taxonID	type	description	language	source
F56B87A7FF98F3775AD2FB3FFCAEF9F7.taxon	etymology	Etymology. The name Orostylis is derived from the Latin “ os ” (mouth), and “ stylis ” (stylet). The name refers to the fact that the male copulatory apparatus connects to the mouth in the type species.	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF98F3775AD2FB3FFCAEF9F7.taxon	diagnosis	Diagnosis. Anterior-facing, cylindrical pharynx and sclerotised stylet in the anterior part of the body. Male genital opening in the oral cavity. Single ovary and vitellarium (O. timucuorum sp. nov. might have two vitellaria). No other female genital structures. Testes paired or unpaired. No visible connection between the female genital system and the environment, except possibly in O. timucuorum sp. nov. Grammatical gender. masculine	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF98F3775AD2FB3FFCAEF9F7.taxon	type_taxon	Type species. Orostylis dohae, sp. nov., described below.	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF98F3715AD2F927FE5AFBF9.taxon	description	(Figs. 2, 3 A, 4 D) Neodalyellida n. gen. n. sp. 2 in Van Steenkiste et al. (2013) Neodalyellida sp. 2 in Stephenson et al. (2019) Localities. Doha, Qatar (25 ° 19 ’ 04 ” N; 51 ° 32 ’ 18 ” E) Sand at about the high-water mark (Mar. 19, 2008): Type locality.	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF98F3715AD2F927FE5AFBF9.taxon	materials_examined	Material. Observations on live specimens. One whole mount designated holotype (FMNH KV. 690), and two serially sectioned specimens (HU XIV. 3.04 – XIV. 3.05).	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF98F3715AD2F927FE5AFBF9.taxon	etymology	Etymology. Species named after the capital of Qatar, where the species was discovered.	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF98F3715AD2F927FE5AFBF9.taxon	description	Description. Animals are about 0.6 mm long (measured on whole mount), with two dark brown, kidney-shaped eyes (Figs. 2 A, 4 D: e). The epidermis is syncytial, strongly ciliated, and 2 – 5 μm thick. The cilia measure 2 – 3 µm. Nuclei in the epidermis are scarce. A thin basal membrane underlies the epidermis, followed by a circular and a longitudinal muscle layer, respectively. Several basophilic, coarse-grained glands are situated in the anterior end of the body, in the area surrounding the mouth. The caudal end of the body contains eosinophilic cement glands. The mouth is located near the anterior end of the body (Fig. 2 A: m) and surrounded by a sphincter. A long, thin-walled oral tube connects the mouth opening to the pharynx. The pharynx (Figs. 2, 4 D: ph) is located in the anterior body half and is between 1 / 4 and 1 / 5 of the body length long. The pharynx is cylindrical in shape, with a conspicuous musculature, including radial, circular, and longitudinal muscles. This is typical for the pharynx doliiformis in neodalyellids, but the radial muscles are very wide and stacked closely together with little or seemingly no space between them. The anterior end of the pharynx bears a “ collar ” (Fig. 2 A: phc) separated by a constriction from the rest of the pharynx. This collar is provided with three pairs of tentacles projecting anteriorly (Figs. 2 A, 4 D: pht). The intestine (Fig. 2 B: i) lies caudally from the ovary (Fig. 2: ov). In live specimens, diatom frustules were observed inside the intestinal tract. The large, paired testes are situated just anterior to the midbody (Figs. 2, 4 D: t), behind the pharynx, on either side of the body. In live specimens, two distinct testes were clearly observed. However, in the serial sections, the separation between both testes was less obvious, creating the impression of a single, large testis. The two testes lie against one another in such a way that the spermatogenic distal ends of the testes point laterally. A pair of vasa deferentia extend anteriorly and connect the testes to the seminal vesicle (Figs. 2, 4 D: sv) in the proximal half of the male copulatory organ. The copulatory organ is situated in the anterior part of the body, alongside the pharynx. It consists of the round to ovoid