identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
FB4187964D16FFF257FEFD26FAD9F8EF.text	FB4187964D16FFF257FEFD26FAD9F8EF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagurixus acanthocarpus Komai & Okuno 2009	<div><p>Pagurixus acanthocarpus Komai &amp; Okuno, 2009</p> <p>(Figs 1A, B, 3A)</p> <p>[Japanese name: Konpeitou-himehonyadokari]</p> <p>Pagurixus acanthocarpus Komai &amp; Okuno, 2009: 146, figs 5–8, 9C [type locality: Kashiwa-jima, Otsuki, Kochi Prefecture, southern Japan, subtidal].— McLaughlin et al. 2010: 31 (list).</p> <p>Material examined. Tanega-shima, Osumi Group: stn <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=131.075&amp;materialsCitation.latitude=30.69" title="Search Plazi for locations around (long 131.075/lat 30.69)">Annou</a> 1, 30°41.4´N, 131°04.5´E, 7 m, 16 Aug. 2012, coll. T. Kawai, 1 female (sl 1.5 mm), RUMF-ZC 2321; stn <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=131.06334&amp;materialsCitation.latitude=30.673334" title="Search Plazi for locations around (long 131.06334/lat 30.673334)">Kurosenohana</a> 3, 30°40.4´N, 131°03.8´E, 7 m, 13–14 Aug. 2012, coll. T. Kawai, 1 male (sl 1.4 mm), 1 ovig. female (sl 1.8 mm), RUMF-ZC 2322; stn <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.94167&amp;materialsCitation.latitude=30.645" title="Search Plazi for locations around (long 130.94167/lat 30.645)">Sumiyoshi</a> 1, 30°38.7´N, 130°56.5´E, 7 m, 4–8 Aug. 2012, coll. T. Kawai, 1 female (sl 2.1 mm), RUMF-ZC 2323; stn <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.93834&amp;materialsCitation.latitude=30.666666" title="Search Plazi for locations around (long 130.93834/lat 30.666666)">Sumiyoshi</a> 2, 30°40.0´N, 130°56.3´E, 7 m, 4–8 Aug. 2012, coll. T. Kawai, 1 male (sl 1.8 mm), 1 ovig. female (sl 1.6 mm), RUMF-ZC 2324. Yaku-shima, Osumi Group: stn <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.47&amp;materialsCitation.latitude=30.231667" title="Search Plazi for locations around (long 130.47/lat 30.231667)">Kurio</a> 1-1, 30°13.9´N, 130°28.2´E, 7 m, 10 Aug. 2012, coll. T. Kawai, 1 male (sl 1.3 mm), RUMF-ZC 2325; stn <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.41333&amp;materialsCitation.latitude=30.273333" title="Search Plazi for locations around (long 130.41333/lat 30.273333)">Kurio</a> 3-1, 30°16.4´N, 130°24.8´E, 7 m, 10 Aug. 2012, coll. T. Kawai, 1 male (sl 1.2 mm), RUMF-ZC 2326. Amakusa, Kumamoto Prefecture, Kyushu main island: stn <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=129.975&amp;materialsCitation.latitude=32.15" title="Search Plazi for locations around (long 129.975/lat 32.15)">Ushibuka</a> 1-1, 32°09.0´N, 129°58.5´E, 3 Dec. 2012, coll. T. Kawai, 1 male (sl 1.7 mm), RUMF-ZC 2327.</p> <p>Coloration in life. The color pattern of the present specimens agrees well with that of P. acanthocarpus described and illustrated by Komai &amp; Okuno (2009, fig. 9C). See Fig. 1A, B.</p> <p>Distribution. Previously known only from Kashiwa-jima in Kochi Prefecture and Izu-oshima in the Izu Islands, Japan; subtidal to 8 m (Komai &amp; Okuno 2009). Now also recorded from Tanega-shima and Yaku-shima, Osumi Group in the northern Ryukyus, and Amakusa in the western coast of Kyushu main island, Japan; 7 m.</p> <p>Habitat. Rocky and coral reefs, on interspaces of rocks with algae and/or corals.</p> <p>Remarks. The present specimens agree well with the original description of P. acanthocarpus in the diagnostic aspects. The dorsal margin of the carpus of each second pereopod is somewhat protuberant, and exclusive of a dorsodistal spine, the margin has one to five (usually one or two), tiny or moderately small spines in males and is unarmed or armed with one or two tiny spines in females. In males, the coxa of the right fifth pereopod (Fig. 3A) has a short but distinct, collar-like “posteromesial protrusion” (see Komai &amp; Osawa 2006; Komai 2010), and the basal margin of its bearing prominent tuft of setae is more clearly demarcated than that of the holotype illustrated by Komai &amp; Okuno (2009, fig. 5I). The fourth pereopods are subequal in size on right and left and in both males and females; the setation is similar on right and left in males, but in females, setae on the mesial surfaces of the dactylus, propodus, carpus, and merus are longer and more pronounced on the left side than on the right side.</p> </div>	https://treatment.plazi.org/id/FB4187964D16FFF257FEFD26FAD9F8EF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Osawa, Masayuki;Kawai, Takashi;Sakamaki, Takashi	Osawa, Masayuki, Kawai, Takashi, Sakamaki, Takashi (2013): New records of Pagurixus Melin, 1939 (Crustacea: Decapoda: Paguridae) from the Ryukyu Islands, southwestern Japan, with description of a new species. Zootaxa 3669 (3): 350-366, DOI: 10.11646/zootaxa.3669.3.9
