Leucotrichia Mosely 1934 and Zumatrichia Mosely 1937 . These eversible head organs have been speculated to produce chemical signals and are concealed by moveable lateral occipital sclerites when they are not extended. There are no experiments to determine the function of these glands, but Roemhild (1980) assumed that they produce sex-pheromone, since they occur only in males at the sexually active stage. There is no mention in the literature about the presence of these eversible head organs in Helicopsychidae species. Helicopsyche (F.) luziae sp. nov. has similar occipital sclerites found in H. planorboides head, however membranous structures were not observed (we did not use lactic acid for the head due the low number of male specimens). So, more morphological and behavioral studies are required to establish the glandular pheromone function of these structures in Helicopsyche species.</p></div>	https://treatment.plazi.org/id/F556879FE75FFF9D0785FE822F9D4ADB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Dumas, Leandro Lourenço;Nessimian, Jorge Luiz	Dumas, Leandro Lourenço, Nessimian, Jorge Luiz (2019): New species of Helicopsyche von Siebold 1856 (Trichoptera: Helicopsychidae) from Brazil, including the redescription of Helicopsyche (Feropsyche) planorboides Machado 1957. Zootaxa 4619 (2): 231-250, DOI: 10.11646/zootaxa.4619.2.2
F556879FE75EFF830785F8D129B04D38.text	F556879FE75EFF830785F8D129B04D38.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helicopsyche (Feropsyche) catoles Souza, Gomes & Calor 2017	<div><p>Helicopsyche (Feropsyche) catoles Souza, Gomes &amp; Calor</p><p>This species was previously recorded for Chapada da Diamantina, a large mountain range in Bahia state, northeastern Brazil. The area includes several vegetation types, like savanna, campo rupestre, forests and Caatinga biome, being one of the highest semiarid areas of Brazil. The type-locality of H. (F.) catoles is a forest fragments at Chapada da Diamantina. Now, we record this species in high areas of the Atlantic Forest biome in Rio de Janeiro state .</p><p>Material examined. BRAZIL: Rio de Janeiro: Nova Friburgo, Macaé de Cima, afluente do <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.83594&amp;materialsCitation.latitude=-21.836973" title="Search Plazi for locations around (long -41.83594/lat -21.836973)">Rio Macaé</a> (ponte), 22°25’57.6” S, 42°32’24.3” W, 17.ii.2016, 1103 m, N Ferreira Jr. &amp; RJP Silva leg., 6 males, 9 females (DZRJ) ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.83594&amp;materialsCitation.latitude=-21.836973" title="Search Plazi for locations around (long -41.83594/lat -21.836973)">Campos dos Goytacazes</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.83594&amp;materialsCitation.latitude=-21.836973" title="Search Plazi for locations around (long -41.83594/lat -21.836973)">Parque Estadual do Desengano</a>, Itacolomi, afluente de 1ª ordem do <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.83594&amp;materialsCitation.latitude=-21.836973" title="Search Plazi for locations around (long -41.83594/lat -21.836973)">Córrego da Malhada Branca</a>, 21°50’13.1” S, 41°50’09.4” W, 24.x. 2016, 950 m, LL Dumas, JL Nessimian &amp; JF Barbosa leg., 3 males, 4 females (DZRJ)</p><p>.</p><p>Distribution. Brazil (Bahia, and Rio de Janeiro).</p></div>	https://treatment.plazi.org/id/F556879FE75EFF830785F8D129B04D38	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Dumas, Leandro Lourenço;Nessimian, Jorge Luiz	Dumas, Leandro Lourenço, Nessimian, Jorge Luiz (2019): New species of Helicopsyche von Siebold 1856 (Trichoptera: Helicopsychidae) from Brazil, including the redescription of Helicopsyche (Feropsyche) planorboides Machado 1957. Zootaxa 4619 (2): 231-250, DOI: 10.11646/zootaxa.4619.2.2