seminal vesicle (Figs. 2, 4 D: sv), followed distally by a small, globular prostate vesicle (Fig. 2 B: pv) filled with eosinophilic gland secretions. The seminal and prostate vesicle are lined with a syncytial, nucleated epithelium and surrounded by circular muscles. In the distal half of the copulatory organ, the seminal vesicle connects directly to a sclerotised stylet (Figs. 2, 4 D: st; 3 A). The stylet is a simple, thin-walled funnel, curved at an angle of about 90 ° at its midpoint. The stylet is 33 µm long and 9 µm wide proximally. It connects to a long male duct, which opens in the oral cavity (Fig. 2 B: oc). The female genital system consists of a large, curved ovary (Fig. 2: ov), positioned at the midbody, dorsally to and in front of the intestine. A female duct or gonopore was not observed. Instead, the oviduct appears to connect directly to the intestinal lumen. It is assumed that eggs are deposited in the intestinal lumen and expelled from the body via the mouth. Eosinophilic glands are present in the region where the ovary connects to the intestine. In the same area, vesicles with sperm were observed in the intestinal wall (Fig. 2 A: isv). Possibly, these vesicles are resorptive and function as a bursa. The posterior half of the body is almost completely taken up by a lobed or branched vitellarium (Fig. 2 B: vit).	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF9EF3705AD2FB00FCC9FEC5.taxon	description	(Figs. 3 B – C, 4 A) Localities. Cala Sant’Andrea, Asinara National Park, Sardinia, Italy (41 ° 00 ’ 48 ” N; 8 ° 14 ’ 56 ” E) Sand from 2 m deep (Sep. 27, 2014): Type locality. Cala dei Ponzesi, Asinara National Park, Sardinia, Italy (41 ° 05 ’ 14 ” N; 08 ° 20 ’ 45 ” E) Sand from the intertidal (Sep. 30, 2014).	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF9EF3705AD2FB00FCC9FEC5.taxon	materials_examined	Material. Observations on live specimens. Two whole mounts, one of which designated holotype (FMNH KV. 691), the other deposited in the HU XIV. 3.06.	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF9EF3705AD2FB00FCC9FEC5.taxon	etymology	Etymology. Species named after the National Park and island of Asinara, Sardinia, where the species was discovered.	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF9EF3705AD2FB00FCC9FEC5.taxon	description	Description. Specimens are transparent, unpigmented. A pair of round eyes (Fig. 4 A: e) is present anteriorly. The overall internal organisation of the specimens corresponds to that of Orostylis dohae sp. nov. The mouth is situated ventrally, near the anterior body end. A horizontally positioned cylindrical pharynx (Fig. 4 A: ph) is situated in the anterior third of the body. The anterior edge of the pharynx bears several small papillae, but long tentacles as in O. dohae sp. nov. were not observed. The single testis (Fig. 4 A: t) is situated ventrally around the midbody. The male copulatory system lies alongside the pharynx. It consists of a large copulatory bulb containing a seminal vesicle (Fig. 4 A: sv) and prostatic vesicle. In live specimens, circular muscles clearly surround the copulatory bulb. The seminal and prostate vesicles enter the stylet separately. The stylet (Figs. 3 B, C; 4 A: st) is a strongly curved, “ raptor-claw ” - shaped, hollow spine, with a “ handle-like ” extension projecting proximally. In one of the specimens (HU XIV. 3.06), the stylet is straighter, with the distal third bent to one side (Fig. 3 C). The total stylet (incl. the handle) measures 64 – 76 μm (x = 70 μm; n = 2). The handle is 15 – 19 µm long (x = 17 μm; n = 2). The stylet shows a longitudinal slit on one side. Based on observations of live specimens, it is likely that the male system opens into the oral cavity, as is the case in O. dohae sp. nov. An ovary (Fig. 4 A: ov) was observed in the midbody. It is large and, together with the intestine, takes up most of the posterior two thirds of the body. A single vitellarium fills the caudal end of the body, with two large branches stretching forwards up to the level of the copulatory organ.	