identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
F55887A5FF2DFF95B6019797FBD92FE6.text	F55887A5FF2DFF95B6019797FBD92FE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptopelis anchietae (Bocage 1873)	<div><p>Leptopelis cf. anchietae (Bocage, 1873)</p> <p>Anchieta’s Tree Frog</p> <p>Material: PEM T 578 (11); SAIAB 187423 (13).</p> <p>Comment: Only tadpoles were collected and are tentatively assigned to Leptopelis anchietae based on tadpole morphology (Channing et al. 2012).</p> </div>	https://treatment.plazi.org/id/F55887A5FF2DFF95B6019797FBD92FE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF2FFF97B59B95B2FD772E7B.text	F55887A5FF2FFF97B59B95B2FD772E7B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sclerophrys funerea (Bocage 1866)	<div><p>Sclerophrys funerea (Bocage, 1866)</p> <p>Somber Toad</p> <p>Material: SAIAB 202005 (12°51’39.0”S 18°17’ 25.02”E).</p> <p>Comment: Monard (1937a) reported two specimens from the Cubango system around the villages of Kakindo [= Caiundo] and Mbale. Our specimen represents the first record from the Cuito system. We follow Ohler and Dubois (2016) in using the senior synonym Sclerophrys Tschudi, 1838 for all African bufonids recently assigned to Amietophrynus (Frost et al. 2006).</p> </div>	https://treatment.plazi.org/id/F55887A5FF2FFF97B59B95B2FD772E7B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF2FFF97B59B9734FBDC2D5E.text	F55887A5FF2FFF97B59B9734FBDC2D5E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sclerophrys gutturalis (Power 1927)	<div><p>Sclerophrys gutturalis (Power, 1927)</p> <p>Guttural Toad</p> <p>Material: PEM A10412 (2), 10777 (29), 10817 (30a), 10834 (32), 10835 (32), 10842 (29), 10974 (35), 10976 (34), 10949 (44a), and 10950 (44); SAIAB 101001 (29), 101025 (30a), 190285 (30d), and 190300 (32).</p> <p>Comment: A common and widespread species recorded from scattered localities across most of Angola (Laurent 1964; Poynton and Haacke 1993; Ruas 1996; Ceríaco et al. 2014). Old historical records referring to Bufo regularis need to be re-examined to confirm their identity and thus fully understand the distribution of the various toad species in Angola (see Ruas 1996).</p> </div>	https://treatment.plazi.org/id/F55887A5FF2FFF97B59B9734FBDC2D5E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF2FFF99B6019411FED42D4E.text	F55887A5FF2FFF99B6019411FED42D4E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sclerophrys lemairii (Boulenger 1901)	<div><p>Sclerophrys lemairii (Boulenger, 1901)</p> <p>Lemaire’s Toad</p> <p>Material: PEM A10413 (7b), 10818-19 (31), 108120- 10829 (30d), and 11527(63); PEM T 555 (7a); SAIAB 101019 (30a), 101031 (31), and 101032 (30d).</p> <p>Comments: This unusual bufonid (Fig. 7c) is adapted for living in flood plains. Previously reported for the Okavango Delta (Botswana), adjacent floodplains in Zambia (Poynton and Broadley 1985; Bittencourt-Silva 2014), and Democratic Republic of the Congo (Boulenger 1901; Laurent 1950, 1964; Schmidt and Inger 1959; Poynton and Broadley 1985). Within Angola, it was previously known from only five localities further north in Angola (i.e., Muita – Laurent 1950; Cazombo, Chimboma, Cuilo, Lake Calundo – Laurent 1964). The new records collect- ed from just north Menongue and around M’Pupa Falls represent the most southerly Angola records and are the first for the Angolan Cubango and Cuito river systems. They bridge the considerable gaps between previous Angolan material and those of the Caprivi region (Channing 2001). Bittencourt-Silva (2014) reported dynamic dichromatism in this species, and this was also noted in males collected during the current surveys (Fig. 7d). However, many features of life history (breeding, call, and tadpoles) of the species remain unknown, and additional natural history data will be presented elsewhere (Conradie in prep.).</p></div> 	https://treatment.plazi.org/id/F55887A5FF2FFF99B6019411FED42D4E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF21FF99B60195A3FC4A2F2B.text	F55887A5FF21FF99B60195A3FC4A2F2B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyperolius angolensis (Steindachner 1867)	<div><p>Hyperolius angolensis (Steindachner, 1867)</p> <p>Angolan Reed Frog</p> <p>Material: PEM A10417 (16), 10778 (29), 10779 (29), 10808 (30a), 10809 (30a), 10917 (43), 10951 (44a), 10952 (44a), 10973 (35), 11522 (45), and 11532 (62); PEM T 556 (6b), 557 (12a), 562 (28), 563(27), 566 (19), 569 (24), 697 (51), 698 (53b), 699 (54), 700 (63), 701 (57), 709 (55), and 710 (58); SAIAB 101002 (29), 101007 (32), 101015 (41b), 101016 (43), 101022 (30a), 101026 (44a), 101035 (30b), 101036 (30b), 101039 (35), 187409 (26), 187410 (10), 187414 (7c), 187422 (12a), 187432 (23), 187433 (28), 187437 (22a), 187439 (21), 188064 (19), 188073 (24a), 190262 (30d),190292‒190293 (35), 190397 (30a), and 200492 (12a).</p> <p>Comment: Frost (2015) considers this species to be part of the unresolved H. parallelus group which is widespread across Angola and adjacent countries. There are regional color patterns, with that in the study area conforming to that of H. angolensis (Schiøtz 1999).</p> </div>	https://treatment.plazi.org/id/F55887A5FF21FF99B60195A3FC4A2F2B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF21FF99B59B9643FC7F2CCE.text	F55887A5FF21FF99B59B9643FC7F2CCE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sclerophrys poweri (Hewitt 1935)	<div><p>Sclerophrys poweri (Hewitt, 1935)</p> <p>Power’s Toad</p> <p>Material: SAIAB 101000 (29).</p> <p>Comment: Only one specimen was collected on the Angola side of the Cubango River near Calai. This represents the first record for Angola, but it is expected to be more widely distributed in southern and eastern Angola. Many of the earlier records of Bufo regularis may be assignable to this species (see Ruas 1996).</p> </div>	https://treatment.plazi.org/id/F55887A5FF21FF99B59B9643FC7F2CCE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF21FF99B59B9461FDC32EAC.text	F55887A5FF21FF99B59B9461FDC32EAC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sclerophrys pusilla (Mertens 1937)	<div><p>Sclerophrys pusilla (Mertens, 1937)</p> <p>Eastern Flat-backed Toad</p> <p>Material: PEM A10283 (6b), 10284 (6b), 10288 (6a), 10289 (6a), 10292 (3), 10322 (12a), 10374 (24), 10418 (20), 10466 (6a), 10467 (22d), 10813‒10816 (30a), 11529 (62), 11531 (62), and 11598 (47); SAIAB 101021 (30a), 101006 (32), 101018 (41), 187406 (22d), 188214 (6d), 190247 (29), and 200501 (12a).</p> <p>Comment: Recorded from mostly eastern Angola (Monard 1937a; Poynton and Haacke 1993; Ruas 1996). Poynton et al. (2016) revised S. pusilla for southern and eastern populations of S. maculata, which is now restricted to West Africa. For further comment see S. gutturalis.</p> </div>	https://treatment.plazi.org/id/F55887A5FF21FF99B59B9461FDC32EAC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF20FF98B59B9531FE0B2E3C.text	F55887A5FF20FF98B59B9531FE0B2E3C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyperolius benguellensis (Bocage 1893)	<div><p>Hyperolius benguellensis (Bocage, 1893)</p> <p>Benguela Reed Frog</p> <p>Material: PEM A10370-10373 (27), 10308 (11), 10398 (22b), 10399 (22b), 10400‒1 (21), 10414 (7c), 10415‒10416 (7c), 10431‒10446 (12a),10447‒52 (5) 10810‒10812 (30a), 10919 (43), and 11521 (45); PEM T 560 (6b) and 564 (8); SAIAB 101008 (32), 101023 (30a), 101036 (30d), 101040 (35), 187416 (7b), 187419 (8), 187421 (12a), 187440 (21), 188218 (12a), 190288 (30d), 200495 (12a), and 200497 (5).</p> <p>Comment: Part of the H. nasutus super group, which has recently been re-evaluated (Channing et al. 2013). Only two species, i.e., H. nasutus and H. benguellensis, are known from southern Angola. We assign our specimens to H. benguellensis based on call differences and the protruding sharp snout. Genetic studies are ongoing.</p> </div>	https://treatment.plazi.org/id/F55887A5FF20FF98B59B9531FE0B2E3C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF20FF98B59B97CBFCEB2D9E.text	F55887A5FF20FF98B59B97CBFCEB2D9E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyperolius cinereus Monard 1937	<div><p>Hyperolius cinereus Monard, 1937</p> <p>Ashy Reed Frog</p> <p>Material: PEM A10296-9 (5), 101300‒10307 (12a), 10314‒5 (12a), 1340 (18), 10342‒3 (18), and 10350‒4 (18); PEM T 558 (11), 559 (4), and 565 (8); SAIAB 187417 (8) and 188069 (22d).</p> <p>Comment: Historically only known from three localities in south-central Angola, i.e., Caluquembe, Bimbe, and Entre Rios (Monard 1937a, Hellmich 1957b). This survey adds seven new localities that extend the distribution of the species 350 km east of the type locality (i.e., Caluquembe) and include the first records for the Cubango-Okavango river system. It is more widespread than previously known but remains endemic to Angola in regions above 1,200 m above sea level. These collections (Fig. 7a), and additional material collected around Lubango and also Lagoa Carumbo in north-eastern Angola, led to the re-description of this poorly known species and the description of a sister taxon, H. raymondi, from northern Angola (Conradie et al. 2013).