identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
F83E3A75F529FF89801FFA6D8753F1BB.text	F83E3A75F529FF89801FFA6D8753F1BB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paromoionchis Dayrat & Goulding & Khalil & Apte & Bourke & Comendador & Tan 2019	<div><p>Paromoionchis Dayrat &amp; Goulding gen. nov.</p><p>urn:lsid:zoobank.org:act: 4506A7F2-CC0B-4F23-8F5E-4DE2AA42B61F</p><p>Type species</p><p>Onchidium tumidum Semper, 1880, designated here.</p><p>Diagnosis</p><p>Body not flattened. No dorsal gills. Dorsal eyes present on notum. Retractable, central papilla (usually with four dorsal eyes) present, often raised above dorsal surface. Eyes at tip of short ocular tentacles. Male opening below right ocular tentacle and to its left. Foot wide. Pneumostome median, on ventral hyponotum. Intestinal loops of type II. Rectal gland absent. Accessory penial gland present or absent. When present, accessory penial gland with muscular sac. Penis with or without hooks.</p><p>Differential diagnosis</p><p>No external diagnostic feature unambiguously distinguishes Paromoionchis gen. nov. from all other genera (which is not surprising because many onchidiid species from different genera are very similar externally). However, Paromoionchis gen. nov. is characterized by a unique combination of internal and external characters: no dorsal gills, male opening below and to the left of the right eye tentacle, no rectal gland and intestinal loops of type II (see Labbé 1934a: 177, fig. 3, for a comparison of digestive types). According to our data, any onchidiid slug with this combination of characters must belong to a species of Paromoionchis gen. nov.</p><p>Etymology</p><p>The name Paromoionchis is a combination of parómoios (παρóμoιoς), which means ʻsimilarʼ in Greek (because members look very similar externally) and onchis, a word derived from the Greek onchos (ὁ ὂγκος) and one of the early names used to refer to onchidiid slugs.</p><p>Gender</p><p>Masculine, the gender of onchis (ICZN Art. 30.1.1), a word derived from the masculine Greek word ὁ ὂγκος (onchos), which means ʻmassʼ or ʻtumor.ʼ As a result (ICZN Art. 31.2), the ending of the specific name tumidum (a Latin adjective) must be changed from neuter (because Onchidium is a name of neuter gender) to masculine ( tumidus).</p><p>Distribution</p><p>The new genus described here is distributed from the western coast of India in the west, all the way to the subtropical waters of Japan (~33° N), Papua New Guinea and the subtropical waters of southeastern Australia (~32° S) in the east (Fig. 6). We did not find Paromoionchis gen. nov. in South Africa, Madagascar or Mauritius, but it is possible that it is present in areas east of Papua New Guinea, such as Fiji and New Caledonia, where we did not collect.</p><p>Habitat</p><p>The five known species of Paromoionchis gen. nov. primarily live on mud, in or next to mangroves, which explains why three species have just been discovered now, because the mangroves of South-East Asia have been very poorly explored. Occasionally, these slugs can also be found in or on muddy logs, coral rubble, sandy mud or even sand with very little mud in it. Paromoionchis tumidus, which is widespread and very common, can be found in nearly all these habitats, even though the mud surface remains where it is most commonly found, like all other species of the genus. Because members of Paromoionchis gen. nov. prefer the mud surface, live animals are often covered with mud.</p><p>Remarks</p><p>A new generic name is needed because no existing name applies to the clade described here. Our remarks are based on the examination of all the type specimens available, especially those of the type species of all genera, the careful analyses of all the original descriptions (especially when no type specimens were available), and our ongoing taxonomic revision of each genus of the family. Three existing generic names are junior synonyms of Onchidella J.E. Gray, 1850, which is not found in the tropical Indo-West Pacific and is characterized by a completely different anatomy (Dayrat 2009; Dayrat et al. 2011b). Seven generic names apply to the clade including all the onchidiid slugs with dorsal gills, i.e., Peronia Fleming, 1822 (Dayrat 2009) . Labella Starobogatov, 1976 is a junior synonym of Onchidium Buchannan, 1800, which applies to a distinct clade including three species (Dayrat et al. 2016). Paraoncidium Labbé, 1934 is a junior synonym of Onchidina Semper, 1882, which applies to a distinct monotypic genus from southeastern Australia (Dayrat &amp; Goulding 2017). Peronina Plate, 1893 applies to a clade including slugs characterized by a pneumostome located at the margin between the dorsal notum and the ventral hyponotum. Platevindex Baker, 1938 applies to a clade including species with a distinctly flattened body and a narrow foot. Semperoncis Starobogatov, 1976 applies to species characterized by a very different anatomy and which are adapted to terrestrial life in the Philippines (Dayrat 2010). And, finally, Melayonchis Dayrat &amp; Goulding, 2017 applies to a distinct clade including slugs with a different anatomy (Dayrat et al. 2017).</p></div>	https://treatment.plazi.org/id/F83E3A75F529FF89801FFA6D8753F1BB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai	Dayrat, Benoît, Goulding, Tricia C., Khalil, Munawar, Apte, Deepak, Bourke, Adam J., Comendador, Joseph, Tan, Shau Hwai (2019): A new genus and three new species of mangrove slugs from the Indo-West Pacific (Mollusca: Gastropoda: Euthyneura: Onchidiidae). European Journal of Taxonomy 500: 1-77, DOI: 10.5852/ejt.2019.500
F83E3A75F52EFFB683F6FE81867DF51D.text	F83E3A75F52EFFB683F6FE81867DF51D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paromoionchis tumidus (Semper 1880) Dayrat & Goulding & Khalil & Apte & Bourke & Comendador & Tan 2019	<div><p>Paromoionchis tumidus (Semper, 1880) comb. nov.</p><p>Figs 7 –25</p><p>Onchidium tumidum Semper, 1880: 262 –263, pl. 20, figs 3–4, pl. 23, fig. 4.</p><p>Onchidium samarense Semper, 1880: 268 –269, pl. 20, figs 9, 13, pl. 23, fig. 7. Syn. nov.</p><p>Onchidium mertoni Simroth, 1918: 294 –296, pl. XX, figs 43–47. Syn. nov.</p><p>Onchidium hongkongense Britton, 1984: 188 –190, figs 6–7. Syn. nov.</p><p>Onchidium samarense – Semper 1882: 268–269, pl. 21, fig. 5.</p><p>Material examined</p><p>Type material SINGAPORE • lectotype (here designated; 28/ 22 mm); ZMB 39019a • 15 paralectotypes; ZMB 39019b • 2 paralectotypes; NHMD 300305 • 1 paralectotype; SMF 333603/1 .</p><p>AUSTRALIA • 2 paralectotypes; Queensland, Mackay; ZMB 39020 .</p><p>Other type material</p><p>PHILIPPINES • lectotype of Onchidium samarense (here designated; 22/ 17 mm); Samar Island, Palapa harbor; ZMB 39025a • 2 paralectotypes of O. samarense (24/20 and 20/ 15 mm); same locality as lectotype; ZMB 39025b .</p><p>INDONESIA • lectotype of Onchidium mertoni (here designated; 15/ 9 mm); Aru Islands, Kobroor, Sungai; 5 Jan. 1908; ZMB 121591a • 4 paralectotypes of O. mertoni (14/8, 14/10, 15/14 and 14/ 10 mm); same data as for lectotype; ZMB 121591 b .</p><p>CHINA • holotype of Onchidium hongkongense (17/ 13 mm); Hong Kong; NHM 1982290 • 15 paratypes; same locality as holotype; NHM 1982291 to 1982292 .</p><p>Notes on type material</p><p>Onchidium tumidum . Lectotype, 28/ 22 mm, designated here (ZMB 39019a). All other syntypes become paralectotypes (the 15 paralectotypes from the same lot are now ZMB 39019b). According to the original description, the type material included 42 specimens from Singapore and an unknown number of specimens from Port Mackay, Queensland, Australia. A total of 21 syntypes were located in museum collections: 19 specimens from Singapore (16 specimens, ZMB 39019; 2 specimens, NHMD 300305; 1 specimen, SMF 333603 /1) and 2 specimens from Mackay (ZMB 39020). There also are two possible syntypes from Australia (ZMH 27480/2). Two similar species of Paromoionchis gen. nov. are found at Port Mackay, P. tumidus and P. daemelii, which anatomically can only be distinguished based on the insertion of the retractor muscle of the penis. In the lectotype designated here from Singapore, the retractor muscle inserts near the heart, exactly as in the species described here. However, in one of the two paralectotypes of P. tumidus from Mackay (ZMB 39020), the retractor muscle is vestigial, as in P. daemelii (in the other paralectotype from Mackay, the male apparatus was destroyed prior to the present investigation and could not be examined). Hence, it was necessary to designate a lectotype from Singapore in order to clarify the application of P. tumidus . Note that the type material was fixed in formalin more than 130 years ago and no DNA sequencing could be attempted.</p><p>Onchidium samarense . Lectotype, 22/ 17 mm, designated here (ZMB 39025a). The two other syntypes become paralectotypes (ZMB 39025b). According to the original description, the type material included only two specimens from the same locality in Samar, Philippines. However, the jar with the type material currently contains three similar-looking specimens (syntypes), all of which were dissected prior to the present study. It is not excluded that the original description was based on only two of those three specimens but it is also possible that Semper himself identified all three specimens as O. samerense [sic] (with a minor typo in the original description). The lectotype still contains all its internal organs, including the male copulatory parts. One paralectotype (24/ 20 mm) is mostly destroyed, with no internal organs left except the digestive gland (a few destroyed pieces of organs are in a vial). The other paralectotype (20/ 15 mm) still contains internal organs, but the male parts are missing. Our observations and comments are mostly based on the only specimen with male parts; hence its designation as a lectotype. Note that the type material was fixed in formalin more than 130 years ago and no DNA sequencing could be attempted. Note also that if, in the future, Onchidium samarense were to be regarded as a valid species name in Paromoionchis gen. nov., the specific name samarense (neuter) would need to become samarensis (masculine).</p><p>Onchidium mertoni . Lectotype, 15/ 9 mm, designated here (ZMB 121591 a). The four other syntypes become paralectotypes (ZMB 121591 b). Simroth mentioned in the original description that all five specimens were very hard. Indeed, it seems that they dried out at some point and they are very poorly preserved. The lectotype designated here is the only specimen that is complete. It was partially dissected for the present study (the penial hooks, identical to those of O. tumidum, are illustrated here). Two paralectotypes (14/8 and 14/ 10 mm) were dissected prior to the present study and are completely empty. Two other paralectotypes (15/14 and 14/ 10 mm) are in very poor condition (the body is extremely hard and the digestive system is partly outside the body through the foot). A lectotype is designated here to clarify the application of the name O. mertoni because several species of Paromoionchis gen. nov. are potentially sympatric in the Aru Islands and so it cannot be excluded that the five original syntypes belong to different species. Note that the type material was fixed in formalin more than 100 years ago and no DNA sequencing could be attempted.