taxonID	type	description	language	source
F92BAE363417FFEFFF5811EDEFF9FBB6.taxon	materials_examined	Holotype: male (CUMZ) THAILAND, Chiang Mai Province, Fang District, Doi Angkhang, near Royal Agricultural Station, 19 º 54 ' 26 " N, 99 ° 2.0 ' 26 " E, ca 1,425 m a. s. l., 12 August 2014, leg. N. Likhitrakarn. Paratypes: 2 males, 2 females (CUMZ) same data with holotype; 1 male, 2 females (CUMZ), THAILAND, Chiang Mai Province, Chiang Dao District, Wat Tham Krab, 19 º 33 ' 31.92 " N, 99 ° 3.0 ' 46.91 " E, ca 620 m a. s. l., 25 October 2015, leg. C. Sutcharit, R. Srisonchai, T. Seesamut and A. Pholyotha.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE363417FFEFFF5811EDEFF9FBB6.taxon	etymology	Etymology. The species epithet is a noun in apposition and refers to the shape of the paraterga which reminds us of an ancient Egyptian flint knife (des).	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE363417FFEFFF5811EDEFF9FBB6.taxon	diagnosis	Diagnosis. Metaterga with 2 transverse rows of setae, 2 + 2 setae in anterior row, 2 + 2 in posterior one; shocking pink paraterga contrasting with a dark brown general body color. D. des sp. n. shares these characters with D. planata, but differs from that species in having wing-shaped, knifelike paraterga, having the lateral sulcus between femorite and postfemorite poorly developed, having a prominent sulcus separating two ridges dorsally on lamina lateralis (ll), having a dentiform lamella on lamina medialis (lm), having male femora 5 and 6 very strongly inflated and stout, and in having the hypoproct with a concave caudal margin.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE363417FFEFFF5811EDEFF9FBB6.taxon	description	Description. Length 26 – 34 mm (male), 30 – 34 mm (female), width of midbody prozona and metazona ca 1.8 and 2.5 mm (male), ca 2.2 and 3.0 mm (female). Live coloration of body dark brownish; paraterga bright pink; surface below paraterga, head and metaterga dark brown; legs, antenna, epiproct and sterna brown; a few basal podomeres brownish pink (Figs. 1 A, 1 B, 2). Coloration in alcohol after twelve months faded to dark brown; metaterga and surface below paraterga dark brownish; paraterga, head, antenna, legs, sterna and epiproct brown. Width of head <collum <segment 2 = 3 <4 = 5 <6 – 16, thereafter body gradually tapering towards telson. Head width 2.0 – 2.2 mm (male), 2.0 – 2.5 mm (female). Clypeolabral region sparsely setose, vertex bare; labrum and genae sparsely setose; epicranial suture distinct, shallow (Figs. 4 A, 4 D). Antennae quite long, surpassing segment 3 when stretched backward dorsally (both sexes) (Fig. 4 D). Collum ca 4.0 mm wide (both sexes), surface smooth and dull, with two transverse rows of inconspicuous setae; 3 + 3 anterior and 1 + 1 posterior setae; anterior margin subtruncate, posterior margin quite convex; paraterga strongly developed, wing-shaped, directed dorsolaterad at about 30 º; with an inconspicuous notch on anterior margin (Fig. 4 A). Post-collum segments with surface of prozona and metazona finely shagreened, quite dull; paraterga smooth; surface below paraterga coarsely microgranulate with distinct wrinkles; sterna smooth (Figs. 2 A – H, 4 G). Suture between prozona and metazona wide, quite deep (Figs. 2 A – F, 4 B, 4 E, 4 G). Metaterga with two transverse rows of setiferous tubercles; metaterga 2 – 18 with 2 + 2 anterior and 2 + 2 posterior tubercles, all tubercles subequal in size; distance between the two mesal tubercles of anterior rows larger than distance between the mesal and the lateral tubercles on each side; distance between the two mesal tubercles of posterior rows smaller than distance between the mesal and the lateral tubercles on each side; metatergum 19 with 2 + 2 anterior and 2 + 2 posterior inconspicuous setae. Transverse sulcus on metaterga quite deep on segments 4 – 17, poorly expressed on segments 3 and 18, entirely missing on segments 2 and 19 (Figs. 2 A, 2 C, 2 F, 4 A – C). Mid-dorsal line