identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
F533EE36FFEBFFDAFF18DC7DFCEA0317.text	F533EE36FFEBFFDAFF18DC7DFCEA0317.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trisinus Ding, Niu & Yin 2025	<div><p>Trisinus cavernosus species group</p><p>Diagnosis. Trisinus species with a large, exaggerated excavation at the middle of the vertex in the male. Other body parts, including the antennae, are simple, unmodified.</p><p>Included species. At present there are two closely related species, Trisinus cavernosus sp. nov. and Trisinus qingliangmontis sp. nov., placed in this newly termed group. Additional species from southern China have been seen (Ding &amp; Yin pers. obs.), and will be dealt with separately.</p></div>	https://treatment.plazi.org/id/F533EE36FFEBFFDAFF18DC7DFCEA0317	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ding, Mei-Lin;Niu, Xiao-Ling;Yin, Zi-Wei	Ding, Mei-Lin, Niu, Xiao-Ling, Yin, Zi-Wei (2025): A new species group of Trisinus Raffray (Coleoptera: Staphylinidae: Pselaphinae), with description of two new species from the Tianmu Mountains, China. Zootaxa 5590 (2): 256-262, DOI: 10.11646/zootaxa.5590.2.5, URL: https://doi.org/10.11646/zootaxa.5590.2.5
F533EE36FFEBFFD8FF18DD39FC790707.text	F533EE36FFEBFFD8FF18DD39FC790707.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trisinus cavernosus Ding & Niu & Yin 2025	<div><p>Trisinus cavernosus sp. nov.</p><p>Chinese common name: ëǎñaȃş</p><p>(Figs 1, 3)</p><p>Type material (56 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Zhejiang, Linan Dist., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=119.504166&amp;materialsCitation.latitude=30.401943" title="Search Plazi for locations around (long 119.504166/lat 30.401943)">W. Tianmu Mt., Gaolingtou</a>, 30°24'7"N, 119°30'15"E, 1000 m, 4.ix.2024, Peng, Yin, Ding &amp; Zhang leg. ṽũ临aex天năü头DZĸDz ’ (SNUC) . PARATYPES: CHINA: Zhejiang: 12 ♂♂, 6 ♀♀, ‘ China: Zhejiang, Linan Dist., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=119.46139&amp;materialsCitation.latitude=30.40611" title="Search Plazi for locations around (long 119.46139/lat 30.40611)">W. Tianmu Mt., Niling to Pingxi</a>, 30°24'22"N, 119°27'41"E, 950–1000 m, 7.ix.2024, Yin, Ding, Zhang leg. ṽũūm临aex天nāü - ṿã ’ ; 1 ♂, 1 ♀, ‘ China: Zhejiang, Linan Dist., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=119.49083&amp;materialsCitation.latitude=30.369444" title="Search Plazi for locations around (long 119.49083/lat 30.369444)">W. Tianmu Mt., Niwuli to Xianrentai</a>, 30°22'10"N, 119°29'27"E, 550–1200 m, 6.ix.2024, Peng, Ding, Zhang leg. ṽũ临aex天n坭坞¶-仙人台 ’ ; 1 ♂, 2 ♀♀, ‘ China: Zhejiang Prov., Hangzhou City, Mt. Wuchaoshan, alt. 200 m, 11-IV-2011, Peng &amp; Zhu leg.’ ; 1 ♂, 8 ♀♀, ‘ China: Zhejiang Prov., Linan City, Mt. East Tianmushan, alt. 370–500 m, 12-IV-2011, Peng &amp; Zhu leg.’ ; 1 ♀, ‘ Longwangshan, Zhejiang Prov., 300–500 m, 23-IV-2004, Jing-wen Zhu leg.’; 2 ♀♀, ‘ China: Zhejiang Prov., Anji, Longwangshan Mt., Qianmutian, 29.vii.2011, 700–1350 m, Yu-Hong Pan leg.’ ; 2 ♂♂, 3 ♀♀, ‘ China: Zhejiang, Anji County, Longwang Shan N. R., Pingxi, creek valley, 1000–1100 m, 09.vi.2012, Hu &amp; Yin leg.’ ; Jiangxi: 2 ♂♂, 1 ♀, ‘ China: Jiangxi, Nanchang City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.78306&amp;materialsCitation.latitude=28.766945" title="Search Plazi for locations around (long 115.78306/lat 28.766945)">Wangjia Mountain</a>, 28°46'1"N, 115°46'59"E, 80 m, 10.x.2024, sifted, Jing-Zhe Du leg. ũ xâ南ēǖdzüĩNJƜữŝẽī ’ ; 5 ♂♂, 1 ♀, ‘ China: Jiangxi, Nanchang City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.78472&amp;materialsCitation.latitude=28.765556" title="Search Plazi for locations around (long 115.78472/lat 28.765556)">Wangjia Mountain</a>, 28°45'56"N, 115°47'5"E, 190 m, 3.xi.,2024, leaf litter, sifted, Ming-Zhe Liu &amp; Jing-Zhe Du leg. ũx南ēdzüĩNJƜữŝẽ 刘ẹẽ ’ ; 3 ♂♂, 3 ♀♀, ‘ China: Jiangxi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.768616&amp;materialsCitation.latitude=28.764166" title="Search Plazi for locations around (long 115.768616/lat 28.764166)">Nanchang Botanical Garden</a>, 28°45'51"N, 115°46'7"E, 325 m, 7.xi.2024, sifted, Jing-Zhe Du leg. ũxâ南ēäń园ữŝẽī ’ (all in SNUC) .