taxonID	type	description	language	source
FA013919F20A3B1943D0DB70FA84C963.taxon	materials_examined	Holotype. ZMMU R- 13171 — Iran, Khorasan Province, 150 km SW from Tabas city, on the road Yazd – Tabas, 33.087534 ° N; 55.912115 ° E; collected by R. A. Nazarov, D. A. Bondarenko & M. Rajabizadeh, 09 May 2010. Paratypes. Five males (ZMMU R- 15624 – 15626, R- 15628, and R- 15630) and four females (ZMMU R- 15627, R- 15629, R- 15631, and R- 12986) with the same data as holotype.	en	Orlova, Valentina F., Rasegar-Pouyani, Eskandar, Rajabizadeh, Khosrow, Nabizadeh, Hossein, Poyarkov, Nikolay A., Melnikov, Daniel A., Nazarov, Roman A. (2023): Taxonomic diversity of racerunners with descriptions of two new Eremias species (Sauria: Lacertidae) from Central Iran. Zootaxa 5369 (3): 336-368, DOI: 10.11646/zootaxa.5369.3.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5369.3.2/52247
FA013919F20A3B1943D0DB70FA84C963.taxon	diagnosis	Diagnosis. Maximum SVL 58.3 mm, TL about two times longer than the body (max. 118 mm). Supraocular plates (preocular and postocular scales) usually separated from frontal and frontoparietal plates by small granular scales; fifth inframaxilare not in contact with infralabial plates. The collar scales are not strongly distinct from surrounding gular scales and usually only five collar scales are larger than the others. The body scales are relatively small, about 60 (54 – 65) scales around middle of the body (Sq); 15 – 18 well developed femoral pores on the left and right sides, separated from each other by 4 – 6 scales; series of femoral pores not reaching the knee bend by 3 – 4 scales. Coloration. The main dorsal background color is beige; dorsal patterns comprise vermiculated light and darkbrown spots; no regular longitudinal dark lines on the dorsum but sometimes dark spots forming narrow longitudinal stripes are more visable on the neck. A light narrow longitudinal line begins from the upper side of tympanic region and continues to the tail, bordering the lateral body surface. Wide, dark, non-contrasting longitudinal stripes without light spots usually visible on the flanks and continue to the tail. Sometimes there is a narrow dark longitudinal line marking the lateral and ventral border of the body. No yellow spot on the tympanic plate. Dorsal surface of the limbs dark brown with round light or yellowish spots. The ventral surface of the body is whitish and the tip of the tail light greyish.	en	Orlova, Valentina F., Rasegar-Pouyani, Eskandar, Rajabizadeh, Khosrow, Nabizadeh, Hossein, Poyarkov, Nikolay A., Melnikov, Daniel A., Nazarov, Roman A. (2023): Taxonomic diversity of racerunners with descriptions of two new Eremias species (Sauria: Lacertidae) from Central Iran. Zootaxa 5369 (3): 336-368, DOI: 10.11646/zootaxa.5369.3.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5369.3.2/52247
FA013919F20A3B1943D0DB70FA84C963.taxon	description	Description of the Holotype. ZMMU R- 13171 (Fig. 8). Adult male, SVL 56.9 mm, TL 114 mm, Pa 20.5 mm, Pp 35.7 mm, Lpil 13.5 mm. Meristic features: G 30, Coll 5, Pfm (right / left) 16 / 15, Dist. Pfm 6, labialia (left / right) 9 / 10, infralabilia (left / right) 6 / 7, ventralia 32. Tail scales smooth on the base, not strongly keeled; Sq. c. cd 29, subdigital lamellae subdigitalia 29. Smooth roundish and relatively small dorsal scales; Sq. 65; Sq. dorsale 153. Head and body moderately depressed. Head length (14.5 mm) 1.74 times its width (8.3 mm) and 1.93 times its height (7.2 mm); nasals swollen; snout longer than postocular part of the head; head wider than neck. Fore and hind limbs relatively long; Pa / SVL 0.34, Pp / SVL 0.63. Hind limb reaches the axilla of forelimb and extends far beyond it. Tail about twice as long as the body, SVL / TL 0.49, cylindrical and depressed at base. Pileus shields smooth, nasals in contact behind the rostral, the suture between them half the length of frontonasal, whose width is more than its length; length of frontonasal is half of the width; prefrontals slightly longer than broad, forming median suture, at the top of posterior edge is placed one tiny triangular plate; frontal the same size as distance from end