identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
FD7487B1FF90FFDCFF205EA73978FDA7.text	FD7487B1FF90FFDCFF205EA73978FDA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius Chamberlin 1929	<div><p>Genus Tyrannochthonius Chamberlin, 1929</p><p>Type species: Chthonius terribilis With, 1906, by original designation.</p></div>	https://treatment.plazi.org/id/FD7487B1FF90FFDCFF205EA73978FDA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF90FFD6FF205D5138E1FDAB.text	FD7487B1FF90FFD6FF205D5138E1FDAB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius acutus Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius acutus sp. nov. ‹ḰẊƟṁae›</p><p>Figs 2–6</p><p>Type material. Holotype ♂ (Ps.- MHBU-GZKL-19-23-01): China, Guizhou Province, Kaili City, Zhouxi Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.927284&amp;materialsCitation.latitude=26.482992" title="Search Plazi for locations around (long 107.927284/lat 26.482992)">Gazuida Cave</a>, under stones within 20–200 m from the cave entrance (Temperature: 16°C, Humidity: 70%) [26°28′58.77″N, 107°55′38.24″E], 716 m a.s.l., 12 October 2019, Zegang Feng &amp; Lingchen Zhao leg. Paratypes: 2 ♂ (Ps.- MHBU-GZKL-19-23-02 &amp; GZKL-19-23-03), 4 ♀ (Ps.- MSWU-GZKL-19-23-04 – GZKL-19-23-07), all with the same data as the holotype .</p><p>Etymology. The specific name is derived from the Latin word “ acutus ”, meaning acute, which refers to the distinctly sharp epistome.</p><p>Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome distinctly pointed, triangular; posterior margin of carapace with 2 setae; tergites II–VI each with 4 setae. Pedipalps slender, femur 6.58–6.67 (♂), 6.63–6.94 (♀) times longer than broad; chela 7.29–7.65 (♂), 7.38–7.79 (♀) times longer than broad; both chelal fingers with intercalary teeth, movable chelal finger teeth distinctly more retrorse than those of the fixed chelal finger.</p><p>Description. Males (holotype and paratypes) (Figs 3A, 4A–F, 5, 6).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 4C, 5A, C): carapace 0.96–1.00 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome distinctly pointed, triangular, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 10–11 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 5C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 4D, 5B): large, about as long as carapace, 2.42 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 20–23 teeth, fourth one largest; movable finger with 24–26 retrorse contiguous small teeth; galea completely vestigial. Serrula exterior with 24–26 and serrula interior with 14–15 blades. Rallum with 7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 5E).</p><p>Pedipalp (Figs 4A, B, E, 5D, 6A, B): long and slender, trochanter 1.10–1.20, femur 6.58–6.67, patella 2.27– 2.35, chela 7.29–7.65, hand 2.46–2.57 times longer than broad; femur 2.50–2.55 times longer than patella; movable chelal finger 1.92–1.93 times longer than hand and 0.65 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissures present on patella (Figs 4E, 5D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming an oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated at same level as est and distal to b (Fig. 6A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 44–46 macrodenticles, long and pointed, plus 13– 14 intercalary microdenticles, 57–60 in total; movable chelal finger with 43–46 macrodenticles (markedly smaller than teeth on fixed chelal finger), markedly retrorse and pointed, plus 15– 16 intercalary microdenticles, 58–62 in total (Fig. 6A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 4B, 6B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4–5: 4: 4: 4: 4: 4: 4: 4–5: 4–5: 4: T2T: 0, tergites VII–IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 12–13: 12–13: 8–9: 7–8: 7: 7: 7–8: 7–8: 0: 2. Anterior genital operculum with 9–12 setae, genital opening slit-like, with 18–20 marginal setae on each side, 47–50 in total (Fig. 4F).</p><p>Legs (Fig. 6C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.85–1.92 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.26–2.59 times longer than tibia. Femoropatella of leg IV 3.53–3.81 times longer than deep; tibia 6.08–6.45 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.67–4.11 times longer than deep (TS= 0.30), tarsus 14.17–15.50 times longer than deep and 2.51–2.58 times longer than metatarsus (TS= 0.33–0.36). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 3B, 4G). Mostly same as males, but a little larger; with same coxal chaetotaxy as males; tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4: 4–5: 5: 5: 4: T2 T: 0; sternal chaetotaxy IV–XII: 11–12: 8: 7–8: 7: 7–8: 8–9: 7: 0: 2; anterior genital operculum with 10–11 setae, posterior margin with 11–12 marginal setae, 22 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.56–3.78 times longer than deep (TS= 0.31– 0.32), tarsus 13.67–14.67 times longer than deep and 2.56–2.59 times longer than metatarsus (TS= 0.33–0.35) .</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 2.07–2.13. Pedipalps: trochanter 0.22–0.24/0.20 (1.10–1.20), femur 1.20–1.25/0.18–0.19 (6.58–6.67), patella 0.47–0.50/0.20–0.22 (2.27–2.35), chela 1.75–1.76/0.23–0.24 (7.29–7.65), hand 0.59/0.23–0.24 (2.46–2.57), movable chelal finger length 1.13–1.14. Chelicera 0.75–0.80/0.31–0.33 (2.42), movable finger length 0.40–0.42. Carapace 0.65–0.68/0.68 (0.96–1.00). Leg I: trochanter 0.20–0.21/0.18 (1.11–1.17), femur 0.72–0.75/0.10 (7.20–7.50), patella 0.39/0.09 (4.33), tibia 0.32–0.35/0.07 (4.57–5.00), tarsus 0.79–0.83/0.06 (13.17–13.83). Leg IV: trochanter 0.28–0.32/0.18–0.20 (1.56–1.60), femoropatella 1.03–1.13/0.27–0.32 (3.53–3.81), tibia 0.71–0.79/0.11–0.13 (6.08–6.45), metatarsus 0.33–0.37/0.09 (3.67–4.11), tarsus 0.85–0.93/0.06 (14.17–15.50).</p><p>Females: body length 2.16–2.29. Pedipalps: trochanter 0.27/0.20–0.22 (1.23–1.35), femur 1.25–1.26/0.18–0.19 (6.63–6.94), patella 0.50/0.22–0.23 (2.17–2.27), chela 1.77–1.87/0.24 (7.38–7.79), hand 0.59–0.62/0.24 (2.46– 2.58), movable chelal finger length 1.16–1.23. Chelicera 0.82–0.86/0.35–0.36 (2.34–2.39), movable finger length 0.44–0.46. Carapace 0.67–0.69/0.72–0.77 (0.90–0.93). Leg I: trochanter 0.17–0.20/0.16–0.18 (1.06–1.11), femur 0.69–0.72/0.09–0.10 (7.20–7.67), patella 0.37–0.39/0.08–0.09 (4.33–4.63), tibia 0.33–0.34/0.07 (4.71–4.86), tarsus 0.78–0.81/0.06 (13.00–13.50). Leg IV: trochanter 0.26–0.28/0.17 (1.53–1.65), femoropatella 0.99–1.03/0.27 (3.67–3.81), tibia 0.70–0.72/0.11–0.12 (6.00–6.36), metatarsus 0.32–0.34/0.09 (3.56–3.78), tarsus 0.82–0.88/0.06 (13.67–14.67).</p><p>Remarks. Tyrannochthonius acutus sp. nov. is similar to T. hispidus sp. nov. in having intercalary teeth on both chelal fingers and a pointed, triangular epistome, but differs by a larger body size (body length 2.07–2.13 (J), 2.16–2.29 (♀) vs. 1.56–1.78 (J), 1.77–1.99 (♀) mm, chela 7.29–7.65 (J), 7.38–7.79 (♀) vs. 6.68–6.72 (J), 6.20–6.79 (♀) times longer than board), the number of blades of rallum (7 vs. 6) and the presence of more fixed chelal finger teeth (57–60 vs. 42–43).</p><p>Tyrannochthonius acutus sp. nov. can be distinguished from T. chixing, T. harveyi and T. zhai by the number of setae at the anterior margin of the carapace (6 vs. 4) and the presence of intercalary teeth on both chelal fingers, from T. akaelus and T. ganshuanensis by the number of setae on tergite II (4 vs. 2) and from T. antridraconis by the presence of more chelal teeth (57–60 vs. 33–36 teeth on the fixed chelal finger; 58–62 vs. 30–36 teeth on the movable chelal finger) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FF90FFD6FF205D5138E1FDAB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF9AFFC8FF205D7A38E1FCEE.text	FD7487B1FF9AFFC8FF205D7A38E1FCEE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius altus Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius altus sp. nov. ‹ṘḾƟṁae›</p><p>Figs 7–11</p><p>Type material. Holotype ♂ (Ps.- MHBU-HBUARA#2022-493-01): China, Guizhou Province, Hezhang County, Shuitangbao Township, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.67693&amp;materialsCitation.latitude=27.015505" title="Search Plazi for locations around (long 104.67693/lat 27.015505)">Caoziping Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.67693&amp;materialsCitation.latitude=27.015505" title="Search Plazi for locations around (long 104.67693/lat 27.015505)">Shenren Cave</a>, under stones and detritus in the deep zone (Temperature: 13°C, Humidity: 90%) [27°0′55.82″N, 104°40′36.98″E], 2042 m a.s.l., 31 July 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun &amp; Wenlong Fan leg. Paratypes: 4 ♂ (Ps.- MHBU-HBUARA#2022-493-02 – HBUARA#2022-493-05), 7 ♀ (Ps.- MSWU-HBUARA#2022-493-06 – HBUARA#2022-493-12), all with the same data as the holotype .</p><p>Etymology. The specific name is derived from the Latin word “ altus ”, meaning high, which refers to the species that live at high altitudes.</p><p>Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome pointed and triangular; posterior margin of carapace with 2 setae; tergites I–II each with 4 setae. Pedipalps slender, femur 8.50–8.79 (♂), 8.06–8.44 (♀) times longer than broad; chela 8.25–8.57 (♂), 7.41–7.42 (♀) times longer than broad; both chelal fingers with intercalary teeth.</p><p>Description. Males (holotype and paratypes) (Figs 7F, 8A, 9A–F, 10, 11).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 9C, 10A): carapace 0.95–0.97 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome pointed and triangular, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 14–15 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 10C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 9D, 10B): large, about as long as carapace, 2.48–2.63 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 18–20 teeth, distal one largest; movable finger with 18–19 retrorse contiguous small teeth; galea completely vestigial. Serrula exterior with 21–22 and serrula interior with 14–15 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 10E).</p><p>Pedipalp (Figs 9A, B, E, 10D, 11A, B): long and slender, trochanter 1.21–1.22, femur 8.50–8.79, patella 2.14– 2.30, chela 8.25–8.57, hand 2.95–2.96 times longer than broad; femur 2.67–2.89 times longer than patella; movable chelal finger 1.69–1.79 times longer than hand and 0.61–0.62 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 9E, 10D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb midway between st and b; b and t situated subdistally, t slightly situated distal to b; est situated distal to t (Fig. 11A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 24–26 macrodenticles, long and pointed, plus 15– 16 intercalary microdenticles, 39–42 in total; movable chelal finger with 34–35 macrodenticles (markedly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 10– 11 intercalary microdenticles, 44–46 in total (Fig. 11A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 9B, 11B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 2–4: 4: 3–4: 4: 4: 4: 5: 4–5: T2T: 0; tergites IX and Ⅹ each with an unpaired median seta. Sternal chaetotaxy III–XII: 16: 12–14: 8–9: 7–9: 8: 8–9: 8–9: 7–8: 0: 2. Anterior genital operculum with 11–12 setae, genital opening slit-like, with 7–12 marginal setae on each side, 27–35 in total (Fig. 9F).</p><p>Legs (Fig. 11C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.90–1.92 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.32–2.47 times longer than tibia. Femoropatella of leg IV 5.18–5.20 times longer than deep; tibia 6.80–6.91 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.50–4.38 times longer than deep (TS= 0.29–0.32), tarsus 13.83–16.33 times longer than deep and 2.80–2.96 times longer than metatarsus (TS= 0.24–0.27). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 8B, 9G). Mostly same as males, but a little larger; with same coxal chaetotaxy as males; tergal chaetotaxy I–XII: 4: 4: 3–4: 4: 4: 4: 4: 4: 4–5: 4: T2T: 0; sternal chaetotaxy IV–XII: 13: 9–10: 9: 7–10: 8–9: 8–9: 6–7: 0: 2; anterior genital operculum with 10–11 setae, posterior margin with 14–15 marginal setae, 25 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.88–4.13 times longer than deep (TS= 0.29–0.30), tarsus 14.67–15.67 times longer than deep and 2.67–3.03 times longer than metatarsus (TS= 0.25–0.30).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 2.36–2.37. Pedipalps: trochanter 0.22–0.23/0.18–0.19 (1.21–1.22), femur 1.23–1.36/0.14–0.16 (8.50–8.79), patella 0.46–0.47/0.20–0.22 (2.14–2.30), chela 1.80–1.98/0.21–0.24 (8.25–8.57), hand 0.62–0.71/0.21–0.24 (2.95– 2.96), movable chelal finger length 1.11–1.20. Chelicera 0.72–0.79/0.29–0.30 (2.48–2.63), movable finger length 0.40–0.43. Carapace 0.59–0.65/0.62–0.67 (0.95–0.97). Leg I: trochanter 0.17–0.19/0.15–0.18 (1.06–1.13), femur 0.71–0.80/0.09 (7.89–8.89), patella 0.37–0.42/0.08 (4.63–5.25), tibia 0.32–0.37/0.06–0.07 (5.29–5.33), tarsus 0.79– 0.86/0.06 (13.17–14.33). Leg IV: trochanter 0.27–0.30/0.16–0.18 (1.67–1.69), femoropatella 1.04–1.14/0.20–0.22 (5.18–5.20), tibia 0.68–0.76/0.10–0.11 (6.80–6.91), metatarsus 0.28–0.35/0.08 (3.50–4.38), tarsus 0.83–0.98/0.06 (13.83–16.33).</p><p>Females: body length 2.61–2.95. Pedipalps: trochanter 0.22–0.23/0.18–0.19 (1.21–1.22), femur 1.35– 1.37/0.16–0.17 (8.06–8.44), patella 0.51/0.21 (2.43), chela 1.93–2.00/0.26–0.27 (7.41–7.42), hand 0.69–0.71/0.26– 0.27 (2.63–2.65), movable chelal finger length 1.19–1.22. Chelicera 0.79–0.82/0.31–0.32 (2.55–2.56), movable finger length 0.44. Carapace 0.67–0.68/0.68–0.70 (0.97–0.99). Leg I: trochanter 0.19/0.16–0.17 (1.12–1.19), femur 0.78–0.79/0.09–0.10 (7.90–8.67), patella 0.41–0.43/0.08–0.09 (4.78–5.13), tibia 0.34–0.36/0.06 (5.67–6.00), tarsus 0.83–0.84/0.06–0.07 (12.00–13.83). Leg IV: trochanter 0.29/0.16 (1.81), femoropatella 1.10–1.14/0.21–0.22 (5.18–5.24), tibia 0.74–0.76/0.11–0.12 (6.33–6.73), metatarsus 0.31–0.33/0.08 (3.88–4.13), tarsus 0.88–0.94/0.06 (14.67–15.67).</p><p>Remarks. Tyrannochthonius altus sp. nov. is similar to T. babaowanensis sp. nov. in having intercalary teeth on both chelal fingers and a pointed, triangular epistome, but differs by the presence of a slender chela (chela 8.25–8.57 (♂) vs. 7.27 (♂) times longer than board), the number of setae on tergite Ⅹ (4–5 vs. 2) and the number of blades of coxal spines (14–15 vs. 8–9).</p><p>Tyrannochthonius altus sp. nov. can be distinguished from T. akaelus, T. ganshuanensis and T. harveyi by the number of setae on the tergites I–II (4 vs. 2), from</p><p>T. chixing and T. zhai by the number of setae at the anterior margin of the carapace (6 vs. 4), a larger body size (body length min. 2.36 mm vs. max. 1.64 mm) and the presence of intercalary teeth on both chelal fingers, from T. antridraconis by the relative position of trichobothrium sb (sb midway between b and st vs. nearer b than st) and the presence of a slender chela (chela 8.25–8.57 (♂) vs. 7.90–8.00 (♂) times longer than board)(Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FF9AFFC8FF205D7A38E1FCEE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF84FFC2FF205CBE38E1FBEA.text	FD7487B1FF84FFC2FF205CBE38E1FBEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius arificus Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius arificus sp. nov. ‹ȐƮƟṁae›</p><p>Figs 12–16</p><p>Type material. Holotype ♂ (Ps.- MHBU-HBUARA#2022-506 - 01): China, Guizhou Province, Anlong County, Dushan Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.600746&amp;materialsCitation.latitude=25.312933" title="Search Plazi for locations around (long 105.600746/lat 25.312933)">Polao Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.600746&amp;materialsCitation.latitude=25.312933" title="Search Plazi for locations around (long 105.600746/lat 25.312933)">Hei Cave</a>, under stones and clods in the deep zone (Temperature: 16°C, Humidity: 65%) [25°18′46.56″N, 105°36′2.69″E], 1067 m a.s.l., 6 August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun &amp; Wenlong Fan leg. Paratypes: 3 ♂ (Ps.- MHBU-HBUARA#2022-506 - 02 –HBUARA#2022-506-04), 3 ♀ (Ps.-MSWU-HBUARA#2022-506-05–HBUARA#2022-506-07), all with the same data as the holotype .</p><p>Etymology. The specific name is derived from the Latin word “ arificus ”, meaning arid, which refers to the species that lives in a dry environment.</p><p>Diagnosis (J ♀). Small-sized troglomorphic species with slightly elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, rounded and obtuse; posterior margin of carapace with 2 setae; tergites I–Ⅴ each with 4 setae. Palpal femur 5.27–5.55 (♂), 5.36–5.50 (♀) times longer than broad; chela 6.29–6.62 (♂), 6.07–6.20 (♀) times longer than broad; both chelal fingers with intercalary teeth.</p><p>Description. Males (holotype and paratypes) (Figs 13A, 14A–F, H, 15, 16).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 14C, D, 15A): carapace 0.93–0.95 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small, rounded and obtuse; with 17–18 setae arranged s3–4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 3–4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 10 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 15C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 14E, 15B): large, about as long as carapace, 2.05–2.20 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on dorsal side. Both fingers well provided with teeth, fixed finger with 11–12 teeth, distal one largest; movable finger with 11–13 retrorse contiguous small teeth; galea completely vestigial (Fig. 15B). Serrula exterior with 18–20 and serrula interior with 11–13 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 15E).