</p><p>Diagnosis. Male. Body length approximately 1.7–1.8 mm. Head sub-rectangular; vertex greatly modified, with large V-shaped cavity between roundly denticulate lateral projections, inside cavity one lingulate protuberance surrounded by four tufts of long setae; antenna simple, lacking modifications. Pronotum in addition to sulci with weak longitudinal discal carinae. Discal striae of elytra extending posteriorly to approximately apical 4/5 of elytral length. Aedeagus greatly asymmetric, ventral stalk composed of one elongate structure with two macrosetae and one broad, membranous part, dorsal lobe thin and serrate at apex. Female. Body length approximately 1.7 mm, head simple, lacking modifications, antenna shorter than for male, genitalia as in Fig. 1J.</p><p>Description. Male. Body (Fig. 1A) length 1.72–1.77 mm; color reddish-brown, with tarsi and mouthparts lighter. Dorsal surface finely punctate, covered with relatively long pubescence.</p><p>Head (Fig. 1B, C) sub-rectangular, almost truncate at base, much wider than long, length 0.43 mm, width across eyes 0.49–0.50 mm; vertex heavily modified, densely and coarsely punctate, lacking foveae (dorsal tentorial pits) and sulci due to modification; with large, roughly V-shaped central cavity between two large, roundly denticulate lateral projections; inside cavity one small, thin, lingulate protuberance, with two setose tufts at middle of anterior margin and two similar tufts at middle of posterior margin of cavity; lateral margins of cavity densely setose; tempora much shorter than eyes; antennal tubercles weakly raised, with setose postantennal foveae; frons broadly impressed between antennal tubercles, impunctate and glabrous at middle, anteriorly fused with clypeus; clypeus finely punctate, anterior margin carinate and moderately raised; lacking ocular-clypeal carinae. Venter with two small gular foveae (posterior tentorial pits) in single pit, lacking median carina. Eyes prominent, each composed of approximately 25 ommatidia. Maxillary palpus with palpomere 1 minute; 2 elongate, curved, pedunculate basally and enlarged apically; 3 short, sub-triangular; 4 fusiform, widest near middle. Antenna relatively short, lacking modifications, length 0.90–0.92 mm; indistinct club loosely formed by slightly enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–10 each submoniliform, 11 largest, slightly longer than 9 and 10 combined (19:16), sub-conical.</p><p>Pronotum (Fig. 1B) slightly shorter than wide, length 0.36–0.37 mm, width 0.40–0.42 mm, widest approximately at middle; semi-circular lateral sulci extending from dorsal surface laterally and anteriorly; disc weakly convex, coarsely punctate, median longitudinal sulcus longer than lateral sulci, with pair of thin discal carinae, lacking median antebasal fovea and mediobasal impression, lateral antebasal foveae connected by transverse antebasal sulcus, lacking antebasal tubercles; outer and inner pair of basolateral foveae present. Prosternum with basisternal (precoxal) portion slightly longer than procoxal rests; with small lateral procoxal foveae; oblique hypomeral grooves present at basal halves, lacking lateral antebasal hypomeral impressions; hypomeral ridges close to coxal cavities.</p><p>Elytra wider than long, length 0.53–0.57 mm, width 0.62–0.66 mm; roundly quadrate, truncate at bases, dorsal surface with relatively long pubescence; each elytron with two distinct, asetose basal foveae; discal striae long, extending from outer basal foveae posteriorly to 4/5 of elytral length; humeri denticulate, with tiny subhumeral foveae, sulcate marginal striae extending from foveae to posterior margins of elytra. Metathoracic wings vestigial.</p><p>Mesoventrite short, laterally fully demarcated from metaventrite by oblique ridges; median mesoventral foveae narrowly separated, originating from shared setose, transverse opening, lateral mesoventral foveae large and setose, forked internally; prepectus broadened, collar-shaped; mesoventral intercoxal process short, apically acute. Metaventrite slightly and broadly impressed at middle, with pair of large, setose lateral mesocoxal foveae and setose lateral metaventral foveae, metaventral intercoxal process narrowly and shallowly emarginate at middle.