identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
FA4EBE79FFF95002FF70FCCA44E1C661.text	FA4EBE79FFF95002FF70FCCA44E1C661.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bettongia ogilbyi subsp. sylvatica Newman-Martin & Travouillon & Warburton & Barham & Blyth 2025	<div><p>Bettongia ogilbyi sylvatica subsp. nov. Newman-Martin, Travouillon &amp; Warburton</p><p>Figs. 13, 14, 17–26, 29, &amp; 30</p></div>	https://treatment.plazi.org/id/FA4EBE79FFF95002FF70FCCA44E1C661	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Newman-Martin, Jake;Travouillon, Kenny J.;Warburton, Natalie;Barham, Milo;Blyth, Alison J.	Newman-Martin, Jake, Travouillon, Kenny J., Warburton, Natalie, Barham, Milo, Blyth, Alison J. (2025): A taxonomic revision of the Bettongia penicillata (Diprotodontia: Potoroidae) species complex and description of the subfossil species Bettongia haoucharae sp. nov. Zootaxa 5690 (1): 1-69, DOI: 10.11646/zootaxa.5690.1.1, URL: https://doi.org/10.11646/zootaxa.5690.1.1
FA4EBE79FFEC5011FF70FD5C413EC477.text	FA4EBE79FFEC5011FF70FD5C413EC477.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bettongia haoucharae Newman-Martin & Travouillon & Warburton & Barham & Blyth 2025	<div><p>Bettongia haoucharae sp. nov.</p><p>The results of this investigation indicate that the taxon identified by Haouchar et al. (2016) is morphologically valid as the taxon is both genetically and morphologically distinct. We have chosen to give the taxon species status with the name Bettongia haoucharae . This taxon is clearly morphologically distinct from the Bettongia penicillata complex, which can be seen in the PCA analyses performed (Fig. 4,5,6,7,8, and 9), where B. haoucharae is more distant morphologically than Bettongia tropica and Bettongia gaimardi are from the rest of the complex. Initially, we suspected that individuals of B. haoucharae could represent Bettongia pusilla or Bettongia anhydra as both of these taxa are known from arid regions and are very cryptic. However, when comparing available specimens of B. pusilla (Supp. Fig. 1), including the type specimen (SAMA P35450), it was shown that the dp2 and dp3 of B. haoucharae are markedly larger than B. pusilla . Additionally, the molars are markedly smaller in B. pusilla . Similarly, B. anhydra was also compared with B. haoucharae, and the P3 of the latter is more buccally angled than the straight molars that are present in B. anhydra . Similary, B. haoucharae was also directly compared with Bettongia lesueur and specimens did not morphologically match.</p><p>Based on the available material, B. haoucharae was likely an arid adapted bettong that could be found throughout the Nullarbor and arid South Australia, through to the Great Victoria Desert until the border of the Gibson Desert. This means that it was a fairly wide-ranging Bettong that occupied the southwestern deserts of Western Australia and South Australia, in the habitats that were formerly believed to be occupied by an arid living Bettongia penicillata . Bettongia haoucharae is fairly abundant in Nullarbor cave deposits alongside Bettongia pusilla and Bettongia lesueur, indicating that it could live in sympatry with both species. No specimens are known from the Roe Plains or any other mesic environments in Western Australia, which could indicate that the species was specialised for arid environments and could not compete with other small potoroids in a mesic setting in Western Australia. However, the species is recorded as far east as Venus Bay in South Australia in the Haouchar et al. (2016) investigation. Whilst no photographs or skins of this taxa survive today, it is possible to speculate at its appearance. Based on the holotype specimen (WAM 75.12.21), and habitat information an artist’s rendition of the taxa in life was created (Fig. 35.)</p><p>According to features found on the forelimbs of the holotype of Bettongia haoucharae the species engaged in partial fossorial behaviours, similar to other Bettongia (Baker et al. 2023) . This is based on the presence of a shorter and more robust humerus with an enlarged insertion of pectoralis superficialis muscle. Bettongia haoucharae, like other Bettongia, likely dug or foraged through the substrate for food. However, without an adult individual or more specimens for analysis, the authors are hesitant to comment on the fossorial abilities of the taxa compared to other Bettongia taxa. In other Bettongia, hypogeous mycorrhizal fungi make up the bulk of their diet (Taylor, 1992; Nuske