FB4187964D14FFF057FEFF45FE94FC50.text	FB4187964D14FFF057FEFF45FE94FC50.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagurixus carinimanus Komai & Osawa 2006	<div><p>Pagurixus carinimanus Komai &amp; Osawa, 2006</p> <p>(Fig. 2A)</p> <p>[New Japanese name: Azuki-himehonyadokari]</p> <p>Pagurixus cf. tweediei.— McLaughlin &amp; Haig 1984: 134, fig. 4.— Rahayu 2000: 394.— Paulay et al. 2003: 490.</p> <p>Pagurixus sp. 3 (slate).— Paulay et al. 2003: 490.</p> <p>Pagurixus carinimanus Komai &amp; Osawa, 2006: 71, figs 29–32, 45B, 48 [type locality: Ankyaba, Amami-oshima, Ryukyu Islands, 1 m].— McLaughlin et al. 2010: 31 (list).—Poupin et al. in press: 44, fig. 23B, C.</p> <p>Material examined. Kuchino-shima, Tokara Group: stn K 1-1, 29°59.1´N, 129°55.7´E, 7 m, 26 Jul. 2012, coll. T. Kawai, 1 female (sl 1.7 mm), RUMF-ZC 2328.</p> <p>Coloration in life. The coloration of the present specimen agrees well with that of P. carinimanus described by Komai &amp; Osawa (2006, fig. 45B). See Fig. 2A.</p> <p>Distribution. Ryukyu Islands, Guam, the Philippines in the western Pacific, and Mayotte and Glorioso Islands in the western Indian Ocean; depth range 1–30 m (Komai &amp; Osawa 2006; Poupin et al. in press). Until now, Amami-oshima was the reliable northern-most locality from which P. carinimanus was known (Komai &amp; Osawa 2006). The present specimen, collected from Kuchino-shima in the Tokara Group, extends the geographical range of the species slightly to the north.</p> <p>Habitat. Coral reef slope.</p> <p>Remarks. Although the present specimen lacks the left cheliped and second pereopod (the left third pereopod is not shown in Fig. 2A), it agrees well with the original description of P. carinimanus in all other morphological characters.</p> </div>	https://treatment.plazi.org/id/FB4187964D14FFF057FEFF45FE94FC50	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Osawa, Masayuki;Kawai, Takashi;Sakamaki, Takashi	Osawa, Masayuki, Kawai, Takashi, Sakamaki, Takashi (2013): New records of Pagurixus Melin, 1939 (Crustacea: Decapoda: Paguridae) from the Ryukyu Islands, southwestern Japan, with description of a new species. Zootaxa 3669 (3): 350-366, DOI: 10.11646/zootaxa.3669.3.9
FB4187964D14FFF657FEFC0EFED7F80A.text	FB4187964D14FFF657FEFC0EFED7F80A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagurixus fasciatus Komai & Myorin 2005	<div><p>Pagurixus fasciatus Komai &amp; Myorin, 2005</p> <p>(Figs 1C, D, 3B–E)</p> <p>[Japanese name: Kashiwajima-himehonyadokari]</p> <p>Pagurixus fasciatus Komai &amp; Myorin, 2005: 2, figs 1–4 [type locality: Kashiwa-jima, Otsuki, Kochi Prefecture, southern Japan, 10–15 m].—Okuno et al. 2006: 156, plate 3E, F.— Okuno &amp; Arima 2006: 36.— Komai 2010: 1287.— McLaughlin et al. 2010: 31 (list).</p> <p>Material examined. Tanega-shima, Osumi Group: stn Kurosenohana 3, 30°40.4´N, 131°03.8´E, 7 m, 13–14 Aug. 2012, coll. T. Kawai, 1 male (sl 0.9 mm), RUMF-ZC 2329; stn Nishino-omote 1, 30°44.6´N, 130°59.4´E, 7 m, 3– 7 Aug. 2012, coll. T. Kawai, 1 male parasitized by peltogastrid rhizocephalan (sl 1.2 mm), RUMF-ZC 2330; stn Sumiyoshi 2, 30°40.0´N, 130°56.3´E, 7 m, 4–8 Aug. 2012, coll. T. Kawai, 1 male (sl 1.7 mm), 1 female (sl 1.4 mm), RUMF-ZC 2331; stn Sumiyoshi 3, 30°41.9´N, 130°57.6´E, 7 m, 4–8 Aug. 2012, coll. T. Kawai, 1 male (sl 1.5 mm), RUMF-ZC 2332. Yaku-shima, Osumi Group: stn Kurio 1-1, 30°13.9´N, 130°28.2´E, 7 m, 10 Aug. 2012, coll. T. Kawai, 1 male (sl 1.1 mm), 1 female (sl 1.1 mm), RUMF-ZC 2333.</p> <p>Coloration in life. The coloration of the present specimens agrees well with that of P. fasciatus described by Komai &amp; Myorin (2005: fig. 1). See Fig. 1C, D.</p> <p>Distribution. Previously known only from Kashiwa-jima (Kochi Prefecture), Izu Peninsula, and Izu and Ogasawara Islands, Japan; intertidal to 30 m (Komai 2010). Now also recorded from Tanega-shima and Yakushima, Osumi Group in the northern Ryukyu Islands; 7 m.</p> <p>Habitat. Rocky reefs, on interspaces of rocks with algae and/or corals.</p> <p>Remarks. The present specimens generally agree well with the original description of P. fasciatus by Komai &amp; Myorin (2005) in both morphology and coloration. Nevertheless, they have variations in the armature on the carpi of the second pereopods. Komai &amp; Myorin (2005) described and illustrated P. fasciatus to have the carpi of the second pereopods unarmed on the dorsal margin, except for a dorsodistal spine. However, in the males of the present Ryukyu material, the dorsal margin of each carpus is slightly protuberant and either unarmed or armed with a tiny spine proximal to the midlength (Fig. 3B, C), this spine is more pronounced in the smallest specimen (sl 0.9 mm, RUMF-ZC 2329); whereas in the females, the dorsal margin always bears a small spine (Fig. 3D, E).</p> <p>At our request, Dr. T. Komai kindly re-examined the holotype and three paratypes of P. fasciatus and two specimens of that species reported by Komai (2010). The holotype male (CBM-ZC 7980 in the collection of the Natural History Museum and Institute, Chiba) has a small dorsal spine on the carpus of the right second pereopod, which is visible only in the mesial view, but such a spine is absent on the left side. In three paratypes (CBM-ZC 7981), the spine is absent or reduced to a minute protuberance in two males, while it is distinct in one female. Two male specimens from the Ogasawara Islands (CBM-ZC 9646 and 9650) lack a dorsal spine on the carpus of each second pereopod. Thus, the armature on the carpus appears to vary in the previous reported material as in the present material from the Ryukyu Islands, and the dorsal spine tends to be reduced in males.</p> <p>In addition to the variation of the armature on the carpi of the second pereopods, a male specimen from Tanega-shima (RUMF-ZC 2332) has a small submedian spine on the dorsal margin of the carpus of the right third pereopod. Such a spine is absent on the carpus of the left third pereopod in the same specimen and on the carpi of both third pereopods in all other specimens. In all the specimens examined, the ventorolateral margin of the merus of the right cheliped has a row of small but distinct spines, although the margin was described as smooth or finely denticulate in the type specimens (Komai &amp; Myorin 2005). The fourth pereopods are subequal in size on right and left and in both males and females; the setation is similar on right and left in males, but in females, setae on the mesial surfaces of the dactylus, propodus, carpus, and merus are longer and more pronounced on the left side than on the right side.</p> </div>	https://treatment.plazi.org/id/FB4187964D14FFF657FEFC0EFED7F80A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Osawa, Masayuki;Kawai, Takashi;Sakamaki, Takashi	Osawa, Masayuki, Kawai, Takashi, Sakamaki, Takashi (2013): New records of Pagurixus Melin, 1939 (Crustacea: Decapoda: Paguridae) from the Ryukyu Islands, southwestern Japan, with description of a new species. Zootaxa 3669 (3): 350-366, DOI: 10.11646/zootaxa.3669.3.9