F556879FE740FF830785FE772F494FB0.text	F556879FE740FF830785FE772F494FB0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helicopsyche (Feropsyche) guara Holzenthal, Blahnik & Calor	<div><p>Helicopsyche (Feropsyche) guara Holzenthal, Blahnik &amp; Calor</p><p>This species was previously recorded for Parque Ecológico Spitzkopf, a small Atlantic Forest area in Santa Catarina state, southern Brazil, and Parque Nacional do Peruaçu, a national park with several limestone caves covered by representative samples of Cerrado biome and seasonal forest, being located in Minas Gerais state, southeastern Brazil. Herein, we record H. (F.) guara for the first time in several Atlantic Forest areas of Rio de Janeiro state.</p><p>Material examined. BRAZIL: Rio de Janeiro: Rio Bonito, Lavras, estrada velha de <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-42.583942&amp;materialsCitation.latitude=-22.667889" title="Search Plazi for locations around (long -42.583942/lat -22.667889)">Lavras</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-42.583942&amp;materialsCitation.latitude=-22.667889" title="Search Plazi for locations around (long -42.583942/lat -22.667889)">Rio Bacaxá</a>, 22°40’04.4” S, 42°35’02.2” W, 22.x.2017, 71 m, LL Dumas, JL Nessimian &amp; JF Barbosa leg., 2 males, 8 females (DZRJ) ; Saquarema, Tinguí, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-42.615776&amp;materialsCitation.latitude=-22.844501" title="Search Plazi for locations around (long -42.615776/lat -22.844501)">Rio Tinguí</a>, 22°50’40.2” S, 42°36’56.8” W, 21.x. 2017, 154 m, LL Dumas, JL Nessimian &amp; JF Barbosa leg., 2 males (DZRJ) ; Nova Iguaçu, Tinguá, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.43667&amp;materialsCitation.latitude=-22.582056" title="Search Plazi for locations around (long -43.43667/lat -22.582056)">Reserva Biológica do Tinguá</a>, estrada de captação de água do <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.43667&amp;materialsCitation.latitude=-22.582056" title="Search Plazi for locations around (long -43.43667/lat -22.582056)">Macuco</a>, afluente do <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.43667&amp;materialsCitation.latitude=-22.582056" title="Search Plazi for locations around (long -43.43667/lat -22.582056)">Rio Tinguá</a>, 22°34’55.4” S, 43°26’12.0” W, 15.ix. 2016, 157 m, LL Dumas, JL Nessimian &amp; JF Barbosa leg., 5 males (DZRJ) ; Nova Iguaçu, Tinguá, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.44542&amp;materialsCitation.latitude=-22.567251" title="Search Plazi for locations around (long -43.44542/lat -22.567251)">Reserva Biológica do Tinguá</a>, afluente do <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.44542&amp;materialsCitation.latitude=-22.567251" title="Search Plazi for locations around (long -43.44542/lat -22.567251)">Rio Tinguá</a>, 22°34’02.1” S, 43°26’43.5” W, 13.ix. 2016, 223 m, LL Dumas, JL Nessimian &amp; JF Barbosa leg., 1 male (DZRJ) ; Cachoeiras de Macacu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.775776&amp;materialsCitation.latitude=-22.451973" title="Search Plazi for locations around (long -43.775776/lat -22.451973)">Reserva Ecológica de Guapiaçu</a>, furo do brejo; 22°27’07.1” S, 43°46’32.8” W, 03.xii.2009, 32 m, LL Dumas &amp; APM Santos leg., 1 male (DZRJ) .</p><p>Distribution. Brazil (Minas Gerais, Rio de Janeiro, and Santa Catarina).</p></div>	https://treatment.plazi.org/id/F556879FE740FF830785FE772F494FB0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Dumas, Leandro Lourenço;Nessimian, Jorge Luiz	Dumas, Leandro Lourenço, Nessimian, Jorge Luiz (2019): New species of Helicopsyche von Siebold 1856 (Trichoptera: Helicopsychidae) from Brazil, including the redescription of Helicopsyche (Feropsyche) planorboides Machado 1957. Zootaxa 4619 (2): 231-250, DOI: 10.11646/zootaxa.4619.2.2