</p> </div>	https://treatment.plazi.org/id/F55887A5FF20FF98B59B97CBFCEB2D9E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF20FF98B6019757FC7C2FE0.text	F55887A5FF20FF98B6019757FC7C2FE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kassina kuvangensis (Monard 1937)	<div><p>Kassina kuvangensis (Monard, 1937)</p> <p>Kuvango Kassina</p> <p>Material: PEM T 571 (10) and 572 (6b); SAIAB 187418 (8).</p> <p>Comment: Described from Vila-da-Ponte [=Kuvango] by Monard (1937a), this continued to be the only known Angolan locality, although it was subsequently recorded from five localities in adjacent Zambia (Channing 2001). No adults were obtained during our surveys, but tadpoles (Fig. 7e) collected just east of the type locality can be assigned to the species (Channing and Broadley 1992) and represent the first Angolan material since the type description.</p></div> 	https://treatment.plazi.org/id/F55887A5FF20FF98B6019757FC7C2FE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF23FF9BB59A94AFFE122F3E.text	F55887A5FF23FF9BB59A94AFFE122F3E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kassina senegalensis (Dumeril and Bibron 1841) Bubbling	<div><p>Kassina senegalensis (Duméril and Bibron, 1841)</p> <p>Bubbling Kassina</p> <p>Material: PEM A10863‒10873 (39); PEM T 573 (24), 574 (6b), and 658 (39); SAIAB 101011 (39), 101014 (40).</p> <p>Comment: Reported from numerous localities within Angola (Schmidt 1936; Monard 1937a; Laurent 1954, 1964; Poynton and Haacke 1996). The species is widely distributed in sub-Saharan Africa with geographic variation in coloration and morphology. The status of the various subspecies proposed (e.g., Laurent 1957) requires a modern appraisal.</p></div> 	https://treatment.plazi.org/id/F55887A5FF23FF9BB59A94AFFE122F3E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF23FF9AB6009710FD98292E.text	F55887A5FF23FF9AB6009710FD98292E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phrynobatrachus mababiensis FitzSimons 1932	<div><p>Phrynobatrachus mababiensis FitzSimons, 1932</p> <p>Mababe Puddle Frog</p> <p>Material: PEM A10278 (6b), 10309 (11), 13010 (11), 10316 (12a), 10384‒10397 (22d), 10402‒3(21), 10426‒10430 (19), 10453‒10462 (3), 10780‒1 (29), 10836‒10841 (29), 10849‒50 (38), 10892‒10899 (39), 10925‒10933 (43), 10943‒4 (40), and 10953‒10957 (44a). SAIAB 101041 (29), 101189 (35), 187403 (18), 187407 (22c), 187412 (6b), 187441 (21), 190301 (39), and 200494 (3).</p> <p>Comment: We assign most of our dwarf puddle frogs to P. mababiensis, although this remains a taxonomically challenging group as multiple cryptic taxa are embedded in the group (Zimkus et al. 2010). Although Channing (2001) illustrated multiple localities in Angola, it has previous only been reported from Lagoa Nutechite (Poynton and Haacke 1993).</p> </div>	https://treatment.plazi.org/id/F55887A5FF23FF9AB6009710FD98292E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF22FF9AB59B90C1FD882B2D.text	F55887A5FF22FF9AB59B90C1FD882B2D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phrynobatrachus natalensis (Smith 1849)	<div><p>Phrynobatrachus natalensis (Smith, 1849)</p> <p>Natal Puddle Frog</p> <p>Material: PEM A10286 (6b), 10287 (3), 10312 (12a), 10319-10321 (12a), 10358 (17), 10376‒10382 (22d), 10404‒10411 (21), 10423‒10425 (19), and 10463-10465 (12a); PEM T 577 (6b) and 576 (10); SAIAB 187405 (22d), 187408 (21), 187412 (6b), 188065 (19) 188065 (19), and 188215 (12a).</p> <p>Comment: Common and widespread across whole of Angola (Bocage 1866, 1879, 1895, 1897; Boulenger 1905; Ferreira 1906; Parker 1936; Schmidt 1936; Monard 1937a; Laurent 1950, 1954, 1964; Hellmich 1954; Poynton and Haacke 1993; Ruas 1996). A taxonomical difficult group in which multiple cryptic taxa are embedded (Zimkus et al. 2010).</p></div> 	https://treatment.plazi.org/id/F55887A5FF22FF9AB59B90C1FD882B2D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF22FF9AB59B92C3FF612CAB.text	F55887A5FF22FF9AB59B92C3FF612CAB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phrynobatrachus parvulus (Boulenger 1905)	<div><p>Phrynobatrachus cf. parvulus (Boulenger, 1905)</p> <p>Dwarf Puddle Frog</p> <p>Material: PEMA 10920‒10924 (43).</p> <p>Comment: Report- ed from Angola by Boulenger (1905), Monard (1937a), Parker (1936), Schmidt (1936), and Laurent (1964). It is easily confused with P. mababiensis, although usually larger and darker in coloration. We provisionally assign our specimens to this taxon due to bars on jaw being pale and not being confluent as in P. mababiensis (Channing 2001).</p> </div>	https://treatment.plazi.org/id/F55887A5FF22FF9AB59B92C3FF612CAB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF22FF9AB60194B3FC622FD5.text	F55887A5FF22FF9AB60194B3FC622FD5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ptychadena grandisonae Laurent 1954	<div><p>Ptychadena cf. grandisonae Laurent, 1954</p> <p>Grandison’s Ridged Frog</p> <p>Material: PEM A11525 (53b).</p> <p>Comment: A grass frog (Fig. 9c) collected on the edge of the upper Longa Riv- er floodplain is tentatively assigned to P. grandisonae. This species has been described from northern Angola (Muita, Laurent 1954) and reported elsewhere in Angola (Laurent 1964; Poynton and Haacke 1993; Ruas 1996). Although members of the genus Ptychadena are notoriously difficult to identify in museum collections, recent work has shown that species on a regional scale can be distinguished by quantitative morphometrics (Dehling and Sinsch 2013).</p> </div>	https://treatment.plazi.org/id/F55887A5FF22FF9AB60194B3FC622FD5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF22FF9AB59B9406FD6E2FD7.text	F55887A5FF22FF9AB59B9406FD6E2FD7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xenopus muelleri (Peters 1844)	<div><p>Xenopus muelleri (Peters, 1844)</p> <p>Tropical Platanna</p> <p>Material: PEM A10789 (29) and 10969‒70 (44a); SA- IAB 101030 (44a). Comment: The taxonomic history of the Xenopus in Angola is confused. Prior to Bocages’ description of X. petersii he referred one specimen from Dombe to X. muelleri (Bocage 1879). Later, in his description of X. petersii, he referred to the same Dombe specimen as X. petersii (Bocage 1895). By inference he thus considered only X. petersii to be present in Angola, although he recognized three varieties (Vars. A‒C, see X. poweri for fuller discussion). Specimens collected west of Calai on the Cubango River and the lower Cuando River represents the first records of X. muelleri for Angola. Both X. poweri and X. muelleri were collected in sympatry in the Cuando River. The identifications are supported by CO1 barcoding (Conradie unpublished data).</p> </div>	https://treatment.plazi.org/id/F55887A5FF22FF9AB59B9406FD6E2FD7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF22FF9AB60191C0FB532A4C.text	F55887A5FF22FF9AB60191C0FB532A4C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xenopus petersii Bocage 1895	<div><p>Xenopus petersii Bocage, 1895</p> <p>Peter’s Platanna</p> <p>Material: PEM A10293 (3), 12094‒5(5), 10276‒7 (6b), and 11526 (53a); PEM T 583 (10) and 584 (24); SAIAB 187443 (21).</p> <p>Comment: Reported from most of Angola (Bocage 1879, 1895; Boulenger 1905; Ferreria 1906; Parker 1936; Schmidt 1936; Monard 1937a; Hellmich 1957; Schmidt and Inger 1959; Laurent 1964; Loumont 1981; Poynton and Haacke 1993; Ruas 1996). Common throughout the north-eastern catchments (Fig. 9b), it was replaced in the Cuando River by X. poweri.</p> </div>	https://treatment.plazi.org/id/F55887A5FF22FF9AB60191C0FB532A4C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF22FF9AB601937FFB672DC4.text	F55887A5FF22FF9AB601937FFB672DC4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xenopus poweri Hewitt 1927	<div><p>Xenopus poweri Hewitt 1927</p> <p>Power’s Platanna</p> <p>Material: PEM A10937‒10942 (43), and 10971 (44a); SAIAB 101029 (44a).</p> <p>Comment: Based on CO1 barcoding genes, Xenopus from the Cuando River are genetically differentiated from those from the Cuito and Cubango River systems (Conradie unpublished data). Eastern populations from the Cuando River are therefore provisionally assigned to X. poweri Hewitt, 1927, and western records from the Cuito and Cubango River to X. petersii Bocage, 1895. Although Schmidt and Inger (1959) assigned X. poweri to Bocage’s “Var. B” and restricted X. petersii to Bocage “Var. A,” preliminary genetic findings do not support this, and most Angolan material should be assigned to X. petersii (Furman et al. 2015; Conradie and Evans work in progress). This includes specimens from Cubal da Ganda (Laurent 1964) and Huila (Schmidt and Inger 1959) referred by them to X. poweri. The survey records for X. poweri are thus the first for Angola. An early record from Cazombo (Laurent 1964) also falls within the newly proposed distribution of X. poweri by Furman et al. (2015), but requires verification. When Furman et al. (2015) validated the specific status of X. poweri (previously confused with X. laevis), they referred eastern X. petersii material from the Okavango system in Botswana to the species.</p> </div>	https://treatment.plazi.org/id/F55887A5FF22FF9AB601937FFB672DC4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF25FF9DB59B91C1FED92A6C.text	F55887A5FF25FF9DB59B91C1FED92A6C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ptychadena guibei Laurent 1964	<div><p>Ptychadena guibei Laurent, 1964</p> <p>Guibe’s Ridged Frog</p> <p>Material: PEM A10934‒10936 (43); SAIAB 101009 (32) and 190310 (44a).