taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
CC99730CDF70555480B1BE618EE76EF6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333160	https://doi.org/10.3897/subtbiol.52.150357.figure1	Figure 1. Charinus santandereanus sp. nov. (female IAvH) A habitus, dorsal view B carapace, dorsal view C anterior part of carapace, frontal view D sternum, ventral view E, F right chelicera, mesal (E) and ectal (F) views G – K right pedipalp: dorsal (G), ventral (H) views I left pedipalp, ventromesal view J tarsus, dorsal view K tibia and tarsus, mesal view. Scale bars: 5 mm (A); 1 mm (B – K).	Figure 1. Charinus santandereanus sp. nov. (female IAvH) A habitus, dorsal view B carapace, dorsal view C anterior part of carapace, frontal view D sternum, ventral view E, F right chelicera, mesal (E) and ectal (F) views G – K right pedipalp: dorsal (G), ventral (H) views I left pedipalp, ventromesal view J tarsus, dorsal view K tibia and tarsus, mesal view. Scale bars: 5 mm (A); 1 mm (B – K).	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
CC99730CDF70555480B1BE618EE76EF6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333161	https://doi.org/10.3897/subtbiol.52.150357.figure2	Figure 2. Charinus santandereanus sp. nov. (female IAvH). Gonopodes: A dorsal view B posterior view. Scale bars: 0.1 mm.	Figure 2. Charinus santandereanus sp. nov. (female IAvH). Gonopodes: A dorsal view B posterior view. Scale bars: 0.1 mm.	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
CC99730CDF70555480B1BE618EE76EF6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333173	https://doi.org/10.3897/subtbiol.52.150357.figure14	Figure 14. A Charinus santandereanus sp. nov., from department of Santander, Cueva el Nitro Villanueva, Santander B Charinus tronchonii (Ravelo, 1977), from Cueva del Cuarzo, Falcón, Venezuela. Phrynus araya Colmenares & Villarreal, 2008, from department of Santander, Cueva Fumador D, E Cueva del Nitro, Santander, views of the cave interior (D) and the rocky outcrop housing the cave entrance (E). Photos: A, C Manuel F. López, B Osvaldo Villarreal D, E Gonzalo Valdivieso.	Figure 14. A Charinus santandereanus sp. nov., from department of Santander, Cueva el Nitro Villanueva, Santander B Charinus tronchonii (Ravelo, 1977), from Cueva del Cuarzo, Falcón, Venezuela. Phrynus araya Colmenares & Villarreal, 2008, from department of Santander, Cueva Fumador D, E Cueva del Nitro, Santander, views of the cave interior (D) and the rocky outcrop housing the cave entrance (E). Photos: A, C Manuel F. López, B Osvaldo Villarreal D, E Gonzalo Valdivieso.	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
CC99730CDF70555480B1BE618EE76EF6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333174	https://doi.org/10.3897/subtbiol.52.150357.figure15	Figure 15. Map showing the records of Amblypygi associated with Colombian caves.	Figure 15. Map showing the records of Amblypygi associated with Colombian caves.	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
3BEE260B388C55D2A349ACCD7A07A950.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333162	https://doi.org/10.3897/subtbiol.52.150357.figure3	Figure 3. Heterophrynus batesii, male from Caquetá (IAvH - I- 7875) A carapace, dorsal view B sternum, ventral view C, D Pedipalpal femur, dorsal (C) and ventral (D) views E, F pedipalpal patella, dorsal (E) and ventral (F) views. Scale bars: 5 mm.	Figure 3. Heterophrynus batesii, male from Caquetá (IAvH - I- 7875) A carapace, dorsal view B sternum, ventral view C, D Pedipalpal femur, dorsal (C) and ventral (D) views E, F pedipalpal patella, dorsal (E) and ventral (F) views. Scale bars: 5 mm.	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
3BEE260B388C55D2A349ACCD7A07A950.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333163	https://doi.org/10.3897/subtbiol.52.150357.figure4	Figure 4. Heterophrynus batesii (IAvH - I- 7875). Reproductive structures A, B Male spermatophore organ, dorsal (A) and ventral (B) views C, D female, gonopods, dorsal (C) and posterior (D) views A – D from SLM- 531. Scale bars: 1 mm.	Figure 4. Heterophrynus batesii (IAvH - I- 7875). Reproductive structures A, B Male spermatophore organ, dorsal (A) and ventral (B) views C, D female, gonopods, dorsal (C) and posterior (D) views A – D from SLM- 531. Scale bars: 1 mm.	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
3BEE260B388C55D2A349ACCD7A07A950.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333174	https://doi.org/10.3897/subtbiol.52.150357.figure15	Figure 15. Map showing the records of Amblypygi associated with Colombian caves.	Figure 15. Map showing the records of Amblypygi associated with Colombian caves.	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
