taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
F83E3A75F529FF89801FFA6D8753F1BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578329/files/figure.png	https://doi.org/10.5281/zenodo.2578329	Fig. 6. Geographic distribution of the five species of Paromoionchis gen. nov. Distinct colors are used for each mitochondrial unit found in P. tumidus (Semper, 1880) comb. nov., P. boholensis gen. et sp. nov. and P. goslineri gen. et sp. nov., and the colors used for each unit are the same as those used in Figs 1–5. Colored areas correspond to hypothetical ranges proposed based on known records (all of which are new here except for the type localities of P. tumidus and P. daemelii (Semper, 1880) comb. nov.).	Fig. 6. Geographic distribution of the five species of Paromoionchis gen. nov. Distinct colors are used for each mitochondrial unit found in P. tumidus (Semper, 1880) comb. nov., P. boholensis gen. et sp. nov. and P. goslineri gen. et sp. nov., and the colors used for each unit are the same as those used in Figs 1–5. Colored areas correspond to hypothetical ranges proposed based on known records (all of which are new here except for the type localities of P. tumidus and P. daemelii (Semper, 1880) comb. nov.).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578331/files/figure.png	https://doi.org/10.5281/zenodo.2578331	Fig. 7. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #1, live animals. A. Dorsal view, 35 mm long [3205], Philippines, Luzon (PNM 041256). B. Dorsal view, 40 mm long [5682], Vietnam (ITBZC IM 00019). C. Dorsal view, 33 mm long [3222], Philippines, Luzon (PNM 041257). D. Dorsal view, 54 mm long [5042], Indonesia, Halmahera (UMIZ 00137). E. Dorsal view, 30 mm long [1705], Australia, Northern Territory (NTM P.57622). F. Dorsal view, 26 mm long [2875], Indonesia, Seram (UMIZ 00130). G. Dorsal view, 17 mm long [2627], Australia, Queensland (MTQ). H. Dorsal view, 27 mm long [1755], Indonesia, Sumatra (UMIZ 00122).	Fig. 7. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #1, live animals. A. Dorsal view, 35 mm long [3205], Philippines, Luzon (PNM 041256). B. Dorsal view, 40 mm long [5682], Vietnam (ITBZC IM 00019). C. Dorsal view, 33 mm long [3222], Philippines, Luzon (PNM 041257). D. Dorsal view, 54 mm long [5042], Indonesia, Halmahera (UMIZ 00137). E. Dorsal view, 30 mm long [1705], Australia, Northern Territory (NTM P.57622). F. Dorsal view, 26 mm long [2875], Indonesia, Seram (UMIZ 00130). G. Dorsal view, 17 mm long [2627], Australia, Queensland (MTQ). H. Dorsal view, 27 mm long [1755], Indonesia, Sumatra (UMIZ 00122).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578333/files/figure.png	https://doi.org/10.5281/zenodo.2578333	Fig. 8. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #1, live animals. A. Dorsal view, 55 mm long [5103], Indonesia, Halmahera (UMIZ 00136). B. Dorsal view, 22 mm long [1732], Indonesia, Sumatra (UMIZ 00121). C. Dorsal view, 22 mm long [5082], Indonesia, Halmahera (UMIZ 00135). D. Dorsal view, 22 mm long [5642], Vietnam (ITBZC IM 00020). E. Dorsal view, 45 mm long [2657], Australia, Queensland (MTQ). F. Dorsal view, 31 mm long [1522], Australia, New South Wales (AM C.468918.005). G. Ventral view, 42 mm long [1686], Australia, Northern Territory (NTM P.57621). H. Ventral view, 47 mm long [2562], Australia, Queensland (MTQ). I. Ventral view, 19 mm long [2961], Indonesia, Lombok (UMIZ 00132). J. Ventral view, 44 mm long [5619], Vietnam (ITBZC IM 00019). K. Ventral view, 33 mm long [3416], Philippines, Bohol (PNM 041260). L. Ventral view, 24 mm long [3371], Philippines, Bohol (PNM 041259).	Fig. 8. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #1, live animals. A. Dorsal view, 55 mm long [5103], Indonesia, Halmahera (UMIZ 00136). B. Dorsal view, 22 mm long [1732], Indonesia, Sumatra (UMIZ 00121). C. Dorsal view, 22 mm long [5082], Indonesia, Halmahera (UMIZ 00135). D. Dorsal view, 22 mm long [5642], Vietnam (ITBZC IM 00020). E. Dorsal view, 45 mm long [2657], Australia, Queensland (MTQ). F. Dorsal view, 31 mm long [1522], Australia, New South Wales (AM C.468918.005). G. Ventral view, 42 mm long [1686], Australia, Northern Territory (NTM P.57621). H. Ventral view, 47 mm long [2562], Australia, Queensland (MTQ). I. Ventral view, 19 mm long [2961], Indonesia, Lombok (UMIZ 00132). J. Ventral view, 44 mm long [5619], Vietnam (ITBZC IM 00019). K. Ventral view, 33 mm long [3416], Philippines, Bohol (PNM 041260). L. Ventral view, 24 mm long [3371], Philippines, Bohol (PNM 041259).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578335/files/figure.png	https://doi.org/10.5281/zenodo.2578335	Fig. 9. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #2, live animals. A. Dorsal view, 38 mm long [1651], Australia, Northern Territory (NTM P.57624). B. Dorsal view, 32 mm long [3237], Philippines, Luzon (PNM 041265). C. Dorsal view, 45 mm long [1638], Australia, Northern Territory (NTM P.57623). D. Dorsal view, 43 mm long [1794], Indonesia, Sumatra (UMIZ 00138). E. Dorsal view, 17 mm long [2960], Indonesia, Lombok (UMIZ 00139). F. Dorsal view, 27 mm long [3610], Philippines, Luzon (PNM 041261). G. Ventral view, same as C. H. Ventral view, same as D. I. Ventral view, 32 mm long [3229], Philippines, Luzon (PNM 041264).	Fig. 9. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #2, live animals. A. Dorsal view, 38 mm long [1651], Australia, Northern Territory (NTM P.57624). B. Dorsal view, 32 mm long [3237], Philippines, Luzon (PNM 041265). C. Dorsal view, 45 mm long [1638], Australia, Northern Territory (NTM P.57623). D. Dorsal view, 43 mm long [1794], Indonesia, Sumatra (UMIZ 00138). E. Dorsal view, 17 mm long [2960], Indonesia, Lombok (UMIZ 00139). F. Dorsal view, 27 mm long [3610], Philippines, Luzon (PNM 041261). G. Ventral view, same as C. H. Ventral view, same as D. I. Ventral view, 32 mm long [3229], Philippines, Luzon (PNM 041264).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578339/files/figure.png	https://doi.org/10.5281/zenodo.2578339	Fig. 10. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #3, preserved specimens, Papua New Guinea, Madang. A. Dorsal view, 13 mm long [5432] (MNHN IM-2013-10478). B. Lateral view, same as A. C. Ventral view, same as A. D. Dorsal view, 14 mm long [5433] (MNHN IM-2013-10479).	Fig. 10. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #3, preserved specimens, Papua New Guinea, Madang. A. Dorsal view, 13 mm long [5432] (MNHN IM-2013-10478). B. Lateral view, same as A. C. Ventral view, same as A. D. Dorsal view, 14 mm long [5433] (MNHN IM-2013-10479).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578341/files/figure.png	https://doi.org/10.5281/zenodo.2578341	Fig. 11. Paromoionchis tumidus (Semper, 1880) comb. nov., external morphology and nervous system. A. Anterior, ventral view, lectotype of Onchidium mertoni Simroth, 1918 (ZMB 121591a). B. Unit #1, posterior, ventral view, Indonesia, Sulawesi [2240] (UMIZ 00125). C. Nervous system, dorsal view, same as B. Abbreviations: a = anus; f = foot; fo = female opening; h = hyponotum; lcg = left cerebral ganglion; lplg = left pleural ganglion; mo = male opening; ol = oral lobe; ot = ocular tentacle; pn = pneumostome; ppg = peripodial groove; rcg = right cerebral ganglion; rplg = right pleural ganglion; vg = visceral ganglion. Scales: A–B = 2 mm; C = 0.5 mm.	