</p><p>Pedipalp (Figs 14A, B, H, 16A, B): long and slender, trochanter 1.15–1.25, femur 5.27–5.55, patella 2.15–2.17, chela 6.29–6.62, hand 2.07–2.15 times longer than broad; femur 2.18–2.23 times longer than patella; movable chelal finger 2.10–2.14 times longer than hand and 0.69–0.70 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 14H, 15D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated distal to b and at same level as est (Fig. 16A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 30–32 macrodenticles, slightly retrorse and pointed, plus 25– 26 intercalary microdenticles, 56–57 in total; movable chelal finger with 12–13 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 10– 11 intercalary microdenticles and 17–19 vestigial, rounded and contiguous basal teeth, 41 in total (Fig. 16A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 14B, 16B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 5: 5: 5: 5: 3: T2T: 0, tergites VI–IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 13–15: 13–14: 10: 9: 9: 9: 9–10: 9: 0: 2. Anterior genital operculum with 10–11 setae, genital opening slit-like, with 11–14 marginal setae on each side, 35–36 in total (Fig. 14F).</p><p>Legs (Fig. 16C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.76–1.89 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.22–2.37 times longer than tibia. Femoropatella of leg IV 2.84–3.06 times longer than deep; tibia 4.78–5.00 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.17 times longer than deep (TS= 0.26–0.37), tarsus 11.75–14.67 times longer than deep and 2.32–2.47 times longer than metatarsus (TS= 0.25–0.34). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 13B, 14G). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; with same tergal chaetotaxy as males; sternal chaetotaxy IV–XII: 14–15: 10: 9–10: 9: 9–10: 9–11: 8–9: 0: 2; anterior genital operculum with 10–11 setae, posterior margin with 14 marginal setae, 24–25 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.00–3.50 times longer than deep (TS= 0.33), tarsus 12.00–14.67 times longer than deep and 2.29–2.44 times longer than metatarsus (TS= 0.27–0.29).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 1.31–1.40. Pedipalps: trochanter 0.15/0.12–0.13 (1.15–1.25), femur 0.58–0.61/0.11 (5.27–5.55), patella 0.26–0.28/0.12–0.13 (2.15–2.17), chela 0.86–0.88/0.13–0.14 (6.29–6.62), hand 0.28–0.29/0.13–0.14 (2.07–2.15), movable chelal finger length 0.60–0.61. Chelicera 0.41–0.44/0.20 (2.05–2.20), movable finger length 0.23–0.24. Carapace 0.41–0.43/0.43–0.46 (0.93–0.95). Leg I: trochanter 0.12/0.10 (1.20), femur 0.36–0.37/0.06–0.07 (5.14– 6.17), patella 0.19–0.21/0.05 (3.80–4.20), tibia 0.18–0.19/0.04 (4.50–4.75), tarsus 0.40–0.45/0.03–0.04 (11.25– 13.33). Leg IV: trochanter 0.16–0.17/0.11–0.12 (1.33–1.55), femoropatella 0.52–0.54/0.17–0.19 (2.84–3.06), tibia 0.40–0.43/0.08–0.09 (4.78–5.00), metatarsus 0.19/0.06 (3.17), tarsus 0.44–0.47/0.03–0.04 (11.75–14.67).</p><p>Females: body length 1.62–1.69. Pedipalps: trochanter 0.16–0.17/0.12–0.14 (1.21–1.33), femur 0.59–0.66/0.11– 0.12 (5.36–5.50), patella 0.30–0.32/0.13–0.15 (2.13–2.31), chela 0.91–0.93/0.15 (6.07–6.20), hand 0.31/0.15 (2.07), movable chelal finger length 0.62–0.63. Chelicera 0.45–0.47/0.21–0.23 (2.04–2.14), movable finger length 0.24–0.25. Carapace 0.45–0.48/0.46–0.49 (0.98). Leg I: trochanter 0.12–0.13/0.10–0.11 (1.18–1.20), femur 0.37– 0.39/0.06–0.07 (5.57–6.17), patella 0.19–0.22/0.06 (3.17–3.67), tibia 0.19–0.20/0.04–0.05 (4.00–4.75), tarsus 0.44– 0.45/0.04 (11.00–11.25). Leg IV: trochanter 0.18/0.11 (1.64), femoropatella 0.50–0.56/0.17–0.19 (2.94–2.95), tibia 0.39–0.45/0.08–0.09 (4.88–5.00), metatarsus 0.18–0.21/0.06 (3.00–3.50), tarsus 0.44–0.48/0.03–0.04 (12.00–14.67).</p><p>Remarks. Tyrannochthonius arificus sp. nov. is similar to T. yanshanensis sp. nov. in having intercalary teeth on both chelal fingers and a small, rounded and obtuse epistome, but differs by the larger body size (body length 1.62–1.69 (♀) vs. 1.29–1.33 (♀) mm; chela 6.07–6.20 (♀) vs. 5.69–5.75 (♀) times longer than board), the number of setae on tergite Ⅴ (4 vs. 5) and tergite Ⅹ (3 vs. 5), more intercalary teeth on the fixed chelal finger (25–26 vs. 14), the number of blades of coxal spines (10 vs. 7–8) and the number of blades of rallum (8 vs. 7).</p><p>Tyrannochthonius arificus sp. nov. can be distinguished from T. akaelus, T. ganshuanensis and T. harveyi by the number of setae on the tergites I–II (4 vs. 2), from</p><p>T. chixing and T. zhai by the number of setae at the anterior margin of the carapace (6 vs. 4), a shorter palp (chela length max. 0.93 vs. min. 1.21 mm; palpal femur length max. 0.66 vs. min. 0.84 mm) and the presence of intercalary teeth on both chelal fingers, from T. antridraconis by a smaller size (body length max. 1.40 vs. min. 1.80 mm; chela length max. 0.88 vs. min. 1.70 mm) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FF84FFC2FF205CBE38E1FBEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF8EFFC4FF205BBA38E1FB92.text	FD7487B1FF8EFFC4FF205BBA38E1FB92.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius babaowanensis Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius babaowanensis sp. nov. (ÃÍẠƟṁae)</p><p>Figs 17–21</p><p>Type material. Holotype ♂ (Ps.-MHBU-GZYJ-20-14-01): China, Guizhou Province, Yinjiang County, Zhongxing Sub-district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.36469&amp;materialsCitation.latitude=28.064741" title="Search Plazi for locations around (long 108.36469/lat 28.064741)">Tianchi Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.36469&amp;materialsCitation.latitude=28.064741" title="Search Plazi for locations around (long 108.36469/lat 28.064741)">Babaowan Cave</a>, on a stone wall in the deep zone (Temperature: 14°C, Humidity: 85%) [28°3′53.07″N, 108°21′52.89″E], 655 m a.s.l., 23 August 2020, Zegang Feng, Hongru Xu &amp; Yanmeng Hou leg.</p><p>Etymology. Named after the type locality, Babaowan Cave.</p><p>Diagnosis (♂). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, pointed, triangular; posterior margin of carapace with 2 setae; tergites I–VI each with 4 setae. Pedipalps slender, femur 7.67 times longer than broad; chela 7.27 times longer than broad; both chelal fingers with intercalary teeth (present in the distal 1/3 of the fingers); chemosensory setae present on dorsum of chelal hand.</p><p>Description. Male (holotype), female unknown (Figs 17F, 18–21).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 19C, 20A): carapace 1.04 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome pointed and small, triangular, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 8–9 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 20C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 19D, 20B): large, about as long as carapace, 2.37 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 14 teeth, distal one largest; movable finger with 13 retrorse contiguous small teeth; galea completely vestigial (Fig. 20B). Serrula exterior with 21 and serrula interior with 14 blades. Rallum with 7 blades, the distal one longer and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 20E).</p><p>Pedipalp (Figs 19A, B, F, 21A, B): long and slender, trochanter 1.22, femur 7.67, patella 2.72, chela 7.27, hand 2.41 times longer than broad; femur 2.35 times longer than patella; movable chelal finger 2.06 times longer than hand and 0.68 times longer than chela. Setae generally long and acuminate; one distal lyrifissures present on patella (Figs 19F, 20D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, b situated at same level as est, t slightly distal to it (Fig. 21A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 42 macrodenticles, slightly retrorse and pointed, plus 15 intercalary microdenticles (present in the distal 1/3 of the fingers, till est), 57 in total; movable chelal finger with 31 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 11 intercalary microdenticles (present in the distal 1/3 of the fingers, till b) and 14 vestigial, rounded and contiguous basal teeth, 56 in total (Fig. 21A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 19B, 21B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4: 5: 5: 5: 2: T2T: 0. Sternal chaetotaxy III–XII: 14: 12: 7: 7: 8: 9: 9: 7: 0: 2. Anterior genital operculum with 13 setae, genital opening slit-like, with 14–15 marginal setae on each side, 42 in total (Fig. 19E).</p><p>Legs (Fig. 21C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.97 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.43 times longer than tibia. Femoropatella of leg IV 4.21 times longer than deep; tibia 7.00 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.75 times longer than deep (TS= 0.33), tarsus 14.00 times longer than deep and 2.80 times longer than metatarsus (TS= 0.32). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Male: body length 2.35. Pedipalps: trochanter 0.22/0.18 (1.22), femur 1.15/0.15 (7.67), patella 0.49/0.18 (2.72), chela 1.60/0.22 (7.27), hand 0.53/0.22 (2.41), movable chelal finger length 1.09. Chelicera 0.64/0.27 (2.37), movable finger length 0.35. Carapace 0.59/0.57 (1.04). Leg I: trochanter 0.19/0.15 (1.27), femur 0.69/0.08 (8.63), patella 0.35/0.07 (5.00), tibia 0.30/0.06 (5.00), tarsus 0.73/0.06 (12.17). Leg IV: trochanter 0.27/0.16 (1.69), femoropatella 1.01/0.24 (4.21), tibia 0.70/0.10 (7.00), metatarsus 0.30/0.08 (3.75), tarsus 0.84/0.06 (14.00).</p><p>Remarks. Female unknown. Tyrannochthonius babaowanensis sp. nov. is similar to T. antridraconis in having intercalary teeth on both chelal fingers and a small, rounded and obtuse epistome, but differs by a larger body size (body length 1.62–1.69 (♂) vs. 1.29–1.33 (♂) mm), the number of setae on tergite VII (5 vs. 4) and tergite Ⅹ (2 vs. 4–5), the proportion of intercalary teeth on the fixed finger (1/3 vs. 2/3), the number of blades of coxal spines (8–9 vs. 12–15) and the number of blades of rallum (7 vs. 8).</p><p>Tyrannochthonius babaowanensis sp. nov. can be distinguished from T. chixing and T. zhai by the larger size (body length 2.35 vs. max. 1.64 mm; chela length 1.60 vs. max. 1.24 mm), the number of setae at the anterior margin of the carapace (6 vs. 4) and the presence of intercalary teeth on both chelal fingers, from T. akaelus, T. ganshuanensis and T. harveyi by the number of setae on the tergites I–II (4 vs. 2) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FF8EFFC4FF205BBA38E1FB92	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF88FFFEFF205B6038E1FEF2.text	FD7487B1FF88FFFEFF205B6038E1FEF2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius breviculus Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius breviculus sp. nov. ‹ÑṚƟṁae›</p><p>Figs 22–26</p><p>Type material. Holotype ♂ (Ps.- MHBU-HBUARA#2021-202 - 01): China, Guizhou Province, Libo County, Maolan Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.061134&amp;materialsCitation.latitude=25.378325" title="Search Plazi for locations around (long 108.061134/lat 25.378325)">Yaochao Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.061134&amp;materialsCitation.latitude=25.378325" title="Search Plazi for locations around (long 108.061134/lat 25.378325)">Yanjia Cave</a>, under stones in the deep zone (Temperature: 13°C, Humidity: 90%) [25°22′41.97″N, 108°3′40.08″E], 609.5 m a.s.l., 1 August 2021, Zegang Feng, Hongru Xu, Liu Fu &amp; Nana Zhan leg. Paratypes: 2 ♂ (Ps.- MHBU-HBUARA#2021-202 - 02 &amp; HBUARA#2021-202-03), 2 ♀ (Ps.-MSWU-HBUARA#2021-202-04 &amp; HBUARA#2021-202-05), all with the same data as the holotype .</p><p>Etymology. The specific name is derived from the Latin word “ breviculus ”, meaning short, which refers to the short pedipalps.</p><p>Diagnosis (J ♀). Small-sized troglomorphic species with slightly elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, rounded and obtuse; posterior margin of carapace with 2 setae; tergites I–Ⅴ each with 4 setae (more variation in females). Palpal femur 5.10–5.33 (♂), 5.00–5.30 (♀) times longer than broad; chela 5.69–5.85 (♂), 5.57–5.64 (♀) times longer than broad; both chelal fingers with intercalary teeth; chemosensory setae present on dorsum of chelal hand.</p><p>Description. Males (holotype and paratypes) (Figs 23A, 24A–C, E, F, H).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 24C, 25A): carapace 0.95–0.98 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small, rounded and obtuse; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 8–9 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 25C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 24E, 25B): large, about as long as carapace, 2.05–2.11 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation both dorsal and ventral side. Both fingers well provided with teeth, fixed finger with 10–11 teeth, distal one largest; movable finger with 13–15 retrorse contiguous small teeth; galea completely vestigial (Fig. 25B). Serrula exterior with 17–18 and serrula interior with 12–14 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 25E).</p><p>Pedipalp (Figs 24A, B, H, 25D, 26A, B): long and slender, trochanter 1.00–1.18, femur 5.10–5.33, patella 2.00–2.09, chela 5.69–5.85, hand 1.85–1.92 times longer than broad; femur 2.09–2.13 times longer than patella; movable chelal finger 2.12–2.13 times longer than hand and 0.69–0.70 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 24H, 25D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb midway between st and b; b and t situated subdistally, t situated distal to b and midway between it and est; est situated distal to b (Fig. 26A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 29–30 macrodenticles, slightly retrorse and pointed, plus 13– 15 intercalary microdenticles, 42–45 in total; movable chelal finger with 9–10 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 8 intercalary microdenticles and 14–16 vestigial, rounded and contiguous basal teeth, 32–33 in total (Fig. 26A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 24B, 26B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 5: 5: 5: 5: 4: T2T: 0, tergites VI–IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 13–14: 11–13: 8–9: 9: 9: 8–9: 9: 9–10: 0: 2. Anterior genital operculum with 10–11 setae, genital opening slit-like, with 10–12 marginal setae on each side, 32–33 in total (Fig. 24F).</p><p>Legs (Fig. 26C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.87–2.00 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.00–2.27 times longer than tibia. Femoropatella of leg IV 2.67 times longer than deep; tibia 4.00–4.43 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 2.80 times longer than deep (TS= 0.36), tarsus 11.33–11.67 times longer than deep and 2.43–2.50 times longer than metatarsus (TS= 0.24–0.29). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 23B, 24D, G). Mostly same as males, but a little larger; with same chaetotaxy of coxae; tergal chaetotaxy I–XII: 3–4: 2–4: 4–5: 4–5: 5: 5: 5: 5: 5: 4: T2T: 0; sternal chaetotaxy IV–XII: 12–13: 7–9: 8–9: 9: 9: 10: 9: 0: 2; anterior genital operculum with 10 setae, posterior margin with 14–16 marginal setae, 24–26 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 2.33–2.80 times longer than deep (TS= 0.36–0.43), tarsus 11.00–12.00 times longer than deep and 2.36–2.57 times longer than metatarsus (TS= 0.24–0.25).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 1.24–1.26. Pedipalps: trochanter 0.11–0.13/0.11 (1.00–1.18), femur 0.48–0.51/0.09–0.10 (5.10–5.33), patella 0.23–0.24/0.11–0.12 (2.00–2.09), chela 0.74–0.76/0.13 (5.69–5.85), hand 0.24–0.25/0.13 (1.85–1.92), movable chelal finger length 0.51–0.53. Chelicera 0.38–0.39/0.18–0.19 (2.05–2.11), movable finger length 0.20. Carapace 0.38–0.40/0.40–0.41 (0.95–0.98). Leg I: trochanter 0.10–0.11/0.09 (1.11–1.22), femur 0.28/0.05–0.06 (4.67–5.60), patella 0.14–0.15/0.05 (2.80–3.00), tibia 0.15–0.16/0.04 (3.75–4.00), tarsus 0.32–0.34/0.03 (10.67–11.33). Leg IV: trochanter 0.13–0.14/0.09–0.10 (1.40–1.44), femoropatella 0.40/0.15 (2.67), tibia 0.28–0.31/0.07 (4.00–4.43), metatarsus 0.14/0.05 (2.80), tarsus 0.34–0.35/0.03 (11.33–11.67).</p><p>Females: body length 1.33–1.44. Pedipalps: trochanter 0.12–0.14/0.11–0.12 (1.08–1.09), femur 0.53–0.55/0.10– 0.11 (5.00–5.30), patella 0.25–0.26/0.13 (1.92–2.00), chela 0.78–0.79/0.14 (5.57–5.64), hand 0.26–0.27/0.14 (1.86– 1.93), movable chelal finger length 0.53–0.55. Chelicera 0.42–0.43/0.20–0.21 (2.05–2.10), movable finger length 0.22–0.23. Carapace 0.40–0.42/0.42–0.45 (0.93–0.95). Leg I: trochanter 0.11/0.09–0.10 (1.10–1.22), femur 0.27– 0.30/0.06 (4.50–5.00), patella 0.14–0.16/0.05 (2.80–3.20), tibia 0.17/0.04 (4.25), tarsus 0.33–0.35/0.04 (8.25–8.75). Leg IV: trochanter 0.12–0.14/0.09–0.10 (1.33–1.40), femoropatella 0.42–0.46/0.14–0.15 (2.80–3.29), tibia 0.32– 0.33/0.07 (4.57–4.71), metatarsus 0.14/0.05–0.06 (2.33–2.80), tarsus 0.33–0.36/0.03 (11.00–12.00).</p><p>Remarks. Tyrannochthonius breviculus sp. nov. is similar to T. yanshanensis sp. nov. in having intercalary teeth on both chelal fingers and a small, rounded and obtuse epistome, but differs by a shorter palp (chela length 0.78–0.79 (♀) vs. 0.91–0.92 (♀) mm; palpal femur 5.00–5.30 (♀) vs. 4.92–5.08 (♀) times longer than board, length 0.53–0.55 (♀) vs. 0.59–0.61 (♀) mm; movable chelal finger 2.04 (♀) vs. 2.17–2.21 (♀) times longer than hand), the number of setae on the tergites VIII‒IX (5 vs. 6), the relative position of trichobothrium sb (sb midway between st and b vs. nearer st than b) and the number of blades of rallum (8 vs. 7).