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF9FF3705AD2FE5CFEEDFB25.taxon	description	(Figs. 3 D, 4 B) Localities. Bahía Larga, Guamá municipality, Santiago de Cuba province, Cuba (19 ° 58 ’ 57 ” N; 76 ° 18 ’ 15 ” W) Intertidal, very fine sand in the upper 10 cm of the sediment, salinity 34 ‰ (Apr. 5, 2017): Type locality. Bahía de Gibara, Holguín province, Cuba (21 ° 06 ’ 00 ” N; 76 ° 08 ’ 09 ” W) Intertidal, on very fine sand with organic material close to the mouth of the river, salinity 32 ‰ (Feb. 17, 2016).	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF9FF3705AD2FE5CFEEDFB25.taxon	materials_examined	Material. Observations on four live specimens, three of which were whole mounted afterwards, one designated holotype (FMNH KV. 692), the others deposited in the HU XIV. 3.07 – XIV. 3.08.	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF9FF3705AD2FE5CFEEDFB25.taxon	etymology	Etymology. The specific name refers to the species’ lack of eye spots (Lat. “ caecus ” = blind).	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF9FF3705AD2FE5CFEEDFB25.taxon	description	Description. General morphology as described for the species above, with the stylet located anteriorly, lateral to the pharynx (Fig. 4 B: ph, st). The body is unpigmented, slender, and 0.25 – 0.42 mm long (measured on two whole-mounted specimens). Eye spots are lacking. The mouth opens subterminally and is surrounded by few tactile hairs. The pharynx (Fig. 4 B: ph) is 1 / 6 – 1 / 3 of the body length in live animals, measuring 104 – 120 µm in two whole mounts and bearing six slender tentacles anteriorly (Fig. 4 B: pht). The paired testes are located at both sides of the pharynx, each one connected to a seminal vesicle. The two seminal vesicles enter a small prostate vesicle. The stylet (Figs. 3 D; 4 B: st) is formed by a broad, proximal piece (Fig. 3 D: bp) and a distal, small, hollow hook (Fig. 3 D: h). The entire stylet is 24 – 31 μm long (x = 29 μm; n = 3) and 14 – 20 μm wide (x = 16 μm; n = 3). The hook is 8 – 15 μm long (x = 11 μm; n = 3) and 3 – 5 μm wide at its base (x = 4 μm; n = 3). The broad piece is partially filled by a granular secretion, either originating from the prostate vesicle or part of the prostate vesicle itself. A single ovary (Fig. 4 B: ov) is situated in the midbody and is surrounded by the gut.	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF9FF3735AD2FB3CFD43FE99.taxon	description	(Figs. 3 G, 4 C) Orostylis distortus n. n. in Armonies (2017) Localities. Sylt, Germany (55 ° 04 ’ 44 ” N; 5 ° 17 ’ 31 ” E) Sublittoral, 8.2 m deep, clean fine sand (Apr. 28, 2015): Type locality.	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF9FF3735AD2FB3CFD43FE99.taxon	materials_examined	Material. Observations on live specimens, one of which whole mounted and designated holotype (FMNH KV. 693).	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF9FF3735AD2FB3CFD43FE99.taxon	etymology	Etymology. The specific name refers to the stylet morphology (Lat. “ distortus ” = distorted).	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF9FF3735AD2FB3CFD43FE99.taxon	description	Description. General morphology as described for the genus, with the stylet located anteriorly, lateral to the pharynx. The body is unpigmented, 0.8 mm (resting) to 1.2 mm long (swimming). The anterior and posterior body ends are more or less rounded. The mouth opens terminally or subterminally and is surrounded by few tactile hairs. Two pigmented eye spots (Fig. 4 C: e) are situated anterior to the pharynx. The pharynx (Fig. 4 C: ph) is cylindrical, with six 30 - µm-long tentacles (Fig. 4 C: pht). The proximal 1 / 3 of the tentacles’ length is anchored in the pharynx, which seems to have no circular muscles in this area and therefore differs in appearance from the distal part of the pharynx. The pharynx is only 1 / 8 of the body length and relatively small compared to the pharynx in other species of the genus. The paired testes are located laterally between the pharynx and the midbody. The copulatory organ lies beside the pharynx or directly behind it in stretched specimens. Its musculature is hardly visible in live specimens. A single, elongated seminal vesicle (Fig. 4 C: sv) and prostatic vesicle (Fig. 4 C: pv) enter the stylet proximally. The stylet (Figs. 3 G; 4 C: st) is S-shaped, 42 µm long and consists of a proximal oval funnel, which rapidly narrows to a coiled, slender, 2 – 3 - µm-wide tube ending in a pointed tip. At its proximal end, a longitudinal slit was observed. A singly ovary is present in the midbody, with the largest egg cells occurring anteriorly. The paired vitellaria stretch from the midbody to the rear body end.	