Fig. 11. Paromoionchis tumidus (Semper, 1880) comb. nov., external morphology and nervous system. A. Anterior, ventral view, lectotype of Onchidium mertoni Simroth, 1918 (ZMB 121591a). B. Unit #1, posterior, ventral view, Indonesia, Sulawesi [2240] (UMIZ 00125). C. Nervous system, dorsal view, same as B. Abbreviations: a = anus; f = foot; fo = female opening; h = hyponotum; lcg = left cerebral ganglion; lplg = left pleural ganglion; mo = male opening; ol = oral lobe; ot = ocular tentacle; pn = pneumostome; ppg = peripodial groove; rcg = right cerebral ganglion; rplg = right pleural ganglion; vg = visceral ganglion. Scales: A–B = 2 mm; C = 0.5 mm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578343/files/figure.png	https://doi.org/10.5281/zenodo.2578343	Fig. 12. Paromoionchis tumidus (Semper, 1880) comb. nov., digestive system. A. Unit #1, dorsal view, Indonesia, Sulawesi [2240] (UMIZ 00125). B. Ventral view (lateral digestive gland removed), same as A. C. Stomach, dorsal view, same as A. D. Dorsal view (digestive gland removed prior to present study, intestinal loops loose), holotype of Onchidium honkongense Britton, 1984 (NHM 1982290). E. Unit #2, dorsal view, Australia, Northern Territory [1638] (NTM P.57623). F. Unit #3, dorsal view, Papua New Guinea, Madang [5433] (MNHN IM-2013-10479). G. Dorsal view, lectotype of Onchidium mertoni Simroth, 1918 (ZMB 121591a). H. Dorsal view, lectotype of Onchidium tumidum (ZMB 39019a). I. Dorsal view, lectotype of Onchidium samarense Semper, 1880 (ZMB 39025a). Abbreviations: ddg = dorsal lobe of digestive gland; i = intestine; oddg = opening of dorsal lobe of digestive gland; oldg = opening of lateral lobe of digestive gland; opdg = opening of posterior lobe of digestive gland; pdg = posterior lobe of digestive gland; r = rectum; st = stomach; st1 = stomach chamber 1; st2 = stomach chamber 2; st3 = stomach chamber 3; st4 = stomach chamber 4. Scales: A–C, F = 2 mm; D, I = 3 mm; E = 5 mm; G = 1 mm; H = 4 mm.	Fig. 12. Paromoionchis tumidus (Semper, 1880) comb. nov., digestive system. A. Unit #1, dorsal view, Indonesia, Sulawesi [2240] (UMIZ 00125). B. Ventral view (lateral digestive gland removed), same as A. C. Stomach, dorsal view, same as A. D. Dorsal view (digestive gland removed prior to present study, intestinal loops loose), holotype of Onchidium honkongense Britton, 1984 (NHM 1982290). E. Unit #2, dorsal view, Australia, Northern Territory [1638] (NTM P.57623). F. Unit #3, dorsal view, Papua New Guinea, Madang [5433] (MNHN IM-2013-10479). G. Dorsal view, lectotype of Onchidium mertoni Simroth, 1918 (ZMB 121591a). H. Dorsal view, lectotype of Onchidium tumidum (ZMB 39019a). I. Dorsal view, lectotype of Onchidium samarense Semper, 1880 (ZMB 39025a). Abbreviations: ddg = dorsal lobe of digestive gland; i = intestine; oddg = opening of dorsal lobe of digestive gland; oldg = opening of lateral lobe of digestive gland; opdg = opening of posterior lobe of digestive gland; pdg = posterior lobe of digestive gland; r = rectum; st = stomach; st1 = stomach chamber 1; st2 = stomach chamber 2; st3 = stomach chamber 3; st4 = stomach chamber 4. Scales: A–C, F = 2 mm; D, I = 3 mm; E = 5 mm; G = 1 mm; H = 4 mm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578345/files/figure.png	https://doi.org/10.5281/zenodo.2578345	Fig. 13. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #1, radula, Indonesia, Bali [3051] (UMIZ 00133). A. Rachidian and innermost lateral teeth. B. Lateral teeth with rachidian teeth. C. Lateral teeth. D. Outermost lateral teeth. Abbreviations: 1llt = first left lateral tooth; 1rlt = first right lateral tooth; 2llt = second left lateral tooth; 2rlt = second right lateral tooth; bls = basal lateral spine; hlt = hook of lateral tooth; mc = median cusp; rt = rachidian tooth. Scales: A = 20 μm; B = 30 μm; C = 50 μm; D = 10 μm.	Fig. 13. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #1, radula, Indonesia, Bali [3051] (UMIZ 00133). A. Rachidian and innermost lateral teeth. B. Lateral teeth with rachidian teeth. C. Lateral teeth. D. Outermost lateral teeth. Abbreviations: 1llt = first left lateral tooth; 1rlt = first right lateral tooth; 2llt = second left lateral tooth; 2rlt = second right lateral tooth; bls = basal lateral spine; hlt = hook of lateral tooth; mc = median cusp; rt = rachidian tooth. Scales: A = 20 μm; B = 30 μm; C = 50 μm; D = 10 μm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578347/files/figure.png	https://doi.org/10.5281/zenodo.2578347	Fig. 14. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #2, radula, Australia, Northern Territory [1638] (NTM P.57623). A. Rachidian and innermost lateral teeth. B. Lateral teeth with rachidian teeth. C. Lateral teeth. D. Outermost lateral teeth. Scales: A = 30 μm; B–C = 50 μm; D = 20 μm.	Fig. 14. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #2, radula, Australia, Northern Territory [1638] (NTM P.57623). A. Rachidian and innermost lateral teeth. B. Lateral teeth with rachidian teeth. C. Lateral teeth. D. Outermost lateral teeth. Scales: A = 30 μm; B–C = 50 μm; D = 20 μm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578349/files/figure.png	https://doi.org/10.5281/zenodo.2578349	Fig. 15. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #3, radula, Papua New Guinea, Madang. A. Rachidian and innermost lateral teeth, [5432] (MNHN IM-2013-10478). B. Lateral teeth with rachidian teeth, same as A. C. Lateral teeth, same as A. D. Outermost lateral teeth, [5433] (MNHN IM-2013-10479). Scales: A = 10 μm; B, D = 20 μm; C = 25 μm.	Fig. 15. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #3, radula, Papua New Guinea, Madang. A. Rachidian and innermost lateral teeth, [5432] (MNHN IM-2013-10478). B. Lateral teeth with rachidian teeth, same as A. C. Lateral teeth, same as A. D. Outermost lateral teeth, [5433] (MNHN IM-2013-10479). Scales: A = 10 μm; B, D = 20 μm; C = 25 μm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578351/files/figure.png	https://doi.org/10.5281/zenodo.2578351	Fig. 16. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #1, reproductive system. A. Posterior hermaphroditic (female) reproductive system, Australia, Northern Territory [1634] (NTM P.57620). B. Male copulatory organs, Philippines, Bohol [3344] (PNM 041258). C. Male copulatory organs, lectotype of Onchidium tumidum (ZMB 39019a). Abbreviations: ag = accessory penial gland; dd = deferent duct; fgm = female gland mass; hd = hermaphroditic duct; hg = hermaphroditic gland; ms = muscular sac (of accessory penial gland); ov = oviduct; ps = penial sheath; rm = retractor muscle; rs = receptaculum seminis; sp = spermatheca; v = vestibule. Scales = 3 mm.	Fig. 16. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #1, reproductive system. A. Posterior hermaphroditic (female) reproductive system, Australia, Northern Territory [1634] (NTM P.57620). B. Male copulatory organs, Philippines, Bohol [3344] (PNM 041258). C. Male copulatory organs, lectotype of Onchidium tumidum (ZMB 39019a). Abbreviations: ag = accessory penial gland; dd = deferent duct; fgm = female gland mass; hd = hermaphroditic duct; hg = hermaphroditic gland; ms = muscular sac (of accessory penial gland); ov = oviduct; ps = penial sheath; rm = retractor muscle; rs = receptaculum seminis; sp = spermatheca; v = vestibule. Scales = 3 mm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578353/files/figure.png	https://doi.org/10.5281/zenodo.2578353	Fig. 17. Paromoionchis tumidus (Semper, 1880) comb. nov., reproductive system. A. Unit #3, posterior hermaphroditic (female) reproductive system, Papua New Guinea, Madang [5433] (MNHN IM- 2013-10479). B. Unit #2, posterior hermaphroditic (female) reproductive system, Australia, Northern Territory [1638] (NTM P.57623). C. Male copulatory organs, same as A. D. Male copulatory organs, same as B. Abbreviations: ag = accessory penial gland; dd = deferent duct; fgm = female gland mass; hd = hermaphroditic duct; hg = hermaphroditic gland; ms = muscular sac (of accessory penial gland); ov = oviduct; ps = penial sheath; rm = retractor muscle; rs = receptaculum seminis; sp = spermatheca; v = vestibule. Scales: A, C = 1 mm; B, D = 3 mm.	Fig. 17. Paromoionchis tumidus (Semper, 1880) comb. nov., reproductive system. A. Unit #3, posterior hermaphroditic (female) reproductive system, Papua New Guinea, Madang [5433] (MNHN IM- 2013-10479). B. Unit #2, posterior hermaphroditic (female) reproductive system, Australia, Northern Territory [1638] (NTM P.57623). C. Male copulatory organs, same as A. D. Male copulatory organs, same as B. Abbreviations: ag = accessory penial gland; dd = deferent duct; fgm = female gland mass; hd = hermaphroditic duct; hg = hermaphroditic gland; ms = muscular sac (of accessory penial gland); ov = oviduct; ps = penial sheath; rm = retractor muscle; rs = receptaculum seminis; sp = spermatheca; v = vestibule. Scales: A, C = 1 mm; B, D = 3 mm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578355/files/figure.png	https://doi.org/10.5281/zenodo.2578355	Fig. 18. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #1, spine of accessory penial gland. A. Indonesia, Lombok [2950] (UMIZ 00131). B. Philippines, Bohol [3344] (PNM 041258). C. Indonesia, Ambon [2832] (UMIZ 00129). D. Australia, Northern Territory [1634] (NTM P.57620). E. Australia, Queensland [2637] (MTQ). F. Distal tip of spine, Indonesia, Halmahera [5102] (UMIZ 00136). G. Distal tip of spine, same as D. Scales: A–D = 200 μm; E = 400 μm; F–G = 20 μm.	Fig. 18. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #1, spine of accessory penial gland. A. Indonesia, Lombok [2950] (UMIZ 00131). B. Philippines, Bohol [3344] (PNM 041258). C. Indonesia, Ambon [2832] (UMIZ 00129). D. Australia, Northern Territory [1634] (NTM P.57620). E. Australia, Queensland [2637] (MTQ). F. Distal tip of spine, Indonesia, Halmahera [5102] (UMIZ 00136). G. Distal tip of spine, same as D. Scales: A–D = 200 μm; E = 400 μm; F–G = 20 μm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578357/files/figure.png	https://doi.org/10.5281/zenodo.2578357	Fig. 19. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #2, spine of accessory penial gland. A. Australia, Northern Territory [1638] (NTM P.57623). B. Philippines, Luzon [3237] (PNM 041265). C. Philippines, Luzon [3172] (PNM 041262). D. Distal tip of spine, same as A. E. Distal tip of spine, same as B. F. Distal tip of spine, same as C. Scales: A = 150 μm; B–C = 200 μm; D–F = 20 μm.	Fig. 19. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #2, spine of accessory penial gland. A. Australia, Northern Territory [1638] (NTM P.57623). B. Philippines, Luzon [3237] (PNM 041265). C. Philippines, Luzon [3172] (PNM 041262). D. Distal tip of spine, same as A. E. Distal tip of spine, same as B. F. Distal tip of spine, same as C. Scales: A = 150 μm; B–C = 200 μm; D–F = 20 μm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578359/files/figure.png	https://doi.org/10.5281/zenodo.2578359	Fig. 20. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #3, spine of accessory penial gland, Papua New Guinea, Madang. A. [5432] (MNHN IM-2013-10478). B. [5433] (MNHN IM-2013-10479). C. Distal tip of spine, same as A. Scales: A = 100 μm; B = 20 μm; C = 10 μm.	Fig. 20. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #3, spine of accessory penial gland, Papua New Guinea, Madang. A. [5432] (MNHN IM-2013-10478). B. [5433] (MNHN IM-2013-10479). C. Distal tip of spine, same as A. Scales: A = 100 μm; B = 20 μm; C = 10 μm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578361/files/figure.png	https://doi.org/10.5281/zenodo.2578361	Fig. 21. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #1, penial hooks. A. Indonesia, Sulawesi [2240] (UMIZ 00125). B. Indonesia, Ambon [2832] (UMIZ 00129). C. Lectotype of Onchidium mertoni Simroth, 1918 (ZMB 121591a). D. Lectotype of Onchidium tumidum (ZMB 39019a). Scales: A = 60 μm; B = 10 μm; C = 4 μm; D = 25 μm.	Fig. 21. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #1, penial hooks. A. Indonesia, Sulawesi [2240] (UMIZ 00125). B. Indonesia, Ambon [2832] (UMIZ 00129). C. Lectotype of Onchidium mertoni Simroth, 1918 (ZMB 121591a). D. Lectotype of Onchidium tumidum (ZMB 39019a). Scales: A = 60 μm; B = 10 μm; C = 4 μm; D = 25 μm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578363/files/figure.png	https://doi.org/10.5281/zenodo.2578363	Fig. 22. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #2, penial hooks. A. Indonesia, Sumatra [1794] (UMIZ 00138). B. Philippines, Luzon [3237] (PNM 041265). C. Same as B. Scales: A = 20 μm; B = 4 μm; C = 6 μm.	Fig. 22. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #2, penial hooks. A. Indonesia, Sumatra [1794] (UMIZ 00138). B. Philippines, Luzon [3237] (PNM 041265). C. Same as B. Scales: A = 20 μm; B = 4 μm; C = 6 μm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578365/files/figure.png	https://doi.org/10.5281/zenodo.2578365	Fig. 23. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #3, penial hooks, Papua New Guinea, Madang. A. [5433] (MNHN IM-2013-10479). B. Same as A. C. [5432] (MNHN IM-2013-10478). D. Same as C. Scales: A, D = 3 μm; B–C = 6 μm.	Fig. 23. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #3, penial hooks, Papua New Guinea, Madang. A. [5433] (MNHN IM-2013-10479). B. Same as A. C. [5432] (MNHN IM-2013-10478). D. Same as C. Scales: A, D = 3 μm; B–C = 6 μm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578367/files/figure.png	https://doi.org/10.5281/zenodo.2578367	Fig. 24. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #1, habitats. A. Australia, Queensland, mangrove by creek with Rhizophora and Avicennia (station 119). B. Indonesia, Ambon, mangrove with large Rhizophora and some open spaces between trees (station 131). C. Indonesia, Lombok, mudflat at the margin of mangrove (station 145). D. Philippines, Bohol, narrow forest on the edge of fish ponds, with tall Rhizophora and Avicennia trees, and different types of mud (station 194). E. Indonesia, Halmahera, logged area by an old Rhizophora forest (station 212). F. Vietnam, Nha Trang, mostly Avicennia trees, mud not deep (station 239).	Fig. 24. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #1, habitats. A. Australia, Queensland, mangrove by creek with Rhizophora and Avicennia (station 119). B. Indonesia, Ambon, mangrove with large Rhizophora and some open spaces between trees (station 131). C. Indonesia, Lombok, mudflat at the margin of mangrove (station 145). D. Philippines, Bohol, narrow forest on the edge of fish ponds, with tall Rhizophora and Avicennia trees, and different types of mud (station 194). E. Indonesia, Halmahera, logged area by an old Rhizophora forest (station 212). F. Vietnam, Nha Trang, mostly Avicennia trees, mud not deep (station 239).