identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
73359F3D798EA3DBA22C6FD474AC4219.text	73359F3D798EA3DBA22C6FD474AC4219.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caucasodesmus Golovatch 1985	<div><p>Caucasodesmus Golovatch, 1985</p><p>Caucasodesmus Golovatch, 1985: 40.</p><p>Type species:</p><p>Caucasodesmus inexpectatus Golovatch, 1985, by original designation.</p></div>	https://treatment.plazi.org/id/73359F3D798EA3DBA22C6FD474AC4219	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sergei I., Golovatch	Sergei I., Golovatch (2011): The millipede genus Caucasodesmus Golovatch, 1985, with the description of a new species from the Crimea, Ukraine (Polydesmida, Diplopoda, Trichopolydesmidae). ZooKeys 93: 1-8, DOI: http://dx.doi.org/10.3897/zookeys.93.1159, URL: http://dx.doi.org/10.3897/zookeys.93.1159
F054D185E0F258E072C2A6E18080054E.text	F054D185E0F258E072C2A6E18080054E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caucasodesmus tauricus	<div><p>Caucasodesmus tauricus sp. n. Figs 18</p><p>Type material:</p><p>Holotype ♂, Ukraine, Crimea, Mt Villya-Burun, Cave Villyaburunskaya, pitfall traps, 12.05.2008-12.10.2010, leg. A. Koval. - Paratype: 1 ♀, same locality, 19.07.2004-17.07.2006, leg. A. Koval.</p><p>Diagnosis:</p><p>Easily distinguished from Caucasodesmus inexpectatus Golovatch, 1985, the type, and only other, known species of this genus, by the abundantly setose metaterga and more elaborate gonopods.</p><p>Description:</p><p>Length of both sexes ca 8 mm, width of midbody pro- and metazona 0.8 and 1.5 mm, respectively. Coloration in alcohol from uniformly pallid to light yellowish.</p><p>Body with 20 segments. Tegument mainly dull, at most slightly shining, texture very delicately alveolate. Head densely pilose throughout; epicranial suture distinct but thin; isthmus between antennae ca 1.5 times broader than length of antennomere 1, still broader than diameter of antennal socket. Antennae rather short, evidently clavate due to a considerably enlarged antennomere 6, slightly overreaching segment 2 dorsally; antennomeres 2, 3 and 6 longest, subequal in length (Figs 1, 2); only antennomere 6 with a large, compact, roundish, distodorsal group of bacilliform sensilla.</p><p>In width, collum &lt;segment 2 = 3 &lt;head = 4 &lt;5=16 (♂) or head = collum = segment 2 = 4 &lt;5=16 (♀), thereafter body gradually tapering towards telson. Paraterga moderately developed, starting from collum, subhorizontal to slightly declivous, set high but always lying slightly below a faintly convex dorsum, devoid of shoulders frontally (Figs 1-3). Caudal corner of postcollum paraterga invariably spiniform, pointed, starting from segment 4 extending increasingly further than rear tergal margin. Lateral edge of paraterga with neither marginal groove nor thickening, with 5-6 clear setigerous indentations. Pore formula normal, ozopores evident, round, located laterally in front of caudalmost incision. Collum and following metaterga beset with numerous medium-sized setae set on minute knobs, polygonal bosses missing (Figs 1-3). Stricture between pro- and metazona wide, shallow and smooth. Limbus very thin, microdenticulate. Pleurosternal carinae absent (Fig. 2). Epiproct short, conical, directed caudoventrally; preapical papillae small (Fig. 4). Hypoproct subtrapeziform, setiferous papillae at caudal corners evident, rather well separated.</p><p>Sterna without modifications, poorly setose. Epigynal ridge very low. Legs rather short (Figs 2, 5), ca 1.2-1.3 (♂) or 0.9-1.0 (♀) times as long as midbody height; ♀ legs slightly slenderer; ♂ legs with clearly enlarged prefemora and femora; tarsi especially long and slender, claw long, ca 1/4 length of tarsus; sphaerotrichomes missing (Fig. 5).</p><p>Gonopod aperture large, transversely oblong-oval, taking up nearly all of ventral part of metazonite 7. Gonopods (Figs 6-8) with large, globose, medially fused coxae carrying rather numerous setae laterally, but no trace of a cannula. Telopodite subfalcate, distally of a rather short prefemoral (= setose) portion split into two branches: exomere (ex) largest and longest, more simple, whereas endomere (en) shorter, more complex in shape; an evident, tooth-like, mesal process (d) at base between both ex and en; no trace of a seminal groove.</p><p>Remarks.</p><p>This species is an unquestioned relict troglobite and, based on its zoogeographical traits, might well represent the first palaeoendemic in the diplopod fauna of the Crimea.</p><p>Only one species of Caucasodesmus has hitherto been known: Caucasodesmus inexpectatus from Cave Nyvjin Lagat (= Nyvdzhinlagat, = Tagardonskaya) in North Ossetia, central Caucasus Major (Golovatch 1984/85). The second congener, Caucasodesmus tauricus sp. n., shares with the type species such remarkable, clearly generic-level apomorphies as the absence of bacilliform sensilla on antennomeres 5 and 7, of a cannula on the gonocoxite, and of a seminal groove on a biramous gononod telopodite. Apparently, both latter characters are functionally correlated to each other, differing from the loss of a cannula alone which is observed in the families Dalodesmidae and Rhachodesmidae . The differences lie in Caucasodesmus inexpectatus showing a far more moniliform body, only three transverse rows of setae on the collum and following metaterga, and less strongly elaborate gonopod telopodites.</p></div>	https://treatment.plazi.org/id/F054D185E0F258E072C2A6E18080054E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sergei I., Golovatch	Sergei I., Golovatch (2011): The millipede genus Caucasodesmus Golovatch, 1985, with the description of a new species from the Crimea, Ukraine (Polydesmida, Diplopoda, Trichopolydesmidae). ZooKeys 93: 1-8, DOI: http://dx.doi.org/10.3897/zookeys.93.1159, URL: http://dx.doi.org/10.3897/zookeys.93.1159