</p><p>Onchidium hongkongense . Holotype, 17/ 13 mm, by original designation (NHM 1982290) and 15 paratypes (NHM 1982291, NHM 1982292). The holotype is largely destroyed due to prior dissection, likely by Britton. Large parts of the notum and of the reproductive organs are missing. Even though it is mostly destroyed, the digestive system is confirmed to be of type II. A few paratypes were checked for the present study and their anatomy matches that of the holotype. Note that the type material was fixed in formalin more than 40 years ago and no DNA sequencing could be attempted. Note also that the specific name hongkongensis (masculine or feminine gender) originally used by Britton is corrected to hongkongense (neuter) for gender agreement with Onchidium . Should Onchidium hongkongense ever become a valid species name in Paromoionchis gen. nov., hongkongense would then need to be changed back to hongkongensis.</p><p>Other material</p><p>AUSTRALIA – New South Wales • 1 spec. (20/15 [1522] mm); Sydney, Pittwater, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=151.3313&amp;materialsCitation.latitude=-33.62205" title="Search Plazi for locations around (long 151.3313/lat -33.62205)">Careel Bay</a>; 33°37.323´S, 151°19.878´E; 24 Nov. 2011; station 40; supratidal zone on margin of salt marsh, mangrove patch on side of creek; AM C.468918.005 • 1 spec. (35/20 [1529] mm); Sydney, Hawkesbury River, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=151.19449&amp;materialsCitation.latitude=-33.51145" title="Search Plazi for locations around (long 151.19449/lat -33.51145)">Cheero Point</a>; 33°30.687´S, 151°11.669´E; 25 Nov. 2011; station 42; open mangrove with old logs; AM C.468924.001 • 1 spec. (33/20 [1528] mm); same data as for preceding; AM C.468923.002 • 1 spec. (32/20 [1530] mm); same data as for preceding; AM C.468925.001 . – Northern Territory • 1 spec. (45/32 [1634] mm); Darwin, near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.9332&amp;materialsCitation.latitude=-12.582983" title="Search Plazi for locations around (long 130.9332/lat -12.582983)">Channel Island</a> Road; 12°34.979´S, 130°55.992´E; 16 Aug. 2012; station 65; sequence of Sonneratia, Rhizophora and Ceriops; NTM P.57620 • 1 spec. (40/25 [1686] mm); Darwin, end of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.88156&amp;materialsCitation.latitude=-12.559283" title="Search Plazi for locations around (long 130.88156/lat -12.559283)">Channel Island</a> Road; 12°33.557´S, 130°52.894´E; 17 Aug. 2012; station 66; sequence of Sonneratia, Rhizophora and Ceriops; NTM P.57621 • 1 spec. (42/38 [1638] mm); same data as for preceding; NTM P.57623 • 2 spec. (30/17 [1705] and 17/12 [1645] mm); Darwin, close to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.9125&amp;materialsCitation.latitude=-12.4797" title="Search Plazi for locations around (long 130.9125/lat -12.4797)">Tiger Brenan Road</a> (small service road); 12°28.782´S, 130°54.750´E; 19 Aug. 2012; station 69; high tidal Ceriops; NTM P.57622 • 1 spec. (36/22 [1651] mm); Darwin, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.9607&amp;materialsCitation.latitude=-12.548217" title="Search Plazi for locations around (long 130.9607/lat -12.548217)">Elizabeth Road</a>; 12°32.893´S, 130°57.642´E; 20 Aug. 2012; station 70; Ceriops and old logs in Rhizophora forest; NTM P.57624 . – Queensland • 1 spec. (45/30 [2562] mm); Cairns, Yorkey’s <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.7128&amp;materialsCitation.latitude=-16.8093" title="Search Plazi for locations around (long 145.7128/lat -16.8093)">Knob</a>; 16°48.558´S, 145°42.768´E; 17 Jun. 2013; station 101; hard, red mud with grasses; MTQ • 1 spec. (45/30 [2602] mm); Townsville, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=146.81715&amp;materialsCitation.latitude=-19.165634" title="Search Plazi for locations around (long 146.81715/lat -19.165634)">Magnetic Island</a>; 19°09.938´S, 146°49.029´E; 24 Jun. 2013; station 109; water on the mud; MTQ • 1 spec. (15/10 [2627] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.26463&amp;materialsCitation.latitude=-20.010967" title="Search Plazi for locations around (long 148.26463/lat -20.010967)">Bowen</a>; 20°00.658´S, 148°15.878´E; 1 Jul. 2013; station 115; back of mangrove across from beach, dense Rhizophora, Avicennia trees with soft mud around; MTQ • 1 spec. (55/30 [2637] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.26242&amp;materialsCitation.latitude=-20.015217" title="Search Plazi for locations around (long 148.26242/lat -20.015217)">Bowen</a>; 20°00.913´S, 148°15.745´E; 1 Jul. 2013; station 116; mangrove away from ocean, small area of open Avicennia mangrove, surrounded by Rhizophora; MTQ • 1 spec. (20/10 [2652] mm); Bowen, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.23909&amp;materialsCitation.latitude=-20.021067" title="Search Plazi for locations around (long 148.23909/lat -20.021067)">Doughty Creek</a>; 20°01.264´S, 148°14.345´E; 2 Jul. 2013; station 117; narrow Avicennia and Rhizophora mangrove, by creek, some muddy areas and some very sandy; MTQ • 1 spec. (35/20 [2657] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.23706&amp;materialsCitation.latitude=-20.024633" title="Search Plazi for locations around (long 148.23706/lat -20.024633)">Bowen</a>; 20°01.478´S, 148°14.224´E; 3 Jul. 2013; station 119; Rhizophora and Avicennia mangrove; MTQ • 1 spec. (30/20 [2701] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.20126&amp;materialsCitation.latitude=-20.14185" title="Search Plazi for locations around (long 149.20126/lat -20.14185)">Mackay</a>; 20°08.511´S, 149°12.076´E; 8 Jul. 2013; station 125; large, dense and sandy mangrove and, by side of river, small strip of mud with Avicennia and Rhizophora; MTQ • 1 spec. (30/25 [1531] mm); Thirsty Sound, Plum Tree, beach in front of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.03093&amp;materialsCitation.latitude=-22.135733" title="Search Plazi for locations around (long 150.03093/lat -22.135733)">Endeavour Park</a>; 22°08.144´S, 150°01.856´E; 14 Sep. 2002; I. Loch, D.L. Beechey and A.C. Miller leg.; sheltered, muddy cobble shore; AM C.575588 .</p><p>BRUNEI DARUSSALAM • 3 spec. (55/30 [1036], 35/20 [1035] and 20/15 [1062] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.04607&amp;materialsCitation.latitude=4.920767" title="Search Plazi for locations around (long 115.04607/lat 4.920767)">Pulau Pyatan</a>, Teluk Brunei; 04°55.246´N, 115°02.764´E; 27 Jul. 2011; station 32; open Avicennia and Rhizophora mangrove, with hard mud; BDMNH .</p><p>INDIA • 1 spec. (26/17 [1119] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=92.77295&amp;materialsCitation.latitude=12.330717" title="Search Plazi for locations around (long 92.77295/lat 12.330717)">Andaman Islands</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=92.77295&amp;materialsCitation.latitude=12.330717" title="Search Plazi for locations around (long 92.77295/lat 12.330717)">Middle Andaman</a>, Shantipur, Kadamtala; 12°19.843´N, 092°46.377´E; 12 Jan. 2011; station 58; open area with hard mud and many old logs, next to a mangrove with medium trees; BNHS 88 .</p><p>INDONESIA – Sumatra • 1 spec. (24/15 [1732] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.85478&amp;materialsCitation.latitude=-5.27125" title="Search Plazi for locations around (long 105.85478/lat -5.27125)">Kualapenet</a>; 05°16.275´S, 105°51.287´E; 17 Oct. 2012; station 77; narrow band of mangrove between ocean and fish ponds; UMIZ 0 0 121 • 2 spec. (35/20 [1754] and 26/16 [1755] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.77&amp;materialsCitation.latitude=-5.8426666" title="Search Plazi for locations around (long 105.77/lat -5.8426666)">Bakauheni</a>; 05°50.560´S, 105°46.200´E; 21 Oct. 2012; station 81; small mangrove, not far from road and next to large harbor, very impacted mangrove; UMIZ 0 0 122 • 1 spec. (38/30 [1794] mm); same data as for preceding; UMIZ 0 0 138 • 1 spec. (20/12 [1798] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.251884&amp;materialsCitation.latitude=-5.5443335" title="Search Plazi for locations around (long 105.251884/lat -5.5443335)">S of Bandar</a> Lampung; 05°32.66´S, 105°15.113´E; 28 Oct. 2012; station 83; high intertidal, fairly dense roots with some Avicennia and Nypa, edge of mangrove by road; UMIZ 0 0 123 . – Sulawesi • 2 spec. (25/15 [2200] and 20/12 [2201] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=125.01328&amp;materialsCitation.latitude=1.6918833" title="Search Plazi for locations around (long 125.01328/lat 1.6918833)">Tamperong</a>; 01°41.513´N, 125°00.797´E; 12 Mar. 2013; station 87; muddy mangrove with small Rhizophora in dense patches; UMIZ 0 0 124 • 1 spec. (27/15 [2240] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=124.543236&amp;materialsCitation.latitude=1.36295" title="Search Plazi for locations around (long 124.543236/lat 1.36295)">Sondaken</a>; 01°21.777´N, 124°32.594´E; 13 Mar. 2013; station 89; sand, small rocks, pieces of wood outside narrow coastal mangrove of mostly Rhizophora; UMIZ 0 0 125 • 1 spec. (30/20 [2345] mm); Makassar, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=119.44628&amp;materialsCitation.latitude=-5.10195" title="Search Plazi for locations around (long 119.44628/lat -5.10195)">Tallo</a> mangrove; 05°06.117´S, 119°26.777´E; 21 Mar. 2013; station 92; small mangrove used as outhouse by village, very impacted with trash; UMIZ 0 0 126 • 1 spec. (20/13 [2355] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=119.59922&amp;materialsCitation.latitude=-4.42395" title="Search Plazi for locations around (long 119.59922/lat -4.42395)">Barru</a>; 04°25.437´S, 119°35.953´E; 22 Mar. 2013; station 93; forest of mostly Avicennia and Rhizophora, with hard and sandy mud; UMIZ 0 0 127. – Ambon • 1 spec. (25/15 [3541] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=119.59922&amp;materialsCitation.latitude=-4.42395" title="Search Plazi for locations around (long 119.59922/lat -4.42395)">Lateri</a>; 03°38.261´S, 128°14.716´E; 12 Feb. 2014; station 128; mudflat next to small creek in low intertidal of mangrove preserve; UMIZ 0 0 128 • 3 spec. (45/30 [2832], 22/12 [2839] and 35/22 [2840] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.24638&amp;materialsCitation.latitude=-3.6372833" title="Search Plazi for locations around (long 128.24638/lat -3.6372833)">Lateri</a>; 03°38.237´S, 128°14.783´E; 14 Feb. 2014; station 131; muddy mangrove with Rhizophora; UMIZ 0 0 129. – Seram • 2 spec. (20/15 [2874] and 30/15 [2875] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.24638&amp;materialsCitation.latitude=-3.6372833" title="Search Plazi for locations around (long 128.24638/lat -3.6372833)">Kawa</a>; 02°58.240´S, 128°07.066´E; 18 Feb. 2014; station 135; mud next to a seawall adjacent to a mangrove; UMIZ 0 0 130. – Lombok • 2 spec. (30/20 [2950] and 20/12 [2952] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=116.0482&amp;materialsCitation.latitude=-8.762466" title="Search Plazi for locations around (long 116.0482/lat -8.762466)">Tanjung Batu village</a>; 08°45.748´S, 116°02.892´E; 24 Mar. 2014; station 145; Avicennia forest; UMIZ 0 0 131 • 1 spec. (20/14 [2961] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=116.5473&amp;materialsCitation.latitude=-8.866" title="Search Plazi for locations around (long 116.5473/lat -8.866)">Seriwe Bay</a>; 08°51.960´S, 116°32.838´E; 25 Mar. 2014; station 146; Avicennia mangrove with hard mud and rocks; UMIZ 0 0 132 • 1 spec. (18/10 [2960] mm); same data as for preceding; UMIZ 0 0 139 . – Bali • 1 spec. (25/14 [3051] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.21995&amp;materialsCitation.latitude=-8.790584" title="Search Plazi for locations around (long 115.21995/lat -8.790584)">Denpasar</a>; 08°47.435´S, 115°13.197´E; 1 Apr. 2014; station 153; large mangrove by road, very soft mud; UMIZ 0 0 133 • 1 spec. (30/20 [3070] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.18005&amp;materialsCitation.latitude=-8.768766" title="Search Plazi for locations around (long 115.18005/lat -8.768766)">Denpasar</a>; 08°46.126´S, 115°10.803´E; 2 Apr. 2014; station 154; large mangrove by road, with shallow mud; UMIZ 0 0 134. – Halmahera • 1 spec. (25/15 [5082] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.65152&amp;materialsCitation.latitude=1.02215" title="Search Plazi for locations around (long 127.65152/lat 1.02215)">Akelamo</a>; 01°01.329´N, 127°39.091´E; 10 Mar. 2015; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.65152&amp;materialsCitation.latitude=1.02215" title="Search Plazi for locations around (long 127.65152/lat 1.02215)">station 207</a>; sandymuddy beach at margin of mangrove near village; UMIZ 0 0 135 • 2 spec. (50/35 [5102] and 55/35 [5103] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.34354&amp;materialsCitation.latitude=0.9241" title="Search Plazi for locations around (long 128.34354/lat 0.9241)">Buli</a>; 00°55.446´N, 128°20.612´E; 16 Mar. 2015; station 212; logged area in front of old Rhizophora forest, by the road; UMIZ 0 0 136 • 1 spec. (55/30 [5042] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.34412&amp;materialsCitation.latitude=0.9227833" title="Search Plazi for locations around (long 128.34412/lat 0.9227833)">Buli</a>; 00°55.367´N, 128°20.647´E; 17 Mar. 2015; station 213; tall and old Rhizophora forests, high intertidal; UMIZ 0 0 137 .