et al. 2018; Zosky et al. 2018; Mitchell et al. 2024). However, it is not known what food source was widely available for B. haoucharae in an arid setting. Although the diet of B. haoucharae is uncertain, it is likely to have been similar to B. lesueur in the region. Future studies could examine the holotype specimen for stomach contents or perform stable carbon and nitrogen isotope analysis on the dentition as well as dental microwear texture analysis. The hind limbs were adapted to saltatorial locomotion as in other macropodines; however, based on their morphology B. haoucharae may have spent more time in a quadrupedal gait due to the shorter proportional length of the limb bones in comparison to other Bettongia, as well as their robust muscle attachments.</p><p>FIGURE 35. Artist’s reconstruction of Bettongia haoucharae in life based on cranial elements, habitat data, and known morphology of related Bettongia taxa. Artwork by Eleanor “Nellie” Pease.</p><p>It is believed that “ B. penicillata ” of the Nullarbor, recognised here as B. haoucharae, went extinct sometime during the 1920s due to the spread of foxes in this region (Richards &amp; Short, 1996). This is a similar extinction window to other small mammalian fauna in the region, such as Perameles papillon (Travouillon &amp; Phillips, 2018) and Dasycercus archeri (Newman-Martin et al., 2023) which were both last seen alive on the Nullarbor in the 1920s.</p><p>The B. penicillata complex</p><p>Another hypothesis that was tested in this investigation was that Bettongia ogilbyi and Bettongia penicillata represent two distinct species. This proposal was based on: (i) Kangaroo Island specimens (which were identified as B. p. penicillata) being genetically distinct from specimens in the southwest of Australia (Haouchar et al. 2016), with Kangaroo Island specimens a 99.3% genetic match to B. penicillata from the mainland (Haouchar et al. 2014);. (ii) if there is a valid distinct taxon present on the Nullarbor ( Bettongia haoucharae), then the subspecies of B. penicillata are geographically separated, indicating that speciation could occur.</p><p>The results of this investigation confirmed that B. p. ogilbyi and B. p. penicillata are indeed morphologically distinct when compared using LDA and PCA (Fig. 4-9) with both cranial and dental characters seperating the taxa. Notably, B. penicillata was found to differ from B. ogilbyi in that; a frontal-squamosal suture contact is present contra to B. ogilbyi where a parietal-alisphenoid contact is present, the auditory bulla is larger and ovate, contra to B. ogilbyi where it is kidney shaped, P3 is more anteriorly angled towards the premaxilla, the P3 is intermittently more bulbous than B. ogilbyi, and there is no StC present on the M1. Many more differences can be observed between B. penicillata and B. ogilbyi cranially, dentally and postcranially, and these can be found in the systematics section. Non-allometry corrected PERMANOVAs demonstrated that the taxa are morphologically distinct (Table 3, 7 &amp; 11); with non-allometry corrected cranial test of just the B. penicillata complex, non-allometry corrected upper dental test for the B. penicillata complex, and allometry corrected lower dental test for the B. penicillata complex. Additional allometry corrected PERMANOVAs demonstrated that the taxa are morphologically distinct (Table 4, 8 &amp; 12); with allometry corrected cranial test of just the B. penicillata complex, allometry corrected upper dental test for the B. penicillata complex, and allometry corrected lower dental test for the B. penicillata complex. Referring to Fig. 2 in Haouchar et al. (2016), it can be seen that B. penicillata is more closely related genetically to Bettongia gaimardi, than to B. ogilbyi . Based on the results of this investigations, which show that B. penicillata and B. ogilbyi are morphologically very different and have many characters separating them, the authors propose that B. ogilbyi be elevated to full species status and should be referred to as Bettongia ogilbyi ogilbyi .</p></div>	https://treatment.plazi.org/id/FA4EBE79FFEC5011FF70FD5C413EC477	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Newman-Martin, Jake;Travouillon, Kenny J.;Warburton, Natalie;Barham, Milo;Blyth, Alison J.	Newman-Martin, Jake, Travouillon, Kenny J., Warburton, Natalie, Barham, Milo, Blyth, Alison J. (2025): A taxonomic revision of the Bettongia penicillata (Diprotodontia: Potoroidae) species complex and description of the subfossil species Bettongia haoucharae sp. nov. Zootaxa 5690 (1): 1-69, DOI: 10.11646/zootaxa.5690.1.1, URL: https://doi.org/10.11646/zootaxa.5690.1.1