</p><p>Legs moderately elongate; mesotibia (Fig. 1D) with small blunt tubercle at apex.</p><p>Abdomen slightly narrower than elytra, widest at lateral margins of tergite 1 (IV), length 0.41–0.51 mm, width 0.57–0.58 mm. Tergite 1 (IV) (Fig. 1E) dorsally longer than 2–4 (V–VII) combined, basal sulcus interrupted by two pairs of basolateral foveae, discal carinae distinct, with complete and oblique inner marginal carinae; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) shorter than 2 and 3 combined along middle, tergite 5 (VIII) transverse, posterior margin roundly emarginate at middle, tergite 2–4 (V–VII) each with one pair of small basolateral foveae. Sternite 2 (IV) densely setose at base, with large, transverse mediobasal and two pair of basolateral foveae, lacking basal impression, with moderately long lateral carinae extending for almost half of sternal length; midlength of sternite 2 longer than 3–5 (V–VII) combined, 3–5 each short at middle, 3 and 4 (VI) with one pair of punctiform basolateral foveae, 5 lacking foveae, sternite 6 (VIII) transverse, posterior margin sinuate at middle, sternite 7 (IX) (Fig. 1F) suboval, weakly sclerotized, apex rounded and with few long setae along apical margin.</p><p>Aedeagus (Fig. 1G–I) 0.27 mm long, weakly sclerotized, dorsoventrally greatly asymmetric; median lobe with large basal capsule and moderately oval foramen, ventral stalk composed of one elongate structure gradually narrowing apically and curved rightward (orientation according to Fig. 1H), with two long macrosetae at middle, and another shorter projection broadest at base and narrowing apically; dorsal lobe much thinner than ventral stalk, narrowed at middle and broadening apically and basally, with serrate apex; parameres fused and reduced to ventral membrane.</p><p>Female. General external morphology similar to male; antenna shorter; head lacking modifications; legs simple; each eye composed of about 18 ommatidia; humeri moderately prominent; metathoracic wings lacking. Measurements (as for male): body length 1.71–1.74 mm, length/width of head 0.36 mm / 0.47 mm, pronotum 0.37– 0.38 mm / 0.41–0.42 mm, elytra 0.49–0.51 mm / 0.63–0.64 mm, abdomen 0.52–0.54 mm / 0.60–0.62 mm; length of antenna 0.80 mm; genitalia transverse (Fig. 1J), 0.24 mm wide, posterior margin convex at middle.</p><p>Comparative notes. This and the following species represent the first members of the genus exhibiting significantly modified male cephalic structures. These species demonstrate apparent close relatedness based on similar external morphological characteristics and aedeagal structures. The primary distinguishing features lie in the configuration of the male vertex. Trisinus cavernosus sp. nov. displays a V-shaped central cavity with greatly projected lateral portions, whereas T. qingliangmontis sp. nov. exhibits a U-shaped central cavity with distinctly impressed lateral portions.</p><p>Distribution. East China: Zhejiang, Jiangxi (Fig. 3A).</p><p>Etymology. The specific epithet ‘ cavernōsus (- a, - um)’ (full of hollows or cavities) is a Latin adjective, referring to the remarkably modified male head with a large cavity of this species.</p></div>	https://treatment.plazi.org/id/F533EE36FFEBFFD8FF18DD39FC790707	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ding, Mei-Lin;Niu, Xiao-Ling;Yin, Zi-Wei	Ding, Mei-Lin, Niu, Xiao-Ling, Yin, Zi-Wei (2025): A new species group of Trisinus Raffray (Coleoptera: Staphylinidae: Pselaphinae), with description of two new species from the Tianmu Mountains, China. Zootaxa 5590 (2): 256-262, DOI: 10.11646/zootaxa.5590.2.5, URL: https://doi.org/10.11646/zootaxa.5590.2.5