</p><p>JAPAN • 1 spec. (22/10 [3761] mm); Ehime Prefecture, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.55342&amp;materialsCitation.latitude=32.960567" title="Search Plazi for locations around (long 132.55342/lat 32.960567)">Misho Bay</a>; 32°57.634´N, 132°33.205´E; 4 Aug. 2014; station o28; mudflats; NSMT Mo 78984 .</p><p>MALAYSIA • 1 spec. (40/25 [963] mm); Peninsular Malaysia, Nibong Tebal, Pulau Burung; 05°12.488´N, 100°25.564´E; 11 Jul. 2011; station 17; soft mud, open mangrove of Rhizophora, with a few Sonneratia; USMMC 0 0 0 57 • 1 spec. (18/12 [928] mm); E Peninsular Malaysia, Balok; 03°53.219´N, 103°21.978´E; 14 Jul. 2011; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.3663&amp;materialsCitation.latitude=3.8869834" title="Search Plazi for locations around (long 103.3663/lat 3.8869834)">station 19</a>; mostly Rhizophora, with some Avicennia, hard mud with shallow pools, patches of soft mud; USMMC 0 0 0 58.</p><p>PAPUA NEW GUINEA • 1 spec. (18/14 [5432] mm); Madang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.79&amp;materialsCitation.latitude=-5.2033334" title="Search Plazi for locations around (long 145.79/lat -5.2033334)">Meiro River</a>, near airport; 05°12.2´S, 145°47.4´E; 5 Nov. 2012; MNHN expedition Papua Niugini leg.; station PM01; Nypa palm swamp; MNHN IM-2013-10478 • 1 spec. (17/12 [5433] mm); same data as for preceding; MNHN IM-2013- 10479 .</p><p>PHILIPPINES – Luzon • 1 spec. (26/18 [3610] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.62452&amp;materialsCitation.latitude=13.968833" title="Search Plazi for locations around (long 120.62452/lat 13.968833)">Lian</a>, Batangas; 13°58.130´N, 120°37.471´E; 5 Jul. 2014; station 179; narrow and impacted mangrove of Avicennia near village, very sandy, little to no mud; PNM 0 41261 • 2 spec. (30/20 [3171] and 25/15 [3192] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.61362&amp;materialsCitation.latitude=14.178567" title="Search Plazi for locations around (long 120.61362/lat 14.178567)">Nasugbu</a>, Batangas; 14°10.714´N, 120°36.817´E; 6 Jul. 2014; station 182; near village, well-preserved and dense forest of Avicennia and Rhizophora; PNM 0 41255 • 1 spec. (22/14 [3172] mm); same data as for preceding; PNM 0 41262 • 2 spec. (30/20 [3200] and 35/22 [3205] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.621&amp;materialsCitation.latitude=13.921984" title="Search Plazi for locations around (long 120.621/lat 13.921984)">Calantagan</a>, Batangas; 13°55.319´N, 120°37.260´E; 7 Jul. 2014; station 183; rocks next to Avicennia and Rhizophora forest; PNM 0 41256 • 1 spec. (30/18 [3202] mm); same data as for preceding; PNM 0 41263 • 1 spec. (35/22 [3222] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.61874&amp;materialsCitation.latitude=13.887967" title="Search Plazi for locations around (long 120.61874/lat 13.887967)">Calantagan</a>, Batangas; 13°53.278´N, 120°37.124´E; 8 Jul. 2014; station 184; narrow forest on the shore, Avicennia and young Rhizophora; PNM 0 41257 • 1 spec. (33/15 [3229] mm); same data as for preceding; PNM 0 41264 • 1 spec. (27/18 [3237] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.62305&amp;materialsCitation.latitude=13.8544" title="Search Plazi for locations around (long 120.62305/lat 13.8544)">Calantagan</a>, Batangas; 13°51.264´N, 120°37.383´E; 8 Jul. 2014; station 185; next to village, impacted, narrow Avicennia mangrove by the shore; PNM 0 41265 . – Bohol • 1 spec. (40/25 [3344] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=124.52808&amp;materialsCitation.latitude=9.858867" title="Search Plazi for locations around (long 124.52808/lat 9.858867)">Mabini</a>; 09°51.532´N, 124°31.685´E; 17 Jul. 2014; station 194; narrow mangrove on edge of fish ponds, tall Rhizophora and Avicennia trees, many old logs; PNM 0 41258 • 1 spec. (26/15 [3371] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=124.56925&amp;materialsCitation.latitude=9.859767" title="Search Plazi for locations around (long 124.56925/lat 9.859767)">Mabini</a>; 09°51.586´N, 124°34.155´E; 18 Jul. 2014; station 196; Avicennia and Sonneratia open forest with sand, algae and coral rubble; PNM 0 41259 • 1 spec. (30/18 [3416] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=123.82315&amp;materialsCitation.latitude=9.738" title="Search Plazi for locations around (long 123.82315/lat 9.738)">Maribojoc</a>; 09°44.280´N, 123°49.389´E; 20 Jul. 2014; station 202; uplifted coral rubble with sand and algae, near Sonneratia trees; PNM 0 41260 .</p><p>VIETNAM • 2 spec. (45/30 [5619] and 40/30 [5682] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.15953&amp;materialsCitation.latitude=12.212967" title="Search Plazi for locations around (long 109.15953/lat 12.212967)">Nha Trang</a>; 12°12.778´N, 109°09.572´E; 27 Jul. 2015; station 237; small strip of mud with a few Rhizophora trees next to a small river, by fish ponds and houses; ITBZC IM 0 0 0 19 • 1 spec. (25/15 [5642] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.16763&amp;materialsCitation.latitude=12.4028" title="Search Plazi for locations around (long 109.16763/lat 12.4028)">Nha Trang</a>; 12°24.168´N, 109°10.058´E; 29 Jul. 2015; station 239; mostly Avicennia and some small Rhizophora, with shallow mud; ITBZC IM 0 0 0 20 .</p><p>Color and morphology of live animals (Figs 7–10)</p><p>Live animals of units #1 and #2 are often abundantly covered with mud, in which case their dorsal color can hardly be seen. The background of the dorsal notum is brown, light to dark. That background can be homogenous or clearly mottled with darker or lighter areas and, occasionally, also with red areas. In addition, in some animals the tip of the dorsal papillae (with and without dorsal eyes) can be bright yellow. The foot varies from gray (light or dark) to yellow or orange. The hyponotum is almost always yellow, from pale yellow to bright yellow and even orange. This variable yellow component can cover the entire hyponotum or just an outer ring (the inner ring being light to dark gray). The color of the foot and of the hyponotum of an individual can change rapidly, especially when disturbed. The ocular tentacles are brown (variable from light to dark) and may or may not be speckled with tiny white dots, exactly like the head. The ocular tentacles are short (just a few millimeters long).</p><p>No live pictures were available for unit #3, so the following description is based on preserved specimens (Fig. 10). It is possible that bright colors (yellow, orange) were lost during preservation on both the ventral and dorsal sides. The background of the dorsal notum is brown, mottled with darker or lighter areas. The foot is light gray. The hyponotum is gray-brown, with a reddish hue on the margin (which could possibly be orange in live animals). The color of the ocular tentacles (retracted, likely short) cannot be determined.</p><p>Generally speaking, the dorsal notum of any given live animal can rapidly change from almost perfectly smooth to covered by many papillae. However, when animals are not disturbed, the dorsum is usually covered by papillae of various sizes. In some animals, larger papillae may be arranged in two longitudinal and lateral ridges (on either side of the median line), but those ridges can appear and disappear rapidly. Some papillae bear from two to four black dorsal eyes at their tip (most papillae bear three eyes). The number of papillae with dorsal eyes is variable (between 10 and 15, on average) and they mostly are on the central part of the notum. Their tip is usually yellow, but not always. A central, much larger papilla, which bears four dorsal eyes (sometimes three), is entirely retractable within the notum. In addition to all these large papillae, the notum is covered by smaller, rounded papillae, which can make it look granular.</p><p>External morphology (Fig. 11 A–B)</p><p>Preserved specimens no longer display the color of live animals. The body is not flattened (although, exceptionally, animals on mud with a thin layer of water can look flattened). The notum is oval. Dorsal gills are absent. The large, central, retractable papilla at the center of the notum can only be seen in live animals. In preserved specimens, it is retracted inside the notum. The hyponotum is horizontal. The width of the hyponotum relative to the total width of the ventral surface (pedal sole and hyponotum) varies among individuals, from approximately one third to half. In the anterior region, the left and right ocular tentacles are superior to the mouth. Eyes are located at the tip of the two ocular tentacles. Inferior to the ocular tentacles, superior to the mouth, the head bears a pair of oral lobes. The latter are smooth, with no transversal protuberance. The male opening (of the copulatory complex) is below and to the left of the right ocular tentacle (i.e., between the two ocular tentacles). The anus is posterior, median, close to the edge of the pedal sole. On the right side (to the left in ventral view), a peripodial groove is present at the junction between the foot and the hyponotum, running longitudinally from the buccal area to the posterior end, a few millimeters from the anus and the pneumostome. The pneumostome is median. Its position on the hyponotum relative to the notum margin and the edge of the pedal sole varies among individuals but averages in the middle. The position of the female pore (at the posterior end of the peripodial groove) does not vary much among individuals.</p><p>Visceral cavity and pallial complex</p><p>The anterior pedal gland is oval and flattened, lying free on the floor of the visceral cavity below the buccal mass. The heart, enclosed in the pericardium, is on the right side of the visceral cavity, slightly posterior to the middle. From the anterior ventricle an anterior vessel exits that supports several anterior organs such as the buccal mass, the nervous system and the copulatory complex. The auricle is posterior. The kidney is more or less symmetrical, the right and left parts being equally developed. The kidney is intricately attached to the respiratory complex. The lung is in two more or less symmetrical parts, left and right.