taxonID	type	description	language	source
FE0687C3FFAB4E1EE5E2FD3CFC90F837.taxon	diagnosis	Diagnosis: Paratelius resembles Scarelus in general appearance and has similar long setae between dense, short pubescence on elytra. Paratelius, like Atelius, bears four longitudinal costae on each elytron. In all known species the phallus is very long, slender and slightly curved (Figs 1 A – 1 J).	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFAB4E1EE5E2FD3CFC90F837.taxon	description	Redescription: Similar to Scarelus in general appearance. Small-bodied, slender, 5.8 – 7.0 mm long. Most species brown to reddish brown colored, some uniformly testaceous; when dark colored, humeral part of elytra lighter and apical half of elytra dark. Head small, free, hypognathous, covered with short pubescence, surface mat, with fine structure. Frons convex; eyes small; antennal tubercles inconspicuous, divided by shallow depression; antennal cavities very close each to other, separated by narrow cranial bridge. Antennae 11 - segmented, compressed, slightly serrate, long, surpassing elytral apex by 3 to 5 antennomeres, covered with dense, recumbent pubescence; apical antennomere longest, narrowed to the apex, pointed. Mandibles stout, quite long, slightly curved, shining, with smooth internal margin. Other mouthparts tiny. Labium and maxillae clearly visible without dissecting, reduced. Labium reaching from gular margin of mouth cavity to one third of its diameter: maxillae reaching to a half of diameter. Labial palpi 2 - segmented, maxillary palpi 4 - segmented, apical palpomere stoutest in both palpi. Labrum projected forwards, triangular. Pronotum trapezoidal, flat, posterior margin wider than frontal one, frontal margin slightly rounded, lateral margins convex, posterior angles acutely projected, posterior margin almost straight. Pronotal disc with single median longitudinal carina with very deep and narrow depression in middle part. Scutellum flat, trapezoidal, shiny, sparsely pubescent, usually emarginate at apex. Elytra flat, with four longitudinal costae. Costae connected by regular transverse ridges, whole elytra densely and shortly pubescent, also bearing sparse, stout, long, erect setae. Legs slender, compressed, densely pubescent. Tarsus with 5 tarsomeres. Male genitalia trilobate, uniform in general shape (Figs 1 A-J). Phallus very long, widest at base, paramerae very short, globular in shape, phallobase longer than paramerae.	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFAB4E1EE5E2FD3CFC90F837.taxon	distribution	Distribution: Malaysia: Sabah State. Distribution is limited to mountain ranges in northeastern Borneo. These mountains were produced by a counterclockwise rotation of Borneo since the Late Miocene (Hall & Blundell, 1996). Based on the endemic occurence in a single mountain range, we may suppose speciation of all Paratelius species occurred in this region. Paratelius has not been recently collected in the lowlands of Borneo, and only the type specimen of P. diversicornis is recorded from Brunei without precise locality data. Although Borneo was connected during the last glacial maximum to Indochina (Voris, 2000) and the generic structure of both faunas is similar, neither Paratelius from Borneo nor related Atelius from Vietnam expanded their ranges over the subaerial shelf of the Asian continent.	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFA84E1CE5E2F894FB3CFD3B.taxon	diagnosis	Diagnosis: P. diversicornis differs from all known Paratelius in the characteristic dark coloration of the elytra.	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFA84E1CE5E2F894FB3CFD3B.taxon	description	Redescription: Male. Body small, head and thorax dark brown; abdomen black; pronotum slightly lighter than elytra; elytra lighter at humeri, dark brown to black apically; eyes and anntennae black. Head small, densely pubescent; eyes small, hemispherically prominent, their interocular distance 3.4 times maximum eye diameter. Antennae slender, compressed, moderately flattened, surpassing apex of elytra by 3 antennomeres. Antennomeres 3 – 11 gradually more slender; apical antennomere eliptical. Elytra slender, parallel-sided, 3.47 times longer than width at humeri, with four well developed primary costae, covered with reddish pubescence. Legs moderately long, slender, flattened, densely pubescent. Male genitalia trilobate; phallus very long, slender, slightly curved; parameres short, reaching to one sixth of phallic length; phallobase short, subtle (Figs 1 G-H). Measurements: BL 6.2 mm, PW 1.2 mm, PL 0.8 mm, WH 1.5 mm, Edist 0.71 mm, Ediam 0.21 mm.	