</p><p>Tyrannochthonius breviculus sp. nov. can be distinguished from T. chixing, T. harveyi and T. zhai by the presence of intercalary teeth on both chelal fingers, from T. akaelus, T. antridraconis and T. ganshuanensis by a shorter palp (chela length max. 0.79 vs. min. 1.21 mm; palpal femur length max. 0.55 vs. min. 0.90 mm) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FF88FFFEFF205B6038E1FEF2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFB2FFFCFF205E9D38E1F826.text	FD7487B1FFB2FFFCFF205E9D38E1F826.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius brevispinus Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius brevispinus sp. nov. ‹ÑḦƟṁae›</p><p>Figs 27–31</p><p>Type material. Holotype ♂ (Ps.- MHBU-HBUARA#2022-518-01): China, Guizhou Province, Danzhai County, Longquan Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.80697&amp;materialsCitation.latitude=26.13776" title="Search Plazi for locations around (long 107.80697/lat 26.13776)">Jinguadong Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.80697&amp;materialsCitation.latitude=26.13776" title="Search Plazi for locations around (long 107.80697/lat 26.13776)">Jingua Cave</a>, under stones about 20 m from the cave entrance (Temperature: 12°C, Humidity: 90%) [26°8′15.94″N, 107°48′25.09″E], 749 m a.s.l., 12 August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun &amp; Wenlong Fan leg. Paratype: 1 ♀ (Ps.- MSWU-HBUARA#2022-518-02), with the same data as the holotype .</p><p>Etymology. The specific name is derived from a combination of the Latin word “ brevis ” and “ spineus ”, meaning short and spined, respectively, which refers to the short coxal spines.</p><p>Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin and finely denticulated, epistome small and represented by a slight bump; posterior margin of the carapace with 2 setae; tergites I–II each with 2 setae. Pedipalps slender, femur 6.64 (♂), 6.67 (♀) times longer than broad; chela 7.39 (♂), 7.53 (♀) times longer than broad; both chelal fingers without intercalary teeth, movable chelal finger teeth distinctly smaller and denser than fixed chelal finger teeth; chemosensory setae present on dorsum of chelal hand; coxal spines short.</p><p>Description. Male (holotype) (Figs 28A, 29A–F, 30, 31).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 29C, 30A): carapace 1.02 times longer than broad, slightly narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and represented by a slight bump, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 8–9 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 30C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 29D, 30B): large, about as long as carapace, 2.36 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 12 teeth, distal one largest; movable finger with 16 retrorse contiguous small teeth; galea represented by a very slight bump on movable finger (Fig. 30B). Serrula exterior with 17 and serrula interior with 13 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 30E).</p><p>Pedipalp (Figs 29A, B, E, 31A, B): long and slender, trochanter 1.33, femur 6.64, patella 2.13, chela 7.39, hand 2.39 times longer than broad; femur 2.74 times longer than patella; movable chelal finger 2.05 times longer than hand and 0.66 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 29E, 30D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb midway between st and b; b and t situated subdistally, t situated distal to b and proximal to est (Fig. 31A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 26 macrodenticles, slightly retrorse and pointed; movable chelal finger with 18 macrodenticles (markedly smaller than teeth on fixed chelal finger), continuous, retrorse and pointed, plus 18 small, rounded and contiguous basal teeth, 36 in total (Fig. 31A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 29B, 31B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 2: 2: 4: 4: 4: 4: 4: 5: 5: 4: T2T: 0. Sternal chaetotaxy III–XII: 12: 12: 7: 8: 9: 9: 9: 9: 0: 2. Anterior genital operculum with 10 setae, genital opening slit-like, with 12–14 marginal setae on each side, 36 in total (Fig. 29F).</p><p>Legs (Fig. 31C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.76 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.32 times longer than tibia. Femoropatella of leg IV 3.43 times longer than deep; tibia 5.67 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.29 times longer than deep (TS= 0.26), tarsus 12.80 times longer than deep and 2.78 times longer than metatarsus (TS= 0.33). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult female (paratype) (Figs 28B, 29G). Mostly same as male, but a little larger; chaetotaxy of coxae: P 3, I 3, II 3–4, III 5, IV 5; with same tergal chaetotaxy as male; sternal chaetotaxy IV–XII: 12: 8: 9: 9: 9: 9: 9: 0: 2; anterior genital operculum with 10 setae, posterior margin with 12 marginal setae, 22 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 4.00 times longer than deep (TS= 0.25), tarsus 12.40 times longer than deep and 2.58 times longer than metatarsus (TS= 0.31).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Male: body length 1.73. Pedipalps: trochanter 0.20/0.15 (1.33), femur 0.93/0.14 (6.64), patella 0.34/0.16 (2.13), chela 1.33/0.18 (7.39), hand 0.43/0.18 (2.39), movable chelal finger length 0.88. Chelicera 0.52/0.22 (2.36), movable finger length 0.28. Carapace 0.49/0.48 (1.02). Leg I: trochanter 0.14/0.13 (1.08), femur 0.51/0.07 (7.29), patella 0.29/0.07 (4.14), tibia 0.25/0.05 (5.00), tarsus 0.58/0.05 (11.60). Leg IV: trochanter 0.22/0.14 (1.57), femoropatella 0.72/0.21 (3.43), tibia 0.51/0.09 (5.67), metatarsus 0.23/0.07 (3.29), tarsus 0.64/0.05 (12.80).</p><p>Female: body length 2.12. Pedipalps: trochanter 0.21/0.16 (1.31), femur 1.00/0.15 (6.67), patella 0.36/0.17 (2.12), chela 1.43/0.19 (7.53), hand 0.47/0.19 (2.47), movable chelal finger length 0.94. Chelicera 0.58/0.24 (2.42), movable finger length 0.30. Carapace 0.55/0.54 (1.02). Leg I: trochanter 0.14/0.14 (1.00), femur 0.56/0.08 (7.00), patella 0.28/0.07 (4.00), tibia 0.26/0.05 (5.20), tarsus 0.62/0.05 (12.40). Leg IV: trochanter 0.23/0.13 (1.77), femoropatella 0.73/0.21 (3.48), tibia 0.51/0.09 (5.67), metatarsus 0.24/0.06 (4.00), tarsus 0.62/0.05 (12.40).</p><p>Remarks. Tyrannochthonius brevispinus sp. nov. is similar to T. oblongus sp. nov. in having a small and slight bump epistome and the same number of setae on tergites I–II (2), but differs by a smaller size (body length 1.73 (♂) vs. 2.18–2.26 (♂) mm; chela 7.39 (♂), 7.53 (♀) vs. 7.27 (♂), 6.78–7.09 (♀) times longer than board, length 1.33 (♂), 1.43 (♀) vs. 1.60 (♂) mm, 1.56 (♀) mm; palpal femur length 0.93 (♂), 1.00 (♀) vs. 1.11–1.17 (♂), 1.09–1.17 (♀) mm), the proportion of movable chelal finger and chelal hand (movable chelal finger 2.05 (♂), 2.00 (♀) vs. 1.74–1.79 (♂), 1.74–1.75 (♀) times longer than hand), the relative position of trichobothrium sb (sb midway between b and st vs. nearer b than st) and the number of blades of rallum (8 vs. 7).</p><p>Tyrannochthonius brevispinus sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on both chelal fingers, from T. harveyi by the number of setae at the anterior margin of the carapace (6 vs. 4), a larger body size (body length min. 1.73 mm vs. max. 1.56 mm; chela length min. 1.33 mm vs. max. 1.30 mm; palpal femur length min. 0.93 mm vs. max. 0.88 mm), the number of setae on tergite III (4 vs. 2) and the number of blades of rallum (8 vs. 6) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FFB2FFFCFF205E9D38E1F826	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFBCFFF5FF205F8838E1FA56.text	FD7487B1FFBCFFF5FF205F8838E1FA56.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius duo Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius duo sp. nov. ‹Ξ⁂Ɵṁae›</p><p>Figs 32–36</p><p>Type material. Holotype ♂ (Ps.- MHBU-HBUARA#2022-305 - 01): China, Guizhou Province, Jiangkou County, Nuxi Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.86204&amp;materialsCitation.latitude=27.827179" title="Search Plazi for locations around (long 108.86204/lat 27.827179)">Mengjiatun Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.86204&amp;materialsCitation.latitude=27.827179" title="Search Plazi for locations around (long 108.86204/lat 27.827179)">Liangfeng Cave</a>, under stones and detritus in the deep zone (Temperature: 13°C, Humidity: 80%) [27°49′37.84″N, 108°51′43.34″E], 408 m a.s.l., 4 July 2022, Yanmeng Hou, Nana Zhan, Long Lin &amp; Jianzhou Sun leg. Paratypes: 2 ♀ (Ps.- MSWU-HBUARA#2022-305 - 02 &amp; HBUARA#2022-305-03), all with the same data as the holotype .</p><p>Etymology. The specific name is derived from the Latin word “ duo ”, meaning dual, which refers to the presence of two well-developed anterior eyes.</p><p>Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace with two anterior corneate eyes, posterior eyes reduced into eyespots; anterior margin of carapace thin, finely denticulated, epistome small, pointed, triangular; posterior margin of carapace with 2 setae; tergites I–II each with 4 setae. Pedipalps slender, femur 5.85 (♂), 5.79–6.08 (♀) times longer than broad; chela 6.94 (♂), 5.85–6.56 (♀) times longer than broad; both chelal fingers with intercalary teeth; chemosensory setae present on dorsum of chelal hand.</p><p>Description. Male (holotype) (Figs 32D, 33A, 34A–F, 35, 36).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 34C, 35A): carapace 1.00 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; anterior pair of eyes well developed, posterior pair of eyes reduced into eyespots; anterior margin slightly serrate; epistome small and pointed, triangular; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 9–11 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 35C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 34D, 35B): large, about as long as carapace, 2.08 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation both dorsal and ventral side. Both fingers well provided with teeth, fixed finger with 15 teeth, distal one largest; movable finger with 15 retrorse contiguous small teeth; galea completely vestigial (Fig. 35B). Serrula exterior with 17 and serrula interior with 13 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 35E).</p><p>Pedipalp (Figs 34A, B, E, 35D, 36A, B): long and slender, trochanter 1.36, femur 5.85, patella 2.27, chela 6.94, hand 2.25 times longer than broad; femur 2.24 times longer than patella; movable chelal finger 2.14 times longer than hand and 0.69 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 34E, 35D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, t situated at same level as est and distal to b (Fig. 36A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 32 macrodenticles, slightly retrorse and pointed, plus 21 intercalary microdenticles, 53 in total; movable chelal finger with 16 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 10 intercalary microdenticles and 14 vestigial, rounded and contiguous basal teeth, 40 in total (Fig. 36A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 34B, 36B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 3: 4: 6: 7: 8: 8: 6: 4: T2T: 0, tergites VI–IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 12: 12: 9: 9: 9: 10: 10: 8: 0: 2. Anterior genital operculum with 12 setae, genital opening slit-like, with 17–18 marginal setae on each side, 47 in total (Fig. 34F).</p><p>Legs (Fig. 36C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 2.09 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.13 times longer than tibia. Femoropatella of leg IV 2.91 times longer than deep; tibia 4.80 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.29 times longer than deep (TS= 0.35), tarsus 11.60 times longer than deep and 2.52 times longer than metatarsus (TS= 0.36). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 33B, 34G). Mostly same as male, but a little larger; with same chaetotaxy of coxae; tergal chaetotaxy I–XII: 4: 4: 4: 4–6: 7–8: 8: 8: 8–9: 8–9: 4: T2T: 0; sternal chaetotaxy IV–XII: 11–13: 12: 11–12: 12: 12: 11–12: 9: 0: 2; anterior genital operculum with 11 setae, posterior margin with 12–14 marginal setae, 23–25 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.00–3.43 times longer than deep (TS= 0.33), tarsus 11.60–11.80 times longer than deep and 2.42–2.46 times longer than metatarsus (TS= 0.31–0.34).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Male: body length 1.73. Pedipalps: trochanter 0.19/0.14 (1.36), femur 0.76/0.13 (5.85), patella 0.34/0.15 (2.27), chela 1.11/0.16 (6.94), hand 0.36/0.16 (2.25), movable chelal finger length 0.77. Chelicera 0.52/0.25 (2.08), movable finger length 0.28. Carapace 0.52/0.52 (1.00). Leg I: trochanter 0.15/0.12 (1.25), femur 0.46/0.07 (6.57), patella 0.22/0.07 (3.14), tibia 0.24/0.05 (4.80), tarsus 0.51/0.04 (12.75). Leg IV: trochanter 0.19/0.14 (1.36), femoropatella 0.67/0.23 (2.91), tibia 0.48/0.10 (4.80), metatarsus 0.23/0.07 (3.29), tarsus 0.58/0.05 (11.60).</p><p>Females: body length 1.93–2.12. Pedipalps: trochanter 0.19–0.21/0.14–0.16 (1.19–1.50), femur 0.79–0.81/0.13– 0.14 (5.79–6.08), patella 0.36/0.15–0.16 (2.25–2.40), chela 1.17–1.18/0.18–0.20 (5.85–6.56), hand 0.38–0.39/0.18– 0.20 (1.95–2.11), movable chelal finger length 0.80–0.82. Chelicera 0.57–0.58/0.28–0.29 (2.00–2.04), movable finger length 0.31–0.32. Carapace 0.56/0.61–0.63 (0.89–0.92). Leg I: trochanter 0.13–0.14/0.12–0.13 (1.08), femur 0.47–0.49/0.07–0.08 (5.88–7.00), patella 0.22–0.24/0.07 (3.14–3.43), tibia 0.24/0.05 (4.80), tarsus 0.48–0.54/0.05– 0.06 (9.00–9.60). Leg IV: trochanter 0.20–0.23/0.14–0.15 (1.33–1.64), femoropatella 0.70/0.23 (3.04), tibia 0.52– 0.53/0.10 (5.20–5.30), metatarsus 0.24/0.07–0.08 (3.00–3.43), tarsus 0.58–0.59/0.05 (11.60–11.80).</p><p>Remarks. Tyrannochthonius duo sp. nov. can be distinguished from other described Chinese troglomorphic Tyrannochthonius species by the presence of a pair of well-developed anterior eyes and more setae on tergites V‒IX (6–9).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FFBCFFF5FF205F8838E1FA56	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFB9FFEFFF20592138E1FA22.text	FD7487B1FFB9FFEFFF20592138E1FA22.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius gracilis Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius gracilis sp. nov. ‹KḦƟṁae›</p><p>Figs 37–41</p><p>Type material. Holotype ♂ (Ps.- MHBU-HBUARA#2022-490-01): China, Guizhou Province, Hezhang County, Pingshan Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.95568&amp;materialsCitation.latitude=27.13096" title="Search Plazi for locations around (long 104.95568/lat 27.13096)">Jiangnan Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.95568&amp;materialsCitation.latitude=27.13096" title="Search Plazi for locations around (long 104.95568/lat 27.13096)">Xianren Cave</a>, under stones, detritus and on the cave walls within 20–200 m from the cave entrance (Temperature: 14°C, Humidity: 85%) [27°7′51.46″N, 104°57′20.45″E], 1446 m a.s.l., 30 July 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun &amp; Wenlong Fan leg. Paratypes: 13 ♂ (Ps.- MHBU-HBUARA#2022- 490-02 – HBUARA#2022-490-14), 15 ♀ (Ps.- MHBU-HBUARA#2022-490-15 – HBUARA#2022-490-29), all with the same data as the holotype; 4 ♂ (Ps.- MSWU- GZC190808-01 – GZC190808-04), 8 ♀ (Ps.- MSWU- GZC190808- 05 – GZC190808-12), 8 August 2019, Zegang Feng, Zhaoyi Li &amp; Chen Zhang leg., all with the same location as the holotype .</p><p>Etymology. The specific name is derived from the Latin word “ gracilis ”, meaning gracile, which refers to the elongated coxal spines.</p><p>Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with 2 setae; tergites I–IV each with 2 setae. Pedipalps slender, femur 8.50–8.88 (♂), 8.05–8.53 (♀) times longer than broad; chela 8.79–9.04 (♂), 8.41–8.81 (♀) times longer than broad; only fixed chelal finger with intercalary teeth.</p><p>Description. Males (holotype and paratypes) (Figs 38A, 39A–F, 40, 41).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 39C, 40A): carapace 0.99–1.00 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, triangular, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exter ior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 11–12 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 40C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 39D, 40B): large, about as long as carapace, 2.50–2.53 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 15–18 teeth, distal one largest; movable finger with 18–21 retrorse contiguous small teeth; galea completely vestigial (Fig. 40B). Serrula exterior with 21–22 and serrula interior with 14–15 blades. Rallum with 6 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 40E).</p><p>Pedipalp (Figs 39A, B, E, 40D, 41A, B): long and slender, trochanter 1.24–1.40, femur 8.50–8.88, patella 2.70–3.05, chela 8.79–9.04, hand 3.08–3.30 times longer than broad; femur 2.47–2.48 times longer than patella; movable chelal finger 1.63–1.76 times longer than hand and 0.60–0.62 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 39E, 40D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, t situated distal to b and slightly proximal to est (Fig. 41A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 28–30 macrodenticles, long and pointed, plus 14– 15 intercalary microdenticles, 42–45 in total; movable chelal finger with 34–37 macrodenticles (smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 5–6 vestigial, rounded and contiguous basal teeth, 40–42 in total, without intercalary teeth (Fig. 41A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 39B, 41B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 2: 2: 2: 2: 4: 4: 4: 4: 5: 4: T2T: 0; tergite IX with an unpaired median seta. Sternal chaetotaxy III–XII: 11–12: 10: 8: 7–8: 6–7: 7: 7: 7: 0: 2. Anterior genital operculum with 10–12 setae, genital opening slit-like, with 9–11 marginal setae on each side, 31–33 in total (Fig. 39F).