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578369/files/figure.png	https://doi.org/10.5281/zenodo.2578369	Fig. 25. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #2, habitats. A. Australia, Northern Territory, large mangrove of Sonneratia, Rhizophora and Ceriops (st. 66). B. Indonesia, Sumatra, small patch of mangrove, very impacted (st. 81). C. Indonesia, Lombok, open Avicennia mangrove (station 146). D. Philippines, Luzon, narrow and very sandy Avicennia mangrove, with no mud (st. 179).	Fig. 25. Paromoionchis tumidus (Semper, 1880) comb. nov. unit #2, habitats. A. Australia, Northern Territory, large mangrove of Sonneratia, Rhizophora and Ceriops (st. 66). B. Indonesia, Sumatra, small patch of mangrove, very impacted (st. 81). C. Indonesia, Lombok, open Avicennia mangrove (station 146). D. Philippines, Luzon, narrow and very sandy Avicennia mangrove, with no mud (st. 179).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578329/files/figure.png	https://doi.org/10.5281/zenodo.2578329	Fig. 6. Geographic distribution of the five species of Paromoionchis gen. nov. Distinct colors are used for each mitochondrial unit found in P. tumidus (Semper, 1880) comb. nov., P. boholensis gen. et sp. nov. and P. goslineri gen. et sp. nov., and the colors used for each unit are the same as those used in Figs 1–5. Colored areas correspond to hypothetical ranges proposed based on known records (all of which are new here except for the type localities of P. tumidus and P. daemelii (Semper, 1880) comb. nov.).	Fig. 6. Geographic distribution of the five species of Paromoionchis gen. nov. Distinct colors are used for each mitochondrial unit found in P. tumidus (Semper, 1880) comb. nov., P. boholensis gen. et sp. nov. and P. goslineri gen. et sp. nov., and the colors used for each unit are the same as those used in Figs 1–5. Colored areas correspond to hypothetical ranges proposed based on known records (all of which are new here except for the type localities of P. tumidus and P. daemelii (Semper, 1880) comb. nov.).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F510FFAC83F4FA0C85BAF202.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578371/files/figure.png	https://doi.org/10.5281/zenodo.2578371	Fig. 26. Paromoionchis daemelii (Semper, 1880) comb. nov., live animals, Australia, New South Wales. A. Dorsal view, 51 mm long [1514] (AM C.468912.002). B. Dorsal view, 56 mm long [1519] (AM C.468917.001). C. Dorsal view, 31 mm long [1515] (AM C.468914.001). D. Dorsal view, 62 mm long [1518] (AM C.468913.001). E. Ventral view, 42 mm long [1512] (AM C.468912.003). F. Ventral view, 36 mm long [1511] (AM C.468910.001). G. Ventral view, same as C.	Fig. 26. Paromoionchis daemelii (Semper, 1880) comb. nov., live animals, Australia, New South Wales. A. Dorsal view, 51 mm long [1514] (AM C.468912.002). B. Dorsal view, 56 mm long [1519] (AM C.468917.001). C. Dorsal view, 31 mm long [1515] (AM C.468914.001). D. Dorsal view, 62 mm long [1518] (AM C.468913.001). E. Ventral view, 42 mm long [1512] (AM C.468912.003). F. Ventral view, 36 mm long [1511] (AM C.468910.001). G. Ventral view, same as C.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F510FFAC83F4FA0C85BAF202.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578373/files/figure.png	https://doi.org/10.5281/zenodo.2578373	Fig. 27. Paromoionchis daemelii (Semper, 1880) comb. nov. A. Digestive system, Australia, New South Wales [1519] (AM C.468917.001). B. Digestive system, lectotype of Onchidium daemelii (ZMB 31640a). C. Posterior hermaphroditic (female) reproductive system, same as A. D. Male copulatory organs, same as A. Abbreviations: ag = accessory penial gland; dd = deferent duct; ddg = dorsal lobe of digestive gland; fgm = female gland mass; hd = hermaphroditic duct; hg = hermaphroditic gland; i = intestine; ms = muscular sac (of accessory penial gland); ov = oviduct; pdg = posterior lobe of digestive gland; ps = penial sheath; r = rectum; rm = retractor muscle; rs = receptaculum seminis; sp = spermatheca; st = stomach; v = vestibule. Scales: A = 4 mm; B–D = 3 mm.	Fig. 27. Paromoionchis daemelii (Semper, 1880) comb. nov. A. Digestive system, Australia, New South Wales [1519] (AM C.468917.001). B. Digestive system, lectotype of Onchidium daemelii (ZMB 31640a). C. Posterior hermaphroditic (female) reproductive system, same as A. D. Male copulatory organs, same as A. Abbreviations: ag = accessory penial gland; dd = deferent duct; ddg = dorsal lobe of digestive gland; fgm = female gland mass; hd = hermaphroditic duct; hg = hermaphroditic gland; i = intestine; ms = muscular sac (of accessory penial gland); ov = oviduct; pdg = posterior lobe of digestive gland; ps = penial sheath; r = rectum; rm = retractor muscle; rs = receptaculum seminis; sp = spermatheca; st = stomach; v = vestibule. Scales: A = 4 mm; B–D = 3 mm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F510FFAC83F4FA0C85BAF202.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578375/files/figure.png	https://doi.org/10.5281/zenodo.2578375	Fig. 28. Paromoionchis daemelii (Semper, 1880) comb. nov., radula, Australia, New South Wales [1519] (AM C.468917.001). A. Rachidian and innermost lateral teeth. B. Lateral teeth with rachidian teeth. C. Lateral teeth. D. Outermost lateral teeth. Scales: A, D = 20 μm; B–C = 30 μm.	Fig. 28. Paromoionchis daemelii (Semper, 1880) comb. nov., radula, Australia, New South Wales [1519] (AM C.468917.001). A. Rachidian and innermost lateral teeth. B. Lateral teeth with rachidian teeth. C. Lateral teeth. D. Outermost lateral teeth. Scales: A, D = 20 μm; B–C = 30 μm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F510FFAC83F4FA0C85BAF202.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578377/files/figure.png	https://doi.org/10.5281/zenodo.2578377	Fig. 29. Paromoionchis daemelii (Semper, 1880) comb. nov., spine of accessory penial gland. A. Australia, New South Wales [1519] (AM C.468917.001). B. Australia, New South Wales [1521] (AM C.468919.001). C. Distal tip of spine, same as A. D. Distal tip of spine, same as B. Scales: A–B = 500 μm; C–D = 50 μm.	Fig. 29. Paromoionchis daemelii (Semper, 1880) comb. nov., spine of accessory penial gland. A. Australia, New South Wales [1519] (AM C.468917.001). B. Australia, New South Wales [1521] (AM C.468919.001). C. Distal tip of spine, same as A. D. Distal tip of spine, same as B. Scales: A–B = 500 μm; C–D = 50 μm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F510FFAC83F4FA0C85BAF202.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578379/files/figure.png	https://doi.org/10.5281/zenodo.2578379	Fig. 30. Paromoionchis daemelii (Semper, 1880) comb. nov., habitats. A. Australia, New South Wales, muddy sand next to small patch of mangrove and rocks on sandy beach (station 39). B. Australia, Queensland, Rhizophora and Avicennia mangrove (station 119).	Fig. 30. Paromoionchis daemelii (Semper, 1880) comb. nov., habitats. A. Australia, New South Wales, muddy sand next to small patch of mangrove and rocks on sandy beach (station 39). B. Australia, Queensland, Rhizophora and Avicennia mangrove (station 119).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F510FFAC83F4FA0C85BAF202.