FB4187964D11FFF557FEFF45FD4DF9D4.text	FB4187964D11FFF557FEFF45FD4DF9D4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagurixus formosus Komai 2010	<div><p>Pagurixus formosus Komai, 2010</p> <p>(Figs 2B, C, 3F)</p> <p>[New Japanese name: Chairo-himehonyardokari]</p> <p>Pagurixus formosus Komai, 2010: 1306, figs 17–19 [type locality: Shiangianwan, Kenting, Pingtung County, southern Taiwan, 3–8 m].— McLaughlin et al. 2010: 31 (list).— Komai &amp; Rahayu 2013: 134.</p> <p>Material examined. Kume-jima: “KUMEJIMA 2009” stn Dive 28 (Oyako-iwa), 26°17.5´N, 126°47.9´E, 0–20 m, 17 Nov. 2009, 1 male (sl 2.7 mm), RUMF-ZC 2334. Okinawa-jima: Benoki, Kunigami, intertidal, 30 Apr. 2010, coll. M. Osawa, 1 male (sl 2.9 mm), RUMF-ZC 2335.</p> <p>Coloration in life. Carapace whitish; shield with several brown patches: 2 pairs on lateral parts, 1 pair on anterior submedian part, and 1 on posterior median part; posterior carapace with numerous, small red spots and 1 pair of longitudinal brown lines on submedian part. Ocular peduncles translucent, with brown tinge proximally and brown marking at base of cornea; ocular acicles brown proximally. Antennular peduncles with subdistal brown band on penultimate segment; ultimate segment entirely dark brown; dorsal flagella pale blue proximally, turning pale red distally. Antennal peduncle generally translucent; second and third segments each with large brown marking; fourth segment with subdistal brown ring; fifth segment marginally brown; flagellum generally translucent, dark brown on margins. Right and left chelipeds brown; meri each with 2 white bands on median and subdistal parts, median band on right merus either interrupted or uninterrupted on dorsal surface. Ambulatory legs brown; meri each with 2 irregular, white bands on median and subdistal parts, median band of each third pereopod interrupted on midline. See Fig. 2B, C.</p> <p>Distribution. Previously known only from Kenting, Pingtung County in the southern Taiwan, and Panglao Island, the Philippines; 2–8 m (Komai &amp; Rahayu 2013). Now also recorded from Japan, Okinawa-jima and Kumejima in the central Ryukyus; intertidal to 20 m.</p> <p>Habitat. Rocky reefs, under rocks and dead coral rubble.</p> <p>Remarks. Komai (2010) described the coloration of P. formosus based on preserved material. The coloration of fresh specimens agrees well with Komai’s (2010) description, particularly in the band pattern of the chelipeds and ambulatory legs. The morphological characters of the present Ryukyu specimens also agree well with the original description, although the dorsomesial margin of the carpus of the left cheliped has a row of small spines rather than tubercles. The male from Kume-jima (RUMF-ZC 2334) has a relatively larger left cheliped (Fig. 2C) and more elongate and slender dactyli of the ambulatory legs compared to the holotype (Komai 2010: fig. 17C–F). The dactylus of the third pereopod is slightly longer than the propodus and about seven times length of its width (Fig. 3F).</p> <p>As discussed by Komai (2010), P. formosus appears closest to P. tweediei in both morphology and coloration. The former species is also closely similar to P. nomurai in the color pattern of the entire body, although the color of the chelipeds and ambulatory legs may differ between the two species. The pereopods are generally brown in P. formosus, instead of bluish gray or bluish brown in P. nomurai. Pagurixus formosus is readily distinguished from P. nomurai in morphology, particularly in the corneas not strongly dilated and the carpus of the left cheliped and dactyli of the ambulatory legs being much stouter.</p> </div>	https://treatment.plazi.org/id/FB4187964D11FFF557FEFF45FD4DF9D4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Osawa, Masayuki;Kawai, Takashi;Sakamaki, Takashi	Osawa, Masayuki, Kawai, Takashi, Sakamaki, Takashi (2013): New records of Pagurixus Melin, 1939 (Crustacea: Decapoda: Paguridae) from the Ryukyu Islands, southwestern Japan, with description of a new species. Zootaxa 3669 (3): 350-366, DOI: 10.11646/zootaxa.3669.3.9