</p><p>Legs (Fig. 41C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.76–1.77 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.37–2.41 times longer than tibia. Femoropatella of leg IV 3.93–4.41 times longer than deep; tibia 6.75–6.83 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.80–4.00 times longer than deep (TS= 0.23–0.26), tarsus 14.14–14.57 times longer than deep and 2.48–2.68 times longer than metatarsus (TS= 0.27). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 37D, 38B, 39G). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 3, II 3–4, III 5, IV 5; with same tergal chaetotaxy as males; sternal chaetotaxy IV–XII: 12–14: 8: 7–8: 7: 7: 7: 7: 0: 2; anterior genital operculum with 10 setae, posterior margin with 10–12 marginal setae, 20–22 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 4.56 times longer than deep (TS= 0.22–0.27), tarsus 13.25–14.57 times longer than deep and 2.49–2.59 times longer than metatarsus (TS= 0.21–0.26).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 1.96–2.06. Pedipalps: trochanter 0.26–0.28/0.20–0.21 (1.24–1.40), femur 1.51–1.53/0.17–0.18 (8.50– 8.88), patella 0.61–0.62/0.20–0.23 (2.70–3.05), chela 2.08–2.11/0.23–0.24 (8.79–9.04), hand 0.74–0.76/0.23–0.24 (3.08–3.30), movable chelal finger length 1.24–1.30. Chelicera 0.80–0.86/0.32–0.34 (2.50–2.53), movable finger length 0.43–0.45. Carapace 0.68–0.71/0.68–0.72 (0.99–1.00). Leg I: trochanter 0.23/0.18–0.19 (1.21–1.28), femur 0.85–0.88/0.09–0.10 (8.80–9.44), patella 0.48–0.50/0.09 (5.33–5.56), tibia 0.38–0.39/0.06–0.08 (4.88–6.33), tarsus 0.90–0.94/0.06–0.07 (13.43–15.00). Leg IV: trochanter 0.32–0.33/0.18–0.20 (1.65–1.78), femoropatella 1.18–1.19/0.27–0.30 (3.93–4.41), tibia 0.81–0.82/0.12 (6.75–6.83), metatarsus 0.38–0.40/0.10 (3.80–4.00), tarsus 0.99–1.02/0.07 (14.14–14.57).</p><p>Females: body length 2.12–2.58. Pedipalps: trochanter 0.27–0.30/0.21 (1.29–1.43), femur 1.61–1.62/0.19–0.20 (8.05–8.53), patella 0.64–0.67/0.23–0.24 (2.67–2.91), chela 2.27–2.29/0.26–0.27 (8.41–8.81), hand 0.78–0.83/0.26– 0.27 (3.00–3.07), movable chelal finger length 1.39–1.43. Chelicera 0.89–0.91/0.36–0.38 (2.39–2.47), movable finger length 0.48–0.49. Carapace 0.74–0.76/0.74–0.76 (0.97–1.03). Leg I: trochanter 0.23–0.24/0.18–0.19 (1.21– 1.33), femur 0.89–0.92/0.09–0.10 (9.20–9.89), patella 0.47/0.09 (5.22), tibia 0.39–0.41/0.07–0.08 (5.13–5.57), tarsus 0.95–0.98/0.07 (13.57–14.00). Leg IV: trochanter 0.35/0.19 (1.84), femoropatella 1.23–1.25/0.27–0.28 (4.39– 4.63), tibia 0.82–0.86/0.12 (6.83–7.17), metatarsus 0.41/0.09 (4.56), tarsus 1.02–1.06/0.07–0.08 (13.25–14.57).</p><p>Remarks. Tyrannochthonius gracilis sp. nov. is similar to T. parcidentatus sp. nov. in having intercalary teeth on the fixed chelal finger only and a pointed, triangular epistome, but differs by a smaller body size but longer appendages (body length 1.96–2.06 (♂), 2.12–2.58 (♀) vs. 2.29–2.62 (♂), 2.64–2.66 (♀) mm; chela length 2.08–2.11 (♂), 2.27–2.29 (♀) vs. 1.89–1.93 (♂), 2.02–2.05 (♀) mm) and more fixed chelal finger teeth (42‒45 vs. 30‒31).</p><p>Tyrannochthonius gracilis sp. nov. can be distinguished from T. akaelus, T. antridraconis and T. ganshuanensis by the presence of intercalary teeth on the fixed chelal finger only, from T. harveyi by the presence of intercalary teeth on the fixed chelal finger, from T. chixing and T. zhai by the number of setae on tergites I–IV (2 vs. 4) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FFB9FFEFFF20592138E1FA22	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFA3FFE2FF205AC738E1F8FE.text	FD7487B1FFA3FFE2FF205AC738E1F8FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius hispidus Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius hispidus sp. nov. ‹SƭƟṁae›</p><p>Figs 42–46</p><p>Type material. Holotype ♂ (Ps.-MHBU-GZJH-19-26-01): China, Guizhou Province, Jianhe County, Yangasha Sub-district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.46168&amp;materialsCitation.latitude=26.76613" title="Search Plazi for locations around (long 108.46168/lat 26.76613)">Chuandong Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.46168&amp;materialsCitation.latitude=26.76613" title="Search Plazi for locations around (long 108.46168/lat 26.76613)">Qilin Cave</a>, under stones within 100–300 m from the cave entrance (Temperature: 15°C, Humidity: 80%) [26°45′58.07″N, 108°27′42.06″E], 743 m a.s.l., 13 October 2019, Zegang Feng &amp; Lingchen Zhao leg. Paratypes: 12 ♂ (Ps.-MHBU-GZJH-19-26-02–GZJH-19-26-13), 21 ♀ (Ps.-MHBU-GZJH-19-26-14–GZJH-19-26-34), all with the same data as the holotype; 15 ♂ (Ps.- MSWU-HBUARA#2022-521 - 01 – HBUARA#2022-521-15), 12 ♀ (Ps.- MSWU-HBUARA#2022-521 - 16 –HBUARA#2022-521-27), 13 August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun &amp; Wenlong Fan leg., all with the same location as the holotype .</p><p>Etymology. The specific name is derived from the Latin word “ hispidus ” (= hirsute), which refers to the increased number of setae at the posterior margin of the carapace.</p><p>Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with 2–4 setae; tergites I–VI each with 4 setae. Pedipalps slender, femur 5.80–5.93 (♂), 5.93– 6.14 (♀) times longer than broad; chela 6.68–6.72 (♂), 6.20–6.79 (♀) times longer than broad; both chelal fingers with intercalary teeth.</p><p>Description. Males (holotype and paratypes) (Figs 43A, 44A–F, 45, 46).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 44C, 45A): carapace 0.94–0.98 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, triangular, with 2 setae flanking base; with 18–20 setae arranged s3–4s: 4: 4: 2–3: 2–4, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 7–10 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 45C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 44D, 45B): large, about as long as carapace, 2.29–2.30 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 10–12 teeth, distal one largest; movable finger with 10–13 retrorse contiguous small teeth; galea completely vestigial (Fig. 45B). Serrula exterior with 18–20 and serrula interior with 14–15 blades. Rallum with 6 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 45E).</p><p>Pedipalp (Figs 44A, B, E, 45D, 46A, B): long and slender, trochanter 1.20–1.31, femur 5.80–5.93, patella 2.00–2.19, chela 6.68–6.72, hand 2.32–2.33 times longer than broad; femur 2.49–2.59 times longer than patella; movable chelal finger 1.77–1.83 times longer than hand and 0.61–0.64 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 44E, 45D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated distal to b and slightly proximal to est (Fig. 46A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 26–28 macrodenticles, long and pointed, plus 15– 16 intercalary microdenticles, 42–43 in total; movable chelal finger with 22–24 macrodenticles (markedly smaller than teeth on fixed chelal finger), retrorse and pointed, plus 9– 10 intercalary microdenticles and 7–8 vestigial, rounded and contiguous basal teeth, 38–42 in total (Fig. 46A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 44B, 46B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4: 4–5: 5: 5: 4: T2T: 0; tergites VIII–IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 13–15: 13–14: 10: 9: 9: 9: 9–11: 9: 0: 2. Anterior genital operculum with 10 setae, genital opening slit-like, with 10–13 marginal setae on each side, 31–34 in total (Fig. 44F).</p><p>Legs (Fig. 46C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.79–1.88 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.27–2.43 times longer than tibia. Femoropatella of leg IV 2.68–2.70 times longer than deep; tibia 5.00–5.38 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.14 times longer than deep (TS= 0.32), tarsus 12.50–12.75 times longer than deep and 2.27–2.32 times longer than metatarsus (TS= 0.26–0.29). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 43B, 44G). Mostly same as males, but a little larger; with same chaetotaxy of coxae as males; tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4: 4: 5: 5: 4: T2T: 0; sternal chaetotaxy IV–XII: 14: 10: 9: 9: 10: 9–10: 8–9: 0: 2; anterior genital operculum with 9–10 setae, posterior margin with 13–14 marginal setae, 23 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.50–3.67 times longer than deep (TS= 0.33–0.36), tarsus 10.00 times longer than deep and 2.27–2.38 times longer than metatarsus (TS= 0.32).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 1.56–1.78. Pedipalps: trochanter 0.17–0.18/0.13–0.15 (1.20–1.31), femur 0.83–0.87/0.14–0.15 (5.80–5.93), patella 0.32–0.35/0.16 (2.00–2.19), chela 1.21–1.27/0.18–0.19 (6.68–6.72), hand 0.42–0.44/0.18–0.19 (2.32– 2.33), movable chelal finger length 0.77–0.78. Chelicera 0.53–0.55/0.23–0.24 (2.29–2.30), movable finger length 0.28–0.29. Carapace 0.48–0.50/0.49–0.53 (0.94–0.98). Leg I: trochanter 0.14–0.15/0.12 (1.17–1.25), femur 0.43– 0.45/0.07–0.08 (5.38–6.43), patella 0.24/0.07 (3.43), tibia 0.21–0.22/0.05–0.06 (3.50–4.40), tarsus 0.50–0.51/0.04– 0.05 (10.20–12.50). Leg IV: trochanter 0.20–0.21/0.12–0.13 (1.62–1.67), femoropatella 0.59–0.62/0.22–0.23 (2.68– 2.70), tibia 0.43–0.45/0.08–0.09 (5.00–5.38), metatarsus 0.22/0.07 (3.14), tarsus 0.50–0.51/0.07 (12.50–12.75).</p><p>Females: body length 1.77–1.99. Pedipalps: trochanter 0.18–0.20/0.14 (1.29–1.43), femur 0.86–0.89/0.14–0.15 (5.93–6.14), patella 0.35–0.36/0.16 (2.19–2.25), chela 1.24–1.29/0.19–0.20 (6.20–6.79), hand 0.44–0.45/0.19– 0.20 (2.25–2.32), movable chelal finger length 0.77–0.82. Chelicera 0.55–0.58/0.24–0.25 (2.29–2.32), movable finger length 0.29. Carapace 0.49/0.53–0.54 (0.91–0.92). Leg I: trochanter 0.12–0.15/0.12–0.13 (1.00–1.15), femur 0.44–0.45/0.07–0.08 (5.50–6.43), patella 0.24–0.26/0.07 (3.43–3.71), tibia 0.21/0.05–0.06 (2.63–3.50), tarsus 0.49–0.51/0.05 (9.80–10.20). Leg IV: trochanter 0.20/0.13 (1.54), femoropatella 0.60–0.62/0.20–0.21 (2.95–3.00), tibia 0.44–0.45/0.08–0.09 (5.00–5.50), metatarsus 0.21–0.22/0.06 (3.50–3.67), tarsus 0.50/0.05 (10.00).</p><p>Remarks. Tyrannochthonius hispidus sp. nov. is similar to T. acutus sp. nov. in having intercalary teeth on both chelal fingers and a pointed, triangular epistome, but differs by a smaller body size (body length 1.56–1.78 (J), 1.77–1.99 (♀) vs. 2.07–2.13 (J), 2.16–2.29 (♀) mm; chela 6.68–6.72 (J), 6.20–6.79 (♀) vs. 7.29–7.65 (J), 7.38–7.79 (♀) times longer than board), fewer fixed chelal finger teeth (42‒43 vs. 57‒60) and the number of blades of rallum (6 vs. 7).</p><p>Tyrannochthonius hispidus sp. nov. can be distinguished from T. chixing, T. harveyi and T. zhai by the presence of intercalary teeth on both chelal fingers, from T. akaelus and T. ganshuanensis by the number of setae on tergites I–II (4 vs. 2), from T. antridraconis by a smaller size (body length max. 1.78 mm vs. min. 1.8 mm; chela 6.68–6.72 times vs. 7.90–8.00 times longer than board, length 1.21–1.27 (J) vs. 1.70–1.76 (J) mm), more fixed chelal finger teeth (42‒43 vs. 33‒36) and the relative position of trichobothrium sb (sb closer to st than to b vs. closer to b) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FFA3FFE2FF205AC738E1F8FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFAAFF98FF205F8838E1FF4B.text	FD7487B1FFAAFF98FF205F8838E1FF4B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius infirmus Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius infirmus sp. nov. ‹NjṄƟṁae›</p><p>Figs 47–51</p><p>Type material. Holotype ♂ (Ps.- MHBU-GZC190804-01): China, Guizhou Province, Pu’an County, Qingshan Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.00544&amp;materialsCitation.latitude=25.468578" title="Search Plazi for locations around (long 105.00544/lat 25.468578)">Xiajiehe Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.00544&amp;materialsCitation.latitude=25.468578" title="Search Plazi for locations around (long 105.00544/lat 25.468578)">Liujiadadong Cave</a>, under stones and clods in the deep zone (Temperature: 13°C, Humidity: 90%) [25°28′6.88″N, 105°0′19.57″E], 1559 m a.s.l., 4 August 2019, Zegang Feng, Zhaoyi Li &amp; Chen Zhang leg. Paratypes: 3 ♂ (Ps.- MHBU-GZC190804-02 – GZC190804-04), 5 ♀ (Ps.- MSWU-GZC190804-05 – GZC190804-09), all with the same data as the holotype .</p><p>Etymology. The specific name is derived from the Latin word “ infirmus ”, meaning weak, which refers to the presence of a pair of weak eyespots.</p><p>Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace with a pair of anterior eyespots only; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with 2 setae; tergites I–Ⅴ each with 4 setae. Pedipalps slender, femur 6.72–6.88 (♂), 6.47–6.94 (♀) times longer than broad; chela 7.27–7.42 (♂), 6.28–6.83 (♀) times longer than broad; both chelal fingers with intercalary teeth.</p><p>Description. Males (holotype and paratypes) (Figs 48A, 49A–F, H, 50, 51).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 49C, D, 50A): carapace 0.97–0.99 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; anterior eyes reduced into eyespots, posterior eyes reduced completely; anterior margin slightly serrate; epistome small and pointed, triangular, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 11–13 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 50C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 49E, 50B): large, about as long as carapace, 2.29–2.30 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 12–14 teeth, distal one largest; movable finger with 12–13 retrorse contiguous small teeth; galea completely vestigial (Fig. 50B). Serrula exterior with 18–21 and serrula interior with 13–15 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 50E).</p><p>Pedipalp (Figs 49A, B, H, 50D, 51A, B): long and slender, trochanter 1.42–1751, femur 6.72–6.88, patella 2.09–2.32, chela 7.27–7.42, hand 2.64–2.67 times longer than broad; femur 2.50–2.63 times longer than patella; movable chelal finger 1.74–1.78 times longer than hand and 0.63–0.64 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 49H, 50D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, t situated midway between it and est; est situated distal to b (Fig. 51A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 28–31 macrodenticles, slightly retrorse and pointed, plus 20– 21 intercalary microdenticles, 48–52 in total; movable chelal finger with 25–26 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 18– 19 intercalary microdenticles and 8–10 vestigial, rounded and contiguous basal teeth, 52–54 in total (Fig. 51A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 49B, 51B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 5: 5–6: 5: 5: 4: T2T: 0; tergites VII–IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 12–13: 12–13: 8–9: 10: 9–10: 8–10: 9–10: 9–10: 0: 2. Anterior genital operculum with 10–11 setae, genital opening slit-like, with 20–23 marginal setae on each side, 53 in total (Fig. 49F).</p><p>Legs (Fig. 51C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.92–2.09 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.44–2.59 times longer than tibia. Femoropatella of leg IV 3.59–3.85 times longer than deep; tibia 6.25–6.58 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.56–3.60 times longer than deep (TS= 0.28), tarsus 12.43–13.83 times longer than deep and 2.42–2.59 times longer than metatarsus (TS= 0.36–0.37). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 48B, 49G). Mostly same as males, but a little larger; with same chaetotaxy of coxae as males; tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4–5: 5–6: 5–7: 6: 4: T2T: 0; sternal chaetotaxy IV–XII: 11–13: 9: 10: 9–10: 9–10: 10–11: 10: 0: 2; anterior genital operculum with 10 setae, posterior margin with 11–13 marginal setae, 21–23 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.44–3.56 times longer than deep (TS= 0.28–0.29), tarsus 12.29–13.50 times longer than deep and 2.53–2.77 times longer than metatarsus (TS= 0.35–0.38).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 2.41–2.54. Pedipalps: trochanter 0.27–0.35/0.19–0.20 (1.42–1.75), femur 1.10–1.21/0.16–0.18 (6.72–6.88), patella 0.44–0.46/0.19–0.22 (2.09–2.32), chela 1.60–1.78/0.22–0.24 (7.27–7.42), hand 0.58–0.64/0.22–0.24 (2.64– 2.67), movable chelal finger length 1.01–1.14. Chelicera 0.69–0.71/0.30–0.31 (2.29–2.30), movable finger length 0.37–0.40. Carapace 0.62–0.68/0.64–0.69 (0.97–0.99). Leg I: trochanter 0.19–0.21/0.15–0.17 (1.24–1.27), femur 0.67–0.75/0.09–0.11 (6.82–7.44), patella 0.32–0.39/0.09 (3.56–4.33), tibia 0.29–0.34/0.06–0.08 (4.25–4.83), tarsus 0.75–0.83/0.06–0.07 (11.86–12.50). Leg IV: trochanter 0.31–0.33/0.17–0.19 (1.74–1.82), femoropatella 0.97– 1.04/0.27 (3.59–3.85), tibia 0.75–0.79/0.12 (6.25–6.58), metatarsus 0.32–0.36/0.09–0.10 (3.56–3.60), tarsus 0.83– 0.87/0.06–0.07 (12.43–13.83).</p><p>Females: body length 2.50–2.67. Pedipalps: trochanter 0.21–0.24/0.18 (1.17–1.33), femur 1.10–1.18/0.17 (6.47–6.94), patella 0.45/0.20 (2.25), chela 1.57–1.64/0.24–0.25 (6.28–6.83), hand 0.59–0.61/0.24–0.25 (2.36–2.54), movable chelal finger length 1.00–1.04. Chelicera 0.69–0.71/0.32–0.33 (2.15–2.16), movable finger length 0.37– 0.39. Carapace 0.66–0.67/0.69–0.70 (0.94–0.97). Leg I: trochanter 0.17–0.19/0.15–0.16 (1.06–1.27), femur 0.64– 0.68/0.10 (6.40–6.80), patella 0.35/0.08–0.09 (3.89–4.38), tibia 0.28–0.32/0.07–0.08 (4.00), tarsus 0.70–0.77/0.07 (10.00–11.00). Leg IV: trochanter 0.29/0.17 (1.71), femoropatella 0.95–0.97/0.25–0.26 (3.73–3.80), tibia 0.69– 0.72/0.11–0.12 (6.00–6.27), metatarsus 0.31–0.32/0.09 (3.44–3.56), tarsus 0.81–0.86/0.06–0.07 (12.29–13.50).