969CE42BD10E5210808F8DCE9C234023.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333174	https://doi.org/10.3897/subtbiol.52.150357.figure15	Figure 15. Map showing the records of Amblypygi associated with Colombian caves.	Figure 15. Map showing the records of Amblypygi associated with Colombian caves.	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
41FEFEB32945504391A32559B52453D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333164	https://doi.org/10.3897/subtbiol.52.150357.figure5	Figure 5. Heterophrynus cervinus from Putumayo (IAvH - I- 7876) A carapace, dorsal view B sternum, ventral view C, D pedipalpal femur, dorsal (C) and ventral (D) views E, F pedipalpal patella and tibia, dorsal (E) an ventral (F) views G, H female gonopodes, dorsal (G) and posterior (H) views. Scale bars: 5 mm (A, C – F); 1 mm (B, G, H).	Figure 5. Heterophrynus cervinus from Putumayo (IAvH - I- 7876) A carapace, dorsal view B sternum, ventral view C, D pedipalpal femur, dorsal (C) and ventral (D) views E, F pedipalpal patella and tibia, dorsal (E) an ventral (F) views G, H female gonopodes, dorsal (G) and posterior (H) views. Scale bars: 5 mm (A, C – F); 1 mm (B, G, H).	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
41FEFEB32945504391A32559B52453D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333174	https://doi.org/10.3897/subtbiol.52.150357.figure15	Figure 15. Map showing the records of Amblypygi associated with Colombian caves.	Figure 15. Map showing the records of Amblypygi associated with Colombian caves.	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
58E8FDFF11DC582FB43E8EDA92A9E441.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333165	https://doi.org/10.3897/subtbiol.52.150357.figure6	Figure 6. Heterophrynus guacharo, male (IAvH - I- 7877) A carapace, dorsal view B sternum, ventral view C, D right chelicera, ectal (C) and mesal (D) views. Scale bars: 5 mm (A, B); 1 mm (C, D).	Figure 6. Heterophrynus guacharo, male (IAvH - I- 7877) A carapace, dorsal view B sternum, ventral view C, D right chelicera, ectal (C) and mesal (D) views. Scale bars: 5 mm (A, B); 1 mm (C, D).	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
58E8FDFF11DC582FB43E8EDA92A9E441.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333166	https://doi.org/10.3897/subtbiol.52.150357.figure7	Figure 7. Heterophrynus guacharo, male (IAvH - I- 7877) A, B pedipalpal femur, dorsal (A) and ventral (B) views C, D pedipalpal patella, dorsal (C) and ventral (D). Reproductive structures: Spermatophore organ, dorsal (G) and ventral (H) views. Scale bars: 5 mm (A – D); 1 mm (G, H).	Figure 7. Heterophrynus guacharo, male (IAvH - I- 7877) A, B pedipalpal femur, dorsal (A) and ventral (B) views C, D pedipalpal patella, dorsal (C) and ventral (D). Reproductive structures: Spermatophore organ, dorsal (G) and ventral (H) views. Scale bars: 5 mm (A – D); 1 mm (G, H).	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
58E8FDFF11DC582FB43E8EDA92A9E441.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333174	https://doi.org/10.3897/subtbiol.52.150357.figure15	Figure 15. Map showing the records of Amblypygi associated with Colombian caves.	Figure 15. Map showing the records of Amblypygi associated with Colombian caves.	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
99C1D0E4D3B05134A9082FDA9C8D6F5D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333167	https://doi.org/10.3897/subtbiol.52.150357.figure8	Figure 8. Heterophrynus yarigui (IAvH - I- 7878) A carapace, dorsal view B sternum, ventral view C, D pedipalpal femur, dorsal (C) and ventral (D) views E, F. pedipalpal patella, dorsal (E) and ventral (F) views. Scale bars: 5 mm.	Figure 8. Heterophrynus yarigui (IAvH - I- 7878) A carapace, dorsal view B sternum, ventral view C, D pedipalpal femur, dorsal (C) and ventral (D) views E, F. pedipalpal patella, dorsal (E) and ventral (F) views. Scale bars: 5 mm.	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
99C1D0E4D3B05134A9082FDA9C8D6F5D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333168	https://doi.org/10.3897/subtbiol.52.150357.figure9	Figure 9. Heterophrynus yarigui (IAvH - I- 7878). Reproductive structures A, B male spermatophore organ, dorsal (A) and ventral (B) views C, D female, gonopods, dorsal (C), and posterior (D) views. Scale bars: 1 mm.	Figure 9. Heterophrynus yarigui (IAvH - I- 7878). Reproductive structures A, B male spermatophore organ, dorsal (A) and ventral (B) views C, D female, gonopods, dorsal (C), and posterior (D) views. Scale bars: 1 mm.	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
99C1D0E4D3B05134A9082FDA9C8D6F5D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333174	https://doi.org/10.3897/subtbiol.52.150357.figure15	Figure 15. Map showing the records of Amblypygi associated with Colombian caves.	Figure 15. Map showing the records of Amblypygi associated with Colombian caves.	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
