taxonID	type	description	language	source
03DC0D07BD59553AB6C4E88B6D7E5382.taxon	description	Fig. 3	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
03DC0D07BD59553AB6C4E88B6D7E5382.taxon	diagnosis	Diagnosis. This species is similar to L. chiriquiensis, L. hispida Thomson & Holzenthal, 2015, and L. viridis, three species that also occur in Panama. The phallus of all four species has a similar appearance, due to the elongate basal supports of the midlength complex and the small pair of membranous, apical lobes (Fig. 3 E, F). Leucotrichia botosaneanui can be distinguished by the small, double-pointed mesoventral process on sternum VII, which is longer and much more prominent in the other species.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
03DC0D07BD59553AB6C4E88B6D7E5382.taxon	materials_examined	Material examined. Panama: Chiriqui Province • 1 male; Cuenca 108; Boquete District; Quebrada Jaramillo; Finca Monterey; 8.7632 ° N, 82.41383 ° W; 1,214 m a. s. l.; 19 - 25 Apr. 2018; K. Collier, leg.; Malaise trap; in alcohol; MUPADI.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
03DC0D07BD59553AB6C4E88B6D7E5382.taxon	distribution	Distribution. Panama, Trinidad, Tobago.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
A77EC3F65FEF538A8C505098568919EF.taxon	description	Figs 2 B, 4	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
A77EC3F65FEF538A8C505098568919EF.taxon	diagnosis	Diagnosis. Leucotrichia chiriquiensis is most similar to L. botosaneanui, L. hispida, and L. viridis, based on a similar appearance of the phallus, as discussed under L. botosaneanui. This species can most swiftly be identified as distinct from the others based on the structural modifications of the antennae and head.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
A77EC3F65FEF538A8C505098568919EF.taxon	materials_examined	Material examined. Panama: Chiriqui Province • 2 males; Cuenca 102; Renacimiento District; La Amistad International Park, Rio Candela, Finca Felix, PSPSCB-PILA-C 102 - 2017 - 021; 8.890557 ° N, 82.61201 ° W; 2,128 m a. s. l.; 25 Jan. 2015; C. Nieto, E. Perez, A. Cornejo, leg.; UV light trap; in alcohol; COZEM • ibid., 2 males; Tierras Altas District; Mount Totumas Cloud Forest and Biological Reserve, Quebrada Norte; 8.873613 ° N, 82.690512 ° W; 1,709 m a. s. l.; 26 Apr. - 10 May 2015; B. Armitage, T. Arefina-Armitage, leg.; Malaise trap; MUPADI • ibid., 5 males; 28 Jan. - 2 Feb. 2018; J. Dietrich, leg.; Malaise trap; MUPADI • ibid., 3 males; 16 - 20 Feb. 2018; MUPADI • ibid., 1 male; 16 - 20 Mar. 2018; MUPADI • ibid., 1 male; 10 - 15 Jul. 2018; MUPADI • ibid., 2 males; 9 - 12 Sep. 2018; MUPADI • ibid., 2 males; 8 - 11 Nov. 2018; UMSP.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
A77EC3F65FEF538A8C505098568919EF.taxon	distribution	Distribution. Panama.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
BBACFDAB10EA57FEBEDA93ACF0CE36E6.taxon	description	Fig. 5	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
BBACFDAB10EA57FEBEDA93ACF0CE36E6.taxon	diagnosis	Diagnosis. Leucotrichia cortadera sp. nov. is similar to L. fulminea Thomson & Holzenthal, 2015, a species endemic to Ecuador. Both species bear a pair of large, distinct sclerotized plates on the phallus apex. Leucotrichia cortadera can be separated by the small spines present on the dorsolateral surface of the phallus apex, which are absent in L. fulminea. Additionally, the inferior appendages are separate in L. fulminea, while they are fused in L. cortadera.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
BBACFDAB10EA57FEBEDA93ACF0CE36E6.taxon	description	Description. Male. Length of forewing 2.1 mm (n = 2). Wings unmodified. Head unmodified, with three ocelli; antennae unmodified. Tibial spur count 1, 3, 4. Color in alcohol brown. Genitalia. Abdominal sternum VII mesoventral process with enlarged apex (Fig. 5 A, D). Sternum VIII with rounded posteroventral production in lateral view (Fig. 5 A); in ventral view, posterior margin concave with broadly rounded mesal projection (Fig. 5 D). Segment IX anterolateral margin convex, posterolateral margin straight with slight irregularity (Fig. 5 B); dorsally, anterior margin concave, posterior margin concave with broadly rounded mesal projection (Fig. 5 C). Tergum X with dorsal sclerite