</p> <p>Comment: Previously only reported from four localities in eastern and northern Angola: Muita (Laurent 1950, 1954), Cazombo (Laurent 1964), Dundo (Laurent 1964), and Cangandala (Ceríaco et al. 2016). Common and widespread in adjacent Zambia and into the panhandle of the Okavango Delta (Channing 2001). The new material extends the distribution into extreme southeast Angola, and about 150 km up the lower Cuando River.</p></div> 	https://treatment.plazi.org/id/F55887A5FF25FF9DB59B91C1FED92A6C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF25FF9DB59B9303FD162C2B.text	F55887A5FF25FF9DB59B9303FD162C2B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ptychadena mascareniensis (Dumeril and Bibron 1841) Mascarene Ridged Frog	<div><p>Ptychadena mascareniensis (Duméril and Bibron, 1841)</p> <p>Mascarene Ridged Frog</p> <p>Material: PEM A10782‒10785 (29), 10843‒10848 (29), 10877 (39), 10945‒6 (41a), 10958 (44a), and 11535‒6 (63); SAIAB 101003 (29) and 190304 (43).</p> <p>Comment: Recent mitochondrial DNA analysis suggests that P. mascareniensis comprises a number of separate species and that the name P. mascareniensis should be restricted to the Madagascar, Seychelles and the Mascarene Islands (Vences et al. 2004). The only name available for clades from mainland Africa is P. nilotica for the Nile River system (Dehling and Sinsch 2013). Further taxonomical work is underway to determine the status of this species in Angola (Ernst work in progress). Widespread in Angola (Bocage 1867; Boulenger 1905; Ferreria 1906; Monard 1937a; Schmidt and Inger 1959; Ruas 1996).</p> </div>	https://treatment.plazi.org/id/F55887A5FF25FF9DB59B9303FD162C2B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF25FF9DB59B95C4FDCD2F09.text	F55887A5FF25FF9DB59B95C4FDCD2F09.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ptychadena mossambica (Peters 1854)	<div><p>Ptychadena cf. mossambica (Peters, 1854)</p> <p>Broad-banded Ridged Frog</p> <p>Material: PEM A10850‒10862 (38), 10874‒10876 (39), 10878‒10880 (39), and 10918 (43); SAIAB 101010 (38) and 101012 (39).</p> <p>Comment: A series of grass frogs (Fig. 7b) collected from the lower Cuando River near the village of Jamba are provisionally assigned to the Ptychadena mossambica complex. Channing (1993) described Ptychadena mapacha from Mapacha in the eastern Caprivi, Namibia, close to our new collection, noting that it was superficially similar to P. mossambica. Morphologically they differ in having a continuous paravertebral fold from head to midbody in P. mossambica, that is interrupted in P. mapacha. Unfortunately, the current series was collected during the dry season and no vocalization was obtained for comparison with that recorded for P. mapacha (Channing 1993). Genetic analysis is underway on the P. mossambica complex to determine the specific identity of the new collection. Neither P. mossambica or P. mapacha, which are both known from the Caprivi area of Namibia, have been recorded from Angola.</p> </div>	https://treatment.plazi.org/id/F55887A5FF25FF9DB59B95C4FDCD2F09	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF25FF9DB60191C1FB12298C.text	F55887A5FF25FF9DB60191C1FB12298C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ptychadena oxyrhynchus (Smith 1849)	<div><p>Ptychadena oxyrhynchus (Smith, 1849)</p> <p>Sharp-nosed Ridged Frog</p> <p>Material: PEM A10282 (6b), 10881‒10891 (39), 10947 (41a), and 10959‒10963 (44a); SAIAB 101013 (39), 188217, 190306 (43), and 190309 (44a).</p> <p>Comment: Common and widespread in Angola and adjacent countries (Bocage 1866, 1895, 1897; Boulenger 1905; Ferreira 1906; Parker 1936; Schmidt 1936; Monard 1937a; Laurent 1950, 1954; Hellmich 1957).</p></div> 	https://treatment.plazi.org/id/F55887A5FF25FF9DB60191C1FB12298C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF25FF9DB60190A3FB482B2B.text	F55887A5FF25FF9DB60190A3FB482B2B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ptychadena subpunctata (Bocage 1866)	<div><p>Ptychadena subpunctata (Bocage, 1866)</p> <p>Spotted Ridged Frog</p> <p>Material: PEM A10359‒10368 (27), 10794‒10807 (30a), 10948 (41a), 10900‒10916 (43), 10964‒10967 (44a), 11528 (63), and 11614‒5 (56); SAIAB 101005 (33a), 101024 (30a), 101028 (43), 101033 (6b), 190260 (30f), 190307 (44a), 190350 (35), and 200502 (27).</p> <p>Comment: Previously known from central and northern Angola (Bocage 1866; Schmidt 1936; Mertens 1938; Laurent 1964; Ruas 1996), the new records fill the large gap in the species’ range in southeast Angola.</p></div> 	https://treatment.plazi.org/id/F55887A5FF25FF9DB60190A3FB482B2B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF25FF9DB60192C4FBAA2D49.text	F55887A5FF25FF9DB60192C4FBAA2D49.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ptychadena taenioscelis Laurent 1954	<div><p>Ptychadena taenioscelis Laurent, 1954</p> <p>Small Ridged Frog</p> <p>Material: PEM A10290-1 (3), 10279‒81 (6b), 10285 (6b), 10419‒22 (19), 10830‒10833 (30d), 10972 (44b), 10975 (35), 11530 11523‒4 (43b), (62), and 11610‒1 (47); PEM T 580‒1 (10); SAIAB 101017 (43), 101034 (30d), and 188217 (6b).</p> <p>Comment: The species has been previously recorded from northern (Laurent 1964) and central Angola (Poynton and Haacke 1993), and the new material extends its range into the grasslands of south-eastern Angola (Fig. 9a).</p></div> 	https://treatment.plazi.org/id/F55887A5FF25FF9DB60192C4FBAA2D49	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF25FF9DB6019466FC502E17.text	F55887A5FF25FF9DB6019466FC502E17.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ptychadena uzungwensis (Loveridge 1932)	<div><p>Ptychadena uzungwensis (Loveridge, 1932)</p> <p>Uzungwe Ridged Frog</p> <p>Material: PEM A10369 (27).</p> <p>Comment: In Angola it was previously restricted to the east and central region (Loveridge 1932; Monard 1937a; Laurent 1954, 1964; Poynton and Haacke 1993; Ruas 1996), but the current specimen extends the range southeast into Kuando-Kubango Province.</p></div> 	https://treatment.plazi.org/id/F55887A5FF25FF9DB6019466FC502E17	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF25FF9CB60197EAFD722A2E.text	F55887A5FF25FF9CB60197EAFD722A2E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amietia angolensis (Bocage 1866)	<div><p>Amietia angolensis (Bocage, 1866)</p> <p>Angolan River Frog</p> <p>Material: PEM A10311 (11), 10313 (12a), 10317‒10318 (12a), and 10375 (22d); PEM T 552 (11) and 553 (4); SAIAB 187112 (12a), 187402 (18), 187411 (14), 187413 (3), 187424 (13), 200499 (18), 200500 (12a), and 200503 (13).</p> <p>Comment: Common and widespread in Angola (Bocage 1866, 1895; Boulenger 1905; Ferreira 1906; Parker 1936; Schmidt 1936; Mertens 1938; Monard 1937a; Laurent 1950, 1954, 1964; Hellmich 1954; Poynton and Haacke 1993; Ruas 1996). A recent revision (Channing and Baptista 2013) restricted A. angolensis to Angola, albeit that few Angolan samples were available for analysis. Subsequently, Larson et al. (2016) identified several well-supported cryptic lineages of river frogs previously assigned to Amietia angolensis in the Albertine Rift region. It is, therefore, possible that cryptic taxa of A. angolensis also occur in Angola, and that A. angolensis may also extend into adjacent western Zambia.</p> </div>	https://treatment.plazi.org/id/F55887A5FF25FF9CB60197EAFD722A2E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF24FF9CB59B9524FE9E2DE8.text	F55887A5FF24FF9CB59B9524FE9E2DE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amnirana darlingi (Boulenger 1902)	<div><p>Amnirana darlingi (Boulenger, 1902)</p> <p>Darling’s White-lipped Frog</p> <p>Material: PEM T 711 (55); SAIAB 187436 (22a).</p> <p>Comment: Only tadpoles were obtained. Previously recorded from central and southern Angola (Monard 1937a; Laurent 1964, Schmidt 1936; Schmidt and Inger 1959; Ruas 1996). Oliver et al. (2015) showed that African species recently assigned to Hylarana are best placed in the genus Amnirana.</p> </div>	https://treatment.plazi.org/id/F55887A5FF24FF9CB59B9524FE9E2DE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF24FF9CB59B97F6FB54288E.text	F55887A5FF24FF9CB59B97F6FB54288E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Crotaphopeltis hotamboeia (Laurenti 1768)	<div><p>Crotaphopeltis hotamboeia (Laurenti, 1768)</p> <p>White-lipped Snake</p> <p>Material: PEM R 20018 (12b); ANG (no number, 6a).</p> <p>Comment: Common and widespread species in Angola (Bocage 1895; Boulenger 1905; Ferreira 1906; Branch and McCartney 1992; Laurent 1950, 1964; Parker 1936). This species is widespread in sub-Saharan Africa. A phylogenetic study is currently underway to investigation different populations (Tolley pers. comm.).</p></div> 	https://treatment.plazi.org/id/F55887A5FF24FF9CB59B97F6FB54288E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF24FF9CB60191A3FAA029AB.text	F55887A5FF24FF9CB60191A3FAA029AB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philothamnus hoplogaster (Gunther 1863)	<div><p>Philothamnus hoplogaster (Günther, 1863)</p> <p>Green Water Snake</p> <p>Material: photograph record (James Kydd: 14°08’56.0”S 18°48’36.0”E).</p> <p>Comment: This is the first record for the Cuito River system. Only recorded from a handful of other locations in Angola (Bocage 1887; Monard 1937b; Laurent 1964; Thys van den Audenaerde 1967).