absent. Paraterga strongly developed, very broad at base, wing-shaped, directed dorsolaterad at about ca 30 º (Fig. 4 F), tapering and curving backwards, ending in sharp point; shoulder present; anterior margin with two distinct denticles, on metaterga 9, 10, 12, 13, 15 – 18 with an additional tiny denticle close to the tip (Fig. 4 H: arrow); calluses on anterior margin of pore-bearing segments distinct (Figs. 2 C, 2 E, 4 H). Ozopore (op) visible from above, ovoid, located on anterior margin of paratergum (Fig. 4 H). Pleurosternal carinae forming complete, tooth-like crests on segment 2, small incomplete ridges on segment 3, missing on remaining segments (Fig. 2 B). Epiproct conoid; tip truncate; apical papillae without tubercles; with 2 pairs of paramedian setae without supporting tubercles; lateral papillae with conspicuous tubercles lying close to the tip (Figs. 4 J – L). Hypoproct subtrapeziform; caudal margin quite concave, with very small setiferous tubercles (Fig. 4 I). Sterna sparsely setose, cross-impression shallow; sternal process between male coxae 4 modified, sparsely setose, subquadrate, tip truncate, two sternal pores on sternal process visible from posterior view (Figs. 2 G, 2 H, 4 M). Legs long and slender, 3 – 3.2 (male) or 3.1 – 3.2 times (female) as long as midbody height (Fig. 4 P). Male femora 5 and 6 distinctly humped ventrally in middle part, femur 5 subequal in width and length to femur 6, delicately and sparsely setose, tarsal brush absent (Figs. 2 I, 4 N, 4 O). Gonopods (Figs. 3, 4 Q – T) long, distal parts strongly condensed. Coxa (cx) about half as long as telopodite, dorsally densely setose, cannula long and slender. Prefemorite (pfe) almost half as long as telopodite, subequal in length to femorite, densely setose. Femorite (fe) elongated, with a prominent deep sulcus on mesal side demarcating the border between femorite and postfemoral part (Fig. 4 Q, arrow), lateral sulcus poorly developed, seminal groove running entirely on mesal surface. Solenophore (sph) well-developed; lamina lateralis (ll) apically with a sulcus separating two ridges, inner ridge large, outer one smaller (Fig. 4 T, arrow); lamina medialis (lm) broadly expanded, distally curving mesad, with dentiform lamella (Fig. 4 S, arrow). Solenomere (sl) quite long, flagelliform, directed straight, twisted distally.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE363417FFEFFF5811EDEFF9FBB6.taxon	distribution	Distribution and habitat. This species is known only from its type locality and another locality in Chiang Mai Province, the two localities lie approximately 50 km apart. The holotype was taken from the limestone hill near the Royal Agricultural Station, Angkhang during the rainy season. Additional specimens were also found in limestone habitats.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE363417FFEFFF5811EDEFF9FBB6.taxon	discussion	Remark. The color of living examples of the new species is clearly aposematic and remarkable for the contrast between dark brownish body and the pink paraterga.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE363410FFE3FF5813FFE854FBF2.taxon	materials_examined	Holotype. male (CUMZ) THAILAND, Phrae Province, Long District, before Sareethai Cave, 18 º 16 ' 43 " N, 100 ° 03 ' 29 " E, ca 264 m a. s. l., 21 October 2014, leg. C. Sutcharit, W. Siriwut, K. Inkhavilay and R. Srisonchai. Paratypes. 27 males, 7 females (CUMZ), 3 males, 2 females (ZMUC) same data with holotype; 5 males, 3 females (CUMZ) same locality together with holotype, 21 July 2008, leg. N. Likhitrakarn; 1 male, 3 females (CUMZ) THAILAND, Lampang Province, Mae Tha District, Nakraua Subdistrict, Wat Tham Phra Sabai, 18 º 05 ' 32 " N, 99 ° 32 ' 03 " E, ca 328 m a. s. l., 21 July 2008, leg. S. Panha, P. Tongkerd and N. Likhitrakarn; 1 male, 5 females (CUMZ) THAILAND, Lampang Province, Mae Tha District, Tham Chakkrabhat monastery (Wat Tham Chakkrabhat), 18 º 06 ' 02 " N, 99 ° 56 ' 48 " E, ca 237 m a. s. l., 8 October 2007, leg. U. Bantaowong, R. Chanabun, P. Pimvichai and T. Krutchuen.