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFA84E1CE5E2F894FB3CFD3B.taxon	distribution	Distribution: Malaysia: Sabah: western part of the Crocker Range; Brunei (type locality).	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFA94E1CE5E2FCE7FED5F944.taxon	materials_examined	Type material: Holotype, male. Borneo, Sabah, km 53 KK-Tambunan, 1650 m, Gn. Emas, 22. iii. – 6. iv. 2000, leg. Bolm. (LMBC); 2 paratypes, males. Same locality data (LMBC).	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFA94E1CE5E2FCE7FED5F944.taxon	diagnosis	Diagnosis: P. emasensis n. sp. is similar to P. nigricornis n. sp. in possessing elytra with bright reddish coloration. These species differ in the dark brown to black head, pronotum, and antennae in P. nigricornis n. sp. and testaceous to reddish brown in P. emasensis n. sp. The slightly more curved phallus in P. emasensis supports separation of these species (Figs. 1 C-D and 1 E-F).	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFA94E1CE5E2FCE7FED5F944.taxon	description	Description: Male. Body small, slender. Head and thorax red; abdomen dark brown; eyes black; scapus, pedicel, antennomeres 3 – 11 and legs light brown, coxae and mouth parts brown. Head small, densely pubescent; eyes small, hemispherically prominent, frontal interocular distance 3.4 times maximum eye diameter. Antennae compressed, slender, flattened, reaching over apex of elytra by 1.5 apical antennomeres. Antennomeres 3 – 11 gradually slender; apical antennomere eliptical. Elytra slender, 3.1 times longer than width at humeri, parallel sided, with four well developed primary costae, covered with red pubescence. Legs moderately long, slender, flattened, densely pubescent. Male genitalia trilobate; phallus very long, slightly turned upward; parameres short reaching one sixth of phallic length. Phallobase small, subtle (Figs 1 C-D). Measurements: BL 6.0 mm, PW 1.3 mm, PL 1.0 mm, WH 1.6 mm, Edist 0.71 mm, Ediam 0.21 mm.	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFA94E1CE5E2FCE7FED5F944.taxon	distribution	Distribution: Malaysia: Sabah.	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFA94E1CE5E2FCE7FED5F944.taxon	etymology	Etymology: The species epithet refers to type locality of this species, Gunung Emas in Sabah.	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFA94E1CE5E2FCE7FED5F944.taxon	discussion	Remarks. The coloration of the antennomeres is slightly variable, and some specimens may have slightly darker apical antennomeres. However, it is notable that several basal antennomeres are brightly coloured in all available specimens of P. emasensis n. sp. DNA was isolated from one paratype, and sequences of several mtDNA and rRNA genes are accessible in the GenBank under the voucher number UPOL 000583 (Bocak et al., 2008).	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFA94E1BE5E2F8C4FC78FD3B.taxon	materials_examined	Type material: Holotype, male. B. N. Borneo, Mt. Kinabalu, Keuokok, 3.300 ft, 26 April 1929, H. M. Pendlebury coll., F. M. S. Museum (BMNH). Other material examined: Male, Borneo, Sabah, km 55 KK- Tambunan, 1650 m, Gn. Emas, 22. iii. – 6. iv. 2000, leg. Bolm (LMBC); male, Borneo, Sabah, Mt. Kinabalu N. P., Feb 2000, M. Snizek (LMBC).	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFA94E1BE5E2F8C4FC78FD3B.taxon	diagnosis	Diagnosis: P. juvencus is similar to P. nigricornis n. sp. in the coloration of pronotum and head. P. juvencus has more robust body with elytron less than 6 times longer than its width in the middle part and darker. The male genitalia of both species are similar (Figs 1 I-J).	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFA94E1BE5E2F8C4FC78FD3B.taxon	description	Redescription: Male. Body small, robust, head, thorax and abdomen black; elytra dark cinnamon brown; antennae, mouth parts and legs black. Head small, densely pubescent; eyes small, hemispherically prominent, frontal distance 3.4 times maximum eye diameter. Antennae slightly serrate, slender, flattened, antennomeres 3 - 10 triangular; last antennomere eliptical. Antennae reaching over apex of elytra by four antennomeres. Elytra wide, 3.1 times longer than width at humeri, parallel-sided, with four well developed primary costae, covered with reddish pubescence. Legs moderately long, slender, flattened, densely pubescent. Male genitalia trilobate; phallus very long, slightly turned upwards; parameres short, reaching one sixth of phallic length; phalobase very small, short and subtle (Figs 1 I-J). Measurements: BL 6.2 mm, PW 1.0 mm, PL 0.8 mm, WH 1.5 mm, Edist 0.71 mm, Ediam 0.21 mm.	