FA4EBE79FFEA5010FF70FEBC47D5C757.text	FA4EBE79FFEA5010FF70FEBC47D5C757.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bettongia ogilbyi (Waterhouse 1841) Newman-Martin & Travouillon & Warburton & Barham & Blyth 2025	<div><p>Bettongia ogilbyi</p><p>Within specimens of Bettongia ogilbyi from the southwest of Western Australia, two morphologies were identified. Two subspecies were initially identified based on the sexual dimorphism present, wherein the females of Bettongia ogilbyi sylvatica were near equal in size to the males of Bettongia ogilbyi ogilbyi . This is visible in the craniodental morphological analysis (Fig. 4, 5, 6, 7, 8, and 9), and supported by the PERMANOVA analysis conducted (Table 1-12) which indicated both taxa represent a subspecies within B. ogilbyi . Of these taxa, the new subspecies was named Bettongia ogilbyi sylvatica after the forest environments where specimens were predominantly found (Supp. 4). PERMANOVA analyses comparing B. o. sylvatica and B. o. ogilbyi demonstrated significant difference cranially (Table 3), as well as for the upper dental (Table 7), and lower dental (Table 11). When allometry corrected, the results showed a similar result based on cranial (Table 4), upper dental (Table 8), and lower dental (Table 12) characters.</p><p>However, our findings are contra to those of previous genetic investigations (Pacioni et al. 2011; Pacioni, 2014), Bettongia ogilbyi is one of the most well studied mammals in Australia, mainly due to the large amount of conservation and translocation efforts undertaken (Short, 2009), with genetic work used in the management of population diversity (Pacioni et al. 2011; Pacioni, 2014). In fact, B. ogilbyi was the first member of Potoroidae with a reference genome (Peel et al. 2021). Prior to this, the majority of genetic work on B. ogilbyi used mitochondrial DNA (mtDNA) sequences such as Cytochrome b (Cytb), as evidenced by Genbank, where (at the time of writing) of the 123 accessions for B. oglibyi, only two use nuclear DNA (nDNA). The use of mtDNA in taxonomy has previously come under scrutiny, and it has been suggested it should not be exclusively used when identifying species (Rubinoff, &amp; Holland, 2005). This is because mtDNA can be of little utility when attempting to identify closely related species (Rubinoff, Cameron, &amp; Will, 2006). Instead, it is suggested that a combination of mtDNA and nDNA should be used for taxonomic purposes (Rubinoff, Cameron, &amp; Will, 2006). Examples of closely related taxa being misidentified due to the exclusive use of mtDNA include rock wallabies ( Petrogale spp.) and short-nose bandicoots ( Isoodon spp.). In the case of the latter, initial genetic work by Pope et al. (2001) using mtDNA could not differentiate between I. obesulus and I. auratus . Later work by Westerman et al. (2012) used a combined mtDNA and nDNA taxonomic approach that could clearly differentiate the species. In the case of rock wallabies ( Petrogale spp.), genetic work using mtDNA was inconsistent with that of nDNA, and clarity could only be brought to the taxonomy of the species once a combined approach was used (Eldridge &amp; Close, 1992). Many genetic investigations traditionally use samples from live B. ogilbyi, and only the work by Pacioni et al. (2015), and Haouchar et al. (2016) including historical museum and subfossil specimens. Whilst Pacioni et al. (2015), and Haouchar et al. (2016) used mtDNA, the use of a wider array of material in these studies allowed for more genetic material to be represented and thus more potential for unidentified taxa.</p><p>Prior to the work of Pacioni et al. (2015), and Haouchar et al. (2016), no outliers were observed in the genetics of the B. penicillata complex. This indicates that the conclusions drawn from a substantial amount of genetic research conflicts with the results of our investigation, where more taxa than B. haoucharae (Clade 1 in Haouchar et al. 2016) were identified. Unfortunately, the genetic data of Pacioni et al. (2015) cannot be matched with physical specimens, as the registration numbers of the material genetically analysed were not recorded. Additionally, it is known that some genetic material from Pacioni et al. (2015) failed to sequence (Supporting information Table S1, Pacioni et al. 2015). Without registration numbers it is not possible to know if material failing to sequence is the reason for taxa being overlooked, or if specimens were not sampled. A lack of Museum registration numbers in Bettongia penicillata genetic reports is not unusual, with other investigations (Pacioni et al., 2011; Haouchar et al. 2016) also lacking this information. This also extends to the genotyped specimen for the Woylie (WAM M65253), which was registered after the Peel et al. (2021) report was published.