small, irregular; ventral sclerite with upper half rounded and bent posteriad; membranous apex suborbicular (Fig. 5 B, C). Subgenital plate with dorsal arm simple, extending dorsad, apex acute (Fig. 5 B); ventral arm simple, apex with emargination, in ventral view slightly restricted mesally, apex with broad emargination (Fig. 5 B, D). Inferior appendage narrow basally, broadest submesally, with single dorsal subapical spine (Fig. 5 B); ventrally entirely fused, broadest mesally, apex rounded with small pointed mesal emargination (Fig. 5 D). Phallus tubular basally, constricted at midlength with median complex bearing spherical " windows "; apex membranous and bearing pair of large, acute sclerotized plates and numerous small apical spines on dorsal and lateral surface (Fig. 5 E, F).	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
BBACFDAB10EA57FEBEDA93ACF0CE36E6.taxon	distribution	Distribution. Panama.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
BBACFDAB10EA57FEBEDA93ACF0CE36E6.taxon	etymology	Etymology. The specific epithet is derived from cortadera, Spanish for " knife, cutting instrument ", referring to the shape of the large sclerotized plates found on the phallus apex.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
00FA4B0400A95D16AFEB75B1FB054561.taxon	description	Fig. 6	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
00FA4B0400A95D16AFEB75B1FB054561.taxon	diagnosis	Diagnosis. Leucotrichia cultrata is similar to L. hispida and L. viridis. The phallus of all three species shares a similar appearance with the basal loop of the median complex extended on basal supports and a ventral " bulge " to the membranous apex. The mesoventral process of sternum VII can be used to separate the three, being long and digitate in L. cultrata, bearing a tuft of prominent apical setae in L. hispida, and enlarged and apically blunt when viewed ventrally in L. viridis.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
00FA4B0400A95D16AFEB75B1FB054561.taxon	materials_examined	Material examined. Panama: Bocas del Toro Province • 2 males; Cuenca 093; Chiriqui Grande District; Quebrada Rambala, Rambala Jungle Lodge, 8.91627 ° N, 82.15469 ° W; 120 m a. s. l.; 9 Aug. 2014; E. Carlson, leg.; UV light trap; in alcohol; MUPADI • ibid., 5 males; 28 Mar. 2015 • ibid., 2 males; 31 Mar. - 11 Apr. 2015; Malaise trap • ibid., 17 males; 12 - 15 Nov. 2017. Chiriqui Province • 1 male; Cuenca 102; Tierras Altas District; Mount Totumas Cloud Forest and Biological Reserve, Quebrada Norte; 8.873613 ° N, 82.690512 ° W; 1,709 m a. s. l.; 10 - 15 Jul. 2018; J. Dietrich, leg.; Malaise trap; in alcohol; MIUP • ibid., 2 males; 8 - 11 Nov. 2018; UMSP • ibid., 1 male; Cuenca 108; Dolega District, Rio Majagua, Banquito de Palmira, Potrerillos; 8.68083 ° N, 82.53250 ° W; 840 m a. s. l.; 28 Feb. - 14 Mar. 2019, Y. Aguirre, T. Rios, leg.; Malaise trap (M 001); in alcohol; MUPADI • ibid., 1 male; Quebrada Jaramillo, Finca Monterey; 8.76320 ° N, 82.41383 ° W; 1,214 m a. s. l.; 8 - 12 May 2018; B. Armitage, T. Arefina-Armitage, leg.; Malaise trap; in alcohol; MUPADI. Panama Oeste Province • 2 males; Cuenca 115; Altos de Campana National Park, Rio Chileno, PSPSCB-PNAC-C 115 - 2018 - 028; 8.71650 ° N, 80.00740 ° W; 497 m a. s. l.; 23 - 31 May 2018; T. Rios, Y. Aguirre, leg.; Malaise trap; in alcohol; COZEM. Veraguas Province • 2 males; Cuenca 097; Santa Fe District; Santa Fe National Park; Rio Calovebora, PSPSCB-PNSF-C 097 - 2017 - 006; 8.55038 ° N, 81.16486 ° W; 461 m a. s. l.; 23 - 27 Apr. 2017, A. Cornejo, T. Rios, E. Alvarez, C. Nieto, leg.; Malaise Trap; in alcohol; COZEM • ibid., 1 male; Cuenca 132; Rio Mulaba, afl. 1 er Brazo, PSPSCB-PNSF-C 132 - 2017 - 008; 8.51706 ° N, 81.12140 ° W; 770 m a. s. l.; 19 - 23 Apr. 2017; T. Rios, E. Alvarez, C. Nieto, leg.; Malaise trap; in alcohol; COZEM.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
00FA4B0400A95D16AFEB75B1FB054561.taxon	distribution	Distribution. Panama.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
436018A884B556C6BF3F5E25D504862B.taxon	description	Fig. 7	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