</p><p>Remarks. Tyrannochthonius infirmus sp. nov. can be distinguished from other described Chinese troglomorphic Tyrannochthonius species by the presence of a pair of anterior eyespots.</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FFAAFF98FF205F8838E1FF4B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFD4FF9EFF205E2F38E1F826.text	FD7487B1FFD4FF9EFF205E2F38E1F826.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius latus Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius latus sp. nov. ‹ŴṞƟṁae›</p><p>Figs 52–56</p><p>Type material. Holotype ♂ (Ps.- MHBU-HBUARA#2021-168 - 01): China, Guizhou Province, Panzhou City, Danxia Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.62685&amp;materialsCitation.latitude=25.674885" title="Search Plazi for locations around (long 104.62685/lat 25.674885)">Zimu Cave</a>, under stones in the deep zone (Temperature: 14°C, Humidity: 90%) [25°40′29.59″N, 104°37′36.64″E], 1787 m a.s.l., 15 July 2021, Zegang Feng, Hongru Xu, Liu Fu &amp; Nana Zhan leg. Paratypes: 3 ♂ (Ps.- MHBU-HBUARA#2021-168 - 02 –HBUARA#2021-168-04), 5 ♀ (Ps.- MSWU-HBUARA#2021-168 - 05 – HBUARA#2021-168-09), all with the same data as the holotype .</p><p>Etymology. The specific name is derived from the Latin word “ latus ”, meaning wide, which refers to the widened chelal hands.</p><p>Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, represented by a bump; posterior margin of carapace with 2 setae; tergites I–VII each with 4 setae. Pedipalps slender, femur 9.29–9.33 (♂), 8.75–8.79 (♀) times longer than broad; chela 6.69–7.13 (♂), 6.15–6.58 (♀) times longer than broad; chelal hand wider, hand 2.80–2.90 (♂), 2.53–2.72 (♀) times longer than broad (hand wide 0.30–0.35 (♂), 0.36–0.40 (♀) mm); both chelal fingers without intercalary teeth.</p><p>Description. Males (holotype and paratypes) (Figs 53A, 54A–F, 55, 56).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 54C, 55A): carapace 0.92–0.96 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, represented by a bump, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 11–12 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 55C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 54D, 55B): large, about as long as carapace, 2.39–2.45 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 18–20 teeth, distal one largest; movable finger with 14–17 retrorse contiguous small teeth; galea completely vestigial (Fig. 55B). Serrula exterior with 20–24 and serrula interior with 14–15 blades. Rallum with 7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 55E).</p><p>Pedipalp (Figs 54A, B, E, 55D, 56A, B): long and slender, trochanter 1.22–1.29, femur 9.29–9.33, patella 2.52–2.58, chela 6.69–7.13, hand 2.80–2.90 times longer than broad; femur 2.71–2.72 times longer than patella; movable chelal finger 1.41–1.46 times longer than hand and 0.59 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 54E, 55D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated at same level as it; est situated distal to b (Fig. 56A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 29–31 macrodenticles, long and pointed, slightly retrorse; movable chelal finger with 30–32 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed (Fig. 56A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 54B, 56B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4: 4: 4–5: 5: 4: T2T: 0; tergite IX with an unpaired median seta. Sternal chaetotaxy III–XII: 13–15: 12–14: 8: 8–9: 9: 9–11: 9–11: 9: 0: 2. Anterior genital operculum with 14 setae, genital opening slit-like, with 13–15 marginal setae on each side, 41–44 in total (Fig. 54F).</p><p>Legs (Fig. 56C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.78–1.87 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.44–2.65 times longer than tibia. Femoropatella of leg IV 4.96–5.32 times longer than deep; tibia 7.00–7.75 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.60–4.10 times longer than deep (TS= 0.27–0.28), tarsus 14.43–16.00 times longer than deep and 2.73–2.81 times longer than metatarsus (TS= 0.26–0.30). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 53B, 54G). Mostly same as males, but a little larger; with same chaetotaxy of coxae as males; tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4: 4: 4–5: 5: 4: T2T: 0; sternal chaetotaxy IV–XII: 13: 9: 9–10: 10–11: 10–11: 10–11: 9: 0: 2; anterior genital operculum with 11 setae, posterior margin with 12 marginal setae, 23 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.70–4.00 times longer than deep (TS= 0.28–0.35), tarsus 12.75–13.88 times longer than deep and 2.76–2.78 times longer than metatarsus (TS= 0.25–0.30).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 2.59–2.80. Pedipalps: trochanter 0.27–0.28/0.21–0.23 (1.22–1.29), femur 1.58–1.68/0.17–0.18 (9.29–9.33), patella 0.58–0.62/0.23–0.24 (2.52–2.58), chela 2.14–2.34/0.30–0.35 (6.69–7.13), hand 0.87–0.98/0.30–0.35 (2.80–2.90), movable chelal finger length 1.27–1.38. Chelicera 0.81–0.86/0.33–0.36 (2.39–2.45), movable finger length 0.45–0.48. Carapace 0.65–0.71/0.71–0.74 (0.92–0.96). Leg I: trochanter 0.20–0.26/0.18–0.19 (1.11–1.37), femur 0.84–0.91/0.11 (7.64–8.27), patella 0.45–0.51/0.10 (4.50–5.10), tibia 0.37–0.41/0.08 (4.63–5.13), tarsus 0.98–1.00/0.07 (14.00–14.29). Leg IV: trochanter 0.33/0.19 (1.74), femoropatella 1.24–1.33/0.25 (4.96–5.32), tibia 0.84–0.93/0.12 (7.00–7.75), metatarsus 0.36–0.41/0.10 (3.60–4.10), tarsus 1.01–1.02/0.07 (14.43–16.00).</p><p>Females: body length 2.91–3.11. Pedipalps: trochanter 0.29–0.32/0.23–0.26 (1.23–1.26), femur 1.67–1.75/0.19– 0.20 (8.75–8.79), patella 0.68–0.70/0.25–0.27 (2.59–2.72), chela 2.37–2.46/0.36–0.40 (6.15–6.58), hand 0.98– 1.01/0.36–0.40 (2.53–2.72), movable chelal finger length 1.39–1.45. Chelicera 0.94–0.97/0.38–0.40 (2.43–2.47), movable finger length 0.52–0.55. Carapace 0.72–0.75/0.77–0.84 (0.89–0.94). Leg I: trochanter 0.23–0.26/0.19–0.20 (1.21–1.30), femur 0.94–0.99/0.11–0.12 (8.25–8.55), patella 0.51–0.53/0.10–0.11 (4.82–5.10), tibia 0.39–0.41/0.07 (5.57–5.86), tarsus 1.02–1.05/0.08 (12.75–13.13). Leg IV: trochanter 0.34–0.36/0.19–0.21 (1.71–1.79), femoropatella 1.32–1.38/0.25–0.26 (5.28–5.31), tibia 0.92–0.93/0.12 (7.67–7.75), metatarsus 0.37–0.40/0.10 (3.70–4.00), tarsus 1.02–1.11/0.08 (12.75–13.88).</p><p>Remarks. Tyrannochthonius latus sp. nov. is similar to T. nanxingensis sp. nov. in lacking intercalary teeth on both chelal fingers, but differs by a longer palp (chela length 2.14–2.34 (♂), 2.37–2.46 (♀) vs. 1.92–1.97 (♂), 1.85 (♀) mm; palpal femur length 1.58–1.68 (♂), 1.67–1.75 (♀) vs. 1.34–1.35 (♂), 1.24 (♀) mm; movable chelal finger 1.41–1.46 vs. 1.78–1.86 times longer than hand), the relative position of trichobothrium sb (sb closer to st than to b vs. closer to b), the number of setae on tergite IV (4 vs. 5) and the number of blades of rallum (7 vs. 8).</p><p>Tyrannochthonius latus sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on fixed chelal fingers, from T. harveyi by the number of setae on tergites I–III (4 vs. 2) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FFD4FF9EFF205E2F38E1F826	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFDEFF94FF205F8838E1FE3A.text	FD7487B1FFDEFF94FF205F8838E1FE3A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius maculosus Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius maculosus sp. nov. ‹AEṀƟṁae›</p><p>Figs 57–61</p><p>Type material. Holotype ♂ (Ps.- MHBU-HBUARA#2022-504 - 01): China, Guizhou Province, Xingren City, Xinlongchang Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.116196&amp;materialsCitation.latitude=25.438032" title="Search Plazi for locations around (long 105.116196/lat 25.438032)">Lianzhuang Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.116196&amp;materialsCitation.latitude=25.438032" title="Search Plazi for locations around (long 105.116196/lat 25.438032)">Daxiao Cave</a>, under stones, detritus and on the cave walls within 200– 400 m from the cave entrance (Temperature: 15°C, Humidity: 85%) [25°26′16.92″N, 105°6′58.31″E], 1473 m a.s.l., 5 August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun &amp; Wenlong Fan leg. Paratypes: 14 ♂ (Ps.-MHBU-HBUARA#2022-504-02–HBUARA#2022-504-15), 6 ♀ (Ps.-MSWU- HBUARA#2022-504-16–HBUARA#2022- 504-21), all with the same data as the holotype .</p><p>Etymology. The specific name is derived from the Latin word “ maculosus ”, meaning speckled, which refers to the speckled tergites.</p><p>Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with 2 setae; tergites I–III each with 4 setae. Pedipalps slender, femur 6.47–6.80 (♂), 6.67–6.80 (♀) times longer than broad; chela 6.81–7.45 (♂), 6.55 (♀) times longer than broad; both chelal fingers with intercalary teeth.</p><p>Description. Males (holotype and paratypes) (Figs 57F, 58A, 59A–F, 60, 61).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 59C, 60A): carapace 1.02–1.04 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, triangular, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 3–4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 9–10 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 60C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 59D, 60B): large, about as long as carapace, 2.22–2.33 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 12–14 teeth, distal one largest; movable finger with 14–17 retrorse contiguous small teeth; galea completely vestigial (Fig. 60B). Serrula exterior with 18–21 and serrula interior with 14–15 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 60E).</p><p>Pedipalp (Figs 59A, B, E, 60D, 61A, B): long and slender, trochanter 1.22–1.35, femur 6.47–6.80, patella 2.22–2.28, chela 6.81–7.45, hand 2.38–2.65 times longer than broad; femur 2.43–2.49 times longer than patella; movable chelal finger 1.83–1.90 times longer than hand and 0.65–0.66 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 59E, 60D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb midway between st and b; b and t situated subdistally, t situated at same level as it; est situated distal to b (Fig. 61C). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 34–35 macrodenticles, slightly retrorse and pointed, plus 24– 25 intercalary microdenticles, 58–60 in total; movable chelal finger with 20–22 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 18– 19 intercalary microdenticles and 12–14 vestigial, rounded and contiguous basal teeth, 52–53 in total (Fig. 61C). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 59B, 61B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 4–5: 4–5: 5: 5: 5–6: 5: 4: T2T: 0; tergite IX with an unpaired median seta. Sternal chaetotaxy III–XII: 14–15: 13–14: 9–10: 8–10: 9–10: 9: 9: 9: 0: 2. Anterior genital operculum with 10 setae, genital opening slit-like, with 17–22 marginal setae on each side, 45–50 in total (Fig. 59F).</p><p>Legs (Fig. 61C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.94–1.97 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.33–2.36 times longer than tibia. Femoropatella of leg IV 3.63–3.79 times longer than deep; tibia 5.82–6.00 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.11–3.63 times longer than deep (TS= 0.21–0.24), tarsus 11.83–12.67 times longer than deep and 2.54–2.62 times longer than metatarsus (TS= 0.28–0.32). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 58B, 59G). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; tergal chaetotaxy I–XII: 4: 3–4: 4: 4–5: 5–6: 5–7: 5–6: 5–6: 5–6: 4: T2T: 0; sternal chaetotaxy IV–XII: 14: 10: 9–10: 9–10: 9–10: 10: 9–10: 0: 2; anterior genital operculum with 12 setae, posterior margin with 14–15 marginal setae, 26–27 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.00–3.22 times longer than deep (TS= 0.24–0.25), tarsus 11.33–12.33 times longer than deep and 2.55–2.83 times longer than metatarsus (TS= 0.34–0.35).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 2.22–2.31. Pedipalps: trochanter 0.22–0.23/0.17–0.18 (1.22–1.35), femur 0.97–1.02/0.15 (6.47–6.80), patella 0.40–0.41/0.18 (2.22–2.28), chela 1.43–1.49/0.20–0.21 (6.81–7.45), hand 0.50–0.53/0.20–0.21 (2.38–2.65), movable chelal finger length 0.95–0.97. Chelicera 0.60–0.63/0.27 (2.22–2.33), movable finger length 0.33–0.35. Carapace 0.59/0.57–0.58 (1.02–1.04). Leg I: trochanter 0.17–0.19/0.14–0.15 (1.21–1.27), femur 0.60–0.61/0.08 (7.50–7.63), patella 0.31/0.07–0.08 (3.88–4.43), tibia 0.27–0.28/0.06 (4.50–4.67), tarsus 0.63–0.66/0.06 (10.50– 11.00). Leg IV: trochanter 0.25–0.27/0.15–0.16 (1.67–1.69), femoropatella 0.87–0.91/0.24 (3.63–3.79), tibia 0.64– 0.66/0.11 (5.82–6.00), metatarsus 0.28–0.29/0.08–0.09 (3.11–3.63), tarsus 0.71–0.76/0.06 (11.83–12.67).</p><p>Females: body length 2.50–2.59. Pedipalps: trochanter 0.22/0.18–0.19 (1.16–1.22), femur 1.00–1.02/0.15 (6.67–6.80), patella 0.40/0.18–0.19 (2.11–2.22), chela 1.44/0.22 (6.55), hand 0.51/0.22 (2.32), movable chelal finger length 0.94. Chelicera 0.64–0.65/0.29–0.30 (2.17–2.21), movable finger length 0.35. Carapace 0.61–0.62/0.62–0.63 (0.98). Leg I: trochanter 0.18/0.15–0.16 (1.13–1.20), femur 0.58–0.61/0.08–0.09 (6.44–7.63), patella 0.30–0.32/0.08 (3.75–4.00), tibia 0.27–0.28/0.06–0.07 (3.86–4.67), tarsus 0.60–0.64/0.06 (10.00–10.67). Leg IV: trochanter 0.27– 0.28/0.15 (1.80–1.87), femoropatella 0.90–0.92/0.23 (3.91–4.00), tibia 0.65–0.70/0.11 (5.91–6.36), metatarsus 0.24–0.29/0.08–0.09 (3.00–3.22), tarsus 0.68–0.74/0.06 (11.33–12.33).</p><p>Remarks. Tyrannochthonius maculosus sp. nov. is similar to T. multidentatus sp. nov. in having intercalary teeth on both chelal fingers and a pointed epistome, but differs by the trait of teeth on movable chelal finger (slightly retrorse and well-spaced vs. markedly retrorse and continuous), the relative position of trichobothrium sb (sb midway between st and b vs. closer to b) and the number of blades of rallum (8 vs. 6).</p><p>Tyrannochthonius maculosus sp. nov. can be distinguished from T. chixing, T. harveyi and T. zhai by the presence of intercalary teeth on both chelal fingers, from T. akaelus and T. ganshuanensis by the number of setae on tergites I–II (4 vs. 2), from T. antridraconis by a shorter palp (chela 6.81–7.45 (♂), 6.55 (♀) vs. 7.90–8.00 (♂), 6.90–7.55 (♀) times longer than broad; palpal femur 6.47–6.80 (♂), 6.67–6.80 (♀) vs. 7.30–8.30 (♂), 7.10–7.50 (♀) times longer than broad) and the relative position of trichobothrium sb (sb midway between st and b vs. closer to b) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FFDEFF94FF205F8838E1FE3A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFD8FF8CFF205ECA3E10FE1E.text	FD7487B1FFD8FF8CFF205ECA3E10FE1E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius multicavus Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius multicavus sp. nov. ‹Ũ栖Ɵṁae›</p><p>Figs 62–69</p><p>Type material. Holotype ♂ (Ps.- MHBU-HBUARA#2021-204 - 01): China, Guizhou Province, Pingtang County, Tangbian Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.763&amp;materialsCitation.latitude=25.645458" title="Search Plazi for locations around (long 106.763/lat 25.645458)">Xinhong Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.763&amp;materialsCitation.latitude=25.645458" title="Search Plazi for locations around (long 106.763/lat 25.645458)">Xiaohui Cave</a>, under stones and on the cave walls in the deep zone (Temperature: 15°C, Humidity: 80%) [25°38′43.65″N, 106°45′46.81″E], 888.6 m a.s.l., 3 August 2021, Zegang Feng, Hongru Xu, Liu Fu &amp; Nana Zhan leg. Paratypes: 1 ♂ (Ps.- MHBU-HBUARA#2021-204 - 02), 2 ♀ (Ps.- MSWU-HBUARA#2021- 204 - 03 &amp; HBUARA#2021-204-04), all with the same data as the holotype; 2 ♂ (Ps.- MHBU-HBUARA#2021-205 - 01 &amp; HBUARA#2021-205-02), 5 ♀ (Ps.- MSWU-HBUARA#2021-205 - 03 –HBUARA#2021-205-07) <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.71893&amp;materialsCitation.latitude=25.6357" title="Search Plazi for locations around (long 106.71893/lat 25.6357)">Tangbian Town</a>, Bailu Village, Weier Cave, under stones in the deep zone [25°38′8.52″N, 106°43′8.16″E], 919.3 m a.s.l., all with the same collection date and collectors as the holotype; 9 ♂ (Ps.- MHBU-HBUARA#2022-513 - 01 –HBUARA#2022- 513-09), 5 ♀ (Ps.- MSWU-HBUARA#2022-513 - 10 –HBUARA#2022-513-14), Luodian County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.68974&amp;materialsCitation.latitude=25.668308" title="Search Plazi for locations around (long 106.68974/lat 25.668308)">Bianyan Town</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.68974&amp;materialsCitation.latitude=25.668308" title="Search Plazi for locations around (long 106.68974/lat 25.668308)">Dahao Village</a>, Da Cave, under stones in the deep zone [25°40′5.91″N, 106°41′23.07″E], 930 m a.s.l., 09 August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun &amp; Wenlong Fan leg. ; 5 ♂ (Ps.- MHBU-HBUARA#2022-514 - 01 – HBUARA#2022-514-05), 2 ♀ (Ps.- MSWU-HBUARA#2022-514 - 06 &amp; HBUARA#2022-514-07), Luodian County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.70645&amp;materialsCitation.latitude=25.632105" title="Search Plazi for locations around (long 106.70645/lat 25.632105)">Bianyan Town</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.70645&amp;materialsCitation.latitude=25.632105" title="Search Plazi for locations around (long 106.70645/lat 25.632105)">Dashang Village</a>, Qixia Cave, under stones in the deep zone [25°37′55.58″N, 106°42′23.24″E], 956 m a.s.l., 10 August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun &amp; Wenlong Fan leg.</p><p>Etymology. The specific name is derived from a combination of the Latin word “ multus ” and “ cavus ”, meaning numerous and cavernous, respectively, which refers to the species found in several caves.