</p><p>Digestive system (Figs 12–15)</p><p>There are no jaws. The left and right salivary glands, heavily branched, join the buccal mass dorsally, on either side of the esophagus. The radula is in between two large postero-lateral muscular masses. Radulae measure up to 5.2 mm in length (unit #1), 4.1 mm (unit #2) and 2.8 mm (unit #3). Each radular row contains a rachidian tooth and two half rows of lateral teeth of similar size and shape. Examples of radular formulae are presented in Table 4. The rachidian teeth are unicuspid: the median cusp is always present; there are no conspicuous cusps on the lateral sides of the base of the rachidian tooth. The length of the rachidian teeth (approximately 25 µm) tend to be approximately half the size of the lateral teeth (approximately 50 µm). The lateral aspect of the base of the rachidian teeth is straight, occasionally slightly convex. The half rows of lateral teeth form an angle of 45° with the rachidian axis. With the exception of the few innermost and few outermost lateral teeth, the size and shape of the lateral teeth do not vary along the half row, nor do they vary among half rows. The lateral teeth seem to be unicuspid with a flattened and curved hook (approximately 50 µm long) with a rounded tip, but there is also a pointed spine on the outer lateral expansion of the base (basal lateral spine). In most cases, the basal lateral spine cannot be observed because it is hidden below the hook of the next, outer lateral tooth. It can only be observed when the teeth are not too close (such as in the innermost and outermost regions) or when teeth are placed in an unusual position. The inner and outer lateral aspects of the hook of the lateral teeth are straight (i.e., not wavy and not with any protuberance).</p><p>The esophagus is narrow and straight, with thin internal folds. The esophagus enters the stomach anteriorly. Only a portion of the posterior aspect of the stomach can be seen in dorsal view because it is partly covered by the lobes of the digestive gland. The dorsal lobe is mainly on the right. The left, lateral lobe is mainly ventral. The posterior lobe covers the posterior aspect of the stomach. The stomach is a U-shaped sac divided into four chambers. The first chamber, which receives the esophagus, is delimited by thin tissue and receives the ducts of the dorsal and lateral lobes of the digestive gland. The second, posterior chamber, delimited by thick muscular tissue, receives the duct of the posterior lobe of the digestive gland. The third, funnel-shaped chamber is delimited by thin tissue with high ridges internally. The fourth chamber is continuous and externally similar to the third, but it bears only low, thin ridges internally. The intestine is long and narrow and the intestinal loops are of type II. There is no rectal gland.</p><p>Nervous system (Fig. 11C)</p><p>The circum-esophageal nerve ring is post-pharyngeal and pre-esophageal. The paired cerebral ganglia are close and the cerebral commissure is short (but its length does vary among individuals). Paired pleural and pedal ganglia are also all distinct. The visceral commissure is short but distinctly present and the visceral ganglion is more or less median. Cerebro-pleural and pleuro-pedal connectives are short and pleural and cerebral ganglia touch each other on either side. Nerves from the cerebral ganglia innervate the buccal area and the ocular tentacles and, on the right side, the penial complex. Nerves from the pedal ganglia innervate the foot. Nerves from the pleural ganglia innervate the lateral and dorsal regions of the mantle. Nerves from the visceral ganglia innervate the visceral organs.</p><p>Reproductive system (Figs 16 –23)</p><p>Sexual maturity is correlated with animal length. Mature individuals have large female organs (with a large female gland mass) and fully-developed male copulatory parts. Immature individuals (&lt;15 mm long) may have inconspicuous (or no) female organs and rudimentary anterior male parts.</p><p>The female organs are located at the posterior end of the visceral cavity, mixed with some male parts (Figs 16A, 17 A–B). The hermaphroditic gland is a single mass, joining the spermoviduct through the hermaphroditic duct (which conveys the eggs and the autosperm). There is a narrow and bent receptaculum seminalis (caecum) along the hermaphroditic duct. The female gland mass contains various glands (mucus and albumen) which can hardly be separated by dissection and of which the exact connections remain uncertain. The hermaphroditic duct becomes the spermoviduct (which conveys eggs, exosperm and autosperm). Proximally, the spermoviduct is not divided (at least externally) and is embedded within the female gland mass. Distally, the spermoviduct branches into the deferent duct (which conveys the autosperm up to the anterior region, running through the body wall) and the oviduct. The free oviduct conveys the eggs up to the female opening and the exosperm from the female opening up to the fertilization chamber. The large, ovate-spherical spermatheca connects to the oviduct through a narrow and short duct. The oviduct is narrow and straight. There is no vaginal gland.</p><p>The male anterior organs consist of the penial complex (penis, penial sheath, vestibule, deferent duct, retractor muscle) and the accessory penial gland (Figs 16 B–C, 17C–D). The penial complex and the accessory penial gland share the same vestibule and the same anterior male opening. The penial gland is a long, tube-like flagellum with a proximal dead end. The length of the flagellum of the penial gland varies among individuals but it is always heavily coiled. Near its distal end (just before the hollow spine), the flagellum is enlarged into a thick muscular sac. Distally, the flagellum ends in a hard, hollow spine protected by a sheath which opens into the vestibule. The hollow spine is narrow, elongated and slightly curved (Figs 18–20). Its base is conical. Its diameter is between 60 and 100 μm. The diameter of the opening at its tip measures between 30 and 60 μm. Its length ranges from 1 mm ([1634] NTM P.57620) to 2 mm ([5619] ITBZC IM 0 0 0 19, [5102] UMIZ 00136) for unit #1, from 1.2 mm ([1638] NTM P.57623) to 1.8 mm ([3237] PNM 0 41265, [3172] PNM 041262) for unit #2 and from 0.8 mm ([5433] MNHN IM-2013-10479) to 1 mm ([5432] MNHN IM-2013-10478) for unit #3, and its shape does vary between individuals (Figs 18–20). There is no disc separating the spine of the penial gland and the vestibule.</p><p>The penial sheath is narrow and elongated (Figs 16 B–C, 17C–D). The penial sheath protects the penis for its entire length. The beginning of the retractor muscle marks the separation between the penial sheath (and the penis inside) and the deferent duct. The retractor muscle is shorter than the penial sheath and inserts on the wall of the body cavity, near the heart. The deferent duct is also highly convoluted, with many loops. Inside the penial sheath, the penis is a narrow, elongated, soft, hollow tube of approximately 200 μm in diameter. Inside the tube-like penis, six longitudinal ridges bear sparse, tiny, conical (but not pointed) hooks which are less than 20 μm long in unit #1, less than 22 μm long in unit #2 and less than 28 μm in unit #3 (Figs 21–23). When the penis is retracted inside the penial sheath, the hooks are inside the tube-like penis; during copulation, the penis is evaginated like a glove and the hooks are outside.</p><p>Distinctive diagnostic features</p><p>Externally, Paromoionchis tumidus (Semper, 1880) cannot be distinguished from other species of Paromoionchis gen. nov. Internally, the presence of penial hooks distinguishes it from other species of the genus (Table 3).</p><p>Distribution (Fig. 6)</p><p>All records here are new, except for the type localities.</p><p>Unit #1. Australia: New South Wales, Northern Territory, Queensland. Brunei Darussalam. Hong Kong (type locality of Onchidium hongkongense). India: Andaman Islands. Indonesia: Ambon, Aru Islands, Bali, Halmahera, Lombok, Seram, Sulawesi, Sumatra. Japan. Malaysia: Peninsular Malaysia. Singapore (type locality of Onchidium tumidum). Philippines: Bohol, Luzon, Samar (type locality of Onchidium samarense). Vietnam.The southernmost locality is in Sydney, New South Wales, Australia (33°37.323´S) and the northernmost locality is in Misho Bay, Ehime Prefecture, Japan (32°57.634´N).</p><p>Unit #2. Australia: Northern Territory. Indonesia: Sumatra, Lombok. Philippines: Luzon.</p><p>Unit #3. Papua New Guinea: Madang.</p><p>Habitat (Figs 24–25)</p><p>Paromoionchis tumidus unit #1 is predominantly found on mud, hard or soft, inside or near mangroves, or on mudflats (Fig. 24). It is also found on old, muddy logs, inside or near mangroves. It occasionally is found on muddy sand, or even rocks and coral rubble, usually in the proximity of some mangrove trees. It is not found on rocky shores. Paromoionchis tumidus unit #2 is found in mangroves, mostly on mud and occasionally on sand (Fig. 25). Paromoionchis tumidus unit #3 is found in Nypa palm swamps and seems rare (only two specimens are known).</p><p>Paromoionchis tumidus is very common across its entire distribution range. It is by far the most abundant species of Paromoionchis gen. nov. and arguably the most abundant onchidiid species in the Indo-West Pacific. Most individuals of P. tumidus are part of unit #1, because P. tumidus unit #2 is rare across its entire distribution (it is only known from a total of nine specimens collected at nine stations) and P. tumidus unit #3 is restricted to two individuals from Papua New Guinea.</p><p>Remarks</p><p>The publication dates of the various sections of the volume on Landmollusken by Carl Semper in the Reisen im Archipel der Philippinen series were clarified by Johnson (1969). The species name Onchidium tumidum was published by Semper with a complete description (text and figures) in 1880.</p><p>The anatomy of the species described here is fully compatible with Semper’s original description of Onchidium tumidum as well as our own observation of the lectotype (and paralectotypes) from Singapore (Table 3). The most important characters are the lack of a rectal gland, a digestive system of type II, an accessory penial gland, a retractor muscle of the penis inserting near the heart, a male opening between the two eye tentacles (not just below the right eye tentacle), and a penis with hooks (Figs 12H, 21D). According to Semper (1880: 263, our translation), the male opening is “almost exactly midway between the two [eye tentacles],” but it actually is closer to the right tentacle. Also, Semper (1880: 263, our translation) described a penis with an “anterior tooth-bearing portion [which] is reduced, namely at most 2 mm long.” It is confirmed here with SEM (Fig. 21D) that the penis of the lectotype bears tiny hooks (&lt;20 μm) and is fully compatible with the species described here.</p><p>The lectotype of O. mertoni is anatomically identical to the species described here, P. tumidus . Simroth did not describe the internal anatomy of O. mertoni, but a description of its lectotype is provided here. Simroth mentioned a male aperture below the right ocular tentacle, but it clearly is to the left of the right tentacle (Fig. 11A); the intestinal loops are of type II; the male apparatus includes an accessory penial gland; the penial retractor muscle inserts to the body wall near the heart; the penis bears tiny hooks which are identical to the hooks of the species described here (Fig. 21C).</p><p>The type material of O. hongkongense is anatomically identical to the species described here and it is characterized by the exact same combination of characters: no rectal gland, intestinal loops of type II (Fig. 12D), an accessory penial gland, a retractor muscle inserting near the heart, a male opening between the two eye tentacles and a penis with tiny hooks (which Britton illustrated and measured as &lt;20 μm).