436018A884B556C6BF3F5E25D504862B.taxon	diagnosis	Diagnosis. This species is similar to L. dianeae Thomson & Holzenthal, 2015 and L. tapantia Thomson & Holzenthal, 2015, two species originally described from Costa Rica that could potentially be collected in Panama (Table 4). In all three species, the posterolateral margin of sternum VIII is notably produced. Additionally, all three share a similar shape to the phallus apex, with bears a pair of apical lobes and lacks any spines or externally sclerotized structures. Leucotrichia extraordinaria can be easily separated by the single peg-like apical seta on the posterolateral production of sternum VIII, a feature not shared with the other two species.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
436018A884B556C6BF3F5E25D504862B.taxon	materials_examined	Material examined. Panama: Bocas del Toro Province • 59 males; Cuenca 093; Chiriqui Grande District; Quebrada Rambala; Rambala Jungle Lodge; 8.91627 ° N, 82.15469 ° W; 120 m a. s. l.; 7 - 9 Oct. 2016; E. Carlson, leg.; Malaise trap; in alcohol; MUPADI • ibid., 18 males; 15 - 20 Nov. 2016; MUPADI • ibid., 13 males; 21 - 31 Dec. 2016; MUPADI; ibid., 6 males; 6 - 12 Feb. 2017; MUPADI • ibid., 18 males; 12 - 15 Jun. 2017; MUPADI • ibid. 3 males; 28 - 30 Jun. 2017; MUPADI. Chiriqui Province • 1 male; Cuenca 108; Boquete District; Quebrada Grande; Valle Escondido; 8.7797 ° N, 82.44016 ° W; 1,122 m a. s. l.; 29 Apr. - 2 May 2018; B. Armitage, T. Arefina-Armitage, leg.; Malaise trap; in alcohol; MUPADI • ibid., 1 male; Quebrada Jaramillo, Finca Monterey; 8.7632 ° N, 82.41383 ° W; 1,214 m a. s. l.; 12 - 19 Aug. 2018; K. Collier, leg.; Malaise trap; in alcohol; MIUP • ibid., 2 males; 14 - 22 Oct. 2018 • ibid., 3 males; Dolega District, Rio Majagua, Potrerillos, Banquito de Palmira; 8.68083 ° N, 82.532528 ° W; 840 m a. s. l.; 28 Feb- 14 Mar. 2019; T. Rios, Y. Aguirre, leg.; Malaise trap; in alcohol; UMSP • ibid., 3 males; Rio Chirigagua, SSE Guayabal; 8.64102 ° N, 82.5578 ° W; 751 m a. s. l.; 19 Jun. 2015; C. Nieto, T. Abrego, E. Perez, A. Tunon, M. Molinar, A. Cornejo, leg.; UV light trap; in alcohol; COZEM. Veraguas Province • 2 males; Cuenca 097; Santa Fe District; Santa Fe National Park; afl. Rio Calovebora; PSPSCB-NPSF-C- 097 - 2017 - 005; 8.54318 ° N, 81.16398 ° W; 515 m a. s. l.; 19 Apr. 2017; A. Cornejo, T. Rios, E. Alvarez, C. Nieto, leg.; UV light trap; in alcohol; COZEM • 1 male; Rio Calovebora, PSPSCB-PNSF-C 097 - 2017 - 006, 8.55038 ° N, 81.16486 ° W; 461 m a. s. l., 23 - 27 Apr. 2017; A. Cornejo, T. Rios, E. Alvarez, C. Nieto, leg.; Malaise Trap; in alcohol; COZEM • ibid., 39 males; Cuenca 132, Rio Mulaba, 2 do Brazo, PSPSCB-NPSF-C- 132 - 2017 - 007; 8.52577 ° N, 81.13045 ° W; 623 m a. s. l.; 19 - 23 Apr. 2017; Malaise trap; COZEM • ibid., 7 males; Rio Piedra de Moler; PSPSCB-NPSF-C- 097 - 2017 - 011; 8.55343 ° N, 81.17675 ° W; 395 m a. s. l.; 20 Apr. 2017; COZEM.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
436018A884B556C6BF3F5E25D504862B.taxon	distribution	Distribution. Mexico, Panama.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
1CF515C5ACD353BEA330D44C7DECF5C4.taxon	description	Figs 2 C, 16	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
1CF515C5ACD353BEA330D44C7DECF5C4.taxon	diagnosis	Diagnosis. Leucotrichia fairchildi is currently the only member of the L. pictipes Group recorded in Panama. Leucotrichia pictipes known distribution includes Mexico, while L. imitator Flint, 1970 and L. sarita Ross, 1944 have been recorded in nearby Costa Rica. All species in this group bear only two ocelli and share similar shapes to the inferior appendage and sternum VII, when viewed laterally. The modified basal antennal segments and setiferous production on the dorsum of the head can be used to easily separate L. fairchildi from each of the three other species.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
1CF515C5ACD353BEA330D44C7DECF5C4.taxon	materials_examined	Material examined. Panama: Bocas del Toro Province • 16 males; Cuenca 093; Chiriqui Grande District; Quebrada Rambala; Rambala Jungle Lodge; 8.91627 ° N, 82.15469 ° W; 120 m a. s. l.; 21 - 31 Dec. 2016; E. Carlson, leg.; Malaise trap; in alcohol; MUPADI • ibid., 3 males; 6 - 12 Feb. 2017 • ibid., 1 male; 12 - 15 Jun. 2017 • 2 males; 28 - 30 Jun. 2017. Comarca Ngaebe Bugle • 3 males; Quebrada Martinez, Bosque Protector Palo Seco, Alto de Valle, detras de las caseta de MiAmbiente; 8.79484 ° N, 82.19391 ° W, 480 m a. s. l.; 16 - 30 Aug. 2019; Y. Aguirre, T. Rios, leg.; Malaise trap; in alcohol; MUPADI • ibid., 2 males; 30 Aug. - 13 Sep. 2019 • ibid., 2 males; 13 - 27 Sep. 2019. Veraguas Province • 1 male; Cuenca 097; Santa Fe District; Santa Fe National Park; Rio Piedra de Moler; PSPSCB-NPSF-C- 097 - 2017 - 011; 8.55343 ° N, 81.17675 ° W; 395 m a. s. l.; 20 Apr. 2017; T. Rios, E. Alvarez, C. Nieto, leg.; UV light trap; in alcohol; COZEM.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