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578329/files/figure.png	https://doi.org/10.5281/zenodo.2578329	Fig. 6. Geographic distribution of the five species of Paromoionchis gen. nov. Distinct colors are used for each mitochondrial unit found in P. tumidus (Semper, 1880) comb. nov., P. boholensis gen. et sp. nov. and P. goslineri gen. et sp. nov., and the colors used for each unit are the same as those used in Figs 1–5. Colored areas correspond to hypothetical ranges proposed based on known records (all of which are new here except for the type localities of P. tumidus and P. daemelii (Semper, 1880) comb. nov.).	Fig. 6. Geographic distribution of the five species of Paromoionchis gen. nov. Distinct colors are used for each mitochondrial unit found in P. tumidus (Semper, 1880) comb. nov., P. boholensis gen. et sp. nov. and P. goslineri gen. et sp. nov., and the colors used for each unit are the same as those used in Figs 1–5. Colored areas correspond to hypothetical ranges proposed based on known records (all of which are new here except for the type localities of P. tumidus and P. daemelii (Semper, 1880) comb. nov.).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F50AFFA783B8FDEC867BF3B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578381/files/figure.png	https://doi.org/10.5281/zenodo.2578381	Fig. 31. Paromoionchis boholensis gen. et sp. nov., unit #1, live animals. A. Holotype, dorsal view, 28 mm long [3288], Philippines, Bohol (PNM 041266). B. Dorsal view, 26 mm long [3609], Philippines, Luzon (PNM 041267). C. Dorsal view, 38 mm long [3417], Philippines, Bohol (PNM 041270). D. Dorsal view, 17 mm long [3283], Philippines, Bohol (PNM 041268). E. Dorsal view, 30 mm long [3372], Philippines, Bohol (PNM 041269). F. Dorsal view, 17 mm long [3619], Philippines, Bohol (PNM 041268). G. Ventral view, same as E. H. Ventral view, 38 mm long [3422], Philippines, Bohol (PNM 041270). I. Ventral view, 32 mm long [3413], Philippines, Bohol (PNM 041270). J. Ventral view, same as C.	Fig. 31. Paromoionchis boholensis gen. et sp. nov., unit #1, live animals. A. Holotype, dorsal view, 28 mm long [3288], Philippines, Bohol (PNM 041266). B. Dorsal view, 26 mm long [3609], Philippines, Luzon (PNM 041267). C. Dorsal view, 38 mm long [3417], Philippines, Bohol (PNM 041270). D. Dorsal view, 17 mm long [3283], Philippines, Bohol (PNM 041268). E. Dorsal view, 30 mm long [3372], Philippines, Bohol (PNM 041269). F. Dorsal view, 17 mm long [3619], Philippines, Bohol (PNM 041268). G. Ventral view, same as E. H. Ventral view, 38 mm long [3422], Philippines, Bohol (PNM 041270). I. Ventral view, 32 mm long [3413], Philippines, Bohol (PNM 041270). J. Ventral view, same as C.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F50AFFA783B8FDEC867BF3B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578383/files/figure.png	https://doi.org/10.5281/zenodo.2578383	Fig. 32. Paromoionchis boholensis gen. et sp. nov., unit #2, live animals. A. Dorsal view, 37 mm long [2129], Indonesia, Sulawesi (UMIZ 00141). B. Dorsal view, 45 mm long [2851], Indonesia, Ambon (UMIZ 00146). C. Dorsal view, 35 mm long [3117], Indonesia, Bali (UMIZ 00140). D. Dorsal view, 18 mm long [2911], Indonesia, Kei (UMIZ 00149). E. Dorsal view, 40 mm long [2937], Indonesia, Kei (UMIZ 00150). F. Dorsal view, 15 mm long [2903], Indonesia, Kei (UMIZ 00149). G. Ventral view, 45 mm long [2884], Indonesia, Seram (UMIZ 00147). H. Ventral view, same as B. I. Ventral view, 45 mm long [2850], Indonesia, Ambon (UMIZ 00146). J. Frontal view, 17 mm long [2901], Indonesia, Kei (UMIZ 00148).	Fig. 32. Paromoionchis boholensis gen. et sp. nov., unit #2, live animals. A. Dorsal view, 37 mm long [2129], Indonesia, Sulawesi (UMIZ 00141). B. Dorsal view, 45 mm long [2851], Indonesia, Ambon (UMIZ 00146). C. Dorsal view, 35 mm long [3117], Indonesia, Bali (UMIZ 00140). D. Dorsal view, 18 mm long [2911], Indonesia, Kei (UMIZ 00149). E. Dorsal view, 40 mm long [2937], Indonesia, Kei (UMIZ 00150). F. Dorsal view, 15 mm long [2903], Indonesia, Kei (UMIZ 00149). G. Ventral view, 45 mm long [2884], Indonesia, Seram (UMIZ 00147). H. Ventral view, same as B. I. Ventral view, 45 mm long [2850], Indonesia, Ambon (UMIZ 00146). J. Frontal view, 17 mm long [2901], Indonesia, Kei (UMIZ 00148).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F50AFFA783B8FDEC867BF3B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578385/files/figure.png	https://doi.org/10.5281/zenodo.2578385	Fig. 33. Paromoionchis boholensis gen. et sp. nov., unit #1, holotype, Philippines, Bohol [3288] (PNM 041266). A. Digestive system. B. Posterior hermaphroditic (female) reproductive system. C. Male copulatory organs. Abbreviations: ag = accessory penial gland; dd = deferent duct; ddg = dorsal lobe of digestive gland; fgm = female gland mass; hd = hermaphroditic duct; hg = hermaphroditic gland; i = intestine; ms = muscular sac (of accessory penial gland); ov = oviduct; pdg = posterior lobe of digestive gland; ps = penial sheath; r = rectum; rm = retractor muscle; rs = receptaculum seminis; sp = spermatheca; st = stomach; v = vestibule. Scales = 2 mm.	Fig. 33. Paromoionchis boholensis gen. et sp. nov., unit #1, holotype, Philippines, Bohol [3288] (PNM 041266). A. Digestive system. B. Posterior hermaphroditic (female) reproductive system. C. Male copulatory organs. Abbreviations: ag = accessory penial gland; dd = deferent duct; ddg = dorsal lobe of digestive gland; fgm = female gland mass; hd = hermaphroditic duct; hg = hermaphroditic gland; i = intestine; ms = muscular sac (of accessory penial gland); ov = oviduct; pdg = posterior lobe of digestive gland; ps = penial sheath; r = rectum; rm = retractor muscle; rs = receptaculum seminis; sp = spermatheca; st = stomach; v = vestibule. Scales = 2 mm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F50AFFA783B8FDEC867BF3B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578387/files/figure.png	https://doi.org/10.5281/zenodo.2578387	Fig. 34. Paromoionchis boholensis gen. et sp. nov., unit #2, Indonesia, Bali [3117] (UMIZ 00140). A. Digestive system. B. Posterior hermaphroditic (female) reproductive system. C. Male copulatory organs. Abbreviations: ag = accessory penial gland; dd = deferent duct; ddg = dorsal lobe of digestive gland; fgm = female gland mass; hd = hermaphroditic duct; hg = hermaphroditic gland; i = intestine; ms = muscular sac (of accessory penial gland); ov = oviduct; pdg = posterior lobe of digestive gland; ps = penial sheath; r = rectum; rm = retractor muscle; rs = receptaculum seminis; sp = spermatheca; st = stomach; v = vestibule. Scales: A–B = 2 mm; C = 3 mm.	Fig. 34. Paromoionchis boholensis gen. et sp. nov., unit #2, Indonesia, Bali [3117] (UMIZ 00140). A. Digestive system. B. Posterior hermaphroditic (female) reproductive system. C. Male copulatory organs. Abbreviations: ag = accessory penial gland; dd = deferent duct; ddg = dorsal lobe of digestive gland; fgm = female gland mass; hd = hermaphroditic duct; hg = hermaphroditic gland; i = intestine; ms = muscular sac (of accessory penial gland); ov = oviduct; pdg = posterior lobe of digestive gland; ps = penial sheath; r = rectum; rm = retractor muscle; rs = receptaculum seminis; sp = spermatheca; st = stomach; v = vestibule. Scales: A–B = 2 mm; C = 3 mm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F50AFFA783B8FDEC867BF3B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578389/files/figure.png	https://doi.org/10.5281/zenodo.2578389	Fig. 35. Paromoionchis boholensis gen. et sp. nov., unit #1, radula, Philippines, Bohol [3372] (PNM 041269). A. Rachidian and innermost lateral teeth. B. Lateral teeth with rachidian teeth. C. Lateral teeth. D. Outermost lateral teeth. Scales: A = 20 μm; B, D = 30 μm; C = 50 μm.	