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE363410FFE3FF5813FFE854FBF2.taxon	etymology	Etymology. The name is a Latin noun in apposition, meaning “ short intromittent organ ”, and referring to the short, flagelliform solenomere.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE363410FFE3FF5813FFE854FBF2.taxon	diagnosis	Diagnosis. Body color shocking pink, paraterga wing-shaped, metaterga 2 – 8 with 2 + 2 spines in anterior row and 2 + 2 spines in posterior one, metaterga 9 – 19 with 2 + 2 spines in anterior row and 3 + 3 spines in posterior one. Similar in these respects to D. taurina (Pocock, 1895), D. purpurosea and D. takensis sp. n. Differs from D. purpurosea and D. takensis sp. n. by having 3 + 3 setiferous tubercles in anterior row of collum, by having the sternal process between male coxae 4 subquadrate with an emarginate tip, by having an emarginate process on lamina medialis (lm), and by having a very short solenomere. Differs from D. taurina by having strongly humped male femora 5 and 6, by having paraterga well developed, extremely elevated, and by having longer spines on metaterga.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE363410FFE3FF5813FFE854FBF2.taxon	description	Description. Length 28 – 30 mm (male), 33 – 35 mm (female), width of midbody prozona and metazona ca 1.7 and 2.0 mm (male), 2.5 and 3.0 mm (female). Live coloration of body shocking pink to purple (Figs. 1 C, 1 D, 5); paraterga vivid pink; surface below paraterga and metaterga brown pink to brown purple; head brown; antenna dark brown; legs, sterna and epiproct pink. Coloration in alcohol after two years faded to pale brown; paraterga, surface below paraterga, metaterga, head, antenna, legs, sterna and epiproct brown to whitish. Width of head <collum = segment 2 = 3 <4 <5 <6 – 17 thereafter body gradually tapering towards telson. Head width ca 2.5 mm (male), 2.8 mm (female). Clypeolabral region sparsely setose, epicranial suture distinct (Figs. 7 A, 7 D). Antennae long and slender, reaching the end of segment 6 (male) or 5 (female) when stretched backward dorsally (Fig. 7 D). Collum width ca 3.0 mm (male) and 3.5 – 4.0 mm (female); surface coarsely microgranulate; with three transverse rows of setiferous tubercles, 3 + 3 anterior, 1 + 1 intermediate and 2 + 2 posterior tubercles; posterior margin concave; paraterga well-developed, wing-liked, elevated at ca 30 º and directed dorsolaterad, tip pointed, anterior margin with two setiferous notches (Fig. 7 A). Post-collum segments with prozonae finely shagreened; surface below paraterga and metazonae coarsely microgranulate; paraterga and sterna quite smooth (Figs. 5 A – H, 7 G). Suture between prozona and metazona conspicuous, shallow, narrow on segments 5 and 6, thereafter much wider (Figs. 5 A – F, 7 B, 7 E, 7 G). Metaterga with two distinct transverse rows of rose-thornlike setiferous spines; metaterga 2 – 8 with 2 + 2 spines in anterior and 2 + 2 spines in posterior row, lateral spines of posterior rows larger than inner spines; metaterga 9 – 18 with 2 + 2 anterior and 3 (4) + 3 (4) posterior spines, lateral spines of posterior row longest, intermediate ones shorter and mesal ones shortest, distance between the two mesal spines of posterior rows smaller than distance between the mesal and the lateral spines on each side; metatergum 19 with 2 + 2 anterior and 3 + 3 posterior spines, all spines equal in size. Transverse sulcus on metaterga conspicuous and quite deep on segments 5 – 17, poorly developed on segments 4 and 18 (Figs. 5 A, 5 C, 5 F, 7 A – C). Mid-dorsal line missing. Paraterga strongly developed, especially in male, winglike, directed dorsolaterad at about 45 – 50 º (Fig. 7 F); shoulder present, obviously rounded, fused to calluses; calluses on anterior margin of pore-bearing segments conspicuous; anterior margin with two conspicuous denticles; metaterga 9, 10, 12, 13, 15 – 17 with a