1CF515C5ACD353BEA330D44C7DECF5C4.taxon	distribution	Distribution. Colombia, Costa Rica, Ecuador, El Salvador, Grenada, Panama, Tobago, Trinidad, Venezuela.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
70331CCB8F305C218D30C9E3D6E91ED4.taxon	description	Fig. 8	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
70331CCB8F305C218D30C9E3D6E91ED4.taxon	diagnosis	Diagnosis. This species is similar to L. botosaneanui, L. chiriquiensis, and L. viridis, based on characteristics of the phallus, as discussed under L. botosaneanui. Leucotrichia hispida can be recognized using the tuft of setae on the posteroventral projection of sternum VIII and the lack of any external spines or sclerites on the apex of phallus.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
70331CCB8F305C218D30C9E3D6E91ED4.taxon	materials_examined	Material examined. Panama: Chiriqui Province • 1 male; Cuenca 102; Renacimiento District; La Amistad International Park, Rio Candela, Finca Felix, PSPSCB-PILA-C 102 - 2017 - 021; 8.890557 ° N, 82.61201 ° W; 1,996 m a. s. l.; 25 Jan. 2015; C. Nieto, E. Perez, A. Cornejo, leg.; UV light trap; in alcohol; COZEM • ibid., 1 male; Rio Chiriqui Viejo, PSPSCB-PNVB-C 108 - 2017 - 016; 8.87550 ° N, 82.55336 ° W; 2,117 m a. s. l.; 5 - 8 Jun. 2017; E. Alvarez, T. Rios, E. Perez, leg.; Malaise trap; COZEM • ibid., 2 males; afl. Rio Chiriqui Viejo, PSPSCB-PILA-C 102 - 2017 - 022; 8.90124 ° N, 82.61817 ° W; 2,354 m a. s. l.; 17 - 21 Jun. 2017; UMSP • ibid., 2 males; Cuenca 108; Quebrada del Guayabo, Volcan Baru National Park, PSPSCB-PNVB-C 108 - 2017 - 018; 8.84939 ° N, 82.49349 ° W; 1,947 m a. s. l.; 5 - 8 Jun. 2017; E. Alvarez, E. Perez, T. Rios, leg.; Malaise trap; in alcohol; MUPADI.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
70331CCB8F305C218D30C9E3D6E91ED4.taxon	distribution	Distribution. Costa Rica, Panama.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
4D444D0ADD445557A999F0D04C293441.taxon	description	Fig. 9	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
4D444D0ADD445557A999F0D04C293441.taxon	diagnosis	Diagnosis. Leucotrichia holzenthali sp. nov., is similar to L. dinamica Bueno-Soria, 2010, a species currently known only from Mexico. Both species bear a pair of large scissor-like sclerites on the apex of the phallus. Leucotrichia holzenthali can be distinguished by the additional pair of ventral sclerites on the phallus apex and the peg-like setae on abdominal sternum VIII, both characteristics that are absent on L. dinamica.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
4D444D0ADD445557A999F0D04C293441.taxon	description	Description. Male. Length of forewing 1.7 mm (n = 2). Wings unmodified. Head unmodified, with three ocelli; antennae unmodified. Tibial spur count 1, 3, 4. Color in alcohol brown. Genitalia. Abdominal sternum VII mesoventral process with enlarged apex (Fig. 9 A, D). Sternum VIII with posteroventral production bearing prominent peg-like setae (Fig. 9 A), in ventral view posterior margin concave (Fig. 9 D). Segment IX anterolateral margin convex, posterolateral margin irregular (Fig. 9 B); dorsally, anterior margin concave, posterior margin broadly concave (Fig. 9 C). Tergum X with dorsal sclerite not apparent; ventral sclerite broadest mesally with slender ventral apex; membranous apex subtriangular in dorsal view (Fig. 9 B, C). Subgenital plate with dorsal arm simple, extending posteriad, apex acute (Fig. 9 B); ventral arm simple, apex with emargination, in ventral view with truncate base and rounded apical emargination (Fig. 9 B, D). Inferior appendage broadest basally, with pointed basal emargination, prominent dorsal subapical seta, apex acute (Fig. 9 B); ventrally broadly fused, broadest basally, apex rounded (Fig. 9 D). Phallus tubular basally, constricted at midlength with median complex bearing spherical " windows "; apex membranous, bearing pair of large scissor-like apical sclerites dorsally and pair of large acute sclerites ventrally (Fig. 9 E, F).	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
