taxonID	type	description	language	source
FE36014CFFA3FFF37CC9DF20FBD2F9C1.taxon	type_taxon	Type species: Amplisegmentum ecuadorensis Webb, 2005: 4. — Gaimari & Webb (2009) [key]; Webb et al. (2013) [catalogue]; Irwin & Winterton (2020) [key].	en	Winterton, Shaun L. (2021): A new species of Amplisegmentum Webb (Diptera: Therevidae) from Venezuela. Zootaxa 4927 (4): 576-582, DOI: 10.11646/zootaxa.4927.4.8
FE36014CFFA3FFF37CC9DF20FBD2F9C1.taxon	diagnosis	Diagnosis. Male frons wider than anterior ocellus; scape bulbous; flagellum turbinate to conical; prosternum with setal pile; scutellum with two pairs of macrosetae; wing cell m 3 open; head thorax and abdomen with extensive elongate admixed white lanceolate and black filiform setae; inner gonocoxal process present; gonocoxites separate medially.	en	Winterton, Shaun L. (2021): A new species of Amplisegmentum Webb (Diptera: Therevidae) from Venezuela. Zootaxa 4927 (4): 576-582, DOI: 10.11646/zootaxa.4927.4.8
FE36014CFFA3FFF37CC9DF20FBD2F9C1.taxon	description	Description. Head. Antennal length equal to head length; flagellum turbinate; flagellum shorter than combined scape and pedicel length; scape bulbous, noticeably thicker than pedicel and base of flagellum; frons only slightly raised around base of antennae, antennae positioned in middle of head and directed anteriorly; head length and height subequal; gena rounded; male frons width (at narrowest point) equal to or wider than anterior ocellus; parafacial with or without setae along eye margin; male postocular macrosetae scattered medially on occiput and concentrated in one row (dorsally) along postocular ridge. Thorax with central depression of prosternum setose; pleuron overlain with dense silver or grey pubescence admixed with pale erect setae; metanepisternum with postspiracular setae present; metakatepisternum with setae absent; scutal chaetotaxy (pairs): supra alar, 1 – 2; post alar, 1; dorsocentral, 2 or 0; scutellar, 2; posterior surface of midcoxa without setae or sometimes just a few setae; midfemoral vestiture as dark, erect filiform seta admixed with pale lanceolate and adpressed scale-like setae; fore- and midfemora anteroventral macrosetae present or absent, present on hind femur as a series along length, all femora with posteroventral macrosetae absent or indistinct; hind femur and tibia approximately equal length to fore and mid legs; hind coxal knob present; wing cell m 3 open, veins M 3 and M 4 separate to margin; wing vein R 2 + 3 smoothly sigmoid to wing margin, wing mostly hyaline or with faint markings present; wing vein R 1 with setae absent distally. Abdomen. Extensive pile of white lanceolate setae on all segments; male with silver velutinous pubescence. Genitalia (Fig. 4). Male with epandrium and gonocoxites with extensive pale, lanceolate setae; gonocoxite with gonocoxal process very small with 1 – 3 large, dark posterolaterally directed setae; inner gonocoxal process present; gonocoxite with posteromedial margins proximal but not fused; ventral lobe large, elongate to rounded; hypoproct (subepandrial sclerite) small or absent; dorsal apodeme of phallus broad; distiphallus narrow, straight or directed ventrally; ventral apodeme narrow. Female with acanthophorite macrosetae well developed (A 1 & A 2); sternite 8 posterior margin emarginate posteromedially; two spherical spermathecae with thickened spermathecal ducts basally; spermathecal sac rounded.	en	Winterton, Shaun L. (2021): A new species of Amplisegmentum Webb (Diptera: Therevidae) from Venezuela. Zootaxa 4927 (4): 576-582, DOI: 10.11646/zootaxa.4927.4.8
FE36014CFFA3FFF37CC9DF20FBD2F9C1.taxon	discussion	Comments. The original circumscription of the genus by Webb (2005 b) was based on a single species (A. ecuadorensis), which is here revised to accommodate the new species and correct some inaccuracies in the original description. Amplisegmentum is easily differentiated from other Therevinae by the following characteristics: antennal scape bulbous; scattered dark macrosetae on the male occiput (in addition to a distinct dorsal row); wing cell m 3 open; prosternum setose medially; metanepisternum with setal patch; two pairs of scutellar macrosetae; posterior surface of mid coxa with, at most, just a couple of setae present. The male gonocoxites are mostly covered in white lanceolate setae, but with a small raised outer gonocoxal process with 2 – 4 large black macrosetae. This condition is also found in the closely related Notiothereva (Webb, 2005 a) and while it is common in Therevidae to have enlarged posterolateral macrosetae on the gonocoxites, it is distinctive here as they are in stark contrast to the abundant white lanceolate setae. Amplisegmentum can be specifically separated from Notiothereva by the presence of scattered macrosetae on the occiput in both sexes (single postocular row in male of Notiothereva), bulbous scape (narrow cylindrical in Notiothereva), mid coxae largely lacking setae (present in Notiothereva) and male eyes separated medially by at least the width of the anterior ocellus (contiguous medially below the medial ocellus in Notiothereva). Note that Webb (2005 b) described that the wing cell m 3 is closed in Amplisegmentum, but it is open in both species (Fig. 4). In addition, the male occiput is described as consisting of a single row of postocular black macrosetae in Webb (2015 b), but there are also black macrosetae scattered medially on the occiput, similar to the condition found in A. venezuelensis sp. n. Included species. Amplisegmentum ecuadorensis Webb; A. venezuelensis sp. n.	en	Winterton, Shaun L. (2021): A new species of Amplisegmentum Webb (Diptera: Therevidae) from Venezuela. Zootaxa 4927 (4): 576-582, DOI: 10.11646/zootaxa.4927.4.8