FB4187964D11FFF457FEF98EFE37FD70.text	FB4187964D11FFF457FEF98EFE37FD70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagurixus pseliophorus Komai & Osawa 2006	<div><p>Pagurixus pseliophorus Komai &amp; Osawa, 2006</p> <p>(Fig. 2E, F)</p> <p>[Japanese name: Samidare-himehonyadokari]</p> <p>Pagurixus pseliophorus Komai &amp; Osawa, 2006: 85, figs 36–40, 46A, 48 [type locality: Ara, Kume-jima, Ryukyu Islands, intertidal].— Okuno &amp; Arima 2006: 37, fig. 3E.— Okuno 2008: 9, fig. 2.— McLaughlin et al. 2010: 32 (list), fig. 14H.</p> <p>Material examined. Okinawa-jima: Benoki, Kunigami, intertidal, 30 Apr. 2010, coll. M. Osawa, 1 ovig. female (sl 2.7 mm), RUMF-ZC 2336; Benoki, Kunigami, intertidal, 6 Sep. 2012, coll. R. Yoshida, 2 males (sl 2.2 mm), 3 females (sl 2.3–3.2 mm), RUMF-ZC 2337; Zatsun, Kunigami, 0.5 m, 29 Aug. 2010, coll. M. Osawa, 1 male (sl 3.2 mm), 1 female (sl 3.0 mm), 2 ovig. females (sl 2.7, 3.4 mm), RUMF-ZC 2338; Zatsun, Kunigami, intertidal, 8 May 2012, coll. M. Osawa, 2 males (sl 2.4, 2.6 mm), 2 females (sl 1.7, 1.8 mm), 1 ovig. female (sl 1.8 mm), RUMF-ZC 2339.</p> <p>Coloration in life. The coloration of the present specimens agrees well with that of P. pseliophorus described by Komai &amp; Osawa (2006, fig. 46A) and a photograph provided by Okuno &amp; Arima (2006, fig. 3E). The color pattern of the carpus of the right cheliped is different between males and females. In females, the carpus has an irregular dark brown band on the median portion, whereas this band occupies the entire or almost the entire portion of the carpus in males. See Fig. 2E, F.</p> <p>Distribution. Presently known only from the Ryukyu and Izu Islands, Izu Peninsula, and Kochi Prefecture in southern Japan; intertidal to 2 m (Okuno 2008).</p> <p>Habitat. Rocky reefs, usually under rocks, sometimes on the sides of large rocks.</p> <p>Remarks. The previous record of P. pseliophorus from the Ryukyu Islands was based on the holotype and two paratypes from Kume-jima, which is currently the southern-most locality of the species. The present specimens were collected from Okinawa-jima, located slightly east to Kume-jima. On the rocky intertidal or shallow subtidal reefs in the northeastern coast of Okinawa-jima, P. pseliophorus is commonly found sympatrically with P. formosus and P. tweediei.</p> </div>	https://treatment.plazi.org/id/FB4187964D11FFF457FEF98EFE37FD70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Osawa, Masayuki;Kawai, Takashi;Sakamaki, Takashi	Osawa, Masayuki, Kawai, Takashi, Sakamaki, Takashi (2013): New records of Pagurixus Melin, 1939 (Crustacea: Decapoda: Paguridae) from the Ryukyu Islands, southwestern Japan, with description of a new species. Zootaxa 3669 (3): 350-366, DOI: 10.11646/zootaxa.3669.3.9
FB4187964D10FFF457FEFD2EFACEF841.text	FB4187964D10FFF457FEFD2EFACEF841.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagurixus tweediei (Forest 1956)	<div><p>Pagurixus tweediei (Forest, 1956)</p> <p>(Fig. 2D)</p> <p>[New Japanese name: Suneobi-himehonyardokari]</p> <p>Eupagurus (Pagurixus) tweediei Forest, 1956: 50, pl. 4 [type locality: Cocos (Keeling) Islands].</p> <p>Pagurus (Pagurixus) tweediei.— Dechancé 1964: 38.— Ball &amp; Haig 1972: 103.— Lewinsohn 1982: 58.</p> <p>? Pagurixus tweediei.— Morgan 1992: 171.</p> <p>? Pagurixus cf. tweediei.— Gherardi &amp; McLaughlin 1994: 644.</p> <p>Pagurixus tweediei.— Davie 2002: 82 (in part).— Komai &amp; Osawa 2006: 25, figs 8–11, 44B, 47.— McLaughlin et al. 2010: 32 (list).</p> <p>Material examined. Kume-jima: “KUMEJIMA 2009” stn Dive 28 (Oyako-iwa), 26°17.5´N, 126°47.9´E, 0–20 m, 17 Nov. 2009, 1 male (sl 2.0 mm), RUMF-ZC 2340; “KUMEJIMA 2009” stn Trawl 29, 26°17.6´N, 126°52.9´E, 90.0– 91.3 m, 13 Nov. 2009, 1 female (sl 2.1 mm), RUMF-ZC 2341. Okinawa-jima: Zatsun, Kunigami, 0.5 m, 29 Aug. 2010, coll. M. Osawa, 1 male (sl 2.5 mm), RUMF-ZC 2342; Zatsun, Kunigami, intertidal, 8 May 2012, coll. M. Osawa, 2 ovig. females (sl 2.0, 2.1 mm), RUMF-ZC 2343; Uza, Yomitan, small cave, intertidal, 22 Nov. 2012, coll. R. Yoshida, 1 male (sl 3.2 mm), 1 female (sl 2.8 mm), RUMF-ZC 2344.</p> <p>Coloration in life. The coloration of the present specimens agrees well with that of P. tweediei described by Komai &amp; Osawa (2006, fig. 44B). See Fig. 2D.</p> <p>Distribution. Previously known with certainty from Cocos (Keeling) Islands in the eastern Indian Ocean, and Ogasawara Islands, Guam, Society Islands in the western Pacific Ocean; intertidal to 7 m (Komai &amp; Osawa 2006). Now also recorded from Okinawa-jima and Kume-jima in the central Ryukyu Islands; intertidal to 91.3 m. One female specimen from Kume-jima (RUMF-ZC 2341) was collected from rather unusual depth (90.0– 91.3 m) for P. tweediei, which typically inhabits intertidal and shallow subtidal areas.</p> <p>Habitat. Rocky and coral reefs, under rocks and dead coral rubble, or in small shoreside cave near coral reefs.</p> <p>Remarks. Until the present report, P. tweediei has not been recorded from the Ryukyu Islands, although this species is widely distributed from the eastern Indian Ocean to the western Pacific Ocean (Komai &amp; Osawa 2006; Osawa &amp; Komai 2007). The previous record of P. tweediei from Japan was based exclusively on the material from the remote Ogasawara Islands (Komai &amp; Osawa 2006).</p> <p>Komai (2010) discussed that the presence of a ventrolateral distal spine on the merus of the third pereopod is one of morphological characters distinguishing P. tweediei from P. formosus. The type material of P. formosus lacks this spine. However, in the present specimens referred to P. tweediei, this character appears to be variable. The spine is absent or reduced to a low protuberance in one male from Kume-jima (RUMF-ZC 2340) and two females from Okinawa-jima (RUMF-ZC 2343), whereas it is present but very small in the other specimens.</p> </div>	https://treatment.plazi.org/id/FB4187964D10FFF457FEFD2EFACEF841	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Osawa, Masayuki;Kawai, Takashi;Sakamaki, Takashi	Osawa, Masayuki, Kawai, Takashi, Sakamaki, Takashi (2013): New records of Pagurixus Melin, 1939 (Crustacea: Decapoda: Paguridae) from the Ryukyu Islands, southwestern Japan, with description of a new species. Zootaxa 3669 (3): 350-366, DOI: 10.11646/zootaxa.3669.3.9