</p><p>Strictly speaking, Onchidium samarense probably should be regarded as a nomen dubium because 1) the jar with the type material includes three specimens while Semper only mentioned two specimens in the original description, 2) the male organs of two of the syntypes are gone (possibly dissected by Semper) and cannot be checked, and 3) important features (e.g., the insertion of the retractor muscle) are not mentioned in the original description. However, our observations are compatible with the original description and it is possible that all three specimens were actually identified as O. samarense by Semper himself. According to Semper (1882: 269, our translation), the penis of O. samarense is “very similar to that of O. tumidum .” Indeed, the male apparatus of the lectotype is very similar to that of the species described here. In particular, the retractor muscle of the penis (not described by Semper) is thin but reaches the heart, following some nerves, as in the species described here. Semper described no ʻcartilaginous teeth.’ However, that can be easily explained by the fact that the hooks inside the penis of O. tumidum are soft and tiny (&lt;20 μm) and, unlike the large and solid hooks found in some other onchidiids, can hardly be seen under a light microscope (in fact, because there are not many hooks, they are hard to find even with SEM). Our collections currently do not include any specimen from Samar, the type locality of O. samarense, but we did collect many species of onchidiids in Luzon, just next to Samar, as well as in Bohol, a bit further south in the Philippines. Given the anatomy of O. samarense (no rectal gland, digestive system of type II, male opening clearly on the left of the right eye tentacle, accessory penial gland and retractor muscle inserting near the heart), it is most likely that O. samarense applies to the same species as O. tumidum . Because its nomenclatural status still remains problematic (it could be regarded as a nomen dubium) and because its written description was published in 1882 and not in 1880, O. samarense is regarded as a junior synonym of O. tumidum .</p><p>Plate (1893) identified five specimens from Ponape (now Pohnpei, Micronesia) and one specimen from Singapore as Onchidium tumidum . Given that their intestinal loops were of type I, the specimens from Ponape were misidentified. It is impossible to determine whether the specimen from Singapore (with intestinal loops of type II) actually belongs to P. tumidus . Bretnall (1919) listed previous records but did not examine any new material. Bretnall also suggested that Onchidium punctatum Quoy &amp; Gaimard, 1832 could possibly refer to the same species as O. tumidum . However, O. punctatum clearly belongs to the genus Peronia . In fact, it was transferred to Scaphis Labbé, 1934 by Labbé (1934a: 203) as Scaphis punctata and Scaphis is a junior synonym of Peronia . Hoffmann (1928) mentioned O. tumidum, O. samarense and O. mertoni but did not examine any new material. Hoffmann also suggested that the record of Onchidium tabularis (Tapparone-Canefri, 1883), listed by Boettger (1923) from the Aru and Kei Islands, was a misidentification for O. mertoni (Boettger provided a record but did not describe any specimens). However, Hoffmann’s claim cannot be checked because the application of Onchidella tabularis Tapparone-Canefri, 1883 (as Oncidiella) is very unclear (the type material could not be located and the original description is not informative).</p><p>Three onchidiid sequences from mainland China were obtained from GenBank, the only ones that are not new in our data set (Table 1). These sequences were misidentified as Paraoncidium reevesii (J.E. Gray, 1850) . Paraoncidium Labbé, 1934 is not a valid name: it is a junior synonym of Onchidina Semper, 1882 . Also, Onchidium reevesi (J.E. Gray, 1850) is actually one of the three valid species of Onchidium (Dayrat et al. 2016) .</p></div>	https://treatment.plazi.org/id/F83E3A75F52EFFB683F6FE81867DF51D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai	Dayrat, Benoît, Goulding, Tricia C., Khalil, Munawar, Apte, Deepak, Bourke, Adam J., Comendador, Joseph, Tan, Shau Hwai (2019): A new genus and three new species of mangrove slugs from the Indo-West Pacific (Mollusca: Gastropoda: Euthyneura: Onchidiidae). European Journal of Taxonomy 500: 1-77, DOI: 10.5852/ejt.2019.500
F83E3A75F510FFAC83F4FA0C85BAF202.text	F83E3A75F510FFAC83F4FA0C85BAF202.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paromoionchis daemelii (Semper 1880) Dayrat & Goulding & Khalil & Apte & Bourke & Comendador & Tan 2019	<div><p>Paromoionchis daemelii (Semper, 1880) comb. nov.</p><p>Figs 26–30</p><p>Onchidium dämelii Semper, 1880: pl. 20, fig. 2.</p><p>Onchidium dämelii – Semper 1882: 270–271, pl. 21, fig. 9.</p><p>Material examined</p><p>Type material</p><p>AUSTRALIA • lectotype (here designated; 17/ 14 mm); New South Wales, Sydney; ZMB 31640a • 1 paralectotype (17/ 17 mm); same locality as lectotype; ZMB 31640b • 1 paralectotype (destroyed, dried); same locality as lectotype; ZMB 39035 • 2 paralectotypes (?); same locality as lectotype; ZMH 27476/2 .</p><p>Notes on type material</p><p>The lectotype, 17/ 14 mm, is designated here (ZMB 31640a). All other syntypes become paralectotypes. According to the original description, the type material included only three specimens. However, five possible syntypes could be located in museum collections, all from Sydney, Australia: 2 specimens, one of which, dissected with male parts remaining inside (17/ 14 mm), is designated as lectotype (ZMB 31640a) and the other one, still entire (17/ 17 mm), is a paralectotype (ZMB 31640b); 1 specimen destroyed, in pieces and completely dried (ZMB 39035); and 2 specimens (ZMH 27476/2), both entire. It is unclear exactly which specimens Semper used for the description, but it is safe to assume that the anatomical details he provided are based on the only two dissected specimens. Two species of Paromoionchis gen. nov. are present in Sydney, P. tumidus and the species described here, which are cryptic externally but distinct internally. Thus, the specimens that were not dissected by Semper could belong either to P. tumidus or to the species treated here. Hence the necessity of designating a lectotype in order to clarify the application of the name Onchidium daemelii .</p><p>Other material</p><p>AUSTRALIA – New South Wales • 1 spec. (37/25 [1511] mm); Sydney, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=151.20177&amp;materialsCitation.latitude=-33.7722" title="Search Plazi for locations around (long 151.20177/lat -33.7722)">Middle Harbour</a>, N of Roseville Bridge, W bank; 33°46.332´S, 151°12.106´E; 23 Nov. 2011; station 38; open mangrove, in old logs on the mud; AM C.468910.001 • 1 spec. (17/10 [1510] mm); same data as for preceding; AM C.468911.001 • 1 spec. (65/35 [1518] mm); Sydney, Pittwater, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=151.28938&amp;materialsCitation.latitude=-33.651783" title="Search Plazi for locations around (long 151.28938/lat -33.651783)">Church Point</a>, next to yacht club; 33°39.107´S, 151°17.363´E; 24 Nov. 2011; station 39; muddy sand next to small patch of mangrove and rocks on sandy beach; AM C.468913.001 • 1 spec. (60/35 [1519] mm); same data as for preceding; AM C.468917.001 • 1 spec. (28/18 [1515] mm); same data as for preceding; AM C.468914.001 • 1 spec. (52/28 [1514] mm); same data as for preceding; AM C.468912.002 • 1 spec. (40/20 [1512] mm); same data as for preceding; AM C.468912.003 • 1 spec. (50/25 [1521] mm); Sydney, Pittwater, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=151.3313&amp;materialsCitation.latitude=-33.62205" title="Search Plazi for locations around (long 151.3313/lat -33.62205)">Careel Bay</a>; 33°37.323´S, 151°19.878´E; 24 Nov. 2011; station 40; supratidal zone on the margin of salt marsh, mangrove patch on side of creek; AM C.468919.001 . – Queensland • 2 spec. (15/12 [1532] and 8/6 [1533] mm); Thirsty Sound, Plum Tree, beach in front of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.03093&amp;materialsCitation.latitude=-22.135733" title="Search Plazi for locations around (long 150.03093/lat -22.135733)">Endeavour Park</a>; 22°08.144´S, 150°01.856´E; 14 Sep. 2002; I. Loch, D.L. Beechey and A.C. Miller leg.; sheltered, muddy cobble shore; AM C.415270 • 1 spec. (8/6 [2668] mm); Bowen, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=148.23706&amp;materialsCitation.latitude=-20.024633" title="Search Plazi for locations around (long 148.23706/lat -20.024633)">Doughty Creek</a>; 20°01.478´S, 148°14.224´E; 3 Jul. 2013; station 119; mangrove of Rhizophora and Avicennia on one side of creek; MTQ .</p><p>Color and morphology of live animals (Fig. 26)</p><p>Live animals are often covered with mud, in which case their dorsal color can hardly be seen. The background of the dorsal notum is brown, occasionally mottled with darker or lighter areas. In addition, in some animals, the tip of dorsal papillae (with and without dorsal eyes) can be bright yellow. The foot is gray. The hyponotum is gray (same color as the foot), yellow, or both (yellow outer ring and gray inner ring). The color of the foot and of the hyponotum of an individual can change rapidly, especially when disturbed. The ocular tentacles are gray or brown, and may or may not be speckled with white dots, like the head. The ocular tentacles are short (just a few millimeters long).</p><p>Digestive system (Figs 27 A–B, 28)</p><p>Radulae measure up to 5.4 mm in length. Examples of radular formulae are presented in Table 4. Reproductive system (Figs 27 C–D, 29)</p><p>The male anterior organs consist of the penial complex (penis, penial sheath, vestibule, deferent duct, retractor muscle) and the accessory penial gland (flagellum and hollow spine). The hollow spine of the accessory penial gland is narrow, elongated, slightly curved. Its base is conical. Its diameter is approximately 80 μm for most of its length, except at its base (200 μm) and tip (60 μm). Its length ranges from 2.5 mm ([1519] AM C.468917.001) to 2.7 mm ([1521] AM C.468919.001), and its shape does vary between individuals (Fig. 29). The penial sheath is narrow and elongated. The retractor muscle is short (shorter than the penial sheath) or even vestigial (its distal end being free in the visceral cavity, with no clear insertion). The deferent duct is highly convoluted, with many loops. Inside the penial sheath, the penis is a narrow, elongated, soft, hollow tube of approximately 100 μm in diameter; internally, the penis bears a few smooth (no hooks) longitudinal ridges.</p><p>Distinctive diagnostic features</p><p>Externally, Paromoionchis daemelii cannot be distinguished from other species of Paromoionchis gen. nov. (Table 3). Also, its internal anatomy (accessory penial gland, vestigial penial retractor muscle, smooth penis) is very similar to that of P. boholensis gen. et sp. nov. The distribution range of P. daemelii overlaps with that of only one species of Paromoionchis gen. nov., P. tumidus (Fig. 6). Both species live in similar habitats and can even be found at the same station. They can only be distinguished internally thanks to a few anatomical details: in P. daemelii, the penis is smooth and the penial retractor muscle is very short or even vestigial, while in P. tumidus the penis bears some tiny hooks and the penial retractor muscle inserts near the heart (Table 3).</p><p>Distribution (Fig. 6)</p><p>Australia: New South Wales (type locality, present study), Queensland (present study).</p><p>Habitat (Fig. 30)</p><p>Paromoionchis daemelii is found on mud or muddy logs, inside or near mangroves, or on muddy sand. It is not common in central Queensland or New South Wales, but its abundance in southern Queensland is unknown.