</p><p>Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and represented by a slight bump; posterior margin of carapace with 2 setae; tergites I–IV each with 2 setae. Pedipalps slender, femur 6.79–7.08 (♂), 6.53–7.21 (♀) times longer than broad; chela 7.53–8.20 (♂), 7.05–8.38 (♀) times longer than broad; both chelal fingers without intercalary teeth.</p><p>Description. Males (holotype and paratypes) (Figs 64F, 66A, 67A–F, 68, 69).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 67C, 68A): carapace 1.02–1.14 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and represented by a slight bump, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 3–4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 8–10 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 68C); intercoxal tubercle absent; without sub-oral seta</p><p>Chelicera (Figs 67D, 68B): large, about as long as carapace, 2.27–2.42 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 11–13 teeth, distal one largest; movable finger with 12–13 retrorse contiguous small teeth; galea represented by a very slight bump on movable finger (Fig. 68B). Serrula exterior with 18–21 and serrula interior with 13–15 blades. Rallum with 7–8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 68E).</p><p>Pedipalp (Figs 67A, B, E, 68D, 69A, B): long and slender, trochanter 1.20–1.47, femur 6.79–7.08, patella 2.40–2.86, chela 7.53–8.20, hand 2.65–2.94 times longer than broad; femur 2.39–2.68 times longer than patella; movable chelal finger 1.65–1.80 times longer than hand and 0.62–0.64 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 67E, 68D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, t situated at same level as est and distal to b (Fig. 69A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 24–27 macrodenticles, slightly retrorse, long and pointed; movable chelal finger with 30–33 macrodenticles (markedly smaller than teeth on fixed chelal finger), continuous and retrorse, almost prostrate (Fig. 69A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 67B, 69B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 2: 2: 2: 2: 2–4: 4: 4: 4: 4: 2: T2T: 0. Sternal chaetotaxy III–XII: 12–15: 11–13: 7–10: 7–10: 7–10: 7–9: 8–10: 7–10: 0: 2. Anterior genital operculum with 9–10 setae, genital opening slit-like, with 13–18 marginal setae on each side, 38–45 in total (Fig. 67F).</p><p>Legs (Fig. 69C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.78–2.07 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.56–2.88 times longer than tibia. Femoropatella of leg IV 3.86–4.10 times longer than deep; tibia 5.90–7.38 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.86–4.43 times longer than deep (TS= 0.23–0.30), tarsus 13.60–15.60 times longer than deep and 2.39–2.71 times longer than metatarsus (TS= 0.23–0.25). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 66B, 67G). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 3, II 3–4, III 5, IV 4–5; tergal chaetotaxy I–XII: 2: 2: 2: 2: 3–4: 4: 4: 4: 4: 2: T2T: 0; sternal chaetotaxy IV–XII: 11–13: 7–9: 7–10: 7–9: 7–9: 9: 7–9: 0: 2; anterior genital operculum with 9–10 setae, posterior margin with 12–14 marginal setae, 21–24 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.25–4.29 times longer than deep (TS= 0.25–0.32), tarsus 14.20–15.40 times longer than deep and 2.45–2.73 times longer than metatarsus (TS= 0.20–0.27).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 1.71–2.01. Pedipalps: trochanter 0.18–0.22/0.14–0.16 (1.20–1.47), femur 0.92–1.02/0.13–0.15 (6.79–7.08), patella 0.36–0.41/0.13–0.15 (2.40–2.86), chela 1.23–1.35/0.13–0.17 (7.53–8.20), hand 0.44–0.50/0.13–0.17 (2.65– 2.94), movable chelal finger length 0.79–0.84. Chelicera 0.46–0.53/0.19–0.22 (2.27–2.42), movable finger length 0.24–0.29. Carapace 0.49–0.53/0.44–0.49 (1.02–1.14). Leg I: trochanter 0.15–0.18/0.13–0.14 (1.07–1.29), femur 0.57–0.63/0.07–0.08 (7.13–8.43), patella 0.29–0.34/0.06–0.07 (4.14–5.33), tibia 0.23–0.26/0.05–0.06 (4.00–5.20), tarsus 0.61–0.70/0.05–0.06 (11.67–13.80). Leg IV: trochanter 0.22–0.26/0.14–0.16 (1.56–1.79), femoropatella 0.82–0.90/0.20–0.22 (3.86–4.10), tibia 0.59–0.63/0.08–0.10 (5.90–7.38), metatarsus 0.27–0.31/0.07 (3.86–4.43), tarsus 0.68–0.78/0.05 (13.60–15.60).</p><p>Females: body length 1.81–2.10. Pedipalps: trochanter 0.17–0.20/0.15–0.16 (1.13–1.27), femur 0.96–1.05/0.14– 0.16 (6.53–7.21), patella 0.37–0.41/0.13–0.17 (2.29–2.92), chela 1.30–1.42/0.16–0.20 (7.05–8.38), hand 0.47– 0.53/0.16–0.20 (2.58–2.94), movable chelal finger length 0.81–0.86. Chelicera 0.50–0.53/0.22–0.24 (2.17–2.32), movable finger length 0.27–0.29. Carapace 0.51–0.56/0.46–0.53 (1.04–1.13). Leg I: trochanter 0.14–0.17/0.13–0.15 (1.00–1.21), femur 0.55–0.61/0.07–0.09 (6.78–8.43), patella 0.28–0.34/0.07 (4.00–4.86), tibia 0.23–0.25/0.05–0.06 (3.83–5.00), tarsus 0.62–0.70/0.05–0.06 (10.67–14.00). Leg IV: trochanter 0.20–0.25/0.13–0.15 (1.43–1.71), femoropatella 0.80–0.88/0.19–0.23 (3.82–4.42), tibia 0.54–0.61/0.08–0.10 (6.00–7.00), metatarsus 0.26–0.30/0.07–0.08 (3.25–4.29), tarsus 0.71–0.77/0.05 (14.20–15.40).</p><p>Remarks. Tyrannochthonius multicavus sp. nov. is similar to T. harveyi in having the same number of setae on tergites I–III (2) and lacking intercalary teeth on both chelal fingers, but differs by the number of setae at the anterior margin of the carapace (6 vs. 4), a larger body size (body length min. 1.71 mm vs. max. 1.56 mm), the relative position of trichobothrium sb (sb closer to b than to st vs. midway between st and b), the number of setae on tergites IX and Ⅹ (2 vs. 4) and the proportion of movable chelal finger and chelal hand (1.65–1.80 vs. 2.29 times).</p><p>Tyrannochthonius multicavus sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on the fixed chelal finger (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the Xiaohui, Weier, Da, and Qixia caves.</p></div>	https://treatment.plazi.org/id/FD7487B1FFD8FF8CFF205ECA3E10FE1E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFC0FF86FF205EEE398EF9C6.text	FD7487B1FFC0FF86FF205EEE398EF9C6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius multidentatus Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius multidentatus sp. nov. ‹SffiƟṁae›</p><p>Figs 70–75</p><p>Type material. Holotype ♂ (Ps.-MHBU-GZFG-20-10-01): China, Guizhou Province, Fenggang County, Longquan Sub-district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.73388&amp;materialsCitation.latitude=27.944696" title="Search Plazi for locations around (long 107.73388/lat 27.944696)">Guanyin Cave</a>, under the stones in the deep zone (Temperature: 16°C, Humidity: 85%) [27°56′40.91″N, 107°44′1.99″E], 684 m a.s.l., 20 August 2020, Zegang Feng, Hongru Xu &amp; Yanmeng Hou leg. Paratypes: 2 ♂ (Ps.- MHBU-GZFG-20-10-02 &amp; GZFG-20-10-03), 4 ♀ (Ps.-MHBU-GZFG-20-10-04–GZFG-20-10-07), all with the same data as the holotype; 2 ♀ (Ps.-MSWU-GZSN-20-11-01 &amp; GZSN-20-11-02), Sinan County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.91362&amp;materialsCitation.latitude=27.876692" title="Search Plazi for locations around (long 107.91362/lat 27.876692)">Yangjiaao Township</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.91362&amp;materialsCitation.latitude=27.876692" title="Search Plazi for locations around (long 107.91362/lat 27.876692)">Guanyinshan Village</a>, Anjialin Cave [27°52′36.09″N, 107°54′49.02″E], 641 m a.s.l., 21 August 2020, with the same collectors as the holotype ; 2 ♀ (Ps.- MSWU-HBUARA#2022-475 - 01 &amp; HBUARA#2022-475-02), Anjialin Cave, 23 July 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun &amp; Wenlong Fan leg.</p><p>Etymology. The specific name is derived from a combination of the Latin word “ multus ” and “ dentatus ”, meaning numerous and toothed, respectively, which refers to the numerous teeth on the chelal fingers.</p><p>Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with 2 setae; tergites I–VI each with 4 setae. Pedipalps slender, femur 7.29–7.69 (♂), 6.59–6.94 (♀) times longer than broad; chela 7.40–7.84 (♂), 6.78–7.23 (♀) times longer than broad; both chelal fingers with intercalary teeth and teeth closely arranged.</p><p>Description. Males (holotype and paratypes) (Figs 72A, 73A–F, 74, 75).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 73C, 74A): carapace 1.02 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, triangular, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 9–12 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 74C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 73D, 74B): large, about as long as carapace, 2.28–2.32 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on dorsal side. Both fingers well provided with teeth, fixed finger with 18–21 teeth, distal one largest; movable finger with 18–21 retrorse contiguous small teeth; galea completely vestigial (Fig. 74B). Serrula exterior with 17–18 and serrula interior with 11–12 blades. Rallum with 6–7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 74E).</p><p>Pedipalp (Figs 73A, B, E, 74D, 75A, B): long and slender, trochanter 1.13–1.19, femur 7.29–7.69, patella 2.12–2.31, chela 7.40–7.84, hand 2.55–2.63 times longer than broad; femur 2.70–2.83 times longer than patella; movable chelal finger 1.82–1.94 times longer than hand and 0.63–0.65 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 73E, 74D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, t situated at same level as it; est situated distal to b (Fig. 75A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced closely along the margin, larger teeth present in the middle of the row, becoming smaller distally and proximally: fixed chelal finger with 34–38 macrodenticles, slightly retrorse and pointed, plus 28– 31 intercalary microdenticles, 62–69 in total; movable chelal finger with 36–40 macrodenticles (slightly smaller than teeth on fixed chelal finger), continuous, markedly retrorse and pointed, plus 17– 18 intercalary microdenticles, 53–58 in total (Fig. 75A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 73B, 75B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4: 4–5: 5–6: 5: 4–5: T2T: 0. Sternal chaetotaxy III–XII: 12: 12–13: 8: 7–8: 7–9: 7–9: 7–9: 7: 0: 2. Anterior genital operculum with 10 setae, genital opening slit-like, with 10–13 marginal setae on each side, 32–34 in total (Fig. 73F).</p><p>Legs (Fig. 75C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.84–1.93 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.58–2.64 times longer than tibia. Femoropatella of leg IV 3.90–3.95 times longer than deep; tibia 6.33 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.57–3.71 times longer than deep (TS= 0.38–0.40), tarsus 14.20 times longer than deep and 2.73–2.84 times longer than metatarsus (TS= 0.41–0.42). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 71D, 72B, 73G). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5–6; tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4: 4: 5–6: 5–6: 4: T2T: 0; sternal chaetotaxy IV–XII: 12–13: 8–10: 7–9: 7–8: 7–9: 7–9: 7: 0: 2; anterior genital operculum with 9–10 setae, posterior margin with 11–12 marginal setae, 20–22 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.50–4.00 times longer than deep (TS= 0.36–0.41), tarsus 12.50–13.17 times longer than deep and 2.68–2.75 times longer than metatarsus (TS= 0.38–0.46).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 1.97–1.99. Pedipalps: trochanter 0.18–0.19/0.16 (1.13–1.19), femur 1.00–1.02/0.13–0.14 (7.29–7.69), patella 0.36–0.37/0.16–0.17 (2.12–2.31), chela 1.48–1.49/0.19–0.20 (7.40–7.84), hand 0.50–0.51/0.19–0.20 (2.55–2.63), movable chelal finger length 0.93–0.97. Chelicera 0.57–0.58/0.25 (2.28–2.32), movable finger length 0.30–0.31. Carapace 0.51–0.53/0.50–0.52 (1.02). Leg I: trochanter 0.15–0.16/0.14 (1.07–1.14), femur 0.57–0.58/0.08 (7.13– 7.25), patella 0.30–0.31/0.07 (4.29–4.43), tibia 0.24–0.25/0.06 (4.00–4.17), tarsus 0.62–0.66/0.05 (12.40–13.20). Leg IV: trochanter 0.23/0.14–0.15 (1.53–1.64), femoropatella 0.82–0.83/0.21 (3.90–3.95), tibia 0.57/0.09 (6.33), metatarsus 0.25–0.26/0.07 (3.57–3.71), tarsus 0.71/0.05 (14.20).</p><p>Females: body length 1.91–2.16. Pedipalps: trochanter 0.21–0.22/0.16–0.18 (1.17–1.31), femur 1.10–1.12/0.16– 0.17 (6.59–6.94), patella 0.40–0.45/0.18–0.20 (2.22–2.28), chela 1.56–1.59/0.22–0.23 (6.78–7.23), hand 0.55– 0.57/0.22–0.23 (2.39–2.55), movable chelal finger length 0.98–1.00. Chelicera 0.63–0.64/0.26–0.28 (2.29–2.42), movable finger length 0.33–0.34. Carapace 0.57–0.59/0.57–0.58 (0.98–1.04). Leg I: trochanter 0.17–0.18/0.14– 0.16 (1.06–1.29), femur 0.61–0.63/0.08–0.09 (6.78–7.75), patella 0.32–0.34/0.08 (4.00–4.25), tibia 0.26–0.27/0.06 (4.33–4.50), tarsus 0.65–0.70/0.06 (10.83–11.67). Leg IV: trochanter 0.23–0.28/0.15–0.16 (1.53–1.75), femoropatella 0.91–0.93/0.22–0.23 (4.00–4.23), tibia 0.60–0.63/0.10 (6.00–6.30), metatarsus 0.28–0.29/0.07–0.09 (3.22– 4.00), tarsus 0.75–0.79/0.06 (12.50–13.17).</p><p>Remarks. Tyrannochthonius multidentatus sp. nov. is similar to T. maculosus sp. nov. in having intercalary teeth on both chelal fingers and a small, pointed epistome, but differs by a smaller body size (body length max. 2.16 mm vs. min. 2.22 mm), the relative position of trichobothrium sb (sb closer to b than to st vs. midway between st and b), the trait of teeth on movable chelal finger (markedly retrorse and continuous, without vestigial proximal half teeth vs. slightly retrorse and well-spaced, with vestigial proximal half teeth) and the number of blades of rallum (6–7 vs. 8).</p><p>Tyrannochthonius multidentatus sp. nov. can be distinguished from T. chixing, T. harveyi and T. zhai by having intercalary teeth on both chelal fingers and the number of setae at the anterior margin of the carapace (6 vs. 4), from T. akaelus and T. ganshuanensis by the number of setae on tergites I–II (4 vs. 2), from T. antridraconis by a shorter palp (chela length max. 1.59 mm vs. min. 1.68 mm; palpal femur length max. 1.12 mm vs. min. 1.18 mm), more chelal finger teeth (fixed chelal finger with 62–69 vs. 33–36 teeth; movable chelal finger with 53–58 vs. 30–36 teeth), the number of blades of rallum (6–7 vs. 8) and coxal spines (9–12 vs. 12–15) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the Guanyin and Anjialin caves.</p></div>	https://treatment.plazi.org/id/FD7487B1FFC0FF86FF205EEE398EF9C6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFCAFFB8FF2059D138E1FA82.text	FD7487B1FFCAFFB8FF2059D138E1FA82.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius nanxingensis Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius nanxingensis sp. nov. ‹南fflƟṁae›</p><p>Figs 76–80</p><p>Type material. Holotype ♂ (Ps.- MHBU-HBUARA#2022-502 - 01): China, Guizhou Province, Panzhou City, Zhuhai Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.87224&amp;materialsCitation.latitude=25.595934" title="Search Plazi for locations around (long 104.87224/lat 25.595934)">Nanxing Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.87224&amp;materialsCitation.latitude=25.595934" title="Search Plazi for locations around (long 104.87224/lat 25.595934)">Feng Cave</a>, on the cave walls (Temperature: 16°C, Humidity: 80%) [25°35′45.36″N, 104°52′20.05″E], 1615 m a.s.l., 4 August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun &amp; Wenlong Fan leg. Paratypes: 2 ♂ (Ps.- MHBU-HBUARA#2022-502 - 02 &amp; HBUARA#2022-502-03), 1 ♀ (Ps.- MSWU-HBUARA#2022- 502 - 04), all with the same data as the holotype .</p><p>Etymology. Named after the village of Nanxing, near the type locality.</p><p>Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, represented by a bump; posterior margin of carapace with 2 setae; tergites I–III each with 4 setae. Pedipalps slender, femur 8.38–8.44 (♂), 7.75 (♀) times longer than broad; chela 7.04–7.01 (♂), 7.12 (♀) times longer than broad; both chelal fingers without intercalary teeth, movable chelal finger teeth slightly closer than fixed chelal finger teeth.</p><p>Description. Males (holotype and paratypes) (Figs 77A, 78A–F, 79, 80).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 78C, 79A): carapace 1.02 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, represented by a bump, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 10–12 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 79C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 78D, 79B): large, about as long as carapace, 2.23–2.37 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on dorsal side. Both fingers well provided with teeth, fixed finger with 14–16 teeth, distal one largest; movable finger with 15–18 retrorse contiguous small teeth; galea completely vestigial (Fig. 79B). Serrula exterior with 20–21 and serrula interior with 14–15 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 79E).</p><p>Pedipalp (Figs 78A, B, E, 79D, 80A, B): long and slender, trochanter 1.30–1.35, femur 8.38–8.44, patella 2.24–2.50, chela 7.04–7.11, hand 2.46–2.56 times longer than broad; femur 2.70–2.85 times longer than patella; movable chelal finger 1.78–1.86 times longer than hand and 0.64–0.65 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 78E, 79D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, t situated at same level as est and distal to b (Fig. 80A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 27–28 macrodenticles, long and pointed, slightly retrorse; movable chelal finger with 30–32 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed (Fig. 80A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 78B, 80B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 5: 4–6: 5: 5–6: 5: 5: 4: T2T: 0; tergites VIII and IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 14: 12–13: 9–10: 9: 9–10: 9: 10–11: 9: 0: 2. Anterior genital operculum with 10–12 setae, genital opening slit-like, with 13–16 marginal setae on each side, 38–41 in total (Fig. 78F).