</p></div> 	https://treatment.plazi.org/id/F55887A5FF24FF9CB60191A3FAA029AB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF24FF9CB6019344FADC2BA9.text	F55887A5FF24FF9CB6019344FADC2BA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philothamnus ornatus Bocage 1872	<div><p>Philothamnus ornatus Bocage, 1872</p> <p>Ornate Green Snake</p> <p>Material: PEM R 20013 (24).</p> <p>Comment: Philothamnus ornata was described by Bocage (1872) from two specimens collected at Huila. Although subsequently recorded from Zambia and Zimbabwe (Broadley et al. 2003), it remained known in Angola from only few additional collections, i.e., Bela Vista (Hellmich 1957), Benguela (Boulenger 1905), Bié (Boulenger 1905), Caconda (Bocage 1895), Caluquembe (Monard 1937b), Cutatu (Monard 1937b). Chimporo (Boulenger 1905), Cunene River (Bocage 1895), and Huambo (Bogert 1940). Our collection fills in the gap between the records from western Angola and the Zambian and Zimbabwean populations.</p> </div>	https://treatment.plazi.org/id/F55887A5FF24FF9CB6019344FADC2BA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF24FF9CB6019546FCD82EB7.text	F55887A5FF24FF9CB6019546FCD82EB7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thelotornis capensis subsp. oatesi (Gunther 1881)	<div><p>Thelotornis capensis oatesi (Günther, 1881)</p> <p>Oates’ Twig Snake</p> <p>Material: PEM R 21484 (52).</p> <p>Comment: This subspecies has a large range from southern Angola to Malawi and western Mozambique (Broadley and Wallach 2002), but with relatively few Angolan records (Bocage 1895; Parker 1936; Monard 1937; Bogert 1940; Laurent 1954, 1964; Thys van den Audenaerde 1967). When reviewing the genus, Broadley (1979) noted few specimens from the southern and eastern regions of the country, while Laurent recorded sympatry between T. kirklandi and T. c. oatseii at Dundo in northeast Angola. The new record is only the second for the south-eastern Angola, following Monard’s (1937b) record from Vila-da-Ponte [=Cuvangu]. The present specimen helps fill the large gap in records for the south-east of Angola. Although Broadley (2001) distinguished T. c. oatesi by coloration of the top of the head and ventral number (&gt;160) these features are variable. The status of T. c. oatesi as a valid taxon was not resolved by a morphological and genetic analysis of the Dispholidini (Eimermacher 2012) and remains problematic.</p> </div>	https://treatment.plazi.org/id/F55887A5FF24FF9CB6019546FCD82EB7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF24FF9CB59B93C0FEF42C4C.text	F55887A5FF24FF9CB59B93C0FEF42C4C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tomopterna cryptotis (Boulenger 1907)	<div><p>Tomopterna cf. cryptotis (Boulenger, 1907)</p> <p>Cryptic Sand Frog</p> <p>Material: PEM A10786‒8 (29), 10790‒3 (29) and 10968 (44a); SAIAB 101004 (29) and 187435 (22c).</p> <p>Comment: Recorded from only a handful of localities in Angola (Boulenger 1907; Poynton and Haacke 1993). Species delineation in the genus is problematic and often dependent upon vocalization and chromosome number. We assign our material conservatively as specimens may represent either T. tandyi, T. cryptotis or even further cryptic diversity.</p> </div>	https://treatment.plazi.org/id/F55887A5FF24FF9CB59B93C0FEF42C4C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF24FF81B6019636FEF62920.text	F55887A5FF24FF81B6019636FEF62920.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limnophis bangweolicus (Mertens 1936)	<div><p>Limnophis bangweolicus (Mertens, 1936)</p> <p>Bangweola Swamp Snake</p> <p>Material: PEM R20489 (44a).</p> <p>Comment: A specimen (Fig. 8b) collected in the floodplain of the lower Cuando River represents the second record for Angola. The previous record was 700 km north at Calundo, Moxico Province (Laurent 1964). The sister species Limnophis bicolor is more widespread in western, central, and northern Angola.</p> </div>	https://treatment.plazi.org/id/F55887A5FF24FF81B6019636FEF62920	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF39FF81B59B9608FAC12B00.text	F55887A5FF39FF81B59B9608FAC12B00.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Boaedon angolensis Bocage 1895	<div><p>Boaedon cf. angolensis Bocage, 1895</p> <p>Angola House Snake</p> <p>Material: PEM R 21846 (50).</p> <p>Comment: This unusual pale yellow-green specimen (Fig. 9g) is referable to the Boaedon lineatus-fuliginosus complex. A specimen of similar coloration was photographed in the upper Cuito River (James Kydd: 13°09’05.0”S 18°28’50.0”E). Kelly et al. (2011) demonstrated complex cryptic diversity with numerous deep lineages in house snakes of the B. lineatus-fuliginosus complex. On morphology, Hughes (1997) referred specimens from East and South Africa to B. capensis Duméril and Bibron, 1854, and this has been adopted in the most subsequent literature (see Uetz 2016). However, southern African specimens fall into a number of well-defined clades (Kelly et al. 2011) that cannot all be accommodated under B. capensis. The name Boaedon lineatus var. angolensis Bocage, 1895 has not been considered for western populations. Laurent (1956) noted that populations from Angola to Katanga could be distinguished morphologically, but took no taxonomic action and the availability of Bocage’s angolensis was not discussed. Unfortunately, neither a type or type locality was nominated for the taxon, and Bocage’s material was also lost in the fire that destroyed the museum in Lisbon.</p> </div>	https://treatment.plazi.org/id/F55887A5FF39FF81B59B9608FAC12B00	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF39FF81B59B9562FF602D2B.text	F55887A5FF39FF81B59B9562FF602D2B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dendroaspis polylepis subsp. polylepis Gunther 1864	<div><p>Dendroaspis polylepis polylepis Günther, 1864</p> <p>Black Mamba</p> <p>Material: Observation only (37).</p> <p>Comment: One specimen was observed crossing the road and disappearing into a tree. Only a handful of records exist for Angola (Schmidt 1933; Monard 1937b; Bogert 1940; Baynham 2010), but it is believed to be more widespread in the southern and central regions of Angola (Branch et al. in prep.).</p></div> 	https://treatment.plazi.org/id/F55887A5FF39FF81B59B9562FF602D2B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF39FF81B59B94C7FE0C2EA9.text	F55887A5FF39FF81B59B94C7FE0C2EA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Naja (Afronaja) mossambica Peters 1854	<div><p>Naja (Afronaja) mossambica Peters, 1854</p> <p>Mozambique Spitting Cobra</p> <p>Material: PEM R20499 (30a) and 20500 (44b).</p> <p>Comment: The new records represent the second (M’Pupa Falls) and third (near village Sashae) for Naja mossambica in Angola. The only other published Angolan record (Maconjo, Broadley 1974) occurs in south-west Angola, although other western records are known (Branch et al. in prep.). This species is expected to be more widely distributed in southern Angola.</p> </div>	https://treatment.plazi.org/id/F55887A5FF39FF81B59B94C7FE0C2EA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF39FF81B59B90DEFEBB2B8E.text	F55887A5FF39FF81B59B90DEFEBB2B8E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Natriciteres olivacea (Peters 1854)	<div><p>Natriciteres olivacea (Peters, 1854)</p> <p>Olive Marsh Snake</p> <p>Material: PEM R 20501 (44a).</p> <p>Comment: A specimen (Fig. 8a) collected in the floodplains of the lower Cuando River represents a southern record of this species for Angola, the other records occurring much further north, i.e., Dondo (Hellmich (1957), Dundo and Muita (Laurent 1950, 1954), and Pungo Andongo (Bocage 1895; Boulenger 1905). Other records from Bela Vista (Hellmich 1956) and Malanje (Bocage 1895) have subsequently been referred to N. bipostocularis (Broadley 1966). There is little variation in scalation among Natriciteres species, and also no molecular phylogeny for the genus. However, the Cuando specimen, despite its relative proximity to the Angolan N. bipostocularis records, still possesses three postoculars and is conservatively referred to N. olivacea.</p> </div>	https://treatment.plazi.org/id/F55887A5FF39FF81B59B90DEFEBB2B8E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF39FF81B601923FFB942C8F.text	F55887A5FF39FF81B601923FFB942C8F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Psammophis mossambicus Peters 1882	<div><p>Psammophis mossambicus Peters, 1882</p> <p>Olive Grass Snake</p> <p>Material: PEM R 20024 (16b) and 21827 (48); ANG (23a).</p> <p>Comment: Species boundaries in the Psammophis sibilans complex remain problematic (Kelly et al. 2008). Our specimens accord with P. mossambicus (Broadley 2002). Historical Angolan records assigned to P. brevirostris, P. leopardinus, P. mossambicus, P. phillipsi, and P. sibilans need to be carefully re-examined, and genetically assessed.</p> </div>	https://treatment.plazi.org/id/F55887A5FF39FF81B601923FFB942C8F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF39FF81B60195A0FC252E6D.text	F55887A5FF39FF81B60195A0FC252E6D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Psammophis subtaeniatus Peters 1882	<div><p>Psammophis subtaeniatus Peters, 1882</p> <p>Yellow-bellied Sand Snake</p> <p>Material: PEM R20506 (44a).