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF9CF37D5AD2F939FE49FE70.taxon	description	(Figs. 3 E, 4 E) Neodalyellida n. gen. n. sp. 1 in Van Steenkiste et al. (2013) Neodalyellida sp. 1 in Stephenson et al. (2019) Localities. Doñana National Park, Andalusia, Spain (36 ° 52 ’ 16 ” N; 06 ° 25 ’ 45 ” W) Sand from Doñana beach sampled at the low water mark, at 10 km north from the Guadalquivir river mouth (Apr. 6, 2008): Type locality. Doñana National Park, Andalusia, Spain (36 ° 47 ’ 57 ” N; 6 ° 23 ’ 21 ” W) Sand from Doñana beach sampled at the low water mark, at the Guadalquivir river mouth (Apr. 6, 2008).	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF9CF37D5AD2F939FE49FE70.taxon	materials_examined	Material. Observations on live specimens. Two whole mounts, one of which designated holotype (FMNH KV. 694), the other one deposited in the HU reference collection HU XIV. 3.09.	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF9CF37D5AD2F939FE49FE70.taxon	etymology	Etymology. Species epithet named after the type locality in Doñana National Park, Spain.	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF9CF37D5AD2F939FE49FE70.taxon	description	Description. Specimens have an elongate body with a pair of small, bean-shaped eyes near the anterior end. Whole-mounted specimens are about 0.5 (holotype) and 0.9 mm (HU XIV. 3.09) long respectively. The pharynx is cylindrical in shape and located just behind the eyes. Based on observations of live specimens and whole mounts, the general internal organisation appears similar to that of Orostylis dohae sp. nov., with two testes just behind the pharynx. The male copulatory organ is situated alongside the pharynx, in the anterior part of the body, with the distal end pointing anteriorly. The copulatory bulb contains an elongate seminal vesicle and connects distally to a sclerotised stylet. This stylet (Figs. 3 E, 4 E) consists of a corkscrew-shaped folded plate or “ gutter ” and is 74 – 94 µm long. The proximal base of the stylet is about 15 µm wide and bears an extension that is folded towards the anterior. As no serial slides were available for this species, it is not clear whether the male gonopore opens into the oral cavity, as is the case in Orostylis dohae sp. nov. No details on the anatomy of the female genital system could be observed on live specimens alone.	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF92F37D5AD2FDF1FB25FB15.taxon	description	(Figs. 3 F, 4 F) Localities. More or less 70 m in front of a cliff at Wimereux beach, Wimereux, France (50 ° 45 ’ 36 ” N; 1 ° 41 ’ 31 ” E) Sand and shell gravel among algae scraped of rocks (Sep. 16, 2015): Type locality. More or less 70 m in front of a cliff at Wimereux beach, Wimereux France (50 ° 45 ’ 36 ” N; 1 ° 41 ’ 31 ” E) Red algae on rocks (Sep. 16, 2015).	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF92F37D5AD2FDF1FB25FB15.taxon	materials_examined	Material. Two whole mounts, one of which designated holotype (FMNH KV. 695), the other one in the HU reference collection (HU XIV. 3.10).	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF92F37D5AD2FDF1FB25FB15.taxon	etymology	Etymology. The species is named after the ancient Celtic people that inhabited the region of the type locality (Lat. “ gallicus ” = Gallic).	