</p><p>Legs (Fig. 80C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.80–1.81 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.62–2.71 times longer than tibia. Femoropatella of leg IV 4.52–5.00 times longer than deep; tibia 7.08–8.09 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 4.33–4.67 times longer than deep (TS= 0.24–0.26), tarsus 13.86–14.57 times longer than deep and 2.43–2.49 times longer than metatarsus (TS= 0.30–0.31). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult female (paratype) (Figs 76E, 77B, 78G). Mostly same as males, but a little larger; with same chaetotaxy of coxae as males; tergal chaetotaxy I–XII: 4: 4: 4: 5: 6: 6: 5: 5: 6: 4: T2T: 0; sternal chaetotaxy IV–XII: 14: 9: 9: 10: 10: 10: 11: 0: 2; anterior genital operculum with 10 setae, posterior margin with 12 marginal setae, 22 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 4.00 times longer than deep (TS= 0.28), tarsus 15.83 times longer than deep and 2.64 times longer than metatarsus (TS= 0.41).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 2.59–2.75. Pedipalps: trochanter 0.26–0.27/0.20 (1.30–1.35), femur 1.34–1.35/0.16 (8.38–8.44), patella 0.47–0.50/0.20–0.21 (2.24–2.50), chela 1.92–1.97/0.27–0.28 (7.04–7.11), hand 0.69/0.27–0.28 (2.46–2.56), movable chelal finger length 1.23–1.28. Chelicera 0.69–0.71/0.30–0.31 (2.23–2.37), movable finger length 0.39. Carapace 0.64–0.67/0.63–0.66 (1.02). Leg I: trochanter 0.19–0.22/0.18 (1.06–1.22), femur 0.83–0.85/0.10 (8.30–8.50), patella 0.46–0.47/0.09–0.10 (4.70–5.11), tibia 0.34/0.08 (4.25), tarsus 0.89–0.92/0.06–0.07 (13.14–14.83). Leg IV: trochanter 0.27–0.34/0.16–0.19 (1.69–1.79), femoropatella 1.22–1.25/0.25–0.27 (4.52–5.00), tibia 0.85–0.89/0.11– 0.12 (7.08–8.09), metatarsus 0.39–0.42/0.09 (4.33–4.67), tarsus 0.97–1.02/0.07 (13.86–14.57).</p><p>Female: body length 2.83. Pedipalps: trochanter 0.27/0.20 (1.35), femur 1.24/0.16 (7.75), patella 0.47/0.22 (2.14), chela 1.85/0.26 (7.12), hand 0.64/0.26 (2.46), movable chelal finger length 1.18. Chelicera 0.65/0.30 (2.17), movable finger length 0.39. Carapace 0.61/0.66 (0.92). Leg I: trochanter 0.19/0.17 (1.12), femur 0.76/0.10 (7.60), patella 0.41/0.09 (4.56), tibia 0.32/0.07 (4.57), tarsus 0.81/0.06 (13.50). Leg IV: trochanter 0.32/0.20 (1.60), femoropatella 1.12/0.25 (4.48), tibia 0.81/0.11 (7.36), metatarsus 0.36/0.09 (4.00), tarsus 0.95/0.06 (15.83).</p><p>Remarks. Tyrannochthonius nanxingensis sp. nov. is similar to T. latus sp. nov. in lacking intercalary teeth on both chelal fingers, but differs by a shorter palp (chela length max. 1.97 mm vs. min. 2.14 mm; palpal femur length max. 1.35 mm vs. min. 1.58 mm; movable chelal finger 1.78–1.86 vs. 1.41–1.46 times longer than hand), the relative position of trichobothrium sb (sb closer to b than to st vs. closer to st), the number of setae on tergite IV (5 vs. 4) and the number of blades of rallum (8 vs. 7).</p><p>Tyrannochthonius nanxingensis sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on fixed chelal fingers, from T. harveyi by the number of setae on tergites I–III (4 vs. 2) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FFCAFFB8FF2059D138E1FA82	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFF4FFBFFF205A9238E1F8FE.text	FD7487B1FFF4FFBFFF205A9238E1F8FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius oblongus Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius oblongus sp. nov. ‹ḢŔƟṁae›</p><p>Figs 81–85</p><p>Type material. Holotype ♂ (Ps.-MHBU-GZGY-20-04-01): China, Guizhou Province, Huaxi District, Gaopo Township, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.81957&amp;materialsCitation.latitude=26.280758" title="Search Plazi for locations around (long 106.81957/lat 26.280758)">Houzi Cave</a>, under stones in the deep zone (Temperature: 16°C, Humidity: 75%) [26°16′50.73″N, 106°49′10.46″E], 1485 m a.s.l., 16 August 2020, Zegang Feng, Hongru Xu &amp; Yanmeng Hou leg. Paratypes: 1 ♂ (Ps.-MHBU-GZGY-20-04-02), with the same data as the holotype; 8 ♂ (Ps.- MHBU-HBUARA#2022-481 - 01 – HBUARA#2022-481-08), 2 ♀ (Ps.- MSWU-HBUARA#2022-481 - 09 &amp; HBUARA#2022-481-10), with the same location as the holotype, 26 July 2022, Yanmeng Hou, Lu Zhang &amp; Jianzhou Sun leg.</p><p>Etymology. The specific name is derived from the Latin word “ oblongus ” (= rectangular), which refers to the presence of a nearly rectangular carapace.</p><p>Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and obtuse, represented by a bump; posterior margin of carapace with 2 setae; tergites I–II each with 2 setae. Pedipalps slender, femur 7.80–7.93 (♂), 7.27–7.80 (♀) times longer than broad; chela 7.27 (♂), 6.78–7.09 (♀) times longer than broad; both chelal fingers without intercalary teeth.</p><p>Description. Males (holotype and paratypes) (Figs 81F, 82A, 83A–F, 84, 85).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 83C, 84A): carapace 1.05–1.09 times longer than broad, only slightly narrowed posteriorly resulting in rectangular appearance; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and obtuse, represented by a bump, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 8–9 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 84C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 83D, 84B): large, about as long as carapace, 2.32–2.36 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 15–17 teeth, distal one largest; movable finger with 17–18 retrorse contiguous small teeth; galea completely vestigial (Fig. 84B). Serrula exterior with 17–19 and serrula interior with 13–15 blades. Rallum with 7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 84E).</p><p>Pedipalp (Figs 83A, B, E, 84D, 85A, B): long and slender, trochanter 1.28–1.33, femur 7.80–7.93, patella 2.41–2.67, chela 7.27, hand 2.59–2.64 times longer than broad; femur 2.78–2.85 times longer than patella; movable chelal finger 1.74–1.79 times longer than hand and 0.63–0.64 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 83E, 84D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, t situated at same level as est and distal to b (Fig. 85A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 23–25 macrodenticles, slightly retrorse, long and pointed; movable chelal finger with 22–23 macrodenticles (markedly smaller than teeth on fixed chelal finger), continuous and retrorse, almost prostrate, plus 8–10 vestigial, rounded and contiguous basal teeth, 30–33 in total (Fig. 85A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 83B, 85B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 2: 2: 3–4: 4: 4: 4: 4: 4: 4–6: 4: T2T: 0. Sternal chaetotaxy III–XII: 14–15: 10–13: 8–9: 8–9: 8–9: 9–10: 10: 7–8: 0: 2. Anterior genital operculum with 10–11 setae, genital opening slit-like, with 12–14 marginal setae on each side, 35–37 in total (Fig. 83F).</p><p>Legs (Fig. 85C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.85–1.97 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.63–2.66 times longer than tibia. Femoropatella of leg IV 4.90–4.95 times longer than deep; tibia 7.20–8.00 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 4.57–5.14 times longer than deep (TS= 0.28), tarsus 14.83–15.00 times longer than deep and 2.47–2.81 times longer than metatarsus (TS= 0.31–0.32). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 82B, 83G). Mostly same as males, but a little larger; with same chaetotaxy of coxae as males; tergal chaetotaxy I–XII: 2: 2: 4: 4: 4: 4: 4: 4: 4: 4: T2T: 0; sternal chaetotaxy IV–XII: 10–11: 8–9: 7–8: 7–9: 8–9: 9: 7: 0: 2; anterior genital operculum with 10–12 setae, posterior margin with 14–15 marginal setae, 24–27 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.88–4.57 times longer than deep (TS= 0.28–0.29), tarsus 13.83–17.00 times longer than deep and 2.66–2.68 times longer than metatarsus (TS= 0.29–0.30).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 2.18–2.26. Pedipalps: trochanter 0.20–0.23/0.15–0.18 (1.28–1.33), femur 1.11–1.17/0.14–0.15 (7.80–7.93), patella 0.40–0.41/0.15–0.17 (2.41–2.67), chela 1.60/0.22 (7.27), hand 0.57–0.58/0.22 (2.59–2.64), movable chelal finger length 1.01–1.02. Chelicera 0.58–0.59/0.25 (2.32–2.36), movable finger length 0.32. Carapace 0.59– 0.62/0.56–0.57 (1.05–1.09). Leg I: trochanter 0.17/0.15–0.16 (1.06–1.13), femur 0.67–0.72/0.08–0.09 (8.00–8.38), patella 0.34–0.39/0.08 (4.25–4.88), tibia 0.29–0.30/0.06 (4.83–5.00), tarsus 0.77–0.79/0.06 (12.83–13.17). Leg IV: trochanter 0.25–0.30/0.15–0.17 (1.67–1.76), femoropatella 0.99–1.03/0.20–0.21 (4.90–4.95), tibia 0.72/0.09–0.10 (7.20–8.00), metatarsus 0.32–0.36/0.07 (4.57–5.14), tarsus 0.89–0.90/0.06 (14.83–15.00).</p><p>Females: body length 2.33–2.53. Pedipalps: trochanter 0.21–0.23/0.16–0.17 (1.24–1.44), femur 1.09–1.17/0.15 (7.27–7.80), patella 0.36–0.42/0.17–0.18 (2.12–2.33), chela 1.56/0.22–0.23 (6.78–7.09), hand 0.56–0.57/0.22–0.23 (2.48–2.55), movable chelal finger length 0.98–0.99. Chelicera 0.59/0.25–0.26 (2.27–2.36), movable finger length 0.31. Carapace 0.57–0.59/0.56 (1.02–1.05). Leg I: trochanter 0.17/0.14–0.15 (1.13–1.21), femur 0.67/0.08 (8.38), patella 0.37/0.07 (5.29), tibia 0.28/0.05–0.07 (4.00–5.60), tarsus 0.70–0.72/0.05–0.06 (12.00–14.00). Leg IV: trochanter 0.27–0.28/0.15–0.16 (1.75–1.80), femoropatella 0.96–0.97/0.20 (4.80–4.85), tibia 0.65–0.68/0.09 (7.22– 7.56), metatarsus 0.31–0.32/0.07–0.08 (3.88–4.57), tarsus 0.83–0.85/0.05–0.06 (13.83–17.00).</p><p>Remarks. Tyrannochthonius oblongus sp. nov. is similar to T. brevispinus sp. nov. in having the same number of setae on tergites I–II (2) and lacking intercalary teeth on both chelal fingers, but differs by a larger body size (body length min. 2.18 mm vs. max. 2.12 mm; chela min. 1.56 mm vs. max. 1.43 mm; palpal femur length min. 1.09 mm vs. max. 1.00 mm; movable chelal finger 1.74–1.79 (♂), 1.74–1.75 (♀) vs. 2.05 (♂), 2.00 (♀) times longer than hand), the relative position of trichobothrium sb (sb nearer b than st vs. midway between b and st) and the number of blades of rallum (7 vs. 8).</p><p>Tyrannochthonius oblongus sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on the fixed chelal finger, from T. harveyi by the number of setae at the anterior margin of the carapace (6 vs. 4) and a slender palp (chela 7.27 (♂), 6.78–7.09 (♀) vs. 7.50 (♂), 7.22 (♀) times longer than board, length min. 1.56 mm vs. max. 1.30 mm; palpal femur 7.80–7.93 (♂), 7.27–7.80 (♀) vs. 6.29 (♂), 5.80 (♀) times longer than board, length min. 1.09 mm vs. max. 0.88 mm) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FFF4FFBFFF205A9238E1F8FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFFFFFB1FF205F8838E1F86E.text	FD7487B1FFFFFFB1FF205F8838E1F86E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius parcidentatus Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius parcidentatus sp. nov. ‹mffiƟṁae›</p><p>Figs 86–90</p><p>Type material. Holotype ♂ (Ps.- MHBU-HBUARA#2022-489 - 01): China, Guizhou Province, Dafang County, Lvtang Township, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.485916&amp;materialsCitation.latitude=27.054644" title="Search Plazi for locations around (long 105.485916/lat 27.054644)">Fengchaner Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.485916&amp;materialsCitation.latitude=27.054644" title="Search Plazi for locations around (long 105.485916/lat 27.054644)">Daniu Cave</a>, under the clods and on the cave walls in the deep zone (Temperature: 15°C, Humidity: 85%) [27°3′16.72″N, 105°29′9.31″E], 1574 m a.s.l., 29 July 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun &amp; Wenlong Fan leg. Paratypes: 1 ♂ (Ps.- MHBU-HBUARA#2022-489 - 02), 3 ♀ (Ps.-MSWU-HBUARA#2022-489-03–HBUARA#2022-489-05), all with the same data as the holotype .</p><p>Etymology. The specific name is derived from a combination of the Latin word “ parcus ” and “ dentatus ”, meaning few and toothed, respectively, which refers to the low number of intercalary teeth on the fixed chelal finger.</p><p>Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with 2 setae; tergites I–III each with 2 setae. Pedipalps slender, femur 9.07–9.13 (♂), 8.41–8.81 (♀) times longer than broad; chela 8.77–9.00 (♂), 7.88–8.42 (♀) times longer than broad; only fixed chelal finger with a few intercalary teeth.</p><p>Description. Males (holotype and paratype) (Figs 87A, 88A–C, E, F, H, 89, 90).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 88C, 89A): carapace 0.98–1.02 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, triangular, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 3–4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 11 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 89C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 88E, 89B): large, about as long as carapace, 2.34–2.45 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 20–21 teeth, distal one largest; movable finger with 15 retrorse contiguous small teeth; galea completely vestigial (Fig. 89B). Serrula exterior with 17–18 and serrula interior with 13–14 blades. Rallum with 7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 89E).</p><p>Pedipalp (Figs 88A, B, H, 89D, 90A, B): long and slender, trochanter 1.28–1.41, femur 9.07–9.13, patella 2.50, chela 8.77–9.00, hand 3.32–3.38 times longer than broad; femur 2.72–2.74 times longer than patella; movable chelal finger 1.53–1.55 times longer than hand and 0.58 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 88H, 89D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb midway between b and st; b and t situated subdistally, t situated distal to b and proximal to est (Fig. 90A).A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 26–28 macrodenticles, long and pointed, slightly retrorse, plus 3– 4 intercalary microdenticles, 30–31 in total; movable chelal finger with 44–46 macrodenticles (smaller and denser than teeth on fixed chelal finger), slightly retrorse and pointed, without intercalary teeth (Fig. 90A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 88B, 90B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 2: 2: 2: 2–3: 4: 4: 4: 5: 5: 3–4: T2T: 0; tergites VIII and IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 14–16: 12–13: 8: 8: 7–8: 7–8: 7–8: 7: 0: 2. Anterior genital operculum with 11 setae, genital opening slit-like, with 12–15 marginal setae on each side, 36–39 in total (Fig. 88F).</p><p>Legs (Fig. 90C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.90–1.93 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.34–2.44 times longer than tibia. Femoropatella of leg IV 4.48–4.63 times longer than deep; tibia 6.64–6.73 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.56–3.67 times longer than deep (TS= 0.28–0.30), tarsus 15.50–15.67 times longer than deep and 2.85–2.91 times longer than metatarsus (TS= 0.32–0.33). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 87B, 88D, G). Mostly same as males, but a little larger; with 17–18 setae of carapace arranged s4s: 4: 4: 2: 2; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; tergal chaetotaxy I–XII: 2: 2: 2: 2: 4: 4: 4: 5: 5: 4: T2T: 0; sternal chaetotaxy IV–XII: 12: 8–9: 8: 7: 7: 7: 7: 0: 2; anterior genital operculum with 10–11 setae, posterior margin with 13–14 marginal setae, 23–25 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.67–4.00 times longer than deep (TS= 0.27–0.28), tarsus 15.00–16.33 times longer than deep and 2.72–2.73 times longer than metatarsus (TS= 0.31–0.37).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 2.29–2.62. Pedipalps: trochanter 0.23–0.24/0.17–0.18 (1.28–1.41), femur 1.36–1.37/0.15 (9.07–9.13), patella 0.50/0.20 (2.50), chela 1.89–1.93/0.21–0.22 (8.77–9.00), hand 0.71–0.73/0.21–0.22 (3.32–3.38), movable chelal finger length 1.10–1.12. Chelicera 0.68–0.71/0.29 (2.34–2.45), movable finger length 0.37–0.39. Carapace 0.61–0.62/0.61–0.62 (0.98–1.02). Leg I: trochanter 0.19/0.15–0.17 (1.12–1.27), femur 0.74–0.79/0.09 (8.22–8.78), patella 0.39–0.41/0.08 (4.88–5.13), tibia 0.32–0.35/0.06–0.07 (5.00–5.33), tarsus 0.78–0.82/0.06 (13.00–13.67). Leg IV: trochanter 0.27–0.30/0.15–0.16 (1.69–2.00), femoropatella 1.11–1.12/0.24–0.25 (4.48–4.63), tibia 0.73– 0.74/0.11 (6.64–6.73), metatarsus 0.32–0.33/0.09 (3.56–3.67), tarsus 0.93–0.94/0.06 (15.50–15.67).</p><p>Females: body length 2.64–2.66. Pedipalps: trochanter 0.23–0.25/0.20 (1.15–1.25), femur 1.41–1.43/0.16–0.17 (8.41–8.81), patella 0.51–0.52/0.20–0.22 (2.36–2.55), chela 2.02–2.05/0.24–0.26 (7.88–8.42), hand 0.75–0.77/0.24– 0.26 (2.96–3.13), movable chelal finger length 1.19. Chelicera 0.75–0.77/0.30–0.31 (2.48–2.50), movable finger length 0.41–0.42. Carapace 0.63–0.65/0.63–0.64 (0.98–1.03). Leg I: trochanter 0.18–0.21/0.16–0.17 (1.13–1.24), femur 0.78–0.80/0.10 (7.80–8.00), patella 0.42/0.08–0.09 (4.67–5.25), tibia 0.35–0.37/0.06 (5.83–6.17), tarsus 0.79–0.84/0.06 (13.17–14.00). Leg IV: trochanter 0.28–0.31/0.17–0.18 (1.65–1.72), femoropatella 1.08–1.16/0.24– 0.25 (4.50–4.64), tibia 0.72–0.75/0.11 (6.55–6.82), metatarsus 0.33–0.36/0.09 (3.67–4.00), tarsus 0.90–0.98/0.06 (15.00–16.33).</p><p>Remarks. Tyrannochthonius parcidentatus sp. nov. is similar to T. gracilis sp. nov. in having intercalary teeth on the fixed chelal finger only and a pointed, triangular epistome, but differs by a larger body size but shorter appendages (body length 2.29–2.62 (♂), 2.64–2.66 (♀) vs. 1.96–2.06 (♂), 2.12–2.58 (♀) mm; chela length 1.89–1.93 (♂), 2.02–2.05 (♀) vs. 2.08–2.11 (♂), 2.27–2.29 (♀) mm; movable chelal finger 1.53–1.55 (♂), 1.55–1.59 (♀) times vs. 1.63–1.76 (♂), 1.67–1.83 (♀) times longer than board), fewer fixed chelal finger teeth (30‒31 vs. 42‒45) and the number of blades of rallum (7 vs. 6).