</p><p>The authors were, however, able to link molecular samples in Pacioni et al., (2011) to Bettongia ogilbyi ogilbyi . In a molecular investigation by Pacioni et al., (2011), it was determined that four genetically distinct indigenous populations were present within the southwest, those being the populations at Perup, Kingston, Dryandra woodland and Tutanning Nature Reserve. Of these populations the individuals at the Tutanning Nature Reserve were noted as being the most genetically distinct. When comparing specimens at the Western Australian Museum, a single specimen was found from Kanyana Wildlife Rehabilitation Centre (WAM M65249) an adult male collected in 2014. This individual is the most recent modern specimen of B. o ogilbyi in the WAM, with the last B. o. ogilbyi from 1973 (pers. obs. Newman-Martin and Travouillon). In 2012 several wild individuals from the Tutanning population were delivered to the Kanyana Wildlife Rehabilitation Centre to establish a breeding population (pers. comm. Tara Jenkins to Newman-Martin &amp; Travouillon, 2024). Upon inquiring into the status of these individuals with the Kanyana Wildlife Rehabilitation Centre, it was discovered that there is a single individual left, an elderly female named “Bad-Leg” (pers. comm. Tara Jenkins to Newman-Martin &amp; Travouillon, 2024). However, in February of 2022 three Tutanning males were sent from the Kanyana Wildlife Rehabilitation Centre to Whiteman Park, though their status is unknown (pers. comm. Tara Jenkins to Newman-Martin &amp; Travouillon, 2024). There have been confirmed observations of wild B. o. ogilbyi in the Tutanning area (pers. comm. Kate Rick to Travouillon, 2024). With only the wild population at Tutanning known, this means that the wild population of B. o. ogilbyi is very low and in critical need of reassessment and protection.</p></div>	https://treatment.plazi.org/id/FA4EBE79FFEA5010FF70FEBC47D5C757	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Newman-Martin, Jake;Travouillon, Kenny J.;Warburton, Natalie;Barham, Milo;Blyth, Alison J.	Newman-Martin, Jake, Travouillon, Kenny J., Warburton, Natalie, Barham, Milo, Blyth, Alison J. (2025): A taxonomic revision of the Bettongia penicillata (Diprotodontia: Potoroidae) species complex and description of the subfossil species Bettongia haoucharae sp. nov. Zootaxa 5690 (1): 1-69, DOI: 10.11646/zootaxa.5690.1.1, URL: https://doi.org/10.11646/zootaxa.5690.1.1
FA4EBE79FFEB5010FF70FD5C4066C14B.text	FA4EBE79FFEB5010FF70FD5C4066C14B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bettongia	<div><p>Bettongia o. francisca</p><p>The subspecies Bettongia penicillata francisca has never been formaly synonomised but is generally recognised as invalid. Our results indicate that B. p. francisca is a valid subspecies, with the taxon displaying significant difference in the one-way PERMANOVAS (Table 1-12). However, our results indicate that the specimens are closer morphologically to Bettongia ogilbyi ogilbyi, with an overlap present in the PCA analysis (Fig. 4 &amp; 5). The authors acknowledge it is possible that this is a case of convergent evolution, wherein the specimens from St. Francis Island have similar dental morphology to B. o. ogilbyi due to similar diet. However, in all PCA analyses conducted, the subspecies did not overlap with B. penicillata, and consistently overlapped with B. o. ogilbyi . As such, we assign the St. Francis Island taxon to Bettongia ogilbyi francisca, until genetic testing can supplement the morphological work presented here. The authors would like to note that modern translocated Bettongia specimens from St. Francis Island were identified herein as B. o. ogilbyi . Notoriously, these translocated populations of Bettongs could not survive (Robinson et al. 1996), indicating that B. o. francisca was uniquely adapted to the conditions present. Additionally, other B. o. francisca subfossil specimens from St. Francis Island were identified and morphologically matched to the holotype specimen, indicating the species did indeed occur on the island as proposed by Finlayson (1957).</p></div>	https://treatment.plazi.org/id/FA4EBE79FFEB5010FF70FD5C4066C14B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Newman-Martin, Jake;Travouillon, Kenny J.;Warburton, Natalie;Barham, Milo;Blyth, Alison J.	Newman-Martin, Jake, Travouillon, Kenny J., Warburton, Natalie, Barham, Milo, Blyth, Alison J. (2025): A taxonomic revision of the Bettongia penicillata (Diprotodontia: Potoroidae) species complex and description of the subfossil species Bettongia haoucharae sp. nov. Zootaxa 5690 (1): 1-69, DOI: 10.11646/zootaxa.5690.1.1, URL: https://doi.org/10.11646/zootaxa.5690.1.1