of the snout, two-thirds times as broad as long, narrow behind; parietals as broad as long; interparietal almost equal to half of the length of frontoparietals; no occipital. Parietals, frontoparietals, and interparietal weakly rugose. Two large supraoculars (preocular and postocular), preocular slightly longer than the postocular; three larger and a few small granules in front of supraoculars; a small, band-like additional supraocular also present behind the postocular; supraoculars separated from frontal and frontoparietal by a single row of granules. Five supraciliaris. Three nasals, the lower in contact with two supralabials and not in contact with rostral; anterior loreal slightly higher than wide, shorter than the second which is wider than high; subocular smooth, bordering the mouth, wedged between sixth / seventh and seventh / eighth supralabials. Temporal scales smooth; no auricular denticulation scales; lower eyelid translucent and covered with small scales. Five pairs of submaxillaries; anterior three pairs completely in contact on the midline, fifth not contacted with infralabials (or separate from infralabials). Thirty gular scales in a straight line between the symphysis of the submaxillary and the collar. Gular fold not distinct. Collar weakly developed, not curved, serrated, and comprising five enlarged plates. Dorsal scales granular, smooth, 65 across midbody. Ventral plates broader than long (except for the outermost series), forming oblique longitudinal rows of 14 plates, in 32 slightly angular transverse rows; first 4 – 5 rows of pectoral scales longer than broad, first one is twice as long as broad. Precloacal region with two enlarged median plates, just anterior to the cloaca. Upper surface of arm with rhombic smooth scales. Scales on upper surface of hind limbs similar to dorsal scales; enlarged plates covering the lower surface of hind limbs; 16 and 15 femoral pores on right and left sides, respectively; the two series separated by six scales; length of the series of femoral pores is three and 3.5 times the distance between these two rows. Femoral pores do not reach the knee bend by three and four scales on the right and left sides, respectively. Subdigital lamellaes unicarinate, in two rows of 29 scales under fourth toe. Two enlarged precloacal plates. Coloration. Unlike most members of Eremias in which the dorsal pattern is formed by dark, narrow, longitudinal stripes, the new species has a vermiculated dorsal pattern (Figs. 7 a, b; 8 a, f). The dorsum has a sand-colored background combined with the dark pattern forming worm-shaped vermiculations. The venter is white; ventral side of the tail is light-greyish.	en	Orlova, Valentina F., Rasegar-Pouyani, Eskandar, Rajabizadeh, Khosrow, Nabizadeh, Hossein, Poyarkov, Nikolay A., Melnikov, Daniel A., Nazarov, Roman A. (2023): Taxonomic diversity of racerunners with descriptions of two new Eremias species (Sauria: Lacertidae) from Central Iran. Zootaxa 5369 (3): 336-368, DOI: 10.11646/zootaxa.5369.3.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5369.3.2/52247
FA013919F20A3B1943D0DB70FA84C963.taxon	distribution	Distribution. The new species is known only from the type locality — Iran, Khorasan Province, 150 km S from Tabas city, on the road from Yazd to Tabas, 33.087 ° N; 55.9121 ° E (Fig. 1).	en	Orlova, Valentina F., Rasegar-Pouyani, Eskandar, Rajabizadeh, Khosrow, Nabizadeh, Hossein, Poyarkov, Nikolay A., Melnikov, Daniel A., Nazarov, Roman A. (2023): Taxonomic diversity of racerunners with descriptions of two new Eremias species (Sauria: Lacertidae) from Central Iran. Zootaxa 5369 (3): 336-368, DOI: 10.11646/zootaxa.5369.3.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5369.3.2/52247