FB4187964D1FFFFD57FEFF45FEE4F96C.text	FB4187964D1FFFFD57FEFF45FEE4F96C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pagurixus vicinus Osawa & Kawai & Sakamaki 2013	<div><p>Pagurixus vicinus sp. nov.</p> <p>(Figs 1E, F, 4–6)</p> <p>[New Japanese name: Irowake-himehonyardokari]</p> <p>Type material. Holotype. Akuseki-jima, Tokara Group: stn A 1-1, 29°26.8´N, 129°35.6´E, 7 m, 23 Jul. 2012, coll. T. Kawai, male (sl 1.4 mm), RUMF-ZC 2345.</p> <p>Paratypes. Kuchino-shima, Tokara Group: stn K 2-1, 30°00.5´N, 129°55.0´E, 7 m, 26 Jul. 2012, coll. T. Kawai, 3 males (sl 0.9–1.4 mm), 1 female (sl 0.9 mm), 2 ovig. female (sl 1.1, 1.2 mm), RUMF-ZC 2346; stn K 2-2, 30°00.5´N, 129°55.0´E, 7 m, 27 Jul. 2012, coll. T. Kawai, 3 females (sl 1.2–1.4 mm), RUMF-ZC 2347. Akusekijima, Tokara Group: stn A 2-2, 29°27.3´N, 129°37.2´E, 7 m, 24 Jul. 2012, coll. T. Kawai, 1 male (sl 0.9 mm), 1 female (sl 1.1 mm), RUMF-ZC 2348; stn A 3-2, 29°28.4´N, 129°36.5´E, 7 m, 24 Jul. 2012, coll. T. Kawai, 1 male (sl 1.0 mm), RUMF-ZC 2349.</p> <p>Additional material. Kuchino-shima, Tokara Group: stn K 1-1, 29°59.1´N, 129°55.7´E, 7 m, 26 Jul. 2012, coll. T. Kawai, 1 female (sl 0.9 mm), RUMF-ZC 2350; stn K 1-2, 29°59.1´N, 129°55.7´E, 7 m, 27 July 2012, coll. T. Kawai, 1 female (sl 1.2 mm), RUMF-ZC 2351. Akuseki-jima, Tokara Group: stn A 3-1, 29°28.4´N, 129°36.5´E, 7 m, 23 Jul. 2012, coll. T. Kawai, 1 ovig. female (sl 1.2 mm), RUMF-ZC 2352. Takara-jima, Tokara Group: stn T 1-1, 29°07.7´N, 129°13.3´E, 7 m, 20 Jul. 2012, coll. T. Kawai, 1 female (sl 1.3 mm), RUMF-ZC 2353; stn T 2- 1, 29°09.5´N, 129°13.3´E, 7 m, 20 Jul. 2012, coll. T. Kawai, 1 female (sl 1.1 mm), RUMF-ZC 2354.</p> <p>Description. Shield (Fig. 4A) approximately 1.1 times longer than wide; anterior margin between rostrum and lateral projections concave; anterolateral margins sloping; posterior margin roundly truncate; dorsal surface smooth, with few tufts of short setae laterally. Rostrum triangular, reaching nearly to midlength of ocular acicles, moderately broad, terminating bluntly or acutely. Lateral projections obtuse, each with submarginal spinule.</p> <p>Ocular peduncles (Fig. 4A) moderately stout, 0.7 length of shield, each with few tufts of short setae on dorsomesial surface; middle portion slightly inflated on mesial surface, subequal to corneal width; corneas not dilated, corneal width 0.3 of peduncular length. Ocular acicles subtriangular, each with submarginal spinule.</p> <p>Antennular peduncles (Fig. 4A) overreaching distal corneal margins by 0.2–0.3 lengths of ultimate segments. Ultimate segment with tuft of few setae near dorsolateral distal angle, ventral surface lacking longitudinal rows of setae. Basal segment with small spine on distal margin of statocyst lobe. Ventral flagellum with few short setae on lateral and mesial margins.</p> <p>Antennal peduncles (Fig. 4A) overreaching distal corneal margins by 0.2–0.3 lengths of fifth segments. Fifth and fourth segments with few tufts of short setae. Third segment with small spine at distomesial angle. Second segment with or without minute spine at dorsomesial distal angle; laterodistal projection not reaching midlength of fourth segment, terminating in simple or bifid spine. First segment unarmed on lateral face; ventromesial distal margin strongly produced, with small spine. Antennal acicle arcuate, terminating in small spine, overreaching base but not reaching distal margin of cornea; mesial margin with row of sparse setae.</p> <p>Third maxilliped moderately stout; ischium with well developed, crista dentata composed of row of small corneous teeth and 1 accessory tooth; merus and carpus unarmed; exopod reaching distal margin of carpus.</p> <p>Male right cheliped (Figs 4B, C, E, 5A, B, 6A, B) large, not particularly elongate. Chela subovate in dorsal view, 1.7–1.9 times as long as wide. Dactylus slightly longer than or equal in length to palm, terminating in small corneous claw; dorsomesial margin not delimited; dorsal, ventral, and mesial surfaces with scattered small granules and short setae, obsolete longitudinal ridge present proximally on mesial surface; cutting edge with 3 blunt calcareous teeth in proximal 0.7 and row of short corneous teeth in remaining margin. Palm 1.0–1.1 times length of carpus; convex dorsal surface generally covered with small granules; dorsolateral margin delimited by weak or distinct granular ridge extending onto fixed finger; dorsomesial margin also delimited by series of short granular ridges or sharp, generally uninterrupted ridge; lateral surface with scattered, coarse granules; mesial and ventral surfaces also with sparse granules. Fixed finger terminating in small calcareous or corneous claw; cutting edge with row of blunt calcareous teeth, median tooth prominent. Carpus nearly equal in length to merus, widened distally, length 1.0–1.1 times of distal width; dorsal surface with several or many, short transverse ridges distally and with sparse, short, bristle-like stiff setae and short simple setae, irregular row of small to moderately large spines present adjacent to proximal half of dorsomesial margin, dorsolateral surface rounded; dorsomesial margin delimited by row of short, somewhat elevated ridges bearing bristle-like stiff setae and with or without small spine near distal end; dorsodistal margin with row of small low tubercles; lateral surface with very short vertical ridges ventrally, ventrolateral margin with row of obsolete tubercles distally; mesial surface nearly smooth, ventromesial margin with row of low tubercles or granules distally; ventral surface convex, with sparse setae distally. Meral-carpal articulation lacking any pronounced clockwise rotation. Merus with median row of short and very short transverse ridges on dorsal surface, dorsodistal margin with few short setae; lateral face nearly smooth, ventrolateral margin distally with row of small sharp or blunt spines; mesial face also nearly smooth, convex ventromesial margin slightly protuberant but unarmed; ventral surface smooth, with short setae. Ischium with smooth ventromesial margin; surfaces unarmed, with sparse short setae.