very small additional denticle near tip (Fig. 7 H); tip pointed, long (Figs. 5 C, 5 E, 7 H). Ozopore (op) conspicuous, visible from above (Fig. 7 H). Pleurosternal carinae forming a tooth-like crest on segment 2, a small tooth on segment 3, absent on remaining segments (both sexes) (Fig. 5 B). Epiproct conical, flattened ventrally; tip subtruncate; apical papillae without tubercles; paramedian setae with supporting by tiny tubercles; lateral papillae inconspicuous, small tubercles (Figs. 7 J – L). Hypoproct subsemicircular; caudal margin quite convex, with a pair of small setigerous tubercles (Fig. 7 I). Sterna sparsely setose, cross-impression shallow; sternal process between male coxae 4 modified, subquadrate, tip emarginate, two sternal pores on sternal process visible from posterior view (Figs. 5 G, 5 H, 7 M, 7 N). Legs very long and slender, ca 3.2 – 3.5 (male) and 2.2 – 2.5 times (female) as long as mid body height (Fig. 7 Q). Male femora 5 and 6 modified; distinctly humped at middle part, (Figs. 5 I, 4 O, 4 P). Gonopods (Figs. 6, 7 R – U) long, distal parts strongly condensed. Coxa (cx) about half as long as telopodite; cannula (ca) long, broadened basally. Prefemorite (pfe) about 2 / 3 as long as telopodite. Femorite (fe) very long and slender, slightly separated from postfemoral part by distinct lateral and mesal sulcus (Figs. 7 R, black arrow; 7 S, arrow), with seminal groove running entirely on mesal surface. Solenophore (sph) condensed: lamina lateralis (ll) with a digitiform, vertical ventral lobe on ventral surface (Figs. 7 R & 7 T: white arrow); lamina medialis (lm) consisting of one process and two lobes; process prominent, directed mesad, spine-shaped, tip emarginate (Fig. 7 U: P arrow); first lobe situated on top, terminating in two lamellae (Fig. 7 U: L 1 arrow); second lobe long, blunt, twisted at base (Fig. 7 U: L 2 arrow). Solenomere (sl) very short, flagelliform, straight, a bit curving distad.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE363410FFE3FF5813FFE854FBF2.taxon	distribution	Distribution and habitat. Known only from the type locality and nearby areas. The holotype and accompanying paratypes were crawling on logs and litter in limestone forest near the main road to Sareethai cave.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE363410FFE3FF5813FFE854FBF2.taxon	discussion	Remark. This species exhibits a remarkable shocking pink color, no doubt aposematic. The specimens present some variation of the sternal process between the male coxae 4, some specimens with a slightly emarginate tip, the others with deep emarginate tip (Figs. 7 M, 7 N). Very similar and probably closely related to D. purpurosea.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE36341CFFE7FF5813BDEE9AFCDE.taxon	materials_examined	Holotype. male (CUMZ) THAILAND, Tak Province, Phobphra District, Nangkruen waterfall, on litters and under decaying bark, 16 º 24 ' 36 " N, 98 ° 41 ' 21 " E, ca 398 m a. s. l., 15 January 2015, leg. R. Srisonchai, T. Seesamut and P. Jirapatrasilp. Paratypes. 12 males, 10 females, 1 juvenile (CUMZ), 2 males, 1 female (ZMUC) same data with holotype; 2 males, 1 female (CUMZ) same locality, 18 January 2011, leg. C. Sutcharit, R. Chanabun, N. Likhitrakarn and T. Krutchuen.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE36341CFFE7FF5813BDEE9AFCDE.taxon	etymology	Etymology. The name is Latin adjective referring to the province where the type locality occurs.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE36341CFFE7FF5813BDEE9AFCDE.taxon	diagnosis	Diagnosis. Paraterga winglike, metaterga 2 – 8 with 2 + 2 anterior and 2 + 2 posterior spines, metaterga 9 – 19 with 2 + 2 anterior and 3 + 3 posterior spines. Sharing these characters with D. taurina, D. purpurosea and D. breviverpa sp. n, but differs by its red live color; by smaller paraterga; by the sternal process between male coxae 4 being subtrapeziform; by lamina lateralis (ll) being subtriangular with a thumblike ventral lobe, and by having one conspicuous process and one lobe on lamina medialis (lm).	