Fig. 35. Paromoionchis boholensis gen. et sp. nov., unit #1, radula, Philippines, Bohol [3372] (PNM 041269). A. Rachidian and innermost lateral teeth. B. Lateral teeth with rachidian teeth. C. Lateral teeth. D. Outermost lateral teeth. Scales: A = 20 μm; B, D = 30 μm; C = 50 μm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F50AFFA783B8FDEC867BF3B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578391/files/figure.png	https://doi.org/10.5281/zenodo.2578391	Fig. 36. Paromoionchis boholensis gen. et sp. nov., unit #2, radula, Indonesia, Ambon [2851] (UMIZ 00146). A. Rachidian and innermost lateral teeth. B. Lateral teeth with rachidian teeth. C. Lateral teeth. D. Outermost lateral teeth. Scales: A, D = 30 μm; B–C = 50 μm.	Fig. 36. Paromoionchis boholensis gen. et sp. nov., unit #2, radula, Indonesia, Ambon [2851] (UMIZ 00146). A. Rachidian and innermost lateral teeth. B. Lateral teeth with rachidian teeth. C. Lateral teeth. D. Outermost lateral teeth. Scales: A, D = 30 μm; B–C = 50 μm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F50AFFA783B8FDEC867BF3B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578393/files/figure.png	https://doi.org/10.5281/zenodo.2578393	Fig. 37. Paromoionchis boholensis gen. et sp. nov., unit #1, spine of accessory penial gland. A. Holotype, Philippines, Bohol [3288] (PNM 041266). B. Philippines, Bohol [3372] (PNM 041269). C. Distal tip of spine, same as B. Scales: A–B = 100 μm; C = 10 μm.	Fig. 37. Paromoionchis boholensis gen. et sp. nov., unit #1, spine of accessory penial gland. A. Holotype, Philippines, Bohol [3288] (PNM 041266). B. Philippines, Bohol [3372] (PNM 041269). C. Distal tip of spine, same as B. Scales: A–B = 100 μm; C = 10 μm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F50AFFA783B8FDEC867BF3B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578395/files/figure.png	https://doi.org/10.5281/zenodo.2578395	Fig. 38. Paromoionchis boholensis gen. et sp. nov., unit #2, spine of accessory penial gland. A. Indonesia, Bali [3117] (UMIZ 00140). B. Indonesia, Kei [2911] (UMIZ 00149). C. Tip of spine, Indonesia, Ambon [2851] (UMIZ 00146). D. Distal tip of spine, same as B. E. Tip of spine, same as A. F. Tip of spine, Indonesia, Halmahera [5140] (UMIZ 00152). Scales: A–B = 200 μm; C–F = 30 μm.	Fig. 38. Paromoionchis boholensis gen. et sp. nov., unit #2, spine of accessory penial gland. A. Indonesia, Bali [3117] (UMIZ 00140). B. Indonesia, Kei [2911] (UMIZ 00149). C. Tip of spine, Indonesia, Ambon [2851] (UMIZ 00146). D. Distal tip of spine, same as B. E. Tip of spine, same as A. F. Tip of spine, Indonesia, Halmahera [5140] (UMIZ 00152). Scales: A–B = 200 μm; C–F = 30 μm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F50AFFA783B8FDEC867BF3B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578397/files/figure.png	https://doi.org/10.5281/zenodo.2578397	Fig. 39. Paromoionchis boholensis gen. et sp. nov., unit #1, habitats. A. Philippines, Bohol, Maribojoc, large porous rocks behind mangrove (station 193, type locality). B. Philippines, Bohol, Mabini, open Avicennia and Sonneratia forest with large gravel of coral pieces, sand and algae on surface (station 196). C. Philippines, Bohol, Maribojoc, uplifted, dead coral flat covered with sand and algae adjacent to a few mangrove trees (station 202).	Fig. 39. Paromoionchis boholensis gen. et sp. nov., unit #1, habitats. A. Philippines, Bohol, Maribojoc, large porous rocks behind mangrove (station 193, type locality). B. Philippines, Bohol, Mabini, open Avicennia and Sonneratia forest with large gravel of coral pieces, sand and algae on surface (station 196). C. Philippines, Bohol, Maribojoc, uplifted, dead coral flat covered with sand and algae adjacent to a few mangrove trees (station 202).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F50AFFA783B8FDEC867BF3B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578399/files/figure.png	https://doi.org/10.5281/zenodo.2578399	Fig. 40. Paromoionchis boholensis gen. et sp. nov., unit #2, habitats. A. Indonesia, Bali, muddy mangrove with Rhizophora and Avicennia trees (station 156). B. Indonesia, Sulawesi, sand, small rocks and pieces of wood outside a narrow coastal mangrove (station 88). C. Indonesia, Ambon, narrow band of old Avicennia trees on sandy mud, with old logs on ground (station 132). D. Indonesia, Kei, back of mangrove with rocks, mud, logs and leaf litter (station 140). E. Indonesia, Halmahera, mostly Rhizophora, with some sandy areas and some open muddy areas (station 219).	Fig. 40. Paromoionchis boholensis gen. et sp. nov., unit #2, habitats. A. Indonesia, Bali, muddy mangrove with Rhizophora and Avicennia trees (station 156). B. Indonesia, Sulawesi, sand, small rocks and pieces of wood outside a narrow coastal mangrove (station 88). C. Indonesia, Ambon, narrow band of old Avicennia trees on sandy mud, with old logs on ground (station 132). D. Indonesia, Kei, back of mangrove with rocks, mud, logs and leaf litter (station 140). E. Indonesia, Halmahera, mostly Rhizophora, with some sandy areas and some open muddy areas (station 219).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F50AFFA783B8FDEC867BF3B0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578329/files/figure.png	https://doi.org/10.5281/zenodo.2578329	Fig. 6. Geographic distribution of the five species of Paromoionchis gen. nov. Distinct colors are used for each mitochondrial unit found in P. tumidus (Semper, 1880) comb. nov., P. boholensis gen. et sp. nov. and P. goslineri gen. et sp. nov., and the colors used for each unit are the same as those used in Figs 1–5. Colored areas correspond to hypothetical ranges proposed based on known records (all of which are new here except for the type localities of P. tumidus and P. daemelii (Semper, 1880) comb. nov.).	Fig. 6. Geographic distribution of the five species of Paromoionchis gen. nov. Distinct colors are used for each mitochondrial unit found in P. tumidus (Semper, 1880) comb. nov., P. boholensis gen. et sp. nov. and P. goslineri gen. et sp. nov., and the colors used for each unit are the same as those used in Figs 1–5. Colored areas correspond to hypothetical ranges proposed based on known records (all of which are new here except for the type localities of P. tumidus and P. daemelii (Semper, 1880) comb. nov.).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F501FFA2838EFB93873FF213.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578401/files/figure.png	https://doi.org/10.5281/zenodo.2578401	Fig. 41. Paromoionchis penangensis gen. et sp. nov., live animals. A. Dorsal view, 29 mm long [5990], Malaysia, Matang (USMMC 00060). B. Dorsal view, 26 mm long [6039], Malaysia, Penang (USMMC 00063). C. Holotype, dorsal view, 25 mm long [6037], Malaysia, Penang (USMMC 00059). D. Dorsal view, 22 mm long [6031], Malaysia, Penang (USMMC 00063). E. Dorsal view, 37 mm long [1167], India, Maharashtra (BNHS 46). F. Dorsal view, 30 mm long [1129], India, Andaman Islands (BNHS 4). G. Ventral view, same as D. H. Ventral view, same as A. I. Ventral view, 27 mm long [1182], India, Maharashtra (BNHS 42).	