F533EE36FFE9FFDDFF18D909FEF0018B.text	F533EE36FFE9FFDDFF18D909FEF0018B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trisinus qingliangmontis Ding & Niu & Yin 2025	<div><p>Trisinus qingliangmontis sp. nov.</p><p>Chinese common name: Oiåėñaȃş</p><p>(Figs 2, 3)</p><p>Type material (11 exx.). HOLOTYPE: CHINA: ♂: ‘ China: Zhejiang Prov., Qingliangfeng Mt., Luminyougu [Dzșṡö], alt. 900 m, 23-VII-2009, Feng, Yuan &amp; Yin leg.’ (SNUC) . PARATYPES: CHINA: 4 ♂♂, 1 ♀, same collecting data as for holotype; 1 ♂, same collecting data as for holotype, except ‘alt. 850 m, 20-VII-2009 ’; 1 ♂, same collecting data as for holotype, except ‘alt. 0.85 km, 22-VII-2009 ’; 1 ♂, 1 ♀, ‘ China: Zhejiang Prov., Linan City, Qianqingtang [千ṃ塘], 16.v.2012, 1000 m, Chen, Ma &amp; Zhao leg. ’; 1 ♀, ‘ China: Zhejiang Prov., Linan City, Qianqing Mt., Lelifeng, 17.v.2012, 1200 m, Chen, Ma &amp; Zhao leg.’ (all in SNUC) .</p><p>Diagnosis. Male. Body length approximately 1.7 mm. Head sub-rectangular; vertex greatly modified, with large U-shaped cavity between distinctly raised lateral projections, areas lateral to projections greatly concave, inside central cavity one lingulate protuberance; antenna simple, lacking modifications. Pronotum in addition to sulci with weak longitudinal discal carinae. Discal striae of elytra extending posteriorly to approximately apical 4/5 of elytral length. Aedeagus greatly asymmetric, ventral stalk composed of one elongate structure with two macrosetae and one broad, membranous part, dorsal lobe thin and serrate at apex, greatly twisted at middle. Female. Body length approximately 1.6–1.7 mm, head simple, lacking modifications, antenna shorter than for male, genitalia as in Fig. 2J.</p><p>Description. Male. Body (Fig. 2A) length 1.65–1.71 mm; color reddish-brown, with tarsi and mouthparts lighter. Dorsal surface finely punctate, covered with relatively long pubescence.</p><p>Head (Fig. 2B, C) sub-rectangular, subtruncate at base, much wider than long, length 0.37–0.39 mm, width across eyes 0.46–0.47 mm; vertex heavily modified, densely and coarsely punctate, lacking foveae (dorsal tentorial pits) and sulci due to modification; with large, roughly U-shaped central cavity between distinctly raised lateral projections, areas lateral to projections broadly and deeply concave; inside central cavity one small, thin, lingulate protuberance; lateral and posterior margin of cavity densely setose; tempora much shorter than eyes; antennal tubercles weakly raised, with setose postantennal foveae; frons distinctly punctate, broadly impressed between antennal tubercles, with moderately long setae at middle, anteriorly fused with clypeus; clypeus finely punctate, anterior margin carinate and moderately raised; lacking ocular-clypeal carinae. Venter with two small gular foveae (posterior tentorial pits) in single pit, lacking median carina. Eyes prominent, each composed of approximately 25 ommatidia. Maxillary palpus with palpomere 1 minute; 2 elongate, curved, pedunculate basally and enlarged apically; 3 short, sub-triangular; 4 fusiform, widest near middle. Antenna relatively short, lacking modifications, length 0.83–0.86 mm; indistinct club loosely formed by slightly enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–10 each submoniliform, 11 largest, slightly longer than 9 and 10 combined (21:17), sub-conical.</p><p>Pronotum (Fig. 2B) slightly shorter than wide, length 0.36–0.38 mm, width 0.40 mm, widest approximately at middle; semi-circular lateral sulci extending from dorsal surface laterally and anteriorly; disc weakly convex, coarsely punctate, median longitudinal sulcus longer than lateral sulci, with pair of thin discal carinae, lacking median antebasal fovea and mediobasal impression, lateral antebasal foveae connected by transverse antebasal sulcus, lacking antebasal tubercles; outer and inner pair of basolateral foveae present. Prosternum with basisternal (precoxal) portion slightly longer than procoxal rests; with small lateral procoxal foveae; oblique hypomeral grooves present at basal halves, lacking lateral antebasal hypomeral impressions; hypomeral ridges close to coxal cavities.</p><p>Elytra wider than long, length 0.51–0.52 mm, width 0.63 mm; roundly quadrate, truncate at bases, dorsal surface with relatively long pubescence; each elytron with two distinct, asetose basal foveae; discal striae long, extending from outer basal foveae posteriorly to 4/5 of elytral length; humeri denticulate, with tiny subhumeral foveae, sulcate marginal striae extending from foveae to posterior margins of elytra. Metathoracic wings vestigial.