</p> <p>Comment: Recorded by Bocage (1896) and Monard (1937b). Broadley (2002) mapped the known Angola material which clusters in the south-west, as far north as Benguela. The current specimen, the first from the Cuito drainage, fills the large gap between populations in Caprivi region of Namibia and Zimbabwe.</p> </div>	https://treatment.plazi.org/id/F55887A5FF39FF81B60195A0FC252E6D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF39FF81B6019702FB832F2B.text	F55887A5FF39FF81B6019702FB832F2B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Psammophylax acutus (Gunther 1888)	<div><p>Psammophylax acutus (Günther, 1888)</p> <p>Sharp-nosed Skaapsteker</p> <p>Material: PEM R 20006 (23) and 21485 (51).</p> <p>Comment: Common and widespread species in Angola (Bocage 1895; Bogert 1940; Ferreira 1906; Laurent 1964 and Monard 1937b). The two from Longa River are the first for south-east Angola. Transferred from Rhamphiophis by Kelly et al. (2008).</p> </div>	https://treatment.plazi.org/id/F55887A5FF39FF81B6019702FB832F2B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF38FF80B601958BFBC12DF2.text	F55887A5FF38FF80B601958BFBC12DF2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Agama aculeata Merrem 1820	<div><p>Agama aculeata Merrem, 1820</p> <p>Ground Agama</p> <p>Material: PEM R 20017 (15).</p> <p>Comment: Recorded from central and southern Angola (Bocage 1895; Boulenger 1905; Monard 1937b; Laurent 1964), but is poorly documented from the south-east.</p></div> 	https://treatment.plazi.org/id/F55887A5FF38FF80B601958BFBC12DF2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF38FF80B59B972CFD9D2F11.text	F55887A5FF38FF80B59B972CFD9D2F11.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bitis arietans (Merrem 1820)	<div><p>Bitis arietans (Merrem, 1820)</p> <p>Puff Adder</p> <p>Material: PEM R 20021 (9) and 21493 (61); road kill, not kept (38). Site: 9, 38, 61.</p> <p>Comment: Widespread species recorded throughout Angola, but recorded only three times during the current surveys.</p></div> 	https://treatment.plazi.org/id/F55887A5FF38FF80B59B972CFD9D2F11	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF38FF80B60191C0FCD82C6C.text	F55887A5FF38FF80B60191C0FCD82C6C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Causus rasmusseni Broadley 2014	<div><p>Causus cf. rasmusseni Broadley, 2014</p> <p>Rasmussen’s Night Adder</p> <p>Material: PEM R 21488 (57).</p> <p>Comment: An adult female from riverine habitat on the Rio Longa, and with a diffuse color pattern (Fig. 9e, f) and only 132 ventrals. It conforms in coloration and low ventral count with the newly-described C. rasmusseni (Broadley 2014) but has a lower subcaudal count (31) than that of the only female (39) in the type series from north-west Zambia. As noted by Rasmussen (2005), the status of night adders in Angola has been confused with many “ rhombeatus ” of earlier authors now assigned to other species, e.g., C. maculatus and C. bilineatus (Piri Dembos; Hellmich 1957a) and C. resimus (Libolo-Luali, Hellmich 1957b). Ventral counts are typically higher (143‒145) for blotched females of C. rhombeatus from the eastern half of Angola (Laurent 1964). A molecular phylogeny of Causus (Tolley et al. in prep.) shows that the Rio Longa specimen is only weakly differentiated from South African C. rhombeatus. However, topotypic material is not available for C. rasmusseni and its taxonomic status thus remains equivocal. If the specific validity of Rasmussen’s night adder is confirmed the Rio Longa specimens would be the first Angolan record for the species, and a southern range extension of approximately 740 km from the Zambian series (Broadley 2014). If, however, it is only a plain color morph of C. rhombeatus it would be the first record for south-east Angola.</p> </div>	https://treatment.plazi.org/id/F55887A5FF38FF80B60191C0FCD82C6C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF38FF80B59B9182FDBD2B28.text	F55887A5FF38FF80B59B9182FDBD2B28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptotyphlops scutifrons (Peters 1854)	<div><p>Leptotyphlops scutifrons complex (Peters, 1854)</p> <p>Peter’s Thread Snake</p> <p>Material: PEM R 20020 (13).</p> <p>Comment: Many earlier records from Angola attributed to L. scutifrons (e.g., Bocage 1895, 186; Boulenger 1905; Werner 1917) were referred to L. (= Namibiana) latifrons (Broadley and Broadley 1999). Monard (1937b) referred seven Angolan specimens, including one from Cuvangu, to this species. These were not discussed by Broadley and Broadley (1999), and although referred to L. scutifrons by Wallach et al. (2014) their identity requires confirmation. Our specimen is the second confirmed locality from Angola, following Broadley and Broadley’s (1999) record of new material from Chitau. We stress, however, that the status of the limited Angolan material, which involves large disjunctions between the known records and from the type locality (Tete, Mozambique), awaits further study, especially as the L. scutifrons complex has been shown to comprise numerous deep lineages that represent cryptic species (Adalsteinsson et al. 2009).</p> </div>	https://treatment.plazi.org/id/F55887A5FF38FF80B59B9182FDBD2B28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF38FF80B59B9289FE942E54.text	F55887A5FF38FF80B59B9289FE942E54.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Python natalensis Smith 1840	<div><p>Python natalensis Smith, 1840</p> <p>Southern Rock Python</p> <p>Material: SARCA 153808 (1), 153811 (30a).</p> <p>Comment: Large pythons are known from multiple localities in Angola (Bocage 1895; Monard 1937b; Bogert 1940; Laurent 1954, 1964; Thys van den Audenaerde 1967; Ceríaco et al. 2014, 2016). Following the revision of the Python sebae complex, Broadley (1984) revived P. natalensis for pythons from East and southern Africa. However, scattered populations of P. sebae have subsequently been recorded in East Africa (Spawls et al. 2002), and the status of pythons in Angola, particularly in the northern Congo drainage, should be re-evaluated. Python natalensis is known from adjacent Namibia and Zambia, and pythons from southern Angola, including our material, are referable to P. natalensis based on head coloration and scalation.</p> </div>	https://treatment.plazi.org/id/F55887A5FF38FF80B59B9289FE942E54	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF38FF83B601974EFEA929CE.text	F55887A5FF38FF83B601974EFEA929CE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zygaspis quadrifrons (Peters 1862) Kalahari	<div><p>Zygaspis quadrifrons (Peters, 1862)</p> <p>Kalahari Round-headed Worm Lizard</p> <p>Material: PEM R 20524 (41b), 20525 (44b), 21491 (59), and 21492 (60).</p> <p>Comment: Along with southern Africa, Angola has a very rich amphisbaenian fauna and 11 species are known, with representatives of three genera (Dalophia, Monopeltis, and Zygaspis). Zygaspis quadrifrons was recorded from Alto Chipata, north-west Angola, by Laurent (1964), but this has subsequently been referred to the new species Z. nigra Broadley and Gans, 1969. The only previous Angolan records are from Kakindo [= Caiundo], Kuando-Kubango Province and Chimporo [=Techimpolo], Cunene Province by Monard (1937b). The new collections (Fig. 8d) from the Cuito and lower Cuando River represent the first records for both river systems and double the total number of Angolan locations.</p> </div>	https://treatment.plazi.org/id/F55887A5FF38FF83B601974EFEA929CE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3BFF83B59B90A3FDD72B0B.text	F55887A5FF3BFF83B59B90A3FDD72B0B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamaeleo dilepis Leach 1819	<div><p>Chamaeleo dilepis Leach, 1819</p> <p>Flap-necked Chameleon</p> <p>Material: SARCA 153813 (30a).</p> <p>Comment: A common and widespread species in Angola (Bocage 1895; Boulenger 1905; Schmidt 1933; Parker 1936; Monard 1937b; Laurent 1950, 1954, 1964; Ceríaco et al. 2016). Although Tilbury’s (2010) recent map of the species shows it limited in Angola to the western and central regions, it is probably more widespread. The gap in south-east Angola is partly filled by the lower Cuito specimen.</p></div> 	https://treatment.plazi.org/id/F55887A5FF3BFF83B59B90A3FDD72B0B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3BFF83B59B92E6FEE92D17.text	F55887A5FF3BFF83B59B92E6FEE92D17.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chondrodactylus pulitzerae (Schmidt 1933) Angolan	<div><p>Chondrodactylus cf. pulitzerae (Schmidt, 1933)</p> <p>Angolan Thick-toed Gecko</p> <p>Material: PEM R 20477‒20479 (30a), 21494‒7 (62).</p> <p>Comment: These geckos are widespread in the southern and western parts of Angola, reaching as far north as Capanda Dam (Ceríaco et al. 2014). The Chondrodactylus turneri complex, of which pulitzerae was recently a synonym, has been shown to have a number of distinct lineages (Ceríaco et al. 2014), with the revival of C. pulitzerae for most Angolan material. Species allocation of our south-east Angolan material is provisional pending molecular analysis.</p> </div>	https://treatment.plazi.org/id/F55887A5FF3BFF83B59B92E6FEE92D17	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3BFF83B60190A3FB952B2B.text	F55887A5FF3BFF83B60190A3FB952B2B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gerrhosaurus nigrolineatus Hallowell 1854	<div><p>Gerrhosaurus cf. nigrolineatus Hallowell, 1854</p> <p>Black-lined Plated Lizard</p> <p>Material: PEM R 20004 (6a), 20481‒2 (30a), 20483‒4 (35), and 21826 (46).