FA013919F20A3B1943D0DB70FA84C963.taxon	etymology	Etymology. The name of this species is derived from the Greek root graphikos, meaning “ drawn ” or “ written, ” because the dorsal pattern resembles artificial hieroglyphs. Natural history. This species inhabits unstabilized sand dunes on gravel plains. The habitat is characterized by scattered vegetation cover consisting of sparse bushes and the desert graminoid plant genus Stipagrostis. Eremias graphica sp. nov. is associated with Stipagrostis, using this plant as cover form predators and as a source of its insect prey, and usually dig burrows under it (Fig. 7 d). These lizards have a relatively short period of daily activity in the summer of approximatley 2 – 3 hours in the morning and the same active time in the evening. The lizards spent the rest of the day in the shadow of the bushes. In the same habitat, we recorded the following sympatric reptile species: Bunopus tuberculatus Blanford, 1874, Phrynocephalus maculatus Anderson, 1872, Rhinogekko misonnei De Witte, 1973, Teratoscincus keyserlingii Strauch, 1863, and Eremias fasciata Blanford, 1874. Comparison. The new species can be distinguished from Eremias fasciata by the following characters: vermicular dorsal patterns versus striped patterns in E. fasciata and yellow tympanic shields (Fig. 9 a, b) in E. fasciata versus a white tympanum in E. graphica (Fig. 9 e, g). Eremias graphica has a wider interspace between the left and right rows of femoral pores (4 – 6 scales) than does E. fasciata (2 – 5 scales). Furthermore, E. fasciata has bright-yellowish coloration on the tip of the tail, whereas E. graphica has a light-greyish tail tip. Eremias graphica can be easy distinguished from Eremias andersoni by a greater SVL (max SVL) of 58.3 mm versus 40 mm, longitudinal stripes on the dorsum versus vermiculated pattern in E. graphica, and fewer ventral scales in E. andersoni — 28 – 29 versus 32 – 35. In addition, E. andersoni has light-bluish coloration at the tip of the tail, whereas E. graphica has a light-greyish tail tip. Eremias graphica differs from E. lineolata by the vermiculated dorsal pattern versus longitudinal dark strips in E. lineolata and the number of scales around the tail segment (Sq. c. cd) of 23 – 29 versus 12 – 17, in E. lineolata. Eremias graphica can be clearly distinguished from E. scripta by a higher number of femoral pores with 8 – 17 versus 15 – 18 and a higher number of gular scales of 26 – 31 versus 15 – 23. Although the dorsal patterns in some subspecies of E. scripta can be vermiculated, the new species clearly differs on the basis or meresitc data. Eremias graphica differs from E. cholistanica by the number of gular scales — 26 – 31 versus 20 – 24, the number of scales around the tail segment — 23 – 29 versus 27 – 35, and by its vermiculated versus striped dorsum.	en	Orlova, Valentina F., Rasegar-Pouyani, Eskandar, Rajabizadeh, Khosrow, Nabizadeh, Hossein, Poyarkov, Nikolay A., Melnikov, Daniel A., Nazarov, Roman A. (2023): Taxonomic diversity of racerunners with descriptions of two new Eremias species (Sauria: Lacertidae) from Central Iran. Zootaxa 5369 (3): 336-368, DOI: 10.11646/zootaxa.5369.3.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5369.3.2/52247
FA013919F2113B1643D0D9AFFE67C9B7.taxon	materials_examined	Holotype. ZMMU R- 12982 Iran, Khorasan Province, 20 km SE of Boshruiyeh, 33.760693 ° N; 57.560714 ° E; elevation 949 m a. s. l.; collected by R. A. Nazarov, D. A. Bondarenko & M. Rajabizadeh, 09 May 2010 (Fig. 11). Paratypes. Eleven specimens including five males (ZMMU R- 15632 – 15633, R- 15635, and R- 15638 – 15639) and one female (ZMMU R- 15634) with the same locality data as the holotype and four males (ZMMU RAN 4201, 4203, 4204, 4208) and one female (ZMMU RAN 4205) from Iran, Khorasan Province, 15 km NE of Boshruiyeh, 33.972851 ° N; 57.517912 ° E; elevation 980 m a. s. l. collected 13 May 2021 by R. A. Nazarov, H. Nabizadeh.	en	Orlova, Valentina F., Rasegar-Pouyani, Eskandar, Rajabizadeh, Khosrow, Nabizadeh, Hossein, Poyarkov, Nikolay A., Melnikov, Daniel A., Nazarov, Roman A. (2023): Taxonomic diversity of racerunners with descriptions of two new Eremias species (Sauria: Lacertidae) from Central Iran. Zootaxa 5369 (3): 336-368, DOI: 10.11646/zootaxa.5369.3.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5369.3.2/52247