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE36341CFFE7FF5813BDEE9AFCDE.taxon	description	Description. Length 24 – 26 mm (male), 25 – 27 mm (female), width of midbody prozona and metazona ca 1.5 and 1.9 mm (male), 1.7 and 2.2 mm (female). Live coloration of body bright red; paraterga, surface below paratega and metaterga red; head, antennae, legs and epiproct brown red (Figs. 1 E, 1 F). Coloration in alcohol after twelve months changed to pale brown; paraterga, surface below paraterga, legs and sterna brown to yellow; head, antenna and epiproct brown. Width of head = collum = segment 2 = 3 = 4 <5 <6 – 16, thereafter body gradually tapering towards telson. Head width ca 2.0 mm (male), 2.5 mm (female). Clypeolabral region, labrum and genae sparsely setose; epicranial suture distinct as brownish stripe (Figs. 10 A, 10 D). Antennae very long and slender, surpassing segments 6 (male) and 5 (female), when stretch backward dorsally (Fig. 10 D). Collum width 2.5 – 3.0 mm (both sexes), with three transverse rows of setiferous tubercles; 4 + 4 anterior, 1 + 1 intermediate and 2 + 2 posterior tubercles, intermediate tubercles larger than other ones, lateral tubercles of posterior row displaced anteriad; paraterga wing-shaped, subhorizontal, elevated at 10 – 15 º, ending in sharp point, with a setiferous notch at about halfway (Fig. 10 A). Post-collum segments with surface metazona and surface below paraterga microgranulate; prozona shagreened; sterna and paraterga smooth (Figs. 8 A – H, 10 G). Suture between prozonae and metazonae conspicuous, wide (Figs. 8 C, 10 B, 10 E, 10 G). Metaterga with two transverse rows of setiferous spines; metaterga 2 – 8 with 2 + 2 anterior and 2 + 2 posterior spines; metaterga 9 – 17 with 2 + 2 anterior and 3 + 3 posterior spines; lateral spines of posterior rows longer than mesal ones; metaterga 18 and 19 with 2 + 2 anterior spines and 3 + 3 posterior tubercles, subequal in size. Transverse sulcus on metaterga visible on segments 6 – 17, incomplete on segments 5 and 18, absent on segments 2, 3, 4 and 19. Mid-dorsal line missing (Figs. 8 A, 8 C, 8 F, 10 A – C). Paraterga strongly developed, especially well in male, winglike, directed dorsolaterad at about ca 45 º (Fig. 10 F); shoulder present, quite narrow; calluses present; anterior margin with two conspicuous denticles; on segments 9, 10, 12, 13, 15 – 18 with a very small additional denticle close to the tip (Fig. 10 H); tip narrow and sharp; posterior margin almost straight (Figs. 8 C, 8 E, 10 H). Ozopore (op) large, ovoid, visible from above (Figs. 8 C, 8 E, 10 H). Pleurosternal carinae forming a complete, toothlike crest only on segment 2 (both sexes), a small crest on segment 3, absent on remaining segments (Fig. 8 B). Epiproct conical, flattened dorsoventrally; tip subtruncate; apical papillae inconspicuous, without tubercles; two pairs of paramedian setae without tubercles, lateral papillae conspicuous, lying close to the tip (Figs. 10 J – K). Hypoproct subsemicircular, caudal margin slightly convex, very small inconspicuous tubercles on caudal edges (Fig. 10 I). Sterna sparsely setose, cross-impression shallow; sternal process between male coxae 4 modified, subtrapeziform, stout, tip rounded, with two pores in posterior view (Figs. 8 G, 8 H, 10 M). Legs very long and slender, ca 3.0 (male), 2.6 times (female) as long as midbody height (Fig. 10 P). Male femora 5 and 6 distinctly humped in ventral part (Fig. 8 I, 10 N, 10 O). Gonopods (Figs. 9, 10 R – T) suberect and long, distal parts strongly condensed. Coxa (cx) about half as long as telopodite, with long sparse setation distoventrally. Prefemorite (pfe) densely setose, almost 1 / 3 as long as telopodite, a bit shorter than femorite. Femorite (fe) elongate and slender; seminal groove running on mesal surface; apically with conspicuous lateral and mesal sulcus demarcating femorite and postfemoral part (Fig. 10 S: arrow). Solenophore (sph) well-developed: lamina lateralis (ll) subtriangular, inner surface subsided; with a huge thumblike ventral lobe, directed in vertical plane (Fig. 10 R, arrow): lamina medialis (lm) with a prominent process, directed almost in vertical plane, tip sharp and a bit curving down (Fig. 10 T: black arrow); with a lamellar lobe curving in horizontal plane, tip projecting into solenomere (Fig. 10 T: white arrow). Solenomere long, flagelliform, straight.