</p> <p>Comment: Bates et al. (2013) revived G. intermedius for eastern populations of G. nigrolineatus, raised G. bulsii to specific status, and noted the unresolved status of G. multilineatus in north-central Angola. The status of these populations and others from western Angola and relationships within the G. nigrolineatus complex in Angola is currently under investigation (M. Bates, pers. comm.).</p> </div>	https://treatment.plazi.org/id/F55887A5FF3BFF83B60190A3FB952B2B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3BFF83B6019748FC7B2EB5.text	F55887A5FF3BFF83B6019748FC7B2EB5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heliobolus lugubris (Smith 1838)	<div><p>Heliobolus lugubris (Smith, 1838)</p> <p>Bushveld Lizard</p> <p>Material: PEM R 21500 (63).</p> <p>Comment: This represents the first record for the south-eastern part of the country. Previously only recorded in Angola from the south-west (Monard 1937b).</p></div> 	https://treatment.plazi.org/id/F55887A5FF3BFF83B6019748FC7B2EB5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3BFF83B59B9428FC6B29CE.text	F55887A5FF3BFF83B59B9428FC6B29CE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lygodactylus bradfieldi Hewitt 1932	<div><p>Lygodactylus bradfieldi Hewitt, 1932</p> <p>Bradfield’s Dwarf Day Gecko</p> <p>Material: PEM R 20492 (32), and 20490‒1 (35). Comment: Pasteur (1965) revived Hewitt’s Namibian species after FitzSimons (1943) had treated it as a subspecies of L. capensis, and Loveridge (1944) had even placed it in the synonymy of the same species. Jacobsen (2011) affirmed features that distinguished the species and confirmed its presence in sympatry with L. capensis in western Limpopo Province, South Africa. Our material conforms to the features highlighted by Jacobsen (2011), and we therefore consider the survey records to represent the first for the Cuito drainage, and possibly the first confirmed records for Angola. Pasteur’s (1965) map did not show L. bradfieldi extending into southern Angola, and although he plotted six localities for L. capensis in south-west Angola, presumably based on records in Bocage (1895) and Monard (1937b), he did not discuss voucher material for any of his localities. Our specimens fill the large gap for the L. capensis complex in south-east Angola. Cryptic diversity within dwarf geckos is now well established (e.g., Travers et al. 2014), and the status of the isolated population of L. capensis in southwest Angola (see above) and its relationship to L. bradfieldi invites further study.</p> </div>	https://treatment.plazi.org/id/F55887A5FF3BFF83B59B9428FC6B29CE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3BFF83B60192C4FC2E2DE9.text	F55887A5FF3BFF83B60192C4FC2E2DE9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tetradactylus ellenbergeri (Angel 1922)	<div><p>Tetradactylus ellenbergeri (Angel, 1922)</p> <p>Ellenberger’s Seps</p> <p>Material: PEM R 20010 (23).</p> <p>Comment: A specimen (Fig. 8c) collected along a dammed section of the Luassingua River, represents only the fifth and most southern record of this species for Angola. Wagner et al. (2012) only listed one record from Angola, overlooking other historical records, e.g., Calundo as T. e. ellenbergeri (Laurent 1964), Dundo as T. e. boulengeri (Laurent 1964), Cuando as T. africanus (Bocage 1895) and Tyiumbwe as T. lundensis (Monard 1937b). Tetradactylus africanus is now restricted to coastal regions of KwaZulu-Natal, South Africa, and adjacent southern Mozambique, while Laurent (1964) treated T. lundensis as a synonym of T. ellenbergeri boulengeri. This is the first record for south-east Angola.</p> </div>	https://treatment.plazi.org/id/F55887A5FF3BFF83B60192C4FC2E2DE9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3AFF82B59B9484FE6B2FC9.text	F55887A5FF3AFF82B59B9484FE6B2FC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acontias kgalagadi subsp. kgalagadi Lamb, Biswas and Bauer 2010	<div><p>Acontias kgalagadi kgalagadi Lamb, Biswas and Bauer, 2010</p> <p>Kalahari Legless Skink</p> <p>Material: PEM R 20474 (44a). Comment: Acontias k. kgalagadi (previously Typhlosaurus lineatus lineatus – see Lamb et al. 2010) occurs mostly in north-eastern South Africa, extending to northern Namibia and Botswana. Although Broadley (1968) reported a single record of A. jappi from Gago Coutinho district in the south-east of Angola, there are no records of A. k. kgalagadi from the country. This is also the first record (Fig. 8f) of this species in Angola and north of the Okavango River (Conradie and Bourquin 2013).</p> </div>	https://treatment.plazi.org/id/F55887A5FF3AFF82B59B9484FE6B2FC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3AFF82B60191C1FC272B6C.text	F55887A5FF3AFF82B60191C1FC272B6C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lubuya ivensii (Bocage 1879)	<div><p>Lubuya ivensii (Bocage, 1879)</p> <p>Iven’s Meadow Skink</p> <p>Material: PEM R 20005 (7c).</p> <p>Comment: A rarely sampled semi-aquatic skink, known from eight localities in Angola, single localities in adjacent Zambia and Democratic Republic of the Congo (Branch and Haagner 1993; Wagner et al. 2012), and a recent isolated record from Lavushi Manda National Park, Zambia (Broadley and Willems 2015). It was previously listed as a near-endemic species to Angola, and can be considered as such pending genetic assessment of the Lavushi Manda population. The new record (Fig. 8g) from the floodplains just north of Menongue is the first record for the Cubango-Okavango system (previously only known to occur in the headwaters of the Zambezi, Kwanza, and Congo drainage systems). Horton (1972) placed Mabuya ivensii into a monotypic genus, Lubuya, that was recently revived due to its basal position in a phylogeny of mabuyine skinks (Metallinou et al. 2016) and it is unique matrotrophic reproduction.</p> </div>	https://treatment.plazi.org/id/F55887A5FF3AFF82B60191C1FC272B6C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3AFF82B59B9541FEC12D2D.text	F55887A5FF3AFF82B59B9541FEC12D2D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Meroles squamulosus (Peters 1854)	<div><p>Meroles squamulosus (Peters, 1854)</p> <p>Rough-scaled Desert Lizard</p> <p>Material: PEM R 20493‒4 (30e) and 20495 (35).</p> <p>Comment: Previously recorded from Angola only from Kapelongo [=Capelongo], Huila Province (Monard 1937b), and Pereira de Eça [=Ondijiva]. Current material includes the third and fourth records for Angola. Previously generically assigned to Ichnotropis, but recently shown to be nested within Meroles (Edwards et al. 2013; Engleder et al. 2013).</p> </div>	https://treatment.plazi.org/id/F55887A5FF3AFF82B59B9541FEC12D2D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3AFF82B6019203FAAC2C0B.text	F55887A5FF3AFF82B6019203FAAC2C0B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mochlus sundevalli (Smith 1849)	<div><p>Mochlus sundevalli (Smith, 1849)</p> <p>Sundevall’s Writhing Skink</p> <p>Material: PEM R 20496 (39), 20497 (42), and 20498 (44a).</p> <p>Comment: Recorded by Bocage (1895), Laurent (1964) and Monard 1937b. Previously placed in the genera Lygosoma Boulenger, 1895 and Riopa Smith, 1937, both of which are now restricted to West Africa.</p> </div>	https://treatment.plazi.org/id/F55887A5FF3AFF82B6019203FAAC2C0B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3AFF82B6019524FB782E09.text	F55887A5FF3AFF82B6019524FB782E09.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Panaspis maculicollis Jacobsen and Broadley 2000	<div><p>Panaspis maculicollis Jacobsen and Broadley, 2000.</p> <p>Speckle-lipped Snake-eyed Skink</p> <p>Material: PEM R 20007 (43).</p> <p>Comment: Only a few records of Panaspis exist for Angola (P. breviceps – Parker 1936; P. cabindae – Parker 1936; Bocage 1895; Ferreira 1906). A recent phylogeny of snake-eyed skinks (Medina et al. 2016) placed Afroablepharus in the synonymy of Panaspis and showed deep divergences for the basal taxa P. breviceps and P. togoensis. The P. wahlbergi-maculicollis complex was also shown to contain numerous cryptic taxa. When describing P. maculicollis Jacobsen and Broadley (2000) recorded no specimens for Angola, and the Cuando specimen of confirmed genetic mono- phyly, represents the first for the country.</p> </div>	https://treatment.plazi.org/id/F55887A5FF3AFF82B6019524FB782E09	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3AFF85B6019726FD592BAE.text	F55887A5FF3AFF85B6019726FD592BAE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trachylepis spilogaster (Peters 1882)	<div><p>Trachylepis cf. spilogaster (Peters, 1882)</p> <p>Kalahari Tree Skink</p> <p>Material: PEM R 20007 (20), 20016 (25), 20019 (12b), 20517 (30a), 20518 (35), 20519 (32), 21483 (49), 21486‒7 (57), 21498‒9 (62), 21828‒21840 (47), and 21842 (6a).