FE36014CFFA1FFF67CC9D892FD3BF86B.taxon	description	(Figs 1 – 4) https: // zoobank. org / urn: lsid: zoobank. org: act: 50 AB 146 A-D 8 E 9 - 4478 - A 7 D 5 - A 6 C 88 D 24 A 973	en	Winterton, Shaun L. (2021): A new species of Amplisegmentum Webb (Diptera: Therevidae) from Venezuela. Zootaxa 4927 (4): 576-582, DOI: 10.11646/zootaxa.4927.4.8
FE36014CFFA1FFF67CC9D892FD3BF86B.taxon	diagnosis	Diagnosis. Parafacial setae absent; palpus with greatly elongate setal pile; male eye slightly wider than ocellar tubercle; dorsocentral setae absent; single supra-alar macroseta present; mid coxa with posterior surface lacking setae; wing hyaline with dark infuscation at apices of basal-medial, subcostal and discal cells; male phallus with dorsal apodeme tapered anteriorly, distiphallus directed slightly dorsally.	en	Winterton, Shaun L. (2021): A new species of Amplisegmentum Webb (Diptera: Therevidae) from Venezuela. Zootaxa 4927 (4): 576-582, DOI: 10.11646/zootaxa.4927.4.8
FE36014CFFA1FFF67CC9D892FD3BF86B.taxon	description	Description. Body length: 7.2 mm (male). Head with greyish-silver pubescence, darker band across frons (Fig. 2) admixed with erect dark filiform setae; frons wider than ocellar tubercle at narrowest point; parafacial without setae; occiput with sparse silver pubescence and elongate, curved, black macrosetae scattered medially and concentrated as a single row along postocular ridge, setae finer and pale laterally to gena; gena with sparse pile of fine white setae and small patch of short dark setae ventrally, adjacent to parafacial; labellum dark; palpus with sparse but greatly elongate, dark, filiform setae; antenna dark, scape with greyish-silver pubescence and with robust, black macrosetae, larger dorsally; pedicel with greyish-silver pubescence and band of small dark setae; flagellum dark brown. Thorax overlain with greyish-silver pubescence, scutum with brown pubescence in broadly tessellate pattern, admixed with elongate, erect, black, filiform setae; scutal macrosetae black; pleuron and scutellum with sparse pile of elongate, white, lanceolate setae; coxae and femora silver pubescent with pale lanceolate and filiform setae, macrosetae black; femora ground colour dark with silver pubescence, short adpressed, lanceolate setae white dorsally, longer and erect lanceolate setae ventrally admixed with black and white filiform setae; legs with all macrosetae black; tibiae dark yellow to brown apically; tarsi brown; haltere brown; wing (Fig. 1) hyaline, venation brown with slight brownish infuscation on membrane along veins, brown infuscate along anterior margin of wing, as well as brown spots distally on basal medial and discal cells. Abdomen with dense grey velutinous pubescence, admixed with white, lanceolate setae on all segments tergites 2 – 5, more elongate laterally, terminalia dark yellow (Fig. 3). Male genitalia (Figs 3, 4) with epandrium quadrangular, posterolateral corners with broad flange, elongate pale lanceolate setae longer laterally; cerci rounded with short filiform setae; gonocoxite broadly rounded, quadrangular posteriorly with small outer gonocoxal process with three large, black, posterolaterally directed setae, remaining setae all white lanceolate; gonocoxal apodeme very small; hypandrium absent; ventral lobe elongate, narrowed distally; inner gonocoxal apodeme elongate, truncate apically with elongate setae; gonostylus robust with strong keel and acuminate apex, filiform seta present medially; phallus with dorsal apodeme of parameral sheath large, flattened in profile; ventral apodeme relatively large, narrow, recurved dorsally; distiphallus narrow and slightly curved dorsally; ejaculatory apodeme narrow, relatively small. Female unknown.	en	Winterton, Shaun L. (2021): A new species of Amplisegmentum Webb (Diptera: Therevidae) from Venezuela. Zootaxa 4927 (4): 576-582, DOI: 10.11646/zootaxa.4927.4.8
FE36014CFFA1FFF67CC9D892FD3BF86B.taxon	etymology	Etymology. The species epithet refers to the collecting locality in Venezuela.	en	Winterton, Shaun L. (2021): A new species of Amplisegmentum Webb (Diptera: Therevidae) from Venezuela. Zootaxa 4927 (4): 576-582, DOI: 10.11646/zootaxa.4927.4.8
FE36014CFFA1FFF67CC9D892FD3BF86B.taxon	discussion	Comments. This species is known only from a single male specimen in relatively poor condition. Amplisegmentum venezuelensis sp. n. is easily distinguished externally from A. ecuadorensis by the lack of parafacial setae and dorsocentral macrosetae, the male frons wider than the anterior ocellus and small wing spots at the apices of the basal medial and discal cells.	en	Winterton, Shaun L. (2021): A new species of Amplisegmentum Webb (Diptera: Therevidae) from Venezuela. Zootaxa 4927 (4): 576-582, DOI: 10.11646/zootaxa.4927.4.8
FE36014CFFA1FFF67CC9D892FD3BF86B.taxon	materials_examined	Type material. Holotype male, VENEZUELA: Mérida: Puente Real [8.4823, - 71.3900], 6. i. 1982, C. Boudón les. (California State Collection of Arthropods).	en	Winterton, Shaun L. (2021): A new species of Amplisegmentum Webb (Diptera: Therevidae) from Venezuela. Zootaxa 4927 (4): 576-582, DOI: 10.11646/zootaxa.4927.4.8