95061BEEAD5E53E3ADCDC690C940A60E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333169	https://doi.org/10.3897/subtbiol.52.150357.figure10	Figure 10. Paraphrynus laevifrons from Provendicia Island (IAvH - I- 7880) A carapace, dorsal view B sternum, ventral view C, D pedipalpal femur, dorsal (C) and ventral (D) views E, F pedipalpal patella, dorsal (E) and ventral (F) views G, H male spermatophore organ, dorsal (G) and ventral (H) views I, J female gonopodes, dorsal (I) and posterior (J) views. Scale bars: 1 mm (A, B); 5 mm (C – F).	Figure 10. Paraphrynus laevifrons from Provendicia Island (IAvH - I- 7880) A carapace, dorsal view B sternum, ventral view C, D pedipalpal femur, dorsal (C) and ventral (D) views E, F pedipalpal patella, dorsal (E) and ventral (F) views G, H male spermatophore organ, dorsal (G) and ventral (H) views I, J female gonopodes, dorsal (I) and posterior (J) views. Scale bars: 1 mm (A, B); 5 mm (C – F).	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
95061BEEAD5E53E3ADCDC690C940A60E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333170	https://doi.org/10.3897/subtbiol.52.150357.figure11	Figure 11. Paraphrynus laevifrons (IAvH - I- 7880). Reproductive structures A, B male spermatophore organ, dorsal (A) and ventral (B) views C, D female, gonopods, dorsal (C), and posterior (D) views. Scale bars: 1 mm.	Figure 11. Paraphrynus laevifrons (IAvH - I- 7880). Reproductive structures A, B male spermatophore organ, dorsal (A) and ventral (B) views C, D female, gonopods, dorsal (C), and posterior (D) views. Scale bars: 1 mm.	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
95061BEEAD5E53E3ADCDC690C940A60E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333174	https://doi.org/10.3897/subtbiol.52.150357.figure15	Figure 15. Map showing the records of Amblypygi associated with Colombian caves.	Figure 15. Map showing the records of Amblypygi associated with Colombian caves.	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
A71D2CE26DC2500298A77F49216F677D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333171	https://doi.org/10.3897/subtbiol.52.150357.figure12	Figure 12. Phrynus araya, female from Santander (IAvH - I- 7883) A carapace, dorsal view B sternum, ventral view C, D pedipalpal femur, dorsal (C) and ventral (D) views E, F pedipalpal patella, dorsal (E) and ventral (F) views. Scale bars: 1 mm.	Figure 12. Phrynus araya, female from Santander (IAvH - I- 7883) A carapace, dorsal view B sternum, ventral view C, D pedipalpal femur, dorsal (C) and ventral (D) views E, F pedipalpal patella, dorsal (E) and ventral (F) views. Scale bars: 1 mm.	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
A71D2CE26DC2500298A77F49216F677D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333172	https://doi.org/10.3897/subtbiol.52.150357.figure13	Figure 13. Phrynus araya (IAvH - I- 7883). Reproductive structures A, B male spermatophore organ, dorsal (A) and ventral (B) views C, D female, gonopods, dorsal (C), and posterior (D) views. Scale bars: 1 mm.	Figure 13. Phrynus araya (IAvH - I- 7883). Reproductive structures A, B male spermatophore organ, dorsal (A) and ventral (B) views C, D female, gonopods, dorsal (C), and posterior (D) views. Scale bars: 1 mm.	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
A71D2CE26DC2500298A77F49216F677D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333173	https://doi.org/10.3897/subtbiol.52.150357.figure14	Figure 14. A Charinus santandereanus sp. nov., from department of Santander, Cueva el Nitro Villanueva, Santander B Charinus tronchonii (Ravelo, 1977), from Cueva del Cuarzo, Falcón, Venezuela. Phrynus araya Colmenares & Villarreal, 2008, from department of Santander, Cueva Fumador D, E Cueva del Nitro, Santander, views of the cave interior (D) and the rocky outcrop housing the cave entrance (E). Photos: A, C Manuel F. López, B Osvaldo Villarreal D, E Gonzalo Valdivieso.	Figure 14. A Charinus santandereanus sp. nov., from department of Santander, Cueva el Nitro Villanueva, Santander B Charinus tronchonii (Ravelo, 1977), from Cueva del Cuarzo, Falcón, Venezuela. Phrynus araya Colmenares & Villarreal, 2008, from department of Santander, Cueva Fumador D, E Cueva del Nitro, Santander, views of the cave interior (D) and the rocky outcrop housing the cave entrance (E). Photos: A, C Manuel F. López, B Osvaldo Villarreal D, E Gonzalo Valdivieso.	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
A71D2CE26DC2500298A77F49216F677D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1333174	https://doi.org/10.3897/subtbiol.52.150357.figure15	Figure 15. Map showing the records of Amblypygi associated with Colombian caves.	Figure 15. Map showing the records of Amblypygi associated with Colombian caves.	2025-05-19	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.		Zenodo	biologists	Villarreal, Osvaldo;Delgado-Santa, Leonardo;Lasso, Carlos A.			