FA4EBE79FFE8506DFF70FD9446CBC72F.text	FA4EBE79FFE8506DFF70FD9446CBC72F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bettongia gouldii Waterhouse 1846	<div><p>Bettongia gouldii</p><p>Only a few specimens were found that could represent individuals of Bettongia gouldii . As the holotype is a juvenile specimen with DP2, DP3, M1 and an M2 that is still in crypt, these specimens, which had worn dentition, could not be reliably compared. When performing the PCA and LDA analysis, it was instead determined that our identified specimens of Bettongia gouldii were representatives of other taxa. Only one specimen was not reliably identified as being from other taxa, that being specimen SAMA M8554, a subfossil skull of a subadult specimen from Cleve, South Australia. When examined in a confusion matrix, SAMA M8554 was identified as B. gaimardi, despite this species not being known from this area. Additionally, when allometry was removed from the PCA and LDA analyses, SAMA M8554 grouped with Bettongia penicillata . While SAMA M8554 does display characters that are different from B. penicillata, such as smaller and less inflated bulla, the authors cannot confidently say that SAMA M8554 is a representative of B. gouldii, or that B. gouldii is a valid taxon. For this reason, the only way to resolve this issue is with a genetic evaluation of the type specimen against other Bettongia . As B. gouldii was never formally synonymised and our investigation of this taxon is inconclusive, we believe that the taxon can be formally classified as nomen dubium.</p><p>Phylogenetic analysis</p><p>The results of our phylogenetic analysis were inconclusive, with all taxa within Bettongia forming a polytomy. This resulting matrix was also unable to separate Macropodidae from Potoroidae indicating that more research is needed into the taxonomy and evolutionary history of Potoroids. One key cranial character that is used for differentiating Potoroidae and Macropodidae is that in the former the squamosal bone contacts frontal bone, unlike in macropodids where a parietal and alisphenoid contact is present (Seebeck and Rose 1989). However, in this investigation it was determined that in some Bettongia ( B. haoucharae, B. o. ogilbyi, B. o. sylva, and B. o. odontoploica) the inverse is true. Additionally, fossil species of Potoroids are known to be problematic as the taxonomy is currently unresolved. Initially the fossil taxa were included in this investigation, particularly in the phylogenetic tree, but were removed as we were unable to construct a conclusive phylogeny. Previous investigations have also noted that the Potoroids need work (Flannery &amp; Archer 1987; Cooke et al. 2015; Travouillon et al. 2016; Travouillon et al. 2022), with the taxonomy of several taxa unresolved within the family. A full taxonomic revision of Potoroidae is considered out of scope for this investigation.</p><p>Conservation implications</p><p>As a result of the taxonomic revision presented in this investigation, Bettongia penicillata is considered extinct. Additionally, the taxonomic changes will have a myriad of effects on the current conservation practices for Bettongia ogilbyi, and its subspecies. As mentioned previously, Bettongia ogilbyi ogilbyi is critically endangered, with a single wild population from Tutanning Nature reserve remaining. Based on museum specimens, the decline of B. o. ogilbyi likely began in the 1960s, with fewer specimens recorded in the WAM. The exact cause of the decline is still unknown. This also indicates that the current populations of Woylies at Perup, Kingston, and Dryandra are Bettongia ogilbyi sylvatica . Likely the Dryandra population of B. o. sylvatica was introduced during an attempt to increase gene flow between the populations (Pacioni et al. 2013). Bettongia ogilbyi sylvatica is critically endangered but recovering due to current