</p> <p>Comment: This form of the T. striata complex was elevated to specific status by Broadley (1969). First recorded from southern Angola by Laurent (1964), based on a single male from Serra do Moco, Luimbale, Huambo. He noted the great disjunction between this specimen and the species’ known distribution and cautioned that further investigation was necessary. His assignment to spilogaster appeared to be influenced by the tricarinate midbody scale count (34), supranasal and prefrontal condition, and speckled ventrum. The new material basically conforms to the descriptions in Laurent (1964) and Broadley (1969, 1977, 2000), in having supranasals in broad contact and 35‒37 tricarinate midbody scale rows (31‒40, mean 34.7; Broadley 1977). Howev- er, the prefrontal condition varies from broad contact to widely separated (usually separated in spilogaster), the ventrum is never speckled although the throat is in adult males, and adult males usually have a red-brown middorsal band (Fig. 9h) that is not recorded in other populations (Laurent 1964; Broadley 1969, 1977, 2000). The Cuito population also has different habits to typical spilogaster, being rarely found on trees although common on the ground, in rock piles, and on village houses. Its relationship with T. angolensis (Monard 1937b), and indeed that taxon’s status with respect to earlier names (Euprepes angolensis Bocage, 1872 and Sepsina angolensis Bocage, 1866), remains problematic (see discussion in Boulenger, 1887). The present series is the subject of a more detailed morphological and genetic assessment.</p> </div>	https://treatment.plazi.org/id/F55887A5FF3AFF85B6019726FD592BAE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3DFF85B6019448FC562E69.text	F55887A5FF3DFF85B6019448FC562E69.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Crocodylus niloticus Laurenti 1768	<div><p>Crocodylus niloticus Laurenti, 1768</p> <p>Nile Crocodile</p> <p>Material: SARCA 153812 (29) and other observational records (35, 43, 44a, 45).</p> <p>Comment: During the 2015 Okavango Expedition and Expedition Okavango evidence was found of crocodiles breeding in the Cuito Riv- er (Shacks 2015), and adults and juveniles were observed at the various sites.</p> </div>	https://treatment.plazi.org/id/F55887A5FF3DFF85B6019448FC562E69	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3DFF84B601978FFDE7288E.text	F55887A5FF3DFF84B601978FFDE7288E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pelomedusa subrufa (Bonnaterre 1789)	<div><p>Pelomedusa subrufa (Bonnaterre, 1789)</p> <p>Marsh Terrapin</p> <p>Material: PEM R 20502-3 (40).</p> <p>Comment: Although the previously monotypic Pelomedusa has been shown to have high levels of cryptic diversity, with the description of six new species (Petzold et al. 2014), Angolan populations remain assigned to P. subrufa.</p> </div>	https://treatment.plazi.org/id/F55887A5FF3DFF84B601978FFDE7288E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3DFF85B59B9541FDB42D2C.text	F55887A5FF3DFF85B59B9541FDB42D2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trachylepis varia (Peters 1867)	<div><p>Trachylepis varia (Peters, 1867)</p> <p>Variable Skink</p> <p>Material: PEM R 20507 (30c), 20508‒9 (35), 20510 (32), 20511‒2 (37), 20513 (38), 20514 (39), 20515 (42), 20516 (44a), 21489 (59), 21825 (46), and 21841 (47).</p> <p>Comment: Most common and widespread species of skink in Angola, with a massive range from Ethiopia to South Africa. The species is known to have both genetic and reproductive diversity, and probably comprises a complex of numerous cryptic taxa.</p></div> 	https://treatment.plazi.org/id/F55887A5FF3DFF85B59B9541FDB42D2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3DFF85B59B94C9FC16288E.text	F55887A5FF3DFF85B59B94C9FC16288E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trachylepis wahlbergi (Peters 1869)	<div><p>Trachylepis wahlbergi (Peters, 1869)</p> <p>Wahlberg’s Skink</p> <p>Material: PEM R 20520‒1 (43).</p> <p>Comment: The Trachylepis striata complex is widespread in southern and eastern Africa, but relationships and taxonomic status of the various subspecies of T. striata, elevated to specific status by Broadley (2000), remain problematic. Subsequently, Castiglia et al. (2006) showed that T. striata and T. wahlbergii share the same karyotype and no significant differentiation in 16S mtDNA gene. They argued that this did not support Broadley’s (2000) action in treating both taxa as distinct species. We retain the current taxonomic arrangement but note that fuller studies on the T. striata complex are required. First recorded from southeast Angola from near Mavinga (Branch and McCarthy 1992), but also known from numerous other localities in south-west Angola.</p> </div>	https://treatment.plazi.org/id/F55887A5FF3DFF85B59B94C9FC16288E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3DFF85B60191A1FB322ACC.text	F55887A5FF3DFF85B60191A1FB322ACC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Typhlacontias rohani Angel 1923	<div><p>Typhlacontias rohani Angel, 1923</p> <p>Rohan’s Legless Skink</p> <p>Material: PEM R 20522‒3 (38), and 20526 (44a).</p> <p>Comment: Angel (1923) described this species from Lwankundu River, a western tributary of the Cuando River. The only other record for Angola is from Chimporo, Cunene Province (Monard 1937b). The new collections (Fig. 8e) from the lower Cuando River represent the third and fourth records for Angola. Haacke (1997) reported the species to be common in north-eastern Namibia, the Caprivi Strip [= Zambezi Region], northern Botswana and western Zimbabwe.</p></div> 	https://treatment.plazi.org/id/F55887A5FF3DFF85B60191A1FB322ACC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3DFF85B60193A4FC402C29.text	F55887A5FF3DFF85B60193A4FC402C29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Varanus niloticus (Linnaeus 1766)	<div><p>Varanus niloticus (Linnaeus, 1766)</p> <p>Water Monitor</p> <p>Material: observations only (3, 32, 43).</p> <p>Comment: Common and widespread along major rivers in Angola. Although only a single Angolan specimen was included in a phylogeographic review of the V. niloticus complex (Dowell et al. 2016), it is likely that all Angolan water monitors are preferable to V. niloticus (of which V. ornatus is now a junior synonym) and not West African V. stellatus (Daudin, 1802), which was recently revalidated (Dowell et al. 2016).</p> </div>	https://treatment.plazi.org/id/F55887A5FF3DFF85B60193A4FC402C29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3CFF84B601948CFC652FD3.text	F55887A5FF3CFF84B601948CFC652FD3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Afrotyphlops mucruso (Peters 1854)	<div><p>Afrotyphlops mucruso / schlegeli</p> <p>– The status of large typhlopids in Angola previously assigned to Megatyphlops (Broadley and Wallach 2009) remains problematic. Broadley and Wallach (2009) report no specimens from south-east Angola, although they record three species in the country: A. mucroso from adjacent western Zambia, with a single record of unstated provenance mapped (Macanda, 10°50’S, 18°52’E) from north-eastern Angola; A. schlegelii from the Caprivi and northern Namibia, extending along the escarpment into the Benguela Province; and A. anomalus in south-central Angola. Afrotyphlops schlegelii may, therefore, occur in upland areas associated with sparse Miombo woodland bordering the river catchments.</p> </div>	https://treatment.plazi.org/id/F55887A5FF3CFF84B601948CFC652FD3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3CFF84B60195C4FBAD2CAB.text	F55887A5FF3CFF84B60195C4FBAD2CAB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hildebrandtia ornata (Peters 1878)	<div><p>Hildebrandtia ornata</p> <p>– Recorded from eastern and western Caprivi (Channing 2001), but no localities known from south-east Angola.</p></div> 	https://treatment.plazi.org/id/F55887A5FF3CFF84B60195C4FBAD2CAB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3CFF84B59B9384FEA82D69.text	F55887A5FF3CFF84B59B9384FEA82D69.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kinixys belliana Gray 1863	<div><p>Kinixys cf. belliana Gray, 1863</p> <p>Bell’s Hinged Tortoise</p> <p>Material: SARCA 153810 (13).</p> <p>Comment: The genus Kinixys has been recently revised (Kindler et al. 2012), with K. belliana now restricted to coastal Kenya and Somalia, and extending through Burundi to Angola. Kinixys spekii has previously been considered to have a wide distribution in southeast Africa and to range into much of Angola. Ceríaco et al. (2014) assign Capanda material to K. spekii, but this was not confirmed by molecular analysis. The only three Angola specimens sampled in the recent revision of Kinixys (Kindler et al. 2012), from Cuemba and Kuito, Bié Province, were all shown to group with East African K. belliana. The current specimen is provisionally assigned to K. belliana, pending further molecular analysis of Angolan material and we caution that the presence of K. spekii in Angola requires confirmation.</p> </div>	https://treatment.plazi.org/id/F55887A5FF3CFF84B59B9384FEA82D69	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3CFF84B59B91A1FEA02A2D.text	F55887A5FF3CFF84B59B91A1FEA02A2D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pelusios bechuanicus FitzSimons 1932	<div><p>Pelusios bechuanicus FitzSimons, 1932</p> <p>Botswanan Hinged Terrapin</p> <p>Material: PEM R 20504 (44a).