</p> <p>Female right cheliped (Figs 4D, F, 5C, D, 6C) relatively slender for genus, stouter but only slightly longer than or equal in length to left. Chela elongate subovate in dorsal view, 1.7–2.1 times longer than wide, with arched lateral margin. Dactylus longer than palm, terminating in small corneous claw; surfaces bearing sparse short setae; dorsal surface with few short ridges proximally, dorsomesial margin weakly or distinctly delimited by blunt ridge of low tubercles; ventral surface nearly smooth; cutting edge with row of small corneous teeth. Palm distinctly shorter than carpus; dorsal surface weakly convex, with slightly elevated row of small tubercles adjacent to dorsomesial margin and irregular double row of tubercles lateral to midline, some low tubercles also present on proximal and distal parts, tubercles occasionally bearing short setae; dorsolateral margin delimited by distinct tuberculate ridge extending onto midlength of fixed finger; dorsomesial margin also delimited by distinct, generally uninterrupted tuberculate ridge; lateral surface with scattered, short ridges; mesial surface with some oblique ridges dorsally; ventral surface slightly protuberant, with sparse short setae. Fixed finger terminating in small corneous claw; cutting edge with row of tiny calcareous teeth and interspersed with short corneous teeth. Carpus subequal in length to merus, widened distally, length 1.7–1.9 times of distal width; dorsal surface laterally with row of short low ridges, mesially with 3 distal spines followed by slightly elevated, short ridges, each spine and ridge occasionally with bristle-like stiff seta or short simple setae, dorsodistal margin with small low tubercles; mesial surface dorsally with longitudinal row of slightly elevated, short ridges each bearing bristle-like stiff seta or simple setae; lateral surface with scattered, short vertical ridges, ventrolateral margin with 2 or 3 small spines distally; mesial surface with few short ridges dorsally and ventrally, ventromesial margin unarmed; ventral surface convex, with few short setae. Meral-carpal articulation lacking any pronounced clockwise rotation. Merus with median row of short transverse ridges on dorsal surface, dorsodistal margin with few short setae; lateral face with scattered, very short vertical ridges, ventrolateral margin with 2 or 3 small spines on distal half; mesial face nearly smooth, ventromesial margin unarmed; ventral surface slightly convex, with few short setae. Ischium nearly smooth, with sparse setae.</p> <p>Left cheliped (Figs 4G, H, 5E, F) similar in both males and females, moderately short and stout. Chela elongate subovate in dorsal view, 2.2–2.6 times longer than wide. Dactylus distinctly longer than palm, terminating in small corneous claw; surfaces with scattered tufts of setae, ventral setae long; dorsal, ventral, and mesial surfaces rounded; mesial surface also with few short ridges proximally; cutting edge with row of sharp corneous teeth. Palm 0.5–0.6 length of carpus; dorsal surface convex, with lateral and mesial rows of small spines or tubercles, mesial row somewhat elevated, dorsolateral surface with some small tubercles; dorsolateral and dorsomesial margins not delimited; lateral and mesial surfaces with short, oblique ridges bearing short to moderately long setae; ventral surface weakly convex, with sparse tufts of setae distally. Fixed finger terminating in small corneous claw, cutting edge with row of small corneous teeth. Carpus somewhat compressed laterally, shorter than merus; length 1.9–2.4 times of width and 1.6–1.8 times of height; dorsal surface flattish, dorsolateral and dorsomesial margins each with 2 or 3 small distal spines followed by short ridges each bearing 1 or 2 bristle-like stiff setae, dorsodistal margin with 1 or 2 small spines flanked by spines in dorsolateral and dorsomesial rows; lateral face nearly perpendicular, dorsally with few short ridges each bearing bristle-like stiff seta and ventrally with scattered, very short ridges, ventrolateral margin with 1–3 small spines distally; mesial face also dorsally with short ridges each bearing bristlelike stiff seta and few simple setae, ventromesial distal margin slightly protuberant; ventral surface convex, with sparse short setae, ventrodistal margin with row of moderately long setae. Merus with median row of short obsolete ridges on dorsal surface, dorsodistal margin unarmed but bearing few short setae; lateral surface with many, very short vertical ridges, ventrolateral margin with row of small spines and moderately long setae in distal half; mesial surface nearly smooth, bearing few short setae, ventromesial margin unarmed but with row of moderately long setae; ventral surface weakly convex, smooth. Ischium unarmed, with sparse short setae.