4D444D0ADD445557A999F0D04C293441.taxon	distribution	Distribution. Panama.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
4D444D0ADD445557A999F0D04C293441.taxon	etymology	Etymology. Named in honor of Dr. Ralph W. Holzenthal, for a long and robust career in caddisfly taxonomy and systematics. Dr. Holzenthal has been a friend and colleague to each of the authors, and an invaluable mentor in particular to the first author.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
3034911A61665EDC9A09618087B3C859.taxon	description	Fig. 10	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
3034911A61665EDC9A09618087B3C859.taxon	diagnosis	Diagnosis. Leucotrichia luma is most similar to L. inflaticornis, a species currently known only from Trinidad. Certain characteristics found on the genitalia of these species make them very similar, such as the symmetrical rows of spines on the phallus apex, a unique arrangement within the genus. Key differences, however, make it possible to separate the two. Inflated antennal segments, a key feature of L. inflaticornis, were not observed in any of the specimens collected in Panama and identified as L. luma. Additionally, L. luma specimens all present three pairs of spines on the phallus apex, while the original description of L. inflaticornis states that there should be four. The first author has observed the holotype specimen of L. inflaticornis and found that the abdomen, including the phallus, was missing and key features of the genitalia cannot be confirmed. Since specimens cannot be compared to the L. inflaticornis holotype, we compare these specimens to the original description and illustration and offer this new species description for the specimens from Panama.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
3034911A61665EDC9A09618087B3C859.taxon	description	Description. Male. Length of forewing 1.8 - 2.0 mm (n = 17). Wings unmodified. Head unmodified, with 3 ocelli; antennae unmodified. Tibial spur count 1, 3, 4. Color in alcohol brown. Genitalia. Abdominal sternum VII without apparent mesoventral process. Sternum VIII with acute posteroventral production, in ventral view posterior margin concave (Fig. 10 A, D). Segment IX anterolateral margin convex, posterolateral margin straight (Fig. 10 B); dorsally, anterior margin concave, posterior margin concave (Fig. 10 C). Tergum X with dorsal sclerite simple, slender; ventral sclerite semi-elliptical with rounded emargination mesally on posterior margin; membranous apex small, suborbicular (Fig. 10 B, C). Subgenital plate with dorsal arm digitate, approximately half the length of ventral arm (Fig. 10 B); ventral arm slender, apex truncate, with irregular ventral margin, in ventral view broadest mesally with rounded apical emargination (Fig. 10 B, D). Inferior appendage with base extending anteriorad, with two dorsal subapical spines, apex curved dorsad (Fig. 10 B); in ventral view broadly fused, with digitate basal projections (Fig. 10 D). Phallus tubular basally, constricted at midlength with median complex bearing basal loop and pair of spherical " windows "; apex membranous and bearing 3 sets of symmetrically arranged stout, dark spines (Fig. 10 E, F).	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
3034911A61665EDC9A09618087B3C859.taxon	distribution	Distribution. Panama.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
3034911A61665EDC9A09618087B3C859.taxon	etymology	Etymology. The specific epithet is derived from luma, Latin for " thorn ", referring to the spines found on the phallus apex.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
4AFA0BB60AA154A1A9F32C6877EBE447.taxon	description	Fig. 11	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
4AFA0BB60AA154A1A9F32C6877EBE447.taxon	diagnosis	Diagnosis. Leucotrichia melleopicta is most similar to L. mutica, also recorded from Panama. These species possess an inferior appendage with a similar shape, and the dorsal sclerite of the phallus of both bears a dorsal sclerite with a distinct apical emargination. Leucotrichia melleopicta can be distinguished by the elongate basal sclerite of the phallus and the poorly developed basal loop (Fig. 11 E), and by the enlarged apex of the mesoventral process on segment VII (Fig. 11 A).	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