FA013919F2113B1643D0D9AFFE67C9B7.taxon	diagnosis	Diagnosis. A medium sized Eremias, maximum SVL 57 mm, TL more than two times longer than the body (max. 122 mm). Supraocular plates usually in contact with frontal and frontoparietal plates, no full row of small granular and roundish scales in between, but sometimes tiny elongate scales between the supraocular and frontal plates are visible; fifth submaxillary small, 5 – 6 times smaller than fourth submaxillary, usually in contact with infralabials or separated by one row of granular scales; collar scales not strongly distinct from surrounding gular scales, usually only a few larger than the others. Scales at midbody 44 – 51; 14 – 19 well developed femoral pores in row, left and right rows separated by 4 – 7 scales; series of femoral pores not reaching the knee bend by 2 – 3 scales. Coloration. Dorsal pattern comprising alternating dark and light longitudinal, regular, contrasting stripes, usually seven dark stripes in total. Two stripes on the sides of the body are darker, wider, and not as contrasting as the stripes on the dorsum. Wide, dark, longitudonal stripes without light spots begin behind the eye and pass along the sides of the body, continuing along the tail. Narrow, dark, longitudinal line between fore and hind limbs marking the ventrolateral border of the body. Juveniles have vermiculated patterns on the middle of the dorsum, formed by dark and light stripes. Sometimes adult specimens have a narrow vermiculate pattern along the vertebral line. No yellow spot on the tympanic plate. Dorsal limb surface dark brown with round light or yellowish spots. The belly is whitish and the tip of the tail light yellowish.	en	Orlova, Valentina F., Rasegar-Pouyani, Eskandar, Rajabizadeh, Khosrow, Nabizadeh, Hossein, Poyarkov, Nikolay A., Melnikov, Daniel A., Nazarov, Roman A. (2023): Taxonomic diversity of racerunners with descriptions of two new Eremias species (Sauria: Lacertidae) from Central Iran. Zootaxa 5369 (3): 336-368, DOI: 10.11646/zootaxa.5369.3.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5369.3.2/52247
FA013919F2113B1643D0D9AFFE67C9B7.taxon	description	Description of Holotype (Fig. 11). ZMMU R- 12982. Adult male, SVL 56.7 mm, TL 112.3 mm, Pa 21.5 mm, Pp 36.7 mm, L. pil. 12.5 mm. Meristic features: G 26, Pfm (right / left) 18 / 18, Dist. Pfm 4, labialia (left / right) 9 / 9, infralabilia (left / right) 7 / 7, ventralia 31. Tail scales smooth on the base, not strongly keeled; Sq. c. cd 23, subdigital lamellae 26. Smooth, roundish dorsal scales; Sq 51, Sq. dorsal 151 Head and body moderately elongated and slightly depressed. Head length (14.5 mm) 1.88 times its width (7.7 mm) and 2.3 times as its height (6.3 mm); nasals smooth or slightly swollen; rostrum approximately the same size as postocular part of the head; head wider than neck. Fore and hind limbs relatively long; Pa / SVL 0.37, Pp / SVL 0.64. Hind limb reaches the axilla of forelimb and extends far beyond it. Tail about twice the length of the body, TL / SVL 0.5, cylindrical and slightly depressed at the base. Pileus (Lpil 12.5 mm) plates slightly swollen. Nasals in broad contact behind the rostral, the suture between them approximately 2 / 5 the length of frontonasal. Frontonasal width 1 / 3 more than its length. Prefrontal slightly longer than broad, forming median suture. Length of frontal same as the distance between rostral to frontal, two times wider in front than behind and narrow in the middle. Frontoparietals somewhat longer than broad, weakly rugose. Interparietal teardrop-shaped, wider in front, with parietal window. Parietals somewhat longer than broad, weakly rugose, surrounded by a row of small elongate scales, different from other dorsal scales, and the next row on the middle of neck formed by triangular scales. No occipital. Two large roundish, approximately equal sized and one small elongate posterior supraoculars. The space in front of the supraoculars occupied by one large plate, another half that size, and several granules. Supraoculars in contact with frontoparietals, but anterior supraocular plates separated from frontal by very tiny and elongate scales (not granular). Five supraciliars, anterior and posterior enlarged, separated by a single series of granules from supraoculars. Three nasals, the lower in contact with first and second supralabials and not touching the rostral. Large loreal slightly higher than broad, with