</p> <p>Comment: A localized species associated with the Okavango Delta in Botswana. A specimen (Fig. 8h) was caught in pushing flood waters of the lower Cuando River. This new record represents the fourth record for Angola. The other records for Angola are from Chonga River, Moxico Province (Laurent 1964), Chimporo Marsh in the Cuanda drainage (Monard 1931) and Vila da Ponte (= Kubango) (Monard 1937b, as P. subniger).</p> </div>	https://treatment.plazi.org/id/F55887A5FF3CFF84B59B91A1FEA02A2D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3CFF84B6019224FB2F2B8B.text	F55887A5FF3CFF84B6019224FB2F2B8B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Poyntonophrynus fenoulheti (Hewitt & Methuen 1912)	<div><p>Poyntonophrynus fenoulheti</p> <p>– A small toad is known from western and eastern Caprivi, but unrecorded from south-east Angola (Channing 2001).</p></div> 	https://treatment.plazi.org/id/F55887A5FF3CFF84B6019224FB2F2B8B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3CFF84B60192A4FADA2C2B.text	F55887A5FF3CFF84B60192A4FADA2C2B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Poyntonophrynus kavangensis (Poynton & Broadley 1988)	<div><p>Poyntonophrynus kavangensis</p> <p>– A rare toad known from scattered localities in the Caprivi region from Rundu to Hwange, including a single record (23 km NW Pereira de Eca) from southern Angola (Poynton and Haacke 1993).</p></div> 	https://treatment.plazi.org/id/F55887A5FF3CFF84B60192A4FADA2C2B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3CFF84B6019444FC6C2D2B.text	F55887A5FF3CFF84B6019444FC6C2D2B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pyxicephalus edulis Peters 1854	<div><p>Pyxicephalus edulis</p> <p>– Recorded from Eastern Caprivi (Channing 2001) and northern Namibia (Ondangwa, Branch pers. obs.)</p></div> 	https://treatment.plazi.org/id/F55887A5FF3CFF84B6019444FC6C2D2B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF3CFF84B59B9406FD3F2E69.text	F55887A5FF3CFF84B59B9406FD3F2E69.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stigmochelys pardalis (Bell 1828)	<div><p>Stigmochelys pardalis (Bell, 1828)</p> <p>Leopard Tortoise</p> <p>Material: SARCA 153814 (44a).</p> <p>Comment: Only known from a handful records from southwest Angola and seems to be rare elsewhere in Angola (Bocage 1895; Mertens 1937; Hellmich 1957; Monard 1937b).</p></div> 	https://treatment.plazi.org/id/F55887A5FF3CFF84B59B9406FD3F2E69	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF31FF89B59B9083FE052A0C.text	F55887A5FF31FF89B59B9083FE052A0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amblyodipsas polylepis (Bocage 1873)	<div><p>Amblyodipsas polylepis</p> <p>– Recorded from the northern Namibia, western Zambia (Broadley 1971) and western Angola (Bocage 1895; Ferreira 1906), but not recorded yet from southeast Angola.</p></div> 	https://treatment.plazi.org/id/F55887A5FF31FF89B59B9083FE052A0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF31FF89B59B9323FF762B0C.text	F55887A5FF31FF89B59B9323FF762B0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amblyodipsas ventrimaculata (Roux 1907)	<div><p>Amblyodipsas ventrimaculata</p> <p>– Broadley (1971) noted this fossorial snake was restricted to Kalahari sands in northern Botswana, adjacent eastern Namibia, and western Zambia. It is a specialist feeder on fossorial reptiles and several suitable prey items (e.g., Zygaspis quadrifrons and Typhlacontias rohani) occur in south-east Angola.</p> </div>	https://treatment.plazi.org/id/F55887A5FF31FF89B59B9323FF762B0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF31FF89B59B91C2FEF729EC.text	F55887A5FF31FF89B59B91C2FEF729EC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atractaspis bibroni Jan 1859	<div><p>Atractaspis bibroni</p> <p>– There are few documented Angolan records for A. bibronii, and all are located in the west or north: Benguela (Boulenger 1895), Catumbela and Dombe (Bocage 1867), Cubal (Mertens 1938), Dundo (Laurent 1964). Broadley (1991) in his revision of the southern African populations of Atractaspis gives no new records for south-east Angola, even though there are numerous records for adjacent Namibia, Botswana, and western Zambia.</p> </div>	https://treatment.plazi.org/id/F55887A5FF31FF89B59B91C2FEF729EC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF31FF89B6019341FAF12B4E.text	F55887A5FF31FF89B6019341FAF12B4E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Colopus wahlbergii Peters 1869	<div><p>Colopus wahlbergii</p> <p>– This small terrestrial gecko is widespread in the Kalahari region, with a single, marginal Angolan record (Angola-Namibia border, 18°E; Haacke 1976). It has subsequently been recorded from the Caprivi (Haacke 1998, Branch unpub. obs.), Zimbabwe (Broadley and Spawls 1991) and extreme western Zambia (Broadley and van Daele 2003), and is, therefore, likely to occur in the sandy areas of the region.</p></div> 	https://treatment.plazi.org/id/F55887A5FF31FF89B6019341FAF12B4E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF31FF89B59B9483FDCF2E6C.text	F55887A5FF31FF89B59B9483FDCF2E6C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Crotaphopeltis barotseensis Broadley 1968	<div><p>Crotaphopeltis barotseensis</p> <p>– A semiaquatic snake, restricted to the Okavango and Kafue region (Rasmussen 1997), but presently unrecorded from Angola.</p></div> 	https://treatment.plazi.org/id/F55887A5FF31FF89B59B9483FDCF2E6C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF31FF89B60191C1FAE4296E.text	F55887A5FF31FF89B60191C1FAE4296E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dalophia longicauda (Werner 1915)	<div><p>Dalophia longicauda</p> <p>– Described from northern Namibia and known to extend through the Caprivi area to western Zimbabwe (Broadley et al. 1976; Gans 2005). Populations are therefore found on either side of the Okavango River and it is expected to occur in southeast Angola.</p></div> 	https://treatment.plazi.org/id/F55887A5FF31FF89B60191C1FAE4296E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF31FF89B59B9703FF682E8C.text	F55887A5FF31FF89B59B9703FF682E8C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Elapsoidea semiannulata Bocage 1882	<div><p>Elapsoidea semiannulata</p> <p>– Recorded in western Zambia, northern Namibia, and west and central Angola (Broadley 1998), but no records from southeast Angolan are known.</p></div> 	https://treatment.plazi.org/id/F55887A5FF31FF89B59B9703FF682E8C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF31FF89B59B9543FE7C2C8C.text	F55887A5FF31FF89B59B9543FE7C2C8C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemirhaggheris nototaenia	<div><p>Hemirhaggheris nototaenia</p> <p>– A dwarf arboreal snake recorded from the western Caprivi and Okavango region, east through Zambia to East Africa (Broadley and Hughes 2000). Earlier records from southwest Angola (Bocage 1895) have been referred to H. viperinus by Broadley and Hughes (2000).</p> </div>	https://treatment.plazi.org/id/F55887A5FF31FF89B59B9543FE7C2C8C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF31FF89B6019001FBFC29AE.text	F55887A5FF31FF89B6019001FBFC29AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ichnotropis grandiceps Broadley 1967	<div><p>Ichnotropis grandiceps</p> <p>– Since its description (Broadley 1967) from the Caprivi area, no new material has been discovered. As noted for Ichnotropis sp. (above), the status of new Angolan material and the validity of I. grandiceps are under investigation.</p> </div>	https://treatment.plazi.org/id/F55887A5FF31FF89B6019001FBFC29AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF31FF89B59B95A3FD992DEC.text	F55887A5FF31FF89B59B95A3FD992DEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Psammophis jallae Peracca 1896	<div><p>Psammophis jallae</p> <p>– Angel (1921) described Psammophis rohani (type: MNHN 20-198; type locality Lumuna River, a tributary of the Luina and Cuando Rivers) from south-east Angola. It was synonymized with P. jallae Peracca, 1886 by Broadley (2002). No other specimens of P. jallae are known from the region.</p> </div>	https://treatment.plazi.org/id/F55887A5FF31FF89B59B95A3FD992DEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF31FF89B59B9223FDBE2BAC.text	F55887A5FF31FF89B59B9223FDBE2BAC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xenocalamus Günther 1868	<div><p>Xenocalamus b. bicolor</p> <p>– Although Broadley (1971) records no Angolan material the species occurs in the Caprivi area and adjacent western Zambia, and it is usually associated with Kalahari sands.</p></div> 	https://treatment.plazi.org/id/F55887A5FF31FF89B59B9223FDBE2BAC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
F55887A5FF31FF89B59B97A2FD8E2FCC.text	F55887A5FF31FF89B59B97A2FD8E2FCC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zygaspis nigra Broadley and Gans 1969	<div><p>Zygaspis nigra</p> <p>– Known from three localities in east-central Angola (Gago Coutinho District, Alto Chicapa, Colanda, seven km E Vila Luso on Moxico road; Broadley and Gans 1975) and adjacent populations in the Caprivi and Zambia, west of the Zambezi River.</p></div> 	https://treatment.plazi.org/id/F55887A5FF31FF89B59B97A2FD8E2FCC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Conradie, Werner;Bills, Roger;Branch, William R.	Conradie, Werner, Bills, Roger, Branch, William R. (2016): The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation (e 126) 10 (2): 6-36, DOI: 10.5281/zenodo.13154851