conservation efforts. Based on available locality data of each taxon (Supp. 4), B. o. ogilbyi occurred in open woodland to scrub habitats, while B. o. sylvatica occurred in denser woodland and forest habitats such as the Jarrah Forest (pers. obs. Newman-Martin &amp; Travouillon). Referring to museum material, B. o. ogilbyi could be found in western Australia, all the way from the southwest to the Roe Plains. Bettongia ogilbyi ogilbyi could also be found far north into Shark Bay (Fig. 36). Museums specimens of B. o. sylvatica have been found to not occur past Esperance to the east or Yanchep to the north (Fig. 36). Both species were found in sympatry on the Swan Coastal Plain and into the Perth Hills.</p><p>FIGURE 36. Distribution of the Bettongia penicillata complex based on the results of this taxonomic review. Distribution map was created using known modern localities as well as verifiable historic and subfossil specimens. Note that temporal changes are not shown, and the distribution of taxa would have changed through time with changes to the environment.</p><p>With the advent of two subspecies within B. ogilbyi, another factor to consider is outbreeding depression. Outbreeding depression occurs when two populations of a taxa are introduced in an attempt to maximise genetic diversity. When the populations reproduce, the alleles from the introduced population may not aid the offspring in survival, and instead lead to a loss of fitness (Frankham et al. 2011). According to Frankham et al. (2011) the probability of outbreeding depression occurring between two populations increases when the populations have at least one of the following characteristics: are distinct species, have fixed chromosomal differences, have exchanged no genes in the last 500 years, or inhabit different environments. As stated previously, the B. ogilbyi subspecies are found in different habitats from one another, with sympatry only occurring on the Swan Coastal Plain and Perth hills. Previously, outbreeding depression has been suggested to be unlikely for B. ogilbyi, due to the species belonging to a single wide range and adapted to many habitats (Pacioni et al. 2018). However, our investigation indicates that B. ogilbyi was not adapted for many habitats and did not occur over a wide range (Fig. 36) as has previously been suggested. For this reason, the authors believe that outbreeding depression cannot be ruled out and caution should be taken when translocating and mixing populations of B. ogilbyi . Whilst it is likely that Bettongia ogilbyi ogilbyi is functionally extinct it is possible that during translocation, some hybrid individuals with Bettongia ogilbyi sylvatica were produced.</p><p>Previous translocations within the indigenous southwest populations (Perup, Kingston, Dryandra woodland and Tutanning nature reserve) have occurred (Pacioni et al. 2013). However, the effects of hybridising the two Bettongia ogilbyi subspecies are yet unknown. It is also possible that an outbreeding depression was the cause of the original population crash of B. ogilbyi in the early 2000s, however, without more material this is just speculation. Additionally, current reintroduction projects across Australia must be labelled as introductions, where B. ogilbyi is being used as an ecological replacement for locally extinct Bettongia taxa. The findings presented here could also help to explain why some reintroduction attempts have failed while other have succeeded. For instance, in this investigation we identified B. ogilbyi specimens from St. Peter Island (NMV C32743 and NMV C32742), which has seen a successful reintroduction (Start et al. 1995). However, B. ogilbyi released on St. Francis Island failed to establish (Start et al. 1995). The subspecies B. o. francisca was identified from this island. This indicates that B. o. francisca was specifically adapted to the conditions of St. Francis Island.</p></div>	https://treatment.plazi.org/id/FA4EBE79FFE8506DFF70FD9446CBC72F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Newman-Martin, Jake;Travouillon, Kenny J.;Warburton, Natalie;Barham, Milo;Blyth, Alison J.	Newman-Martin, Jake, Travouillon, Kenny J., Warburton, Natalie, Barham, Milo, Blyth, Alison J. (2025): A taxonomic revision of the Bettongia penicillata (Diprotodontia: Potoroidae) species complex and description of the subfossil species Bettongia haoucharae sp. nov. Zootaxa 5690 (1): 1-69, DOI: 10.11646/zootaxa.5690.1.1, URL: https://doi.org/10.11646/zootaxa.5690.1.1