two additional plates 2.5 – 3 times smaller anteriorly and posteriorly. Subocular smooth with broad contact with mouth, wedged between sixth and seventh supralabials. Six supralabials before subocular and three more posteriorly, the seventh infralabial on each side. Temporal scales smooth; no auricular denticulation scales; lower eyelid translucent and covered with small scales. Five pairs of submaxillaries, three anterior pairs in full contact with each other on the midline, 4 th and 5 th pairs in contact with 5 th, 6 th, and 7 th infralabials. Between 6 th and 7 th infralabials and 4 th and 5 th submaxillaries, there is one small additional scale. Twenty six gular scales in a straight line between the symphysis of the submaxillary and the collar. Collar weakly developed and formed by seven slightly enlarged scales. Dorsal scales granular, smooth, 51 across middle of the body and 151 along middle of dorsum. Ventral plates, broader than long, forming oblique longitudinal rows of 16 plates, in 31 slightly angular transverse rows. Precloacal region with one enlarged median plate, just above the cloaca; 18 femoral pores on each side, separated by four scales; length of the series of femoral pores is four times the distance between these two rows. Femoral pores do not reach the knees bend by two ventral femoral scales on the right and left sides. Upper surface of arm with rhombic, smooth scales. Scales on upper surface of hind limbs similar in size to the dorsal scales, enlarged plates covering the lower surface of hind limbs. Subdigital lamellae weakly carinate, 29 scales under fourth toe. Three scales around the toes. Tail expanded at the base, covered by weakly keeled elongate scales, around 9 – 10 tail segments with 23 scales. Coloration and patterns. The dorsal pattern of Eremias pseudofasciata comprises alternating dark and light longitudinal contrasting stripes, with eight dark strips on the dorsum. On the body sides, the dark stripes are wider and not as contrasting as those on the dorsum. A wide, dark, longitudinal stripe begins behind the eye, passes along the sides of the body, and continues along the entire tail. A narrow dark longitudinal line is present between the fore and hind limbs, marking the ventrolateral border of the body. A white tympanic plate is present. The dorsal surface of the limbs is dark brown with round light or yellowish spots. The tip of the tail slightly yellowish. The ventral surface of the body is white.	en	Orlova, Valentina F., Rasegar-Pouyani, Eskandar, Rajabizadeh, Khosrow, Nabizadeh, Hossein, Poyarkov, Nikolay A., Melnikov, Daniel A., Nazarov, Roman A. (2023): Taxonomic diversity of racerunners with descriptions of two new Eremias species (Sauria: Lacertidae) from Central Iran. Zootaxa 5369 (3): 336-368, DOI: 10.11646/zootaxa.5369.3.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5369.3.2/52247
FA013919F2113B1643D0D9AFFE67C9B7.taxon	distribution	Distribution. Eremias pseudofasciata is only known from the type locality and a nearby location (see Paratypes).	en	Orlova, Valentina F., Rasegar-Pouyani, Eskandar, Rajabizadeh, Khosrow, Nabizadeh, Hossein, Poyarkov, Nikolay A., Melnikov, Daniel A., Nazarov, Roman A. (2023): Taxonomic diversity of racerunners with descriptions of two new Eremias species (Sauria: Lacertidae) from Central Iran. Zootaxa 5369 (3): 336-368, DOI: 10.11646/zootaxa.5369.3.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5369.3.2/52247
FA013919F2113B1643D0D9AFFE67C9B7.taxon	etymology	Etymology. Since this new population more closely resembles Eremias fasciata on the basis of morphology and molecular analysis (Fig. 5 – 6), we named this new species “ pseudofasciata, ” i. e., false fasciata. Natural history. Eremias pseudofasciata inhabits weakly stabilized, isolated sand dunes on a gravel plain. The vegetation in this habitat is represented by the genera Salsola and Astragalus, and the families Brassicaceae and Poaceae (Fig. 10 b). In the same habitat, we recorded the following sympatric reptile species: Eremias fasciata Blanford, 1874, E. lineolata (Nikolsky, 1897), Phrynocephalus mystaceus (Pallas, 1776), Crossobamon eversmanni (Wiegmann, 