4AFA0BB60AA154A1A9F32C6877EBE447.taxon	materials_examined	Material examined. Panama: Bocas del Toro Province • 6 males; Cuenca 093; Chiriqui Grande District; Quebrada Rambala; Rambala Jungle Lodge; 8.91627 ° N, 82.15469 ° W; 120 m a. s. l.; 28 Mar. 2015; B. Armitage, T. Arefina- Armitage, leg.; UV light trap; in alcohol; MUPADI • ibid., 68 males; 31 Mar. - 11 Apr. 2015; E. Carlson, leg.; Malaise trap • ibid., 38 males; 7 - 9 Oct. 2016 • ibid., 36 males; 15 - 20 Nov. 2016 • ibid., 66 males; 21 - 31 Dec. 2016 • ibid., 55 males; 6 - 12 Feb. 2017 • ibid., 32 males; 12 - 15 Jun. 2017 • ibid., 17 males; 28 - 20 Jun. 2017. Comarca Ngaebe Bugle • 1 male; Cuenca 093; Palo Seco Forest Preserve; Quebrada Martinez; Alto de Valle, detras de las caseta de MiAmbiente; 8.79484 ° N, 82.19047 ° W; 480 m a. s. l.; 5 - 19 May 2019; T. Rios, Y. Aguirre, leg.; Malaise trap; in alcohol; MUPADI • ibid., 4 males; 24 May- 6 Jun. 2019 • ibid., 5 males; 22 Sep. - 11 Oct. 2019 • ibid., 2 males; Willie Mazu, 8.79361 ° N, 82.19391 ° W, 538 m a. s. l.; 13 - 27 Sep. 2019 • ibid., 1 male; 11 - 30 Oct. 2019. Chiriqui Province • 27 males; Cuenca 108, Boquete District; Quebrada Jaramillo; Finca Monterey; 8.7632 ° N, 82.41383 ° W; 1,214 m a. s. l.; 19 - 25 Apr. 2018, K. Collier, leg.; Malaise trap; in alcohol; MUPADI • ibid., 41 males; 8 - 12 May 2018 • ibid., 54 males; 16 - 20 Jun. 2018 • ibid., 5 males; 12 - 19 Aug. 2018 • ibid., 52 males; 14 - 22 Oct. 2018 • ibid., 6 males; 15 - 22 Nov. 2018 • ibid., 48 males; Dolega District, Rio Majagua, Potrerillos, Banquito de Palmira; 8.68083 ° N, 82.532528 ° W; 840 m a. s. l.; 28 Feb- 14 Mar. 2019; T. Rios, Y. Aguirre, leg.; Malaise trap; in alcohol; UMSP. Veraguas Province • 2 males; Cuenca 097, Santa Fe District, Santa Fe National Park, afl. Rio Calovebora; PSPSCB-NPSF-C- 097 - 2017 - 005; 8.54318 ° N, 81.16398 ° W; 515 m a. s. l.; 19 - 23 Apr. 2017; A. Cornejo, T. Rios, E. Alvarez, C. Nieto, leg.; Malaise trap; in alcohol; COZEM • ibid., 1 male; Rio Llanito, PSPSCB-NPSF-C- 097 - 2017 - 012; 8.56553 ° N, 81.18817 ° W; 340 m a. s. l.; 21 Apr. 2017; UV light trap; COZEM • ibid., 1 male; Rio Pedra Moler; PSPSCB-NPSF-C- 097 - 2017 - 011; 8.55343 ° N, 81.17675 ° W; 395 m a. s. l.; 20 Apr. 2017 • ibid., 104 males; Cuenca 132, Rio Mulaba, 2 do Brazo, PSPSCB-NPSF-C- 097 - 2017 - 007; 8.52577 ° N, 81.13045 ° W; 623 m a. s. l.; 19 - 23 Apr. 2017; Malaise trap; COZEM • ibid., 41 males; Rio Mulaba, afl. 1 er Brazo; PSPSCB-NPSF-C- 097 - 2017 - 008; 8.51706 ° N, 81.1214 ° W; 770 m a. s. l.; 19 - 23 Apr. 2017; COZEM.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
4AFA0BB60AA154A1A9F32C6877EBE447.taxon	distribution	Distribution. Mexico, Panama, Venezuela.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
DB6C7A0C7B2F5D739547ED458BE8D007.taxon	description	Fig. 12	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
DB6C7A0C7B2F5D739547ED458BE8D007.taxon	diagnosis	Diagnosis. This species is similar to L. melleopicta; both species display ranges that include Panama. As discussed under L. melleopicta, these species share similarities in the dorsal sclerite of the phallus apex, although this sclerite is much smaller than that seen in L. melleopicta (Fig. 12 E), and the general shape of the inferior appendage. Leucotrichia mutica can be recognized separately from L. melleopicta by the tapering mesoventral process on segment VII (Fig. 12 A), and by the prominent basal loop of the phallus (Fig. 12 D, E).	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