</p><p>Remarks</p><p>The publication dates of the various sections of the volume on Landmollusken by Carl Semper in the Reisen im Archipel der Philippinen series were clarified by Johnson (1969). The species name Onchidium daemelii was first published by Semper in 1880 with one figure (pl. 20, fig. 2) but no written description. Because Onchidium daemelii was published before 1931, ICZN Article 12.2.7 applies and the name is available (Semper’s figures are regarded as an indication accompanying the name Onchidium daemelii). Also, the specific name was originally spelled dämelii . However, according to ICZN Article 32.5.2.1., the correct spelling is daemelii . Both daemelii (e.g., Labbé 1934a) and damelii (e.g., Kenny &amp; Smith 1987, 1988) are spelling mistakes.</p><p>According to our current data, there are only two species of Paromoionchis gen. nov. in New South Wales (Fig. 6): P. tumidus and P. daemelii . They cannot be distinguished externally but they differ anatomically (Table 3). Both species are characterized by the lack of a rectal gland, a digestive system of type II, a male opening clearly to the left of the right eye tentacle (Semper described a male opening under the right eye tentacle, but it is distinctly below and left of it) and an accessory penial gland. The retractor muscle of the penis of O. daemelii, described as “very thin” by Semper, is vestigial in the lectotype, whereas the retractor muscle of P. tumidus is not vestigial and inserts near the heart. No “teeth” (the term he used to refer to penial hooks) are mentioned by Semper in the original description of O. daemelii, while the penis of P. tumidus bears some hooks. Therefore, the combination of characters found in the lectotype of O. daemelii and in Semper’s original description (retractor muscle vestigial and soft penis with no hooks) is only compatible with the species described here, not with P. tumidus, which justifies the present application of P. daemelii .</p><p>Onchidium daemelii was recorded from New South Wales (Lendenfeld 1886; Tenison-Woods 1888) and even New Guinea (Tapparone-Canefri 1883) but it is not possible to determine whether it was identified properly without re-examining the material which these authors examined (which may or may not have been deposited). Bretnall (1919), Hoffmann (1928) and Labbé (1934a) mentioned Onchidium daemelii without adding any new material. Finally, Kenny &amp; Smith (1987) published an ecological study on a species they identified as Onchidium damelii in a mangrove on Magnetic Island, Queensland. However, given that P. daemelii is rare in northern and central Queensland and that its identification requires detailed study of the internal anatomy, Kenny &amp; Smith likely studied P. tumidus rather than P. daemelii (or a mix of both species).</p></div>	https://treatment.plazi.org/id/F83E3A75F510FFAC83F4FA0C85BAF202	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai	Dayrat, Benoît, Goulding, Tricia C., Khalil, Munawar, Apte, Deepak, Bourke, Adam J., Comendador, Joseph, Tan, Shau Hwai (2019): A new genus and three new species of mangrove slugs from the Indo-West Pacific (Mollusca: Gastropoda: Euthyneura: Onchidiidae). European Journal of Taxonomy 500: 1-77, DOI: 10.5852/ejt.2019.500
F83E3A75F50AFFA783B8FDEC867BF3B0.text	F83E3A75F50AFFA783B8FDEC867BF3B0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paromoionchis boholensis Dayrat & Goulding & Khalil & Apte & Bourke & Comendador & Tan 2019	<div><p>Paromoionchis boholensis Dayrat &amp; Goulding gen. et sp. nov.</p><p>urn:lsid:zoobank.org:act: 104C35EF-FBF9-4EC2-A5A8-89AC30009270</p><p>Figs 31–40</p><p>Etymology</p><p>Paromoionchis boholensis gen. et sp. nov. is named after Bohol Island, where the type locality is.</p><p>Material examined</p><p>Holotype</p><p>PHILIPPINES • holotype (28/18 [3288] mm); Bohol, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=123.8498&amp;materialsCitation.latitude=9.727417" title="Search Plazi for locations around (long 123.8498/lat 9.727417)">Maribojoc</a>; 09°43.645´N, 123°50.988´E; 15 Jul. 2014; station 193; small island at end of boardwalk, sandy mud and rocks in back of mangrove; PNM 0 41266.</p><p>Other material</p><p>INDONESIA – Sulawesi • 2spec.(35/22[2128]and37/20[2129]mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=124.86217&amp;materialsCitation.latitude=1.6009166" title="Search Plazi for locations around (long 124.86217/lat 1.6009166)">Wori</a>; 01°36.055´N, 124°51.730´E; 9 Mar. 2013; station 84; tall mangrove forest of Sonneratia and Avicennia, with old logs; UMIZ 0 0 141 • 1 spec. (9/6 [2175] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=125.02053&amp;materialsCitation.latitude=1.7225833" title="Search Plazi for locations around (long 125.02053/lat 1.7225833)">Bahoi</a>; 01°43.355´N, 125°01.232´E; 10 Mar. 2013; station 85; sand, small rocks and pieces of wood, near narrow coastal mangrove; UMIZ 0 0 142 • 1 spec. (20/13 [2199] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=125.01328&amp;materialsCitation.latitude=1.6918833" title="Search Plazi for locations around (long 125.01328/lat 1.6918833)">Tamperong</a>; 01°41.513´N, 125°00.797´E; 12 Mar. 2013; station 87; muddy mangrove with small and dense Rhizophora; UMIZ 0 0 143 • 1 spec. (12/8 [2316] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=124.779015&amp;materialsCitation.latitude=1.698" title="Search Plazi for locations around (long 124.779015/lat 1.698)">Mantehang</a>; 01°41.880´N, 124°46.741´E; 15 Mar. 2013; station 91; Sonneratia at low intertidal and Rhizophora at high intertidal; UMIZ 0 0 144 • 1 spec. (20/13 [2360] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=119.593834&amp;materialsCitation.latitude=-4.362167" title="Search Plazi for locations around (long 119.593834/lat -4.362167)">Panikkiang Island</a>; 04°21.730´S, 119°35.630´E; 25 Mar. 2013; station 94; Rhizophora, Avicennia, Sonneratia and old logs; UMIZ 0 0 145. – Ambon • 3 spec. (40/25 [2849], 45/30 [2850] and 45/25 [2851] mm); Wai; 03°34.652´S, 128°19.526´E; 15 Feb. 2014; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.32544&amp;materialsCitation.latitude=-3.5775332" title="Search Plazi for locations around (long 128.32544/lat -3.5775332)">station 132</a>; narrow band of old Avicennia trees on sandy mud, old logs on ground; UMIZ 0 0 146. – Seram • 1 spec. (45/28 [2884] mm); Piru; 03°04.072´S, 128°11.362´E; 19 Feb. 2014; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.18936&amp;materialsCitation.latitude=-3.0678666" title="Search Plazi for locations around (long 128.18936/lat -3.0678666)">station 136</a>; Sonneratia mangrove next to fish market, next to beach of palms and ferns, with cattle roaming around; UMIZ 0 0 147. – Kei Islands • 2 spec. (10/8 [2896] and 17/8 [2901] mm); Un; 05°38.273´S, 132°45.738´E; 23 Feb. 2014; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.7623&amp;materialsCitation.latitude=-5.637883" title="Search Plazi for locations around (long 132.7623/lat -5.637883)">station 137</a>; Bruguiera and Rhizophora, some muddy areas and some with coral rubble; UMIZ 0 0 148 • 2 spec. (15/10 [2903] and 18/8 [2911] mm); Un; 05°38.273´S, 132°45.738´E; 25 Feb. 2014; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.7623&amp;materialsCitation.latitude=-5.637883" title="Search Plazi for locations around (long 132.7623/lat -5.637883)">station 140</a>; back of mangrove, on rocks, on mud, inside logs and under leaf litter; UMIZ 0 0 149 • 3 spec. (40/22 [3565], 30/20 [2935] and 40/22 [2937] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.75186&amp;materialsCitation.latitude=-5.599283" title="Search Plazi for locations around (long 132.75186/lat -5.599283)">Fiditan</a>; 05°35.957´S, 132°45.112´E; 28 Feb. 2014; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.75186&amp;materialsCitation.latitude=-5.599283" title="Search Plazi for locations around (long 132.75186/lat -5.599283)">station 144</a>; rocks behind muddy Rhizophora mangrove; UMIZ 0 0 150 . – Bali • 1 spec. (35/17 [3117] mm); Gilimanuk; 08°10.156´S, 114°26.652´E; 4 Apr. 2014; station 156; muddy mangrove with Rhizophora and Avicennia trees; UMIZ 0 0 140. – Halmahera • 1 spec. (25/16 [5019] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.59828&amp;materialsCitation.latitude=0.7578833" title="Search Plazi for locations around (long 127.59828/lat 0.7578833)">Sofifi</a>; 00°45.473´N, 127°35.897´E; 8 Mar. 2015; station 205; Sonneratia mangrove, with dense roots and hard mud; UMIZ 0 0 151 • 2 spec. (47/30 [5140] and 35/22 [5146] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.527084&amp;materialsCitation.latitude=1.4485166" title="Search Plazi for locations around (long 127.527084/lat 1.4485166)">Gamkonora</a>; 01°26.911´N, 127°31.625´E; 21 Mar. 2015; station 219; mostly Rhizophora mangrove with some sandy areas and some open muddy spaces; UMIZ 0 0 152 .</p><p>PHILIPPINES – Luzon • 1 spec. (25/18 [3609] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.62452&amp;materialsCitation.latitude=13.968833" title="Search Plazi for locations around (long 120.62452/lat 13.968833)">Lian</a>, Batangas; 13°58.130´N, 120°37.471´E; 5 Jul. 2014; station 179; narrow and impacted mangrove of Avicennia near village, very sandy, little to no mud; PNM 0 41267 . – Bohol • 2 spec. (16/9 [3283] and 17/10 [3619] mm); same data as for holotype; PNM 0 41268 • 3 spec. (20/15 [3369], 35/18 [3372] and 27/20 [3411] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=124.56925&amp;materialsCitation.latitude=9.859767" title="Search Plazi for locations around (long 124.56925/lat 9.859767)">Mabini</a>; 09°51.586´N, 124°34.155´E; 18 Jul. 2014; station 196; open Avicennia and Sonneratia forest with sand, algae and coral rubble; PNM 0 41269 • 5 spec. (30/22 [3412], 30/23 [3413], 40/17 [3417], 40/20 [3422] and 42/25 [3423] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=123.82315&amp;materialsCitation.latitude=9.738" title="Search Plazi for locations around (long 123.82315/lat 9.738)">Maribojoc</a>; 09°44.280´N, 123°49.389´E; 20 Jul. 2014; station 202; uplifted coral rubble with sand and algae, near Sonneratia trees; PNM 0 41270 .</p><p>Color and morphology of live animals (Figs 31–32)</p><p>Live animals are often covered with mud, in which case their dorsal color can hardly be seen. In unit #1, the background of the dorsal notum is brown, occasionally mottled with darker or lighter areas, while in unit #2 it ranges from very light brown (almost white) to dark brown, mottled or not. In some animals, there is a reddish hue on the margin of the dorsal notum (unit #1). In addition, in most animals the tip of the dorsal papillae (with and without dorsal eyes) can be bright yellow. The foot is orange (unit #1) or varies from gray to yellow and orange (unit #2). The hyponotum is also orange, often with a darker ring on the margin which may be bright red (unit #1) or homogenously gray, yellow, or orange, but can also display a mix of two or three of those colors (unit #2). The color of both the foot and the hyponotum of an individual can change very rapidly, especially when disturbed. The ocular tentacles are reddish brown and may or may not be speckled with white dots, like the head. The ocular tentacles are short (just a few millimeters long).</p><p>Digestive system (Figs 33A, 34A, 35–36)</p><p>Radulae measure up to 4.5 mm (unit #1) and 4 mm (unit #2) in length. Examples of radular formulae are presented in Table 4.