</p><p>Mesoventrite short, laterally fully demarcated from metaventrite by oblique ridges; median mesoventral foveae narrowly separated, originating from shared setose, transverse opening, lateral mesoventral foveae large and setose, forked internally; prepectus broadened, collar-shaped; mesoventral intercoxal process short, apically acute. Metaventrite slightly and broadly impressed at middle, with pair of large, setose lateral mesocoxal foveae and setose lateral metaventral foveae, metaventral intercoxal process narrowly and shallowly emarginate at middle.</p><p>Legs moderately elongate; mesotibia (Fig. 2D) with small blunt tubercle at apex.</p><p>Abdomen slightly narrower than elytra, widest at lateral margins of tergite 1 (IV), length 0.44–0.50 mm, width 0.54–0.55 mm. Tergite 1 (IV) (Fig. 2E) dorsally longer than 2–4 (V–VII) combined, basal sulcus interrupted by two pairs of basolateral foveae, discal carinae distinct, with complete and oblique inner marginal carinae; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) shorter than 2 and 3 combined along middle, tergite 5 (VIII) transverse, posterior margin roundly emarginate at middle, tergite 2–4 (V–VII) each with one pair of small basolateral foveae. Sternite 2 (IV) densely setose at base, with large, transverse mediobasal and two pair of basolateral foveae, lacking basal impression, with moderately long lateral carinae extending for almost half of sternal length; midlength of sternite 2 longer than 3–5 (V–VII) combined, 3–5 each short at middle, 3 and 4 (VI) with one pair of punctiform basolateral foveae, 5 lacking foveae, sternite 6 (VIII) transverse, posterior margin sinuate at middle, sternite 7 (IX) (Fig. 2F) suboval, weakly sclerotized, apex rounded and with few long setae along apical margin.</p><p>Aedeagus (Fig. 2G–I) 0.26 mm long, moderately sclerotized, dorsoventrally greatly asymmetric; median lobe with large basal capsule and suboval foramen, ventral stalk composed of one elongate structure gradually narrowing apically and curved rightward (orientation according to Fig. 2H), with two long macrosetae at middle, and another shorter projection broadest at base and narrowing apically; dorsal lobe much thinner than ventral stalk, narrowed at middle and broadening apically and basally, greatly twisted at middle, with serrate apex; parameres fused and reduced to ventral membrane.</p><p>Female. General external morphology similar to male; antenna shorter; head lacking modifications; legs simple; each eye composed of about 18 ommatidia; humeri moderately prominent; metathoracic wings lacking. Measurements (as for male): body length 1.61–1.65 mm, length/width of head 0.31–0.33 mm / 0.44 mm, pronotum 0.36 mm / 0.40 mm, elytra 0.45–0.46 mm / 0.60–0.61 mm, abdomen 0.46–0.52 mm / 0.58–0.59 mm; length of antenna 0.71–0.72 mm; genitalia transverse (Fig. 2J), 0.20 mm wide, posterior margin convex at middle.</p><p>Comparative notes. The primary distinction between this species and T. cavernosus sp. nov. described above is evident in the structure of their cephalic modifications. In this species, the lateral sides of the ridges encircling the central cavity are markedly concave. This feature contrasts sharply with T. cavernosus sp. nov., which exhibits laterally greatly prominent projections.</p><p>Distribution. East China: Zhejiang (Fig. 3B).</p><p>Etymology. The species epithet is a toponym, referring to the type locality of this species, Qingliangfeng Nature Reserve.</p></div>	https://treatment.plazi.org/id/F533EE36FFE9FFDDFF18D909FEF0018B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ding, Mei-Lin;Niu, Xiao-Ling;Yin, Zi-Wei	Ding, Mei-Lin, Niu, Xiao-Ling, Yin, Zi-Wei (2025): A new species group of Trisinus Raffray (Coleoptera: Staphylinidae: Pselaphinae), with description of two new species from the Tianmu Mountains, China. Zootaxa 5590 (2): 256-262, DOI: 10.11646/zootaxa.5590.2.5, URL: https://doi.org/10.11646/zootaxa.5590.2.5