DB6C7A0C7B2F5D739547ED458BE8D007.taxon	materials_examined	Material examined. Panama: Bocas del Toro Province • 17 males; Cuenca 093; Chiriqui Grande District; Quebrada Rambala; Rambala Jungle Lodge; 8.91627 ° N, 82.15469 ° W; 120 m a. s. l.; 7 - 9 Oct. 2016; E. Carlson, leg.; Malaise trap; in alcohol; MUPADI • ibid., 10 males; 15 - 20 Nov. 2016 • ibid., 6 males; 21 - 31 Dec. 2016 • ibid., 6 males; 6 - 12 Feb. 2017 • ibid., 14 males; 12 - 15 Jun. 2017 • ibid., 47 males; 28 - 30 Jun. 2017. Comarca Ngaebe Bugle • 1 male; Quebrada Martinez, Bosque Protector Palo Seco, Alto de Valle; detras de las caseta de MiAmbiente, M 0002; 8.79424 ° N, 82.1904724 ° W; 480 m a. s. l.; 24 May- 6 Jun. 2018; Y. Aguirre, T. Rios, leg.; Malaise trap; in alcohol; MUPADI. Chiriqui Province • 1 male; Cuenca 108; Boquete District; Quebrada Grande; Valle Escondido; 8.7797 ° N, 82.44016 ° W; 1,122 m a. s. l.; 11 Mar. 2018; T. Arefina-Armitage, leg.; UV light trap • 6 males; 29 Apr. - 2 May 2018; Malaise trap • ibid., 1 male; 21 May 2018; UV light trap • ibid., 1 male; 8.783645 ° N, 82.444287 ° W; 1,147 m a. s. l.; 27 - 30 May 2018; Malaise trap • ibid., 1 male; 17 - 20 Jun. 2018 • 6 males; 23 Jul. 2018; UV light trap • ibid., 2 males; 10 Nov. 2018. Veraguas Province • 1 male; Cuenca 097; Santa Fe District; Santa Fe National Park; afl. Rio Calovebora; PSPSCB-NPSF-C- 097 - 2017 - 005; 8.54318 ° N, 81.16398 ° W; 515 m a. s. l.; 19 - 23 Apr. 2017; Malaise trap; in alcohol; COZEM.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
DB6C7A0C7B2F5D739547ED458BE8D007.taxon	distribution	Distribution. Colombia, Panama.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
56DF00270A455E3A86B1666B30D04600.taxon	description	Fig. 13	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
56DF00270A455E3A86B1666B30D04600.taxon	diagnosis	Diagnosis. Leucotrichia rhomba is similar to L. brochophora Flint, 1991 and L. padera Flint, 1991, two species recorded from Colombia, but not currently known from Panama. All three species share a similar appearance of the phallus apex, with no spines and either few or no externally sclerotized structures. The anterolateral margin of segment IX is also produced in all three species. Leucotrichia rhomba can be recognized by the elongate mesoventral process of segment VII, which is enlarged, rhomboid, and rugose in ventral view (Fig. 13 A, B), and by the obovoid dorsal sclerite of the phallus apex (Fig. 13 E).	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
56DF00270A455E3A86B1666B30D04600.taxon	materials_examined	Material examined. Panama: Bocas del Toro Province • 6 males; Cuenca 093; Chiriqui Grande District; Quebrada Rambala; Rambala Jungle Lodge; 8.91627 ° N, 82.15469 ° W; 120 m a. s. l.; 12 - 15 Jun. 2017; E. Carlson, leg.; Malaise trap; in alcohol; MUPADI • ibid., 1 male; 28 - 30 Jun. 2017 • ibid., 2 males; 15 - 20 Nov. 2016. Chiriqui Province • 5 males; Cuenca 108; Boquete District; Quebrada Jaramillo; Finca Monterey; 8.7632 ° N, 82.41383 ° W; 1,214 m a. s. l.; 16 - 20 Jun. 2017; K. Collier, leg.; Malaise trap • ibid., 1 male; 14 -- 22 Oct. 2018. • ibid., 5 males; Dolega District, Rio Majagua, Potrerillos, Banquito de Palmira; 8.68083 ° N, 82.532528 ° W; 840 m a. s. l.; 28 Feb. - 14 Mar. 2019; T. Rios, Y. Aguirre, leg.; MUPADI. Veraguas Province • 14 males; Cuenca 132; Santa Fe District, Santa Fe National Park, Rio Mulaba, 2 do Brazo, PSPSCB-NPSF-C- 097 - 2017 - 007; 8.52577 ° N, 81.13045 ° W; 623 m a. s. l.; 19 - 23 Apr. 2017; A. Cornejo, T. Rios, E. Alvarez, C. Nieto, leg.; Malaise trap; in alcohol; COZEM.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
56DF00270A455E3A86B1666B30D04600.taxon	distribution	Distribution. Costa Rica, Panama.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
0AE63BE727355777955239625A0FE4E0.taxon	description	Figs 2 A, 14	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
0AE63BE727355777955239625A0FE4E0.taxon	materials_examined	Other material examined. Panama: Chiriqui Province • 1 male; Cuenca 102, Renacimiento District; La Amistad International Park, Rio Candela, Finca Felix, PSPSCB-PILA-C 102 - 2017 - 021; 8.90614 ° N, 82.72882 ° W; 1,799 m a. s. l., 1 - 5 Sep. 2017; E. Alvarez, T. Rios, E. Perez, leg.; Malaise trap; in alcohol; COZEM. Veraguas Province • 1 male; Cuenca 132, Santa Fe National Park, Rio Mulaba, 2 do Brazo, PSPSCB-NPSF-C- 097 - 2017 - 007; 8.52577 ° N, 81.13045 ° W; 623 m a. s. l.; 19 - 23 Apr. 2017; A. Cornejo, T. Rios, E. Alvarez, C. Nieto, leg.; Malaise trap; in alcohol; MUPADI.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