</p><p>Reproductive system (Figs 33 B–C, 34B–C, 37–38)</p><p>The male anterior organs consist of the penial complex (penis, penial sheath, vestibule, deferent duct, retractor muscle) and the accessory penial gland (flagellum and hollow spine). The hollow spine of the accessory penial gland is narrow, elongated, slightly curved. Its base is conical. Its diameter is approximately 50 to 70 μm for most of its length and 100–130 μm at its base (unit #1) and approximately 70 to 80 μm for most of its length and 150–200 μm at its base (unit #2). Its length ranges from 1 mm ([3288] PNM 0 41266, holotype) to 1.2 mm ([3372] PNM 041269) in unit #1 and from 1.1 mm ([5019] UMIZ 00151) and 1.3 mm ([3117] UMIZ 00140) to 1.8 mm ([2911] UMIZ 0 0 149, [5140] UMIZ 00152) in unit #2, and its shape does vary between individuals (Fig. 38). The penial sheath is narrow and elongated. The retractor muscle is vestigial, i.e., with its distal end being free in the visceral cavity, with no clear insertion (unit #1), or absent or vestigial (unit #2). The deferent duct is highly convoluted, with many loops. Inside the penial sheath, the penis is a narrow, elongated, soft, smooth (no hooks) and hollow tube of approximately 200 μm in diameter.</p><p>Distinctive diagnostic features</p><p>Externally, the color of the foot and hyponotum can help one to identify Paromoionchis boholensis gen. et sp. nov., but unfortunately it is not fully reliable. Specimens with a bright orange foot and hyponotum are only found in P. boholensis gen. et sp. nov., especially in unit #1 but also in unit #2; the ventral side of P. tumidus, which is sympatric with P. boholensis (Fig. 6), can be orange but not bright orange. However, specimens with a more yellowish or greyish foot and hyponotum cannot be identified externally. The internal anatomy of P. boholensis gen. et sp. nov. (accessory penial gland, vestigial penial retractor muscle, penis with no hooks) is similar to that of P. daemelii . However, P. boholensis gen. et sp. nov. and P. daemelii do not overlap geographically, at least based on the present data. Thus, within its distribution range P. boholensis gen. et sp. nov. is the only species with this combination of internal characters. Indeed, the internal characters of the two species of Paromoionchis gen. nov. with which P. boholensis gen. et sp. nov. is sympatric ( P. goslineri gen. et sp. nov. and P. tumidus) are different (Table 3). It must be noted that the known distribution of species of Paromoionchis gen. nov. may change as new records are found in the future and so the use of geographic data should only be used with caution for identification.</p><p>Distribution (Fig. 6)</p><p>Philippines (unit #1): Bohol (type locality), Luzon. Indonesia (unit #2): Ambon, Bali, Halmahera, Kei Islands, Seram, Sulawesi.</p><p>Habitat (Figs 39–40)</p><p>Unit #1 of Paromoionchis boholensis gen. et sp. nov. is found on sandy mud or sand with very little mud, in mangroves or near mangrove trees and is rare (it was found at only four stations). Unit #2 is found in open or dense mangroves, on soft or hard mud, as well as on muddy sand and is common (but not as common as P. tumidus unit #1).</p></div>	https://treatment.plazi.org/id/F83E3A75F50AFFA783B8FDEC867BF3B0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai	Dayrat, Benoît, Goulding, Tricia C., Khalil, Munawar, Apte, Deepak, Bourke, Adam J., Comendador, Joseph, Tan, Shau Hwai (2019): A new genus and three new species of mangrove slugs from the Indo-West Pacific (Mollusca: Gastropoda: Euthyneura: Onchidiidae). European Journal of Taxonomy 500: 1-77, DOI: 10.5852/ejt.2019.500
F83E3A75F501FFA2838EFB93873FF213.text	F83E3A75F501FFA2838EFB93873FF213.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paromoionchis penangensis Dayrat & Goulding & Khalil & Apte & Bourke & Comendador & Tan 2019	<div><p>Paromoionchis penangensis Dayrat &amp; Goulding gen. et sp. nov.</p><p>urn:lsid:zoobank.org:act: 840ABD3E-61AC-4B1E-9D83-B15A8099F0BA</p><p>Figs 41–44</p><p>Etymology</p><p>Paromoionchis penangensis gen. et sp. nov. is named after Penang Island, Malaysia, in the Strait of Malacca, which is the type locality.</p><p>Material examined</p><p>Holotype</p><p>MALAYSIA • holotype (26/14 [6037] mm); Peninsular Malaysia, Penang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.19285&amp;materialsCitation.latitude=5.4153666" title="Search Plazi for locations around (long 100.19285/lat 5.4153666)">Pantai Acheh</a>; 05°24.922´N, 100°11.571´E; 1 Aug. 2016; station 261; Avicennia mangrove, with both very soft mud and hard mud; USMMC 00059.</p><p>Other material</p><p>INDIA – Andaman Islands • 1 spec. (18/10 [1086] mm); Middle Andaman, Rangat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=92.89653&amp;materialsCitation.latitude=12.457517" title="Search Plazi for locations around (long 92.89653/lat 12.457517)">Yerrata</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=92.89653&amp;materialsCitation.latitude=12.457517" title="Search Plazi for locations around (long 92.89653/lat 12.457517)">Saban</a>; 12°27.451´N, 092°53.792´E; 10 Jan. 2011; station 56; open, impacted mangrove patch by a creek, near village, with medium trees and old logs; BNHS 92 • 2 spec. (18/8 [1100] and 9/6 [1101] mm); Middle Andaman, Rangat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=92.84396&amp;materialsCitation.latitude=12.48255" title="Search Plazi for locations around (long 92.84396/lat 12.48255)">Shyamkund</a>; 12°28.953´N, 092°50.638´E; 11 Jan. 2011; station 57; by a large river, deep mangrove with tall trees, small creeks and plenty of old logs, next to a road and a small cemented bridge over a creek; BNHS 53 • 2 spec. (20/10 [1117] and 22/12 [1118] mm); Middle Andaman, Shantipur, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=92.77295&amp;materialsCitation.latitude=12.330717" title="Search Plazi for locations around (long 92.77295/lat 12.330717)">Kadamtala</a>; 12°19.843´N, 092°46.377´E; 12 Jan. 2011; station 58; open area with hard mud and many old logs, next to a mangrove with medium trees; BNHS 11 • 2 spec. (30/15 [1129] and 12/7 [1130] mm); South Andaman, Bamboo Flat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=92.70962&amp;materialsCitation.latitude=11.792183" title="Search Plazi for locations around (long 92.70962/lat 11.792183)">Shoal Bay</a>; 11°47.531´N, 092°42.577´E; 13 Jan. 2011; station 59; open mangrove with medium trees, hard mud, dead logs, next to a road and a small cemented bridge for creek; BNHS 4. – Maharashtra • 1 spec. (35/20 [1167] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=73.29372&amp;materialsCitation.latitude=17.263184" title="Search Plazi for locations around (long 73.29372/lat 17.263184)">Watad</a>; 17°15.791´N, 73°17.623´E; 23 Dec. 2011; station 46; Avicennia mangrove, by field, with deep and very watery mud; BNHS 46 • 3 spec. (30/18 [1177], 20/14 [1175] and 15/10 [1173] mm); same data as for preceding; BNHS 98 • 2 spec. (16/11 [1176] and 27/21 [1182] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=73.295784&amp;materialsCitation.latitude=17.0734" title="Search Plazi for locations around (long 73.295784/lat 17.0734)">Aare Ware</a>; 17°04.404´N, 73°17.747´E; 24 Dec. 2011; station 47; mangrove with soft mud and some areas with pools, mostly Avicennia with a few small Rhizophora; BNHS 42 .</p><p>MALAYSIA – Peninsular Malaysia • 2 spec. (30/20 [5990] and 30/20 [5991] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.448616&amp;materialsCitation.latitude=4.933183" title="Search Plazi for locations around (long 100.448616/lat 4.933183)">Kuala Gula</a>; 04°55.991´N, 100°26.917´E; 29 Jul. 2016; station 259; mostly Avicennia, a few Bruguiera and Rhizophora, along a creek, both soft and hard mud; USMMC 0 0 0 60 • 2 spec. (20/14 [957] and 15/10 [958] mm); Nibong Tebal, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.42606&amp;materialsCitation.latitude=5.208133" title="Search Plazi for locations around (long 100.42606/lat 5.208133)">Pulau Burung</a>; 05°12.488´N, 100°25.564´E; 11 Jul. 2011; station 17; soft mud, open mangrove of Rhizophora, with a few Sonneratia; USMMC 0 0 0 61 • 1 spec. (48/35 [6020] mm); Nibong Tebal, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.42606&amp;materialsCitation.latitude=5.208133" title="Search Plazi for locations around (long 100.42606/lat 5.208133)">Pulau Burung</a>; 05°12.488´N, 100°25.564´E; 30 Jul. 2016; station 260; soft mud, open mangrove of Rhizophora, with a few Sonneratia; USMMC 0 0 0 62 • 3 spec. (25/12 [6031], 25/18 [6033] and 25/16 [6039] mm); same data as for holotype; USMMC 0 0 0 63 .</p><p>Color and morphology of live animals (Fig. 41)</p><p>Live animals are most often covered with mud, in which case their dorsal color can hardly be seen. The background of the dorsal notum is brown, occasionally mottled with darker or lighter areas. In addition, in some animals, the tip of dorsal papillae (with and without dorsal eyes) can be yellow. The foot is gray, occasionally with a light yellow hue. The hyponotum is uniform gray or gray (inner ring) and yellow (outer ring). The color of both the foot and the hyponotum of an individual can change rapidly, especially when disturbed. The ocular tentacles are brown and may or may not be speckled with white dots, like the head. The ocular tentacles are short (just a few mm long).</p><p>Digestive system (Figs 42A, 43)</p><p>Radulae measure up to 3.2 mm in length. Examples of radular formulae are presented in Table 4. Reproductive system (Fig. 42 B–C)</p><p>In the posterior (female) organs, the distal portion of the oviduct and of the duct to the spermatheca is wider than in other species, which makes sense given the wide penis. The male anterior organs consist of the penial complex (penis, penial sheath, vestibule, deferent duct, retractor muscle). An accessory penial gland is absent. The penial sheath is large (at least ten times as large as the deferent duct). The retractor muscle is strong, long and inserts near the heart. The deferent duct is convoluted, with many loops. Inside the penial sheath, the penis is a large (wider than long), smooth (no hooks), muscular mass.</p><p>Distinctive diagnostic features</p><p>Externally, Paromoionchis penangensis gen. et sp. nov. cannot be reliably distinguished from other species of Paromoionchis gen. nov. Its distribution only overlaps with that of P. tumidus . Our data suggest that the tips of the dorsal papillae of P. penangensis gen. et sp. nov. tend to be paler yellow, while they tend to be brighter yellow in P. tumidus . However, the internal anatomy of P. penangensis gen. et sp. nov., especially the large penis inside the large penial sheath, is very distinct from that of all other species and reliably distinguishes it from P. tumidus .</p><p>Distribution (Fig. 6)</p><p>Malaysia: Peninsular Malaysia, Strait of Malacca (type locality). India: Andaman Islands (Bay of Bengal), Maharashtra (W coast of India).</p><p>Habitat (Fig. 44)</p><p>Paromoionchis penangensis gen. et sp. nov. is found on soft and hard mud, in mangroves or in open areas near mangroves. This species was only found at three stations in the Strait of Malacca, three stations in the Andaman Islands (Bay of Bengal), and three stations in Maharashtra (W coast of India). However, at each station it was found to be quite abundant.</p></div>	https://treatment.plazi.org/id/F83E3A75F501FFA2838EFB93873FF213	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai	Dayrat, Benoît, Goulding, Tricia C., Khalil, Munawar, Apte, Deepak, Bourke, Adam J., Comendador, Joseph, Tan, Shau Hwai (2019): A new genus and three new species of mangrove slugs from the Indo-West Pacific (Mollusca: Gastropoda: Euthyneura: Onchidiidae). European Journal of Taxonomy 500: 1-77, DOI: 10.5852/ejt.2019.500