Fig. 41. Paromoionchis penangensis gen. et sp. nov., live animals. A. Dorsal view, 29 mm long [5990], Malaysia, Matang (USMMC 00060). B. Dorsal view, 26 mm long [6039], Malaysia, Penang (USMMC 00063). C. Holotype, dorsal view, 25 mm long [6037], Malaysia, Penang (USMMC 00059). D. Dorsal view, 22 mm long [6031], Malaysia, Penang (USMMC 00063). E. Dorsal view, 37 mm long [1167], India, Maharashtra (BNHS 46). F. Dorsal view, 30 mm long [1129], India, Andaman Islands (BNHS 4). G. Ventral view, same as D. H. Ventral view, same as A. I. Ventral view, 27 mm long [1182], India, Maharashtra (BNHS 42).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F501FFA2838EFB93873FF213.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578403/files/figure.png	https://doi.org/10.5281/zenodo.2578403	Fig. 42. Paromoionchis penangensis gen. et sp. nov., holotype, Malaysia, Penang [6037] (USMMC 00059). A. Digestive system. B. Posterior hermaphroditic (female) reproductive system. C. Male copulatory organs. Abbreviations: dd = deferent duct; ddg = dorsal lobe of digestive gland; fgm = female gland mass; hd = hermaphroditic duct; hg = hermaphroditic gland; i = intestine; ov = oviduct; pdg = posterior lobe of digestive gland; ps = penial sheath (penis inside indicated by dotted line); r = rectum; rm = retractor muscle; rs = receptaculum seminis; sp = spermatheca; st = stomach. Scales: A = 2 mm; B–C = 1 mm.	Fig. 42. Paromoionchis penangensis gen. et sp. nov., holotype, Malaysia, Penang [6037] (USMMC 00059). A. Digestive system. B. Posterior hermaphroditic (female) reproductive system. C. Male copulatory organs. Abbreviations: dd = deferent duct; ddg = dorsal lobe of digestive gland; fgm = female gland mass; hd = hermaphroditic duct; hg = hermaphroditic gland; i = intestine; ov = oviduct; pdg = posterior lobe of digestive gland; ps = penial sheath (penis inside indicated by dotted line); r = rectum; rm = retractor muscle; rs = receptaculum seminis; sp = spermatheca; st = stomach. Scales: A = 2 mm; B–C = 1 mm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F501FFA2838EFB93873FF213.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578405/files/figure.png	https://doi.org/10.5281/zenodo.2578405	Fig. 43. Paromoionchis penangensis gen. et sp. nov., radula. A–D. Holotype, Malaysia, Penang [6037] (USMMC 00059). A. Entire radula. B. Rachidian and innermost lateral teeth. C. Lateral teeth with rachidian teeth. D. Lateral teeth. E–F. Western Peninsular Malaysia [6020] (USMMC 00062). E. Lateral teeth with rachidian teeth. F. Outermost lateral teeth. Scales: A = 0.5 mm; B, F = 20 μm; C–E = 30 μm.	Fig. 43. Paromoionchis penangensis gen. et sp. nov., radula. A–D. Holotype, Malaysia, Penang [6037] (USMMC 00059). A. Entire radula. B. Rachidian and innermost lateral teeth. C. Lateral teeth with rachidian teeth. D. Lateral teeth. E–F. Western Peninsular Malaysia [6020] (USMMC 00062). E. Lateral teeth with rachidian teeth. F. Outermost lateral teeth. Scales: A = 0.5 mm; B, F = 20 μm; C–E = 30 μm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F501FFA2838EFB93873FF213.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578407/files/figure.png	https://doi.org/10.5281/zenodo.2578407	Fig. 44. Paromoionchis penangensis gen. et sp. nov., habitats. A. Malaysia, Penang, open Avicennia mangrove with different types of mud (station 261, type locality). B. Western Peninsular Malaysia, Rhizophora and Sonneratia mangrove with soft mud, next to a landfill (station 17). C. India, Maharashtra, Avicennia mangrove with soft mud and a wall of large rocks (station 45). D. India, Maharashtra, Avicennia mangrove with a few small Rhizophora, soft mud and pools (station 47).	Fig. 44. Paromoionchis penangensis gen. et sp. nov., habitats. A. Malaysia, Penang, open Avicennia mangrove with different types of mud (station 261, type locality). B. Western Peninsular Malaysia, Rhizophora and Sonneratia mangrove with soft mud, next to a landfill (station 17). C. India, Maharashtra, Avicennia mangrove with soft mud and a wall of large rocks (station 45). D. India, Maharashtra, Avicennia mangrove with a few small Rhizophora, soft mud and pools (station 47).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F501FFA2838EFB93873FF213.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578329/files/figure.png	https://doi.org/10.5281/zenodo.2578329	Fig. 6. Geographic distribution of the five species of Paromoionchis gen. nov. Distinct colors are used for each mitochondrial unit found in P. tumidus (Semper, 1880) comb. nov., P. boholensis gen. et sp. nov. and P. goslineri gen. et sp. nov., and the colors used for each unit are the same as those used in Figs 1–5. Colored areas correspond to hypothetical ranges proposed based on known records (all of which are new here except for the type localities of P. tumidus and P. daemelii (Semper, 1880) comb. nov.).	Fig. 6. Geographic distribution of the five species of Paromoionchis gen. nov. Distinct colors are used for each mitochondrial unit found in P. tumidus (Semper, 1880) comb. nov., P. boholensis gen. et sp. nov. and P. goslineri gen. et sp. nov., and the colors used for each unit are the same as those used in Figs 1–5. Colored areas correspond to hypothetical ranges proposed based on known records (all of which are new here except for the type localities of P. tumidus and P. daemelii (Semper, 1880) comb. nov.).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F504FFDF83A4FDF28593F3CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578409/files/figure.png	https://doi.org/10.5281/zenodo.2578409	Fig. 45. Paromoionchis goslineri gen. et sp. nov. unit #1, live animals, Philippines, Luzon. A. Dorsal view, 30 mm long [3232] (PNM 041273). B. Dorsal view, 22 mm long [3221] (PNM 041272). C. Holotype, dorsal view, 25 mm long [3233] (PNM 041271). D. Ventral view, same as B. E. Ventral view, same as A.	Fig. 45. Paromoionchis goslineri gen. et sp. nov. unit #1, live animals, Philippines, Luzon. A. Dorsal view, 30 mm long [3232] (PNM 041273). B. Dorsal view, 22 mm long [3221] (PNM 041272). C. Holotype, dorsal view, 25 mm long [3233] (PNM 041271). D. Ventral view, same as B. E. Ventral view, same as A.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F504FFDF83A4FDF28593F3CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578411/files/figure.png	https://doi.org/10.5281/zenodo.2578411	Fig. 46. Paromoionchis goslineri gen. et sp. nov. unit #2, live animals. A. Dorsal view, 33 mm long [3120], Indonesia, Bali (UMIZ 00156). B. Dorsal view, 32 mm long [5073], Indonesia, Halmahera (UMIZ 00158). C. Dorsal and ventral views, 18 mm long [2241], Indonesia, Sulawesi (UMIZ 00153). D. Dorsal view, 24 mm long [3079], Indonesia, Bali (UMIZ 00155). E. Ventral view, 26 mm long [5072], Indonesia, Halmahera (UMIZ 00158). F. Ventral view, 16 mm long [5891], Indonesia, Timor (UMIZ 00157) G. Ventral view, 26 mm long [3066], Indonesia, Bali (UMIZ 00154).	