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE36341CFFE7FF5813BDEE9AFCDE.taxon	distribution	Distribution and habitat. Known only from the type locality. Almost all specimens of this species were found on humid litter surrounding the waterfall nearby a concreted natural board trail. Interestingly, we saw some specimens which were crawling on plastic garbages. The waterfall is a popular tourist attraction located just opposite the main road connecting to human habitation.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE36341CFFE7FF5813BDEE9AFCDE.taxon	discussion	Remark. The noticeable red body color is clearly aposematic.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE363418FFFBFF581457E926F87D.taxon	materials_examined	Holotype. male (CUMZ) THAILAND, Phitsanulok Province, Noen Maprang District, near Pra Tham Mans Monastery (Tham Wangdaeng), 16 º 41 ' 40 " N, 100 ° 40 ' 42 " E, ca 62 m a. s. l., 22 August 2014, leg. S. Panha, C. Sutcharit and U. Bantaowong. Paratypes. 22 males, 5 females (CUMZ), 2 males, 1 female (ZMUC) same data with holotype; 10 males, 17 females (CUMZ) same locality together with holotype, 23 July 2008, leg. N. Likhitrakarn; 2 females (CUMZ) same locality together with holotype, 8 September 2009, leg. U. Bantaowong and R. Chanabun; 3 males, 6 females (CUMZ) THAILAND, Phitsanulok Province, Noen Maprang District, near Pa Ma Muang monastery, 16 º 34 ' 00 " N, 100 ° 40 ' 38 " E, ca 92 m a. s. l., 23 July 2008, leg. C. Sutcharit and P. Tongkerd.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE363418FFFBFF581457E926F87D.taxon	etymology	Etymology. The name is a Latin noun in apposition, meaning “ shark fin ” and referring to the shape of a process on lamina medialis.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE363418FFFBFF581457E926F87D.taxon	diagnosis	Diagnosis. Metaterga with 2 + 2 anterior and 2 + 2 posterior spines, paraterga wing-like, shocking pink. Similar in these respect to D. planata, but differing by pink brown general body color, epiproct with digitiform apical papillae, sternal process between male coxae 4 being subquadrate, lamina medialis (lm) being highly elevated with a shark finlike process. Descriptions. Length 24 – 29 mm (male), 28 – 30 mm (female); width of midbody prozona and metazona ca 2.0 and 2.0 mm (male), 2.2 and 2.5 mm (female). Live coloration vivid pink; paraterga shocking pink; surface below below paraterga and metaterga brown pink; head and antenna dark brown; legs and epiproct pink; sterna brown (Figs. 1 G, 1 H). Coloration in alcohol after one year faded to pale brown; paraterga, legs and sterna yellow brown; surface below paraterga, metaterga, head, antenna and epiproct brown. Width of head <collum <segment 2 <3 = 4 <5 – 16, thereafter body gradually tapering towards telson. Head broad ca 2.0 mm (male), 2.5 mm (female). Clypeolabral region sparsely setose; vertex bare so on; labrum and genae sparsely setose; epicranial suture visible as dark line, quite deep (Figs. 13 A, 13 D). Antennae very long and slender, reaching back to segments 5 (male) and 4 (female) when expanded dorsally (Fig. 13 D). Collum width ca 3.0 mm (both sexes), surface coarsely microgranulate, with three transverse rows of small setiferous tubercles, 3 (4) + 3 (4) anterior, 1 + 1 intermediate and 2 + 2 posterior tubercles; paraterga wing-shaped, elevated at about 30 º (male) and 20 º (female), directed dorsolaterad, tip pointed, with two distinct setiferous notches on anterior margin (Fig. 13 A). Post-collum segments with surface of prozona finely shagreened; metazona coarsely microgranulate and shining; surface below paraterga finely microgranulate with inconspicuous wrinkles; paraterga and sterna quite smooth (Figs. 8 A – H, 13 G). Suture between prozonae and metazonae conspicuous, quite deep and wide, narrow only in segments 2 – 5, thereafter much wider (Figs. 11 C, 11 E, 13 B, 13 E, 13 G). Metaterga with two transverse rows of setiferous spines; 2 + 2 anterior and 2 + 2 posterior spines; spines of anterior rows on metaterga 2 – 16 equal in size, lateral spines of posterior rows longer than mesal ones; metaterga 17 – 19 with all spines equal in size. Transverse sulcus on metaterga distinctly expressed on segments 5 – 17, quite deep, inconspicuous on segments 4 and 18, missing on segments 2, 3 and 19 (Figs. 11 A, 11 C, 11 F, 13 A – C). Mid-dorsal line absent. Paraterga strongly developed, especially in male, wing-shaped, directed dorsolaterad at ca 45 º (Fig. 13 F), thereafter curving backward; shoulder present; calluses conspicuous, broad anteriorly and continuing slightly narrowed near tip; lateral margin with two distinct denticles, segments 9, 10, 12,13, 15 – 18 with an additional very small denticle near the tip (Fig. 13 H: arrow); tip of paraterga pointed and sharp, curving dorsolaterad; posterior margin of paraterga curving at base and almost straight near tip (Figs. 11 C, 11 E, 13 H). Ozopore (op) visible from above, ovoid (Figs. 11 C, 11 E, 13 H). Pleurosternal carinae forming complete crests on segment 2 (both sexes), small ridge on segment 3 and absent on following segments (Fig. 2 B). Epiproct conical; tip concave; apical papillae conspicuous, digitiform; with tiny tubercles supporting two pairs of paramedian setae; lateral papillae distinct (Figs. 13 J – K). Hypoproct subsemicircular, with prominent setigerous tubercles on convex caudal margin (Fig. 13 I). Sterna sparsely setose, cross-impression shallow; sternal process between male coxae 4 modified, subquadrate, tip round, sternal pores visible from posterior view (Figs. 11 G, 11 H, 13 M). Legs very long and slender, ca 2.5 (male), 2.0 times (female) as long as midbody height (Fig. 13 P). Male femora 5 and 6 distinctly humped ventrally in middle part, femur 6 a bit bigger than 5 (Figs. 11 I, 13 N, 13 O). Gonopods (Figs. 12, 13 Q – T) long and slender, suberect, distal parts strongly condensed. Coxa (cx) about 1 / 3 as long as telopodite. Prefemorite (pfe) about 1 / 3 as long as telopodite, quite stout. Femorite (fe) slender, very elongate, with seminal groove running entirely on mesal surface, postfemoral part demarcated by deep mesal and lateral sulcus (Fig. 13 R: black & white arrows). Solenophore strongly developed: lamina lateralis (ll) swollen, without a lobe: lamina medialis (lm) broad and highly elevated, with one process and two lobes; process shark finlike, tip obtuse, directed mesad (Figs. 13 Q, 13 S, P arrow); first lobe lamellalike, terminating in two lamellae, outer one smaller than the inner one (Figs. 13 Q, 13 S, L 1 arrow); second lobe lamellalike, thick and broad (Figs. 13 Q, 13 S, L 2 arrow). Solenomere flagelliform, curving distad.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE363418FFFBFF581457E926F87D.taxon	distribution	Distribution and habitat. This new species occurs in a small limestone area in the west of Thung Salaeng Luang National Park, northern Thailand and was seen crawling on humid rocks, litter and vegetation under a shading tree.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
F92BAE363418FFFBFF581457E926F87D.taxon	discussion	Remark. This species shows a bizarre pink body color which is clearly aposematic. It may be closely related to D. planata with which it shares the same patterns of color and metatergal spines.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2016): Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae). Zootaxa 4170 (1): 93-113, DOI: 10.11646/zootaxa.4170.1.4