F83E3A75F504FFDF83A4FDF28593F3CB.text	F83E3A75F504FFDF83A4FDF28593F3CB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paromoionchis goslineri Dayrat & Goulding & Khalil & Apte & Bourke & Comendador & Tan 2019	<div><p>Paromoionchis goslineri Dayrat &amp; Goulding gen. et sp. nov.</p><p>urn:lsid:zoobank.org:act: E375A628-4AE6-4E10-B424-D4BBF58F8941</p><p>Figs 45–51</p><p>Etymology</p><p>Paromoionchis goslineri gen. et sp. nov. is dedicated to Dr. Terry Gosliner, Senior Curator at the California Academy of Sciences, San Francisco, California, USA, who has been exploring the marine life of the Batangas region for many years, where this new species was found, and, more importantly, for years ago providing the first author with a great post-doctoral opportunity to focus on alpha-taxonomy.</p><p>Material examined</p><p>Holotype</p><p>PHILIPPINES • holotype (25/15 [3233] mm); Luzon, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.62305&amp;materialsCitation.latitude=13.8544" title="Search Plazi for locations around (long 120.62305/lat 13.8544)">Calantagan</a>, Batangas; 13°51.264´N, 120°37.383´E; 8 Jul. 2014; station 185; next to village, impacted narrow mangrove forest of Avicennia by the shore; PNM 0 41271.</p><p>Other material</p><p>INDONESIA – Sulawesi • 1 spec. (20/14 [2210] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=125.02053&amp;materialsCitation.latitude=1.7225833" title="Search Plazi for locations around (long 125.02053/lat 1.7225833)">Bahoi</a>; 01°43.355´N, 125°01.232´E; 12 Mar. 2013; station 88; sand, small rocks and pieces of wood outside narrow coastal mangrove; UMIZ 0 0 161 • 1 spec. (18/10 [2241] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=124.543236&amp;materialsCitation.latitude=1.36295" title="Search Plazi for locations around (long 124.543236/lat 1.36295)">Sondaken</a>; 01°21.777´N, 124°32.594´E; 13 Mar. 2013; station 89; mostly Rhizophora, with sand, small rocks and pieces of wood outside narrow coastal mangrove; UMIZ 0 0 153 . – Bali • 3 spec. (22/15 [3060], 22/15 [3066] and 27/20 [3068] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.18005&amp;materialsCitation.latitude=-8.768766" title="Search Plazi for locations around (long 115.18005/lat -8.768766)">Denpasar</a>; 08°46.126´S, 115°10.803´E; 2 Apr. 2014; station 154; large mangrove by road, with shallow mud; UMIZ 0 0 154 • 4 spec. (20/12 [3072], 17/10 [3074], 22/14 [3078] and 24/14 [3079] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.443436&amp;materialsCitation.latitude=-8.170983" title="Search Plazi for locations around (long 114.443436/lat -8.170983)">Gilimanuk</a>; 08°10.259´S, 114°26.606´E; 3 Apr. 2014; station 155; from high intertidal with water pools and many mounds up to shore with sand and rocks; UMIZ 0 0 155 • 2 spec. (22/10 [3118] and 37/20 [3120] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.4442&amp;materialsCitation.latitude=-8.169267" title="Search Plazi for locations around (long 114.4442/lat -8.169267)">Gilimanuk</a>; 08°10.156´S, 114°26.652´E; 4 Apr. 2014; station 156; muddy mangrove with Rhizophora and Avicennia trees; UMIZ 0 0 156. – Timor • 2 spec. (12/7 [5890] and 15/7 [5891] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=123.63493&amp;materialsCitation.latitude=-10.145534" title="Search Plazi for locations around (long 123.63493/lat -10.145534)">Oesapa</a>; 10°08.732´S, 123°38.096´E; 11 Jul. 2016; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=123.63493&amp;materialsCitation.latitude=-10.145534" title="Search Plazi for locations around (long 123.63493/lat -10.145534)">station 250</a>; sandy part of mangrove, with Sonneratia and Avicennia; UMIZ 0 0 157. – Halmahera • 2 spec. (24/16 [5072] and 32/23 [5073] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.64173&amp;materialsCitation.latitude=0.8558" title="Search Plazi for locations around (long 127.64173/lat 0.8558)">Dodinga</a>; 00°51.348´N, 127°38.504´E; 9 Mar. 2015; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.64173&amp;materialsCitation.latitude=0.8558" title="Search Plazi for locations around (long 127.64173/lat 0.8558)">station 206</a>; back of mangrove, high intertidal, with ferns and mounds; UMIZ 0 0 158 • 1 spec. (35/22 [5145] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.527084&amp;materialsCitation.latitude=1.4485166" title="Search Plazi for locations around (long 127.527084/lat 1.4485166)">Gamkonora</a>; 01°26.911´N, 127°31.625´E; 21 Mar. 2015; station 219; mostly Rhizophora, with some sandy and open muddy areas; UMIZ 0 0 159. – Ambon • 1 spec. (12/7 [3555] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.32544&amp;materialsCitation.latitude=-3.5775332" title="Search Plazi for locations around (long 128.32544/lat -3.5775332)">Wai</a>; 03°34.652´S, 128°19.526´E; 15 Feb. 2014; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.32544&amp;materialsCitation.latitude=-3.5775332" title="Search Plazi for locations around (long 128.32544/lat -3.5775332)">station 132</a>; narrow band of old Avicennia trees on sandy mud, with old logs; UMIZ 0 0 160 .</p><p>PHILIPPINES – Luzon • 2 spec. (22/15 [3221] and 20/16 [6049] mm); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.61874&amp;materialsCitation.latitude=13.887967" title="Search Plazi for locations around (long 120.61874/lat 13.887967)">Calantagan</a>, Batangas; 13°53.278´N, 120°37.124´E; 8 Jul. 2014; station 184; narrow forest on the shore, with Avicennia and young Rhizophora; PNM 0 41272 • 1 spec. (28/20 [3232] mm); same data as for holotype; PNM 0 41273 .</p><p>Color and morphology of live animals (Figs 45–46)</p><p>Live animals are most often covered with mud, in which case their dorsal color can hardly be seen. The background of the dorsal notum is gray-brown, mottled with darker and lighter areas. In addition, in some animals, the tip of dorsal papillae (with and without dorsal eyes) can be lighter (pale yellow or white). The foot and the hyponotum are dark or light gray. The color of both the foot and the hyponotum of an individual can change rapidly, especially when disturbed. The ocular tentacles are gray-brown and may or may not be speckled with white dots, like the head. The ocular tentacles are short (just a few millimeters long). The tip of dorsal papillae is usually white or pale yellow, but not always (in any case generally covered with mud).</p><p>Digestive system (Figs 47A, 48A, 49–50)</p><p>Radulae measure up to 2.6 mm (unit #1) and 2.2 mm (unit #2) in length. Examples of radular formulae are presented in Table 4.</p><p>Reproductive system (Figs 47 B–C, 48B–C)</p><p>The male anterior organs consist of the penial complex (penis, penial sheath, vestibule, deferent duct, retractor muscle). An accessory penial gland is absent. The penial sheath is narrow and elongated. In unit #1, the retractor muscle is very short (much shorter than the penial sheath), inserting on the body wall near the nervous system, or vestigial (its distal end being free in the visceral cavity, with no clear insertion). In unit #2, the retractor muscle is long (as long as the penial sheath), inserting near the heart. The deferent duct is also highly convoluted, with many loops. Inside the penial sheath, the penis is a narrow, elongated, soft, smooth (no hooks) and hollow tube of approximately 200 μm in diameter.</p><p>Distinctive diagnostic features</p><p>Externally, Paromoionchis goslineri gen. et sp. nov. cannot be distinguished from other species of Paromoionchis gen. nov. The ventral side (foot and hyponotum) is gray, i.e., never yellow or orange. Unfortunately, a gray ventral side can occasionally be found in all other species of the genus, so the use of that color trait is not fully reliable for identification. However, the internal anatomy of P. goslineri gen. et sp. nov. (no accessory penial gland, thin penis with no hooks) is very distinctive and can be used for a fully reliable identification. The only other species of Paromoionchis gen. nov. without an accessory penial gland, P. penangensis gen. et sp. nov., differs greatly from P. goslineri gen. et sp. nov. anatomically because its penis is very large. Units #1 and #2 of P. goslineri gen. et sp. nov. differ slightly with respect to the penial retractor: it is short and thin, inserting near the nervous system, or even vestigial in unit #1, while it is as long as the penial sheath, inserting near the heart in unit #2. However, given that only four specimens could be dissected in unit #1, it is very possible that intermediates may be found in the future, especially considering that units #1 and #2 are widely distant geographically.</p><p>Distribution (Fig. 6)</p><p>Philippines (unit #1): Luzon (type locality). Indonesia (unit #2): Ambon, Bali, Halmahera, Sulawesi and Timor.</p><p>Habitat (Fig. 51)</p><p>Paromoionchis goslineri gen. et sp. nov. unit #1 is found on mud, in Avicennia forests near the shore and is rare (only four specimens are known from two stations). Unit #2 is found on soft and hard mud, in mangroves or in open areas near mangroves and is rare (except in Bali, where several specimens were found at a few stations).</p></div>	https://treatment.plazi.org/id/F83E3A75F504FFDF83A4FDF28593F3CB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai	Dayrat, Benoît, Goulding, Tricia C., Khalil, Munawar, Apte, Deepak, Bourke, Adam J., Comendador, Joseph, Tan, Shau Hwai (2019): A new genus and three new species of mangrove slugs from the Indo-West Pacific (Mollusca: Gastropoda: Euthyneura: Onchidiidae). European Journal of Taxonomy 500: 1-77, DOI: 10.5852/ejt.2019.500
F83E3A75F572FFD482ADFE58813CF466.text	F83E3A75F572FFD482ADFE58813CF466.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paromoionchis Dayrat & Goulding & Khalil & Apte & Bourke & Comendador & Tan 2019	<div><p>Identification key</p><p>A key is provided here to help identify the five species of Paromoionchis gen. nov. Because species cannot be distinguished externally, the key is based on internal characters of reproductively mature specimens.</p><p>1. Accessory penial gland absent.........................................................................................................2</p><p>– Accessory penial gland present.........................................................................................................3</p><p>2. Penis large, within large penial sheath ..... P. penangensis gen. et sp. nov. (W India to Malacca Strait)</p><p>– Penis thin, within thin penial sheath ........ P. goslineri gen. et sp. nov. (Indonesia and Philippines)</p><p>3. Penial retractor muscle reaches heart; thin penis with hooks ............................................................... .... P. tumidus (Semper, 1880) (Andaman Islands to subtropical waters of SE Australia and S Japan)</p><p>– Penial retractor muscle very short, vestigial, or absent; thin penis with no hooks............................4</p><p>4. Accessory penial gland spine &lt;1.8 mm in length .............................................................................. .................................................................. P. boholensis gen. et sp. nov. (Indonesia and Philippines)</p><p>– Accessory penial gland spine&gt; 2.5 mm in length ............ P. daemelii gen. et sp. nov. (SE Australia)</p></div>	https://treatment.plazi.org/id/F83E3A75F572FFD482ADFE58813CF466	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Dayrat, Benoît;Goulding, Tricia C.;Khalil, Munawar;Apte, Deepak;Bourke, Adam J.;Comendador, Joseph;Tan, Shau Hwai	Dayrat, Benoît, Goulding, Tricia C., Khalil, Munawar, Apte, Deepak, Bourke, Adam J., Comendador, Joseph, Tan, Shau Hwai (2019): A new genus and three new species of mangrove slugs from the Indo-West Pacific (Mollusca: Gastropoda: Euthyneura: Onchidiidae). European Journal of Taxonomy 500: 1-77, DOI: 10.5852/ejt.2019.500