Fig. 46. Paromoionchis goslineri gen. et sp. nov. unit #2, live animals. A. Dorsal view, 33 mm long [3120], Indonesia, Bali (UMIZ 00156). B. Dorsal view, 32 mm long [5073], Indonesia, Halmahera (UMIZ 00158). C. Dorsal and ventral views, 18 mm long [2241], Indonesia, Sulawesi (UMIZ 00153). D. Dorsal view, 24 mm long [3079], Indonesia, Bali (UMIZ 00155). E. Ventral view, 26 mm long [5072], Indonesia, Halmahera (UMIZ 00158). F. Ventral view, 16 mm long [5891], Indonesia, Timor (UMIZ 00157) G. Ventral view, 26 mm long [3066], Indonesia, Bali (UMIZ 00154).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F504FFDF83A4FDF28593F3CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578413/files/figure.png	https://doi.org/10.5281/zenodo.2578413	Fig. 47. Paromoionchis goslineri gen. et sp. nov. unit #1, holotype, Philippines, Luzon [3233] (PNM 041271). A. Digestive system. B. Posterior hermaphroditic (female) reproductive system. C. Male copulatory organs. Abbreviations: dd = deferent duct; ddg = dorsal lobe of digestive gland; fgm = female gland mass; hd = hermaphroditic duct; hg = hermaphroditic gland; i = intestine; ov = oviduct; pdg = posterior lobe of digestive gland; ps = penial sheath; r = rectum; rm = retractor muscle; rs = receptaculum seminis; sp = spermatheca; st = stomach. Scales: A–B = 2 mm; C = 1 mm.	Fig. 47. Paromoionchis goslineri gen. et sp. nov. unit #1, holotype, Philippines, Luzon [3233] (PNM 041271). A. Digestive system. B. Posterior hermaphroditic (female) reproductive system. C. Male copulatory organs. Abbreviations: dd = deferent duct; ddg = dorsal lobe of digestive gland; fgm = female gland mass; hd = hermaphroditic duct; hg = hermaphroditic gland; i = intestine; ov = oviduct; pdg = posterior lobe of digestive gland; ps = penial sheath; r = rectum; rm = retractor muscle; rs = receptaculum seminis; sp = spermatheca; st = stomach. Scales: A–B = 2 mm; C = 1 mm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F504FFDF83A4FDF28593F3CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578415/files/figure.png	https://doi.org/10.5281/zenodo.2578415	Fig. 48. Paromoionchis goslineri gen. et sp. nov. unit #2, Indonesia, Sulawesi [2241] (UMIZ 00153). A. Digestive system. B. Posterior hermaphroditic (female) reproductive system. C. Male copulatory organs. Abbreviations: dd = deferent duct; ddg = dorsal lobe of digestive gland; fgm = female gland mass; hd = hermaphroditic duct; hg = hermaphroditic gland; i = intestine; ov = oviduct; pdg = posterior lobe of digestive gland; ps = penial sheath; r = rectum; rm = retractor muscle; rs = receptaculum seminis; sp = spermatheca; st = stomach. Scales: A = 2 mm; B–C = 1 mm.	Fig. 48. Paromoionchis goslineri gen. et sp. nov. unit #2, Indonesia, Sulawesi [2241] (UMIZ 00153). A. Digestive system. B. Posterior hermaphroditic (female) reproductive system. C. Male copulatory organs. Abbreviations: dd = deferent duct; ddg = dorsal lobe of digestive gland; fgm = female gland mass; hd = hermaphroditic duct; hg = hermaphroditic gland; i = intestine; ov = oviduct; pdg = posterior lobe of digestive gland; ps = penial sheath; r = rectum; rm = retractor muscle; rs = receptaculum seminis; sp = spermatheca; st = stomach. Scales: A = 2 mm; B–C = 1 mm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F504FFDF83A4FDF28593F3CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578417/files/figure.png	https://doi.org/10.5281/zenodo.2578417	Fig. 49. Paromoionchis goslineri gen. et sp. nov. unit #1, radula, Philippines, Luzon [6049] (PNM 041272). A. Rachidian and innermost lateral teeth. B. Lateral teeth with rachidian teeth. C. Lateral teeth. D. Outermost lateral teeth. Scales: A = 20 μm; B–C = 50 μm; D = 10 μm.	Fig. 49. Paromoionchis goslineri gen. et sp. nov. unit #1, radula, Philippines, Luzon [6049] (PNM 041272). A. Rachidian and innermost lateral teeth. B. Lateral teeth with rachidian teeth. C. Lateral teeth. D. Outermost lateral teeth. Scales: A = 20 μm; B–C = 50 μm; D = 10 μm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F504FFDF83A4FDF28593F3CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578419/files/figure.png	https://doi.org/10.5281/zenodo.2578419	Fig. 50. Paromoionchis goslineri gen. et sp. nov. unit #2, radula, Indonesia, Bali [3078] (UMIZ 00155). A. Rachidian and innermost lateral teeth. B. Lateral teeth with rachidian teeth. C. Lateral teeth. D. Outermost lateral teeth. Scales: A–B = 20 μm; C = 30 μm; D = 10 μm.	Fig. 50. Paromoionchis goslineri gen. et sp. nov. unit #2, radula, Indonesia, Bali [3078] (UMIZ 00155). A. Rachidian and innermost lateral teeth. B. Lateral teeth with rachidian teeth. C. Lateral teeth. D. Outermost lateral teeth. Scales: A–B = 20 μm; C = 30 μm; D = 10 μm.	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F504FFDF83A4FDF28593F3CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578421/files/figure.png	https://doi.org/10.5281/zenodo.2578421	Fig. 51. Paromoionchis goslineri gen. et sp. nov., habitats. A. Philippines, Luzon, impacted mangrove next to a village, narrow band of Avicennia by the shore (station 185, type locality). B. Indonesia, Sulawesi, mostly Rhizophora, with sand, small rocks and pieces of wood outside narrow coastal mangrove (station 89). C. Indonesia, Bali, short mangrove shrubs, high intertidal, muddy area with pools (station 155). D. Indonesia, Bali, short mangrove shrubs high in intertidal, muddy area with pools (station 156). E. Indonesia, Halmahera, Rhizophora trees in open mangrove with sandy and muddy areas (station 219).	Fig. 51. Paromoionchis goslineri gen. et sp. nov., habitats. A. Philippines, Luzon, impacted mangrove next to a village, narrow band of Avicennia by the shore (station 185, type locality). B. Indonesia, Sulawesi, mostly Rhizophora, with sand, small rocks and pieces of wood outside narrow coastal mangrove (station 89). C. Indonesia, Bali, short mangrove shrubs, high intertidal, muddy area with pools (station 155). D. Indonesia, Bali, short mangrove shrubs high in intertidal, muddy area with pools (station 156). E. Indonesia, Halmahera, Rhizophora trees in open mangrove with sandy and muddy areas (station 219).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
F83E3A75F504FFDF83A4FDF28593F3CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/2578329/files/figure.png	https://doi.org/10.5281/zenodo.2578329	Fig. 6. Geographic distribution of the five species of Paromoionchis gen. nov. Distinct colors are used for each mitochondrial unit found in P. tumidus (Semper, 1880) comb. nov., P. boholensis gen. et sp. nov. and P. goslineri gen. et sp. nov., and the colors used for each unit are the same as those used in Figs 1–5. Colored areas correspond to hypothetical ranges proposed based on known records (all of which are new here except for the type localities of P. tumidus and P. daemelii (Semper, 1880) comb. nov.).	Fig. 6. Geographic distribution of the five species of Paromoionchis gen. nov. Distinct colors are used for each mitochondrial unit found in P. tumidus (Semper, 1880) comb. nov., P. boholensis gen. et sp. nov. and P. goslineri gen. et sp. nov., and the colors used for each unit are the same as those used in Figs 1–5. Colored areas correspond to hypothetical ranges proposed based on known records (all of which are new here except for the type localities of P. tumidus and P. daemelii (Semper, 1880) comb. nov.).	2019-02-22	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai		Zenodo	biologists	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai			