</p> <p>Ambulatory legs (Fig. 6D–K) similar on right and left (right slightly longer than left) and between males and females, moderately slender. Dactyli 0.7–0.9 length of propodi, 4.6–5.2 (second pereopods) or 5.1–6.4 (third pereopods) times longer than wide, terminating in large corneous claws; dorsal surfaces with few short setae; lateral and mesial faces also with sparse tufts of short setae adjacent to dorsal and ventral margins; ventral margins each with 5–7 slender corneous spines increasing in size distally. Propodi not tapering distally, 3.7–4.3 (second pereopods) and 3.9–5.1 (third pereopods) times longer than wide; dorsal surfaces nearly smooth, with row of sparse short setae; lateral and mesial faces also with few short setae dorsally and ventrally; ventral margins weakly curved, each with 1–4 (usually 1 or 2, right second pereopod), 0–2 (usually 1 or 2, left second), 1–7 (usually 1 or 2, right third), and 1 or 2 (left third) corneous spinules; ventrodistal margin with 1 or 2 corneous spines larger than spines on ventral margin. Carpi each with small dorsodistal spine; dorsal surfaces unarmed but slight protuberant, bearing sparse short setae; lateral and mesial faces also with few short setae; ventrodistal margins rounded, not produced. Meri each with row of short low ridges bearing sparse short setae on dorsal margin; lateral and mesial surfaces nearly smooth; ventral margins somewhat inflated on distal third, ventrolateral margins each with small subdistal spine (second pereopods) or unarmed (third pereopods).</p> <p>Fourth pereopods (Fig. 4I) subequal in size on right and left and between males and females; in males, setation similar on right and left, but in females, setae somewhat longer on left. Dactyli moderately broad, not curved, terminating in small corneous claws; dorsal margins with sparse setae. Propodi each with tuft of few setae on dorsal margin; mesial faces nearly flat; propodal rasp composed of single row of corneous scales. Carpi with sparse setae on margins.</p> <p>Coxae of fifth pereopods in males (Fig. 4K) unequal and dissimilar. Right coxa with short but distinct, collarlike posteromesial protrusion; prominent tuft of long stiff setae arising from ventrodistal margin of protrusion, reaching to left coxa; papilla-like vas deferens present at top of posteromesial protrusion. Left coxa with gonopore encircled by short setae, no protrusion of vas deference seen. Female with unpaired left gonopore.</p> <p>Anterior lobe of sixth thoracic sternite (Fig. 4J) subrectangular, with row of short to long setae anteriorly; anterolateral angles rounded. Eighth thoracic sternite (Fig. 4K) composed of 2 subequal, closely-set rounded lobes; ventral surface of each lobe flattened, with sparse short setae anteriorly.</p> <p>Pleon without distinctive features. Telson (Fig. 4L) with terminal margins oblique or nearly horizontal and each bearing 2–4 small spines; dorsal surface adjacent to terminal margins slightly elevated, well calcified.</p> <p>Coloration in life. Carapace translucent; shield with pair of small brown spots posterior to base of ocular peduncles and brown marking on posterior part; posterior carapace without distinct color markings. Ocular peduncles translucent, with brown tinge proximally and irregular brown markings on lateral and mesial surfaces; ocular acicles whitish. Antennular peduncles with submedian brown band on penultimate segment; ultimate segment with 2 brown rings on proximal and distal portions, median portion and proximal and distal marginal portions colorless; flagella pale orange. Antennal peduncle generally translucent; second, third, and fourth segments each with submedian brown marking; fifth segment marginally brown; flagellum translucent, light brown on margins. Right cheliped generally white; palm and carpus with some brown spots or blotches on dorsal and mesial surfaces; merus dorsally with narrow, subdistal brown ring and subproximal brown blotch. Left cheliped white; dactylus and fixed finger with tingle of pale brown; palm and carpus with some brown spots on dorsal and mesial surfaces; merus dorsally with narrow, subdistal brown ring and subproximal brown blotch. Ambulatory legs banded with white and dark brown; dactyli dark brown in proximal 0.4–0.5 (in holotype, small white blotch present on dorsal margin), remaining portion white; propodi each with broad, dark brown band occupying about proximal 0.7, band sometimes bearing white blotch dorsally; carpi with brown blotches on proximal 0.7; meri each with 2 complete or incomplete bands distally and proximally; ischia each with brown blotches. See Fig. 1E, F.</p> <p>Distribution. Presently known only from Akuseki-jima, Kuchino-shima, and Takara-jima, Tokara Group in the northern Ryukyus; 7 m.</p> <p>Habitat. Coral reef slopes.</p> <p>Etymology. The specific name is derived from the Latin, vicinus (= near), referring to the similarity in both morphology and coloration between the new species and P. fasciatus.</p> <p>Remarks. Male specimens of the new species appear to have variations in the degree of the marginal delimitation on the right chela and the shape of the anterior lobe of the sixth thoracic sternite. The dorsolateral margin of the right chela is delimited by a weak granular ridge in the holotype, whereas the ridge is distinct in other males. The dorsomesial margin of the right palm in the holotype has a series of short granular ridges, instead of a sharp, generally uninterrupted ridge observed in the others. The anterior lobe of the sixth thoracic sternite is broader in the holotype compared to the other males. However, all males of the type series are undoubtedly conspecific because of the same color pattern.