1834), Teratoscincus keyserlingii Strauch, 1863 and Bunopus tuberculatus Blanford, 1874. Comparison. Eremias pseudofasciata sp. nov. is most closely related to E. fasciata (Fig. 12) and can be distinguished by the following characters: lower number of femoral pores, with 14 – 19 versus 15 – 22 in E. fasciata, supraocular plates in a contact with frontal and frontoparietals versus separate supraocular from frontal, and full row of small granular and roundish scales in E. fasciata, colour of the tympanic plate yellow in E. fasciata versus white in E. pseudofasciata, contrasting longitudinal dark stripes in E. pseudofasciata versus no contrasting stripes in E. fasciata. Additionally, E fasciata has a dark, wide contrasting stripe on the upper part of the temporal region beginning at the posterior margin of the eye and ending above the ear opening, with 2 – 3 bluish-grey light roundish spots on it, while E. pseudofasciata has two dark stripes on the temporal region without light spots. One of these stripes is wider and begins at the posterior side of the eye and continues along the body to the tail, the second stripe starts at the posterior margin of the suboculars and runs to the middle of the ear opening. Rounded light spots on the femur and tibia are noticeably smaller in E. pseudofasciata in contrast to E. fasciata (Fig. 12). Eremias pseudofasciata sp. nov. clearly differs from E. graphica sp. nov. by lower number of Sq — 44 – 51 versus 54 – 65, by its striped dorsal pattern versus vermiculate, and by its number of femoral pores — 14 – 19 versus 8 – 17 in E. graphica. Eremias pseudofasciata differs from E. lineolata by the mean number of femoral pores 16.07 (14 – 19) versus 13.6 (9 – 17), the distance between the series of femoral pores being four times the distance in E. pseudofasciata versus two times the distance between right and left rows in E. lineolata, the number of scales around the 9 th – 10 th tail segment — 21 – 26 versus 12 – 17, supraocular plates separated from frontals and frontoparietals by full row of small granular and roundish scales in E. lineolata versus contact between supraocular and frontal in E. pseudofasciata, the number of subdigital lamellae on the fourth toe — 24 – 30 versus 19 – 24, color of the tail venter slightly yellow versus orange-reddish in E. lineolata, and two dark stripes on the temporal region without any light spots in E. pseudofasciata versus two dark contrasting stripes in the temporal region with the upper being wider and containing light spots in E. lineolata. E. pseudofasciata can be distinguished from E. andersoni by its larger SVL (max SVL) of 57 mm versus 40 mm, smaller number of ventral scales — 28 – 29 versus 31 – 33 in E. andersoni, and length of the series of femoral pores four times the distance between these two rows in E. pseudofasciata versus three times in E. andersoni. In addition, E. andersoni has light-bluish coloration on the tip of tail versus a light-yellowish tail tip in E. pseudofasciata. Eremias pseudofasciata can be clearly distinguished from E. scripta by the greater number of femoral pores — 8 – 17 versus 14 – 18, the row of femoral pores almost reaching the knee in E. pseudofasciata but noticeably shorter in E. scripta, and gular scales 26 – 29 versus 15 – 23. In addition, the new species is characterised by smooth caudal scales, whereas for E. scripta has clearly visible keels on subcaudal scales. Eremias pseudofasciata differs from E. cholistanica by its larger number of gular scales — 26 – 29 versus 20 – 24 and the number of scales around the tail segment 21 – 26 versus 27 – 35. From all other Eremias species inhabiting the region, the new species can be distinguished by a combination of the above-described characters.	en	Orlova, Valentina F., Rasegar-Pouyani, Eskandar, Rajabizadeh, Khosrow, Nabizadeh, Hossein, Poyarkov, Nikolay A., Melnikov, Daniel A., Nazarov, Roman A. (2023): Taxonomic diversity of racerunners with descriptions of two new Eremias species (Sauria: Lacertidae) from Central Iran. Zootaxa 5369 (3): 336-368, DOI: 10.11646/zootaxa.5369.3.2, URL: https://www.mapress.com/zt/article/download/zootaxa.5369.3.2/52247