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFA94E1BE5E2F8C4FC78FD3B.taxon	distribution	Distribution: Malaysia: Sabah: Mt. Kinabalu, Crocker Range.	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFAE4E1BE5E2FCEBFB6EF9EA.taxon	materials_examined	Type material: Holotype, male. Borneo, Sabah, km 53, KK-Tambunan, 1650 m, Gn. Emas, 22. iii. – 6. iv. 2000, leg. Bolm (LMBC). Paratype: Same locality data (LMBC).	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFAE4E1BE5E2FCEBFB6EF9EA.taxon	diagnosis	Diagnosis: P. nigricornis n. sp. is similar to P. juvencus in its dark colored pronotum. It differs mainly in the bright reddish coloration of the elytra and more slender body (the elytron about 8 times longer than the width in the middle part). The differences in male genitalia are very subtle (Figs 1 E-F).	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFAE4E1BE5E2FCEBFB6EF9EA.taxon	description	Description: Male. Body small; head, and thorax black; abdomen brown; elytra bright reddish brown; eyes black; scapus and pedicel brown; anntennae completely black; legs brown. Head small, densely pubescent; eyes small, hemispherically prominent, their distance 2.53 times maximum eye diameter. Antennae slender, flattened, reaching over apex of elytra by 3 antennomeres. Antennomeres 3 – 11 gradually slender; apical antennomere eliptical. Elytra slender, 3.16 times longer than width at humeri, parallel sided, with four well developed primary costae, covered with reddish pubescence. Legs moderately long, slender, flattened, densely pubescent. Male genitalia trilobate; phallus very long, slightly turned upwards; parameres short, subtle, reaching one fifth of phallic length; phallobase short, subtle (Figs 1 E-F). Measurements: BL 7.0 mm, PW 1.4 mm, PL 1.0 mm, WH 1.9 mm, Edist 0.71 mm, Ediam 0.28 mm.	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFAE4E1BE5E2FCEBFB6EF9EA.taxon	distribution	Distribution: Malaysia: Sabah.	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFAE4E1BE5E2FCEBFB6EF9EA.taxon	etymology	Etymology: The specific epithet refers to the black coloration of antennae.	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFAE4E1BE5E2FCEBFB6EF9EA.taxon	discussion	Remark. DNA was isolated from the paratype, and sequences of several mtDNA and rRNA genes are accessible in the GenBank under the voucher number UPOL 000 L 15 (Bocak et al., 2008).	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFAE4E1AE5E2F9B5FA4DFE13.taxon	materials_examined	Type material: Holotype, male. Borneo, Sabah, Crocker Mt., Gunung Emas env., 15. - 27. iv. 1993, leg. Jenis & Strba (LMBC). Paratype: male, Borneo, Sabah, Mt. Kinabalu, 28 May 1999, M. Snizek lgt. (LMBC).	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFAE4E1AE5E2F9B5FA4DFE13.taxon	diagnosis	Diagnosis: Paratelius snizeki n. sp. is similar to P. emasensis n. sp. in the testaceous to reddish brown coloration of the cranium and pronotum. These species differ in the shape of antennomeres, which are more robust and acutely serrate in P. snizeki. Male genitalia differ in the relatively shorter phallus of P. snizeki (Figs 1 A-B and 1 C-D).	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFAE4E1AE5E2F9B5FA4DFE13.taxon	description	Description: Male. Body small, moderately slender; head, thorax, elytra reddish brown; abdomen dark brown; eyes black; antennae, mouth parts and legs brown. Head small, hypognathous; eyes small, hemispherically prominent; frontal interocular distance 3.04 times maximum eye diameter. Antennae slender, flattened, surpassing by 3 antennomeres apex of elytra. Antennomeres 3 – 11 gradually more slender; apical antennomere eliptical. Elytra slender, 3.69 times longer than width at humeri, parallel-sided, with four well developed primary costae, covered with dense brown pubescence. Legs moderately long, slender, flattened, densely pubescent. Male genitalia trilobate; phallus very long, slightly turned upwards; parameres short, reaching one fifth of phallic length; phallobase small and subtle (Figs 1 A-B). Measurements: BL 5.8 mm, PW 1.0 mm, PL 0.8 mm, WH 1.3 mm, Edist 0.64 mm, Ediam 0.21 mm.	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFAE4E1AE5E2F9B5FA4DFE13.taxon	distribution	Distribution: Malaysia: Sabah.	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
FE0687C3FFAE4E1AE5E2F9B5FA4DFE13.taxon	etymology	Etymology: The specific epithet is a patronym in honor of M. Snizek, the collector of the type specimen.	en	Malohlava, Vladimír, Bocak, Ladislav (2009): A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae). Zootaxa 2306 (1): 44-50, DOI: 10.11646/zootaxa.2306.1.4, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4