</p> <p>McLaughlin &amp; Haig (1984) divided the genus Pagurixus into two informal species groups based on the presence or absence of distinct row(s) of setae on the ventral surface of the ultimate segment of the antennular peduncle. Later, Komai &amp; Osawa (2006) named them “ P. boninensis ” and “ P. anceps ” species groups. Pagurixus vicinus sp. nov. is referred to the P. anceps species group, which is characterized by the absence of longitudinal rows or series of setae on the ventral surface of the antennular ultimate segment. In this species group, P. vicinus sp. nov. closely resembles P. fasciatus. Shared characters are: ocular peduncles relatively short and exceeded by antennular and antennal peduncles; right cheliped with distinct dorsomesial spines but unarmed dorsolaterally on carpus in males and females; male right cheliped massive, without distinctly delimited dorsomesial margin on dactylus but bearing delimited granular ridge on dorsomesial margin of palm; left cheliped with dorsolateral and ventrolateral spines on carpus and ventrolateral row of spines on merus; third and fourth pereopods generally similar on right and left; coxa of right fifth pereopod in males with short but distinct, collar-like posteromesial protrusion; and female with unpaired left gonopore. In addition to these morphological features, the color patterns of the new species and P. fasciatus are also very similar, particularly in the possession of a broad dark brown band occupying about proximal 0.7 of the propodus of each ambulatory leg. However, P. vicinus sp. nov. is distinguishable from P. fasciatus by the armature on the left cheliped and right second pereopod based on the comparisons of the present specimens of the two species and with the illustrations of the type specimens of P. fasciatus by Komai &amp; Myorin (2005). The carpus of the left cheliped is armed with fewer spines on the ventrolateral margin in the new species than in P. fasciatus (one to three versus four to six spines). The ventral margins of the dactylus and propodus of the right second pereopod also have fewer spines in P. vicinus sp. nov. than in P. fasciatus, i.e., five or six versus seven to nine spines on the dactylus and one to four (usually one or two) versus five to ten spines on the propodus, respectively. Additionally, in females, the dorsal margin of the carpus of each second pereopod is slightly protuberant but is always unarmed (exclusive of a dorsodistal spine) in the new species, whereas it bears a small submedian spine in P. fasciatus. In fresh condition, the two species are readily discriminated by the color patterns of the antennular peduncles and the dactyli of the ambulatory legs (Fig. 1C–F). The ultimate segment of the antennular peduncle has two brown rings on the proximal and distal portions in P. vicinus sp. nov., instead of being entirely brown in P. fasciatus. The dactyli of the ambulatory legs are dark brown on the proximal two-fifths to half and white on the remaining portion in the new species, whereas they are mostly dark brown and pale-colored on the midline in P. fasciatus. The shield of P. vicinus sp. nov. also lacks a posterolateral pair of brown spots, which are present in P. fasciatus.</p> <p>General morphology and coloration also link P. vicinus sp. nov. to P. acanthocarpus. However, the new species differs from P. acanthocarpus in having apparently weaker, granulation and sculpture on the dorsal surface and margins of the right chela and armature on the dorsal surface of the carpus of the right cheliped (cf. Komai &amp; Okuno 2009: figs 6A, 7A). Additionally, there are no marked sexual differences on the left chela and third pereopods in the new species, unlike in P. acanthocarpus. The palm of the left cheliped of P. acanthocarpus has a much more strongly arched and ridged lateral margin and a more prominently elevated dorsal surface in females than in males. The right and left third pereopods of P. acanthocarpus are generally similar in males, but in females, the left propodus is much stouter than the right and the left carpus bears a distinct row of setae on the laterodistal margin, which is absent on the right carpus. In males, the dorsal margin of the carpus of each second pereopod is unarmed (exclusive of a dorsodistal spine) in P. vicinus sp. nov., but it has at least one additional spine in P. acanthocarpus. Some details of the color pattern are also different between the two species (Fig. 1A, B, E, F). For instance, the posterolateral pair of brown spots on the shield as seen in P. acanthocarpus is absent in the new species. The dark brown band on the propodus of each ambulatory leg is broad and occupies about proximal 0.7 portion of the segment in P. vicinus sp. nov., whereas it is narrow and present only on about proximal 0.3 portion in P. acanthocarpus.</p> </div>	https://treatment.plazi.org/id/FB4187964D1FFFFD57FEFF45FEE4F96C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Osawa, Masayuki;Kawai, Takashi;Sakamaki, Takashi	Osawa, Masayuki, Kawai, Takashi, Sakamaki, Takashi (2013): New records of Pagurixus Melin, 1939 (Crustacea: Decapoda: Paguridae) from the Ryukyu Islands, southwestern Japan, with description of a new species. Zootaxa 3669 (3): 350-366, DOI: 10.11646/zootaxa.3669.3.9