0AE63BE727355777955239625A0FE4E0.taxon	diagnosis	Diagnosis. Leucotrichia ruiteri sp. nov. is most similar to L. hispida, as both species bear prominent setae on the posteroventral production of sternum VIII, a bilobed phallus apex, and a similar shaped inferior appendage. Leucotrichia ruiteri can be separated by the single, elongate seta on sternum VIII compared to the cluster of setae present on L. hispida. Additionally, the forewings of L. ruiteri are modified with a large setae-filled pocket, while those of L. hispida are unmodified.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
0AE63BE727355777955239625A0FE4E0.taxon	description	Description. Male. Length of forewing 2.0 - 2.1 mm (n = 4). Forewing with large pocket filled with scales (Fig. 2 A); hindwing unmodified. Head unmodified, with three ocelli; antennae unmodified. Tibial spur count 1, 3, 4. Color in alcohol brown. Genitalia. Abdominal sternum VII slender, elongate (Fig. 14 A, D). Sternum VIII with posteroventral production bearing prominent apical seta (Fig. 14 A); in ventral view posterior margin concave (Fig. 14 D). Segment IX anterolateral margin convex, posterolateral margin straight with slight irregularity (Fig. 14 B); dorsally, anterior margin concave, posterior margin broadly convex (Fig. 14 C). Tergum X with dorsal sclerite with irregular dorsal margin; ventral sclerite semi-elliptic with knoblike projection mesally on posterior margin; membranous apex subquadrate (Fig. 14 B, C). Subgenital plate with dorsal arm simple, with slight preapical emargination on dorsal margin, apex truncate (Fig. 14 B); ventral arm narrowing apically, ventral margin slightly sinuate, in ventral view subovate with narrow basal projection and small rounded apical emargination (Fig. 14 B, D). Inferior appendage broadest mesally, with small dorsal subapical peg-like seta, apex rounded (Fig. 14 B); ventrally slender, with digitate basal projections, apex slightly hooked on inner margin (Fig. 14 D). Phallus tubular basally, constricted at midlength with typical median complex bearing basal loop and pair of spherical " windows "; apex membranous with internal sclerotized structures and two apical lobes extending dorsad (Fig. 14 E, F).	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
0AE63BE727355777955239625A0FE4E0.taxon	distribution	Distribution. Panama.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
0AE63BE727355777955239625A0FE4E0.taxon	etymology	Etymology. This species is named in honor and memory of Dave Ruiter, a passionate and enthusiastic caddisfly researcher and good friend, who recently passed away.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
CABCDD3CEF2154EEB73FB810E2F06FA9.taxon	description	Fig. 15	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
CABCDD3CEF2154EEB73FB810E2F06FA9.taxon	diagnosis	Diagnosis. Due to the similar overall appearance of the phallus, L. viridis is most similar to L. botosaneanui, L. hispida, and L. chiriquiensis, as discussed under L. botosaneanui. Leucotrichia viridis has two dorsal spines on the inferior appendage, while the other species each bear only a single spine.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
CABCDD3CEF2154EEB73FB810E2F06FA9.taxon	materials_examined	Material examined. Panama: Bocas del Toro Province • 21 males; Cuenca 093, Chiriqui Grande District, Quebrada Rambala, Rambala Jungle Lodge; 8.91627 ° N, 82.15469 ° W; 120 m a. s. l.; 9 Aug. 2014; E. Carlson, leg.; Malaise trap; in alcohol; MUPADI • ibid., 3 males; 28 Mar. 2015; UV light trap • ibid., 3 males; 31 Mar. - 11 Apr. 2015; Malaise trap. Chiriqui Province • 1 male; Cuenca 108; Dolega District; Rio Majagua; Potrerillos, Banquito de Palmira; 8.68083 ° N, 82.53253 ° W; 840 m a. s. l.; 28 Feb. - 14 Mar. 2019; T. Rios, Y. Aguirre, leg.; Malaise trap; in alcohol; MUPADI. - Veraguas Province • 3 males; Cuenca 132; Santa Fe District, Santa Fe National Park, Rio Mulaba, afl. 1 er Brazo; PSPSCB-NPSF-C- 097 - 2017 - 008; 8.51706 ° N, 81.1214 ° W; 770 m a. s. l.; E. Alvarez, E. Perez, T. Rios, leg.; 19 - 23 Apr. 2017; in alcohol; COZEM.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
CABCDD3CEF2154EEB73FB810E2F06FA9.taxon	distribution	Distribution. El Salvador, Guatemala, Mexico, Panama.	en	Thomson, Robin E., Armitage, Brian J., Harris, Steven C. (2022): The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama. ZooKeys 1111: 425-466, DOI: http://dx.doi.org/10.3897/zookeys.1111.77371, URL: http://dx.doi.org/10.3897/zookeys.1111.77371