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF92F37D5AD2FDF1FB25FB15.taxon	description	Description. Based on observations of the whole mounts, the general morphology is as described for the genus, with the copulatory organ located anteriorly in the body, lateral to the pharynx. The number of testes is unknown. The stylet (Figs. 3 F, 4 F) is a strongly curved, “ raptor-claw ” - shaped hollow spine, and is 45 – 51 μm long. The proximal part is funnel-shaped and has an asymmetric aperture with a rounded flap. The distal third of the stylet is thin and tapers into a sharp tip. Laterally, the stylet shows a narrow, longitudinal slit. Additional details of the male copulatory system and female genital system could not be observed on the available specimens.	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF92F37F5AD2FB2CFD1AFDB9.taxon	description	(Figs 3 H, 5, S 1 – S 3) Localities. Small beach facing the intracoastal waterway near the boat launch and pier, Whitney Marine Laboratory, FL (29 ° 40 ’ 14 ” N; 81 ° 12 ’ 57 ” W) Muddy sand with oyster shells and shell fragments (May 21, 2019): Type locality. Bear Island, NC (approximately 34 ° 38 ’ 51 ” N; 77 ° 07 ’ 02 ” W) Muddy sand-flat near tidal drainage stream (Jun., 2018). Outer sandy beach on Bogue Banks, Emerald Isle Site, NC (34 ° 38 ’ 42 ” N; 77 ° 05 ’ 23 ” W) Surface sediment halfway between MTL and LTL (Mar. 15, 2013). “ Anne McCrary’s Mudflat ” (muddy sand-flat facing the intracoastal waterway near Wilmington, NC — approximately 34 ° 13 ’ 11 ” N; 77 ° 48 ’ 49 ” W) (Jan. and Feb., 1970) by R. M. Rieger. Bogue Inlet, NC. Shallow subtidal in “ Amphioxus sand ” (date unknown) by R. M. Rieger.	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF92F37F5AD2FB2CFD1AFDB9.taxon	materials_examined	Material. Five specimens studied alive; two turned into whole mounts; one designated holotype (FMNH KV. 696); the other reference material (HU XIV. 3.19); three specimens examined by confocal microscopy; three live-drawings made by R. M. Rieger.	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF92F37F5AD2FB2CFD1AFDB9.taxon	etymology	Etymology. The specific epithet is in honour of the people (called “ Timucua ” by the Spanish) who first inhabited the region near the type locality.	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
F56B87A7FF92F37F5AD2FB2CFD1AFDB9.taxon	description	Description. Living, sexually mature specimens from the type locality measure 705 – 895 µm long; all specimens possess a pair of pigmented eyes, as well as the general characters of the genus (Fig. 5 A). The pharynx averages 16 % of the body length (Fig. 5 A: ph; n = 3). The pharynx bears 6 club-tipped, distal tentacles (Figs. 5 A inset, S 1). Radial pharyngeal musculature is thickly arranged and pseudostriated (Fig. 5 B: rm). The male organ is directed forward and lies just behind or beside the pharynx; testes are paired (Figs. 5 A: t, S 2). The ovary is V-shaped, with paired germinal zones, sometimes giving the impression of two ovaries (Figs. 5 A, C: ov). The vitellarium is either very large, or paired, situated immediately behind the ovary and extending posteriorly (Fig. 5 C: vit). A bursal tissue lies in the ventral midline at approximately the level of the ovaries (Fig. 5 A, D: bu). Confocal microscopy reveals a musculature associated with this that appears to lead to a mid-ventral pore (Figs. 5 D, E: red arrow). The 58 - µmlong stylet ranges in length from 47 to 69 µm (n = 5), possessing a flared base with two distinct areas (Figs. 3 H, 5 F – G; 5 H: sb, st). The stylet appears to have separate pathways for sperm and prostatic secretions. The stylet has a lateral slit that extends at least partway along it (Fig. 5 F, arrows). Squeezing apparently flattens a stylet that, at rest, comprises a base with an elongate, semi-circular tip whose point rests in a small “ cup ” (Fig. 5 H: green arrow); the base and tip are in nearly orthogonal planes (Fig. 5 H: red arrows). Connection of the male canal with the pharynx tube could not be confirmed with the material at hand.	en	GOBERT, STEFAN, ARMONIES, WERNER, DIEZ, YANDER L., JOUK, PHILIPPE, MONNENS, MARLIES, REVIS, NATHALIE, REYGEL, PATRICK, SMITH, JULIAN III, STEENKISTE, NIELS VAN, ARTOIS, TOM (2022): Orostylis gen. nov., a new genus of Dalytyphloplanida with seven new species (Platyhelminthes: Rhabdocoela). Zootaxa 5115 (1): 29-46, DOI: 10.11646/zootaxa.5115.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5115.1.2