</p><p>Tyrannochthonius parcidentatus sp. nov. can be distinguished from T. akaelus, T. antridraconis and T. ganshuanensis by the presence of intercalary teeth on the fixed chelal finger only, from T. harveyi by the presence of intercalary teeth on the fixed chelal finger, from T. chixing and T. zhai by the number of setae on tergites I–III (2 vs. 4) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FFFFFFB1FF205F8838E1F86E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFF9FFA8FF205F883985F86E.text	FD7487B1FFF9FFA8FF205F883985F86E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius pictus Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius pictus sp. nov. ‹AEṞƟṁae›</p><p>Figs 91–96</p><p>Type material. Holotype ♂ (Ps.- MHBU-HBUARA#2022-515 - 01): China, Guizhou Province, Dushan County, Baiquan Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.56324&amp;materialsCitation.latitude=25.74105" title="Search Plazi for locations around (long 107.56324/lat 25.74105)">Jiabao Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.56324&amp;materialsCitation.latitude=25.74105" title="Search Plazi for locations around (long 107.56324/lat 25.74105)">Gaoluo Cave</a>, under stones near the cave entrance and in the deep zone (Temperature: 13°C, Humidity: 90%) [25°44′27.78″N, 107°33′47.64″E], 967 m a.s.l., 11 August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun &amp; Wenlong Fan leg. Paratypes: 5 ♂ (Ps.- MHBU-HBUARA#2022-515 - 02 –HBUARA#2022-515- 06), 7 ♀ (Ps.- MSWU-HBUARA#2022-515 - 07 –HBUARA#2022-515-13), all with the same data as the holotype; 16 ♂ (Ps.- MHBU-HBUARA#2022-516 - 01 –HBUARA#2022-516-16), 13 ♀ (Ps.- MSWU-HBUARA#2022-516 - 17 –HBUARA#2022-516-29), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.56771&amp;materialsCitation.latitude=25.74222" title="Search Plazi for locations around (long 107.56771/lat 25.74222)">Jiabao Village</a>, Cangjun Cave [25°44′31.99″N, 107°34′3.76″E], 948 m a.s.l., with the same collection date and collectors as the holotype .</p><p>Etymology. The specific name is derived from the Latin word “ pictus ” (= figured), which refers to the dorsal surface of the cheliceral palm possessing rounded whitish patches.</p><p>Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, without epistome; posterior margin of carapace with 2 setae; tergite I with 2 setae, tergites III–VIII each with 4 setae. Pedipalps slender, femur 7.29–7.71 (♂), 7.07–7.29 (♀) times longer than broad; chela 8.06–8.50 (♂), 7.24–8.00 (♀) times longer than broad; both chelal fingers without intercalary teeth, movable chelal finger teeth smaller and denser than fixed chelal finger teeth.</p><p>Description. Males (holotype and paratypes) (Figs 93A, 94A–F, 95, 96).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 94C, 95A): carapace 1.02–1.08 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; without epistome; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 8–9 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 95C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 94D, 95B): large, about as long as carapace, 2.36–2.52 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 13–16 teeth, distal one largest; movable finger with 15–17 retrorse contiguous small teeth; galea represented by a very slight bump on movable finger (Fig. 95B). Serrula exterior with 17–20 and serrula interior with 13–15 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 95E).</p><p>Pedipalp (Figs 94A, B, E, 95D, 96A, B): long and slender, trochanter 1.24–1.31, femur 7.29–7.71, patella 2.27–2.40, chela 8.06–8.50, hand 2.83–3.00 times longer than broad; femur 2.83–2.97 times longer than patella; movable chelal finger 1.76–1.79 times longer than hand and 0.62–0.63 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 94E, 95D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated distal to b; est situated distal to t (Fig. 96A).A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 30–32 macrodenticles, slightly retrorse, long and pointed; movable chelal finger with 40–43 macrodenticles (smaller and denser than teeth on fixed chelal finger), slightly retrorse and pointed (Fig. 96A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 94B, 96B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 2: 2–4: 4: 4: 4: 4: 4: 4: 4–5: 4: T2T: 0. Sternal chaetotaxy III–XII: 12–15: 12–13: 8–9: 8–9: 9: 9: 9–10: 9: 0: 2. Anterior genital operculum with 9–11 setae, genital opening slit-like, with 15–17 marginal setae on each side, 41–46 in total (Fig. 94F).</p><p>Legs (Fig. 96C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.81–2.00 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.31–2.37 times longer than tibia. Femoropatella of leg IV 3.87–4.00 times longer than deep; tibia 6.50–7.33 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.50–3.63 times longer than deep (TS= 0.25–0.34), tarsus 13.17–15.00 times longer than deep and 2.59–2.72 times longer than metatarsus (TS= 0.24–0.31). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 91F, 92F, 93B, 94G). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3–4, I 3, II 4, III 5, IV 5; tergal chaetotaxy I–XII: 2: 3–4: 4: 4: 4: 4: 4: 4: 4–5: 4: T2T: 0; sternal chaetotaxy IV–XII: 11–12: 8–10: 9: 8–9: 8–9: 8–10: 9–10: 0: 2; anterior genital operculum with 10 setae, posterior margin with 12–15 marginal setae, 22–25 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.13–3.86 times longer than deep (TS= 0.30–0.36), tarsus 11.83–15.40 times longer than deep and 2.72–2.88 times longer than metatarsus (TS= 0.25–0.30).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 1.80–2.09. Pedipalps: trochanter 0.20–0.21/0.16–0.17 (1.24–1.31), femur 0.98–1.08/0.13–0.14 (7.29–7.71), patella 0.34–0.38/0.15–0.16 (2.27–2.40), chela 1.45–1.57/0.18–0.19 (8.06–8.50), hand 0.51–0.55/0.18–0.19 (2.83–3.00), movable chelal finger length 0.91–0.98. Chelicera 0.52–0.55/0.21–0.23 (2.36–2.52), movable finger length 0.28–0.30. Carapace 0.47–0.54/0.46–0.51 (1.02–1.08). Leg I: trochanter 0.16–0.17/0.14–0.15 (1.07–1.21), femur 0.58–0.65/0.08 (7.25–8.13), patella 0.32–0.33/0.07 (4.57–4.71), tibia 0.27–0.29/0.05–0.06 (3.50–5.40), tarsus 0.64–0.68/0.05–0.06 (11.17–13.60). Leg IV: trochanter 0.24–0.26/0.14–0.15 (1.67–1.86), femoropatella 0.87– 0.95/0.22–0.24 (3.87–4.00), tibia 0.61–0.66/0.09–0.10 (6.50–7.33), metatarsus 0.28–0.29/0.08 (3.50–3.63), tarsus 0.73–0.79/0.05–0.06 (13.17–15.00).</p><p>Females: body length 2.18–2.25. Pedipalps: trochanter 0.20–0.23/0.16–0.17 (1.25–1.44), femur 1.02– 1.08/0.14–0.15 (7.07–7.29), patella 0.35–0.39/0.16–0.17 (2.19–2.44), chela 1.46–1.56/0.19–0.21 (7.24–8.00), hand 0.54–0.56/0.19–0.21 (2.67–2.84), movable chelal finger length 0.88–0.99. Chelicera 0.55–0.57/0.24 (2.29–2.38), movable finger length 0.29–0.32. Carapace 0.53–0.55/0.51–0.54 (1.00–1.04). Leg I: trochanter 0.16/0.14–0.15 (1.07–1.14), femur 0.58–0.66/0.08 (7.25–8.25), patella 0.31–0.33/0.07–0.08 (4.00–4.71), tibia 0.27–0.29/0.05–0.06 (4.67–5.80), tarsus 0.64–0.69/0.05–0.06 (10.67–13.60). Leg IV: trochanter 0.24–0.27/0.14–0.16 (1.56–1.93), femoropatella 0.86–0.92/0.22–0.23 (3.74–4.00), tibia 0.58–0.63/0.09–0.10 (5.90–7.00), metatarsus 0.25–0.29/0.07–0.08 (3.13–3.86), tarsus 0.71–0.79/0.05–0.06 (11.83–15.40).</p><p>Remarks. Tyrannochthonius pictus sp. nov. is similar to T. planus sp. nov. in lacking an epistome on the carapace as well as intercalary teeth on both chelal fingers, but differs by having a greater number of teeth on the chelal fingers (fixed chelal finger with 30–32 vs. 20–21 teeth; movable chelal finger with 40–43 vs. 25–26 teeth) and the number of blades of rallum (8 vs. 7).</p><p>Tyrannochthonius pictus sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on the fixed chelal finger, from T. harveyi by the number of setae at the anterior margin of the carapace (6 vs. 4), a slender palp (chela 8.06–8.50 (♂), 7.24–8.00 (♀) times vs. 7.50 (♂), 7.22 (♀) times longer than board, length min. 1.45 mm vs. max. 1.30 mm; palpal femur 7.29–7.71 (♂), 7.07–7.29 (♀) times vs. 6.29 (♂), 5.80 (♀) times longer than board, length min. 0.98 mm vs. max. 0.88 mm) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the Gaoluo and Cangjun caves.</p></div>	https://treatment.plazi.org/id/FD7487B1FFF9FFA8FF205F883985F86E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFE0FFA1FF205F8838E1F92A.text	FD7487B1FFE0FFA1FF205F8838E1F92A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius pinguis Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius pinguis sp. nov. ‹ợaeƟṁae›</p><p>Figs 97–101</p><p>Type material. Holotype ♂ (Ps.- MHBU-HBUARA#2022-510 - 01): China, Guizhou Province, Ziyun County, Daying Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.31135&amp;materialsCitation.latitude=25.486408" title="Search Plazi for locations around (long 106.31135/lat 25.486408)">Daying Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.31135&amp;materialsCitation.latitude=25.486408" title="Search Plazi for locations around (long 106.31135/lat 25.486408)">Zharou Cave</a>, under the stones and clods in the deep zone (Temperature: 15°C, Humidity: 90%) [25°29′11.07″N, 106°18′40.83″E], 1102 m a.s.l., 8 August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun &amp; Wenlong Fan leg. Paratypes: 1 ♂ (Ps.- MHBU-HBUARA#2022-510 - 02), 3 ♀ (Ps.- MSWU-HBUARA#2022-510 - 03 –HBUARA#2022-510-05), all with the same data as the holotype .</p><p>Etymology. The specific name is derived from the Latin word “ pinguis ”, meaning fat, which refers to the enlarged abdomen present in this species.</p><p>Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, without epistome; posterior margin of carapace with 2 setae; tergites I–IV each with 2 setae. Pedipalps slender, femur 8.07–8.14 (♂), 7.79–8.33 (♀) times longer than broad; chela 7.45–7.95 (♂), 7.52 (♀) times longer than broad; both chelal fingers without intercalary teeth, movable chelal finger teeth markedly smaller than fixed chelal finger teeth.</p><p>Description. Males (holotype and paratype) (Figs 98A, 99A–F, 100, 101).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 99C, 100A): carapace 1.06 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; without epistome; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 3–4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 8–9 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 100C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 99D, 100B): large, about as long as carapace, 2.39–2.41 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 10–11 teeth, distal one largest; movable finger with 13–15 retrorse contiguous small teeth; galea completely vestigial (Fig. 100B). Serrula exterior with 19–22 and serrula interior with 14–15 blades. Rallum with 7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 100E).</p><p>Pedipalp (Figs 99A, B, E, 100D, 101A, B): long and slender, trochanter 1.18–1.25, femur 8.07–8.14, patella 2.71–2.88, chela 7.45–7.95, hand 2.80–2.95 times longer than broad; femur 2.46–2.48 times longer than patella; movable chelal finger 1.61–1.68 times longer than hand and 0.60–0.62 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 99E, 100D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, t situated at same level as est and distal to b (Fig. 101A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 22–23 macrodenticles, slightly retrorse, long and pointed; movable chelal finger with 31–33 macrodenticles (markedly smaller than teeth on fixed chelal finger), continuous and retrorse, almost prostrate (Fig. 101A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 99B, 101B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 2: 2: 2: 2: 4: 4: 4: 4: 4: 2: T2T: 0. Sternal chaetotaxy III–XII: 12–13: 10–12: 8: 7–8: 7: 7–8: 7: 7: 0: 2. Anterior genital operculum with 10 setae, genital opening slit-like, with 15–16 marginal setae on each side, 41 in total (Fig. 99F).</p><p>Legs (Fig. 101C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.76–1.92 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.82–2.85 times longer than tibia. Femoropatella of leg IV 4.42–4.64 times longer than deep; tibia 6.90–7.11 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 4.13 times longer than deep (TS= 0.24–0.30), tarsus 14.67–14.83 times longer than deep and 2.67–2.70 times longer than metatarsus (TS= 0.25–0.28). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 98B, 99G). Mostly same as males, but a little larger; with same chaetotaxy of coxae as males; tergal chaetotaxy I–XII: 2: 2: 2: 2: 4: 4: 4: 4: 4–5: 2: T2T: 0; sternal chaetotaxy IV–XII: 11–12: 8: 7: 7: 7: 7: 7: 0: 2; anterior genital operculum with 10 setae, posterior margin with 10–12 marginal setae, 20–22 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.88–4.38 times longer than deep (TS= 0.29), tarsus 15.50–16.20 times longer than deep and 2.61–2.66 times longer than metatarsus (TS= 0.21–0.28).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 1.94–2.13. Pedipalps: trochanter 0.20/0.16–0.17 (1.18–1.25), femur 1.13–1.14/0.14 (8.07–8.14), patella 0.46/0.16–0.17 (2.71–2.88), chela 1.49–1.51/0.19–0.20 (7.45–7.95), hand 0.56/0.19–0.20 (2.80–2.95), movable chelal finger length 0.90–0.94. Chelicera 0.53–0.55/0.22–0.23 (2.39–2.41), movable finger length 0.29–0.30. Carapace 0.53–0.55/0.50–0.52 (1.06). Leg I: trochanter 0.18/0.15–0.16 (1.13–1.20), femur 0.67–0.71/0.08 (8.38–8.88), patella 0.37–0.38/0.07 (5.29–5.43), tibia 0.27–0.28/0.06 (4.50–4.67), tarsus 0.77–0.79/0.06 (12.83–13.17). Leg IV: trochanter 0.27–0.28/0.16–0.17 (1.59–1.75), femoropatella 1.02–1.06/0.22–0.24 (4.42–4.64), tibia 0.64–0.69/0.09– 0.10 (6.90–7.11), metatarsus 0.33/0.08 (4.13), tarsus 0.88–0.89/0.06 (14.67–14.83).</p><p>Females: body length 1.84–2.24. Pedipalps: trochanter 0.19–0.23/0.15–0.17 (1.27–1.35), femur 1.09–1.25/0.14– 0.15 (7.79–8.33), patella 0.45–0.50/0.15–0.17 (2.94–3.00), chela 1.43–1.58/0.19–0.21 (7.52), hand 0.52–0.61/0.19– 0.21 (2.74–2.90), movable chelal finger length 0.88–0.96. Chelicera 0.52–0.55/0.22–0.24 (2.29–2.36), movable finger length 0.28–0.30. Carapace 0.52–0.54/0.50–0.51 (1.04–1.06). Leg I: trochanter 0.14–0.19/0.15–0.16 (0.93– 1.19), femur 0.68–0.75/0.08 (8.50–9.38), patella 0.34–0.40/0.07–0.08 (4.86–5.00), tibia 0.27–0.29/0.05–0.06 (4.83–5.40), tarsus 0.75–0.81/0.06 (12.50–13.50). Leg IV: trochanter 0.27–0.28/0.14–0.16 (1.75–1.93), femoropatella 0.95–1.10/0.21 (4.52–5.24), tibia 0.63–0.70/0.09–0.10 (7.00), metatarsus 0.31–0.35/0.08 (3.88–4.38), tarsus 0.81–0.93/0.05–0.06 (15.50–16.20).</p><p>Remarks. Tyrannochthonius pinguis sp. nov. is similar to T. umidus sp. nov. in lacking an epistome on the carapace as well as intercalary teeth on both chelal fingers, but differs by the presence of more robust chela (chela 7.45–7.95 (♂), 7.52 (♀) vs. 8.44–9.06 (♂), 7.75–8.33 (♀) times longer than board; movable chelal finger 1.61–1.68 (♂), 1.57–1.69 (♀) vs. 1.79–1.83 (♂), 1.73–1.92 (♀) times longer than hand), the relative position of trichobothrium sb (sb 1.67–1.70 vs. 1.20–1.22 times as far from st as from b) and the number of blades of coxal spines (8–9 vs. 12–13).</p><p>Tyrannochthonius pinguis sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on the fixed chelal finger, from T. harveyi by the number of setae at the anterior margin of the carapace (6 vs. 4) and a larger body size (body length min. 1.84 mm vs. max. 1.56 mm; chelal length min. 1.43 mm vs. max. 1.30 mm; palpal femur length min. 1.09 mm vs. max. 0.88 mm) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FFE0FFA1FF205F8838E1F92A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFEDFF5BFF2059FA38E1F9B2.text	FD7487B1FFEDFF5BFF2059FA38E1F9B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius planus Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius planus sp. nov. ‹ṬËƟṁae›</p><p>Figs 102–106</p><p>Type material. Holotype ♂ (Ps.- MHBU-GZC190801-01 - 01): China, Guizhou Province, Sandu County, Zhonghe Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.93352&amp;materialsCitation.latitude=25.71335" title="Search Plazi for locations around (long 107.93352/lat 25.71335)">Sandong Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.93352&amp;materialsCitation.latitude=25.71335" title="Search Plazi for locations around (long 107.93352/lat 25.71335)">Yanggong Cave</a>, under the stones in the deep zone (Temperature: 13°C, Humidity: 90%) [25°42′48.06″N, 107°56′00.66″E], 800 m a.s.l., 1 August 2019, Zegang Feng, Zhaoyi Li &amp; Chen Zhang leg. Paratypes: 1 ♂ (Ps.- MHBU-GZC190801-01 - 02), 2 ♀ (Ps.- MSWU-GZC190801-01 - 03 &amp; GZC190801-01 - 04), all with the same data as the holotype .</p><p>Etymology. The specific name is derived from the Latin word “ planus ”, meaning flat, which refers to the relatively straight anterior margin of the carapace.</p><p>Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated and relatively straight, without epistome; posterior margin of carapace with 2 setae; tergites II–VIII each with 4 setae. Pedipalps slender, femur 7.36–7.38 (♂), 6.79– 7.29 (♀) times longer than broad; chela 8.11–8.38 (♂), 7.68–7.82 (♀) times longer than broad; both chelal fingers without intercalary teeth, movable chelal finger teeth smaller and denser than fixed chelal finger teeth.</p><p>Description. Males (holotype and paratype) (Figs 103A, 104A–F, 105, 106).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 104C, 105A): carapace 0.98–1.00 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; without epistome; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 8–9 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 105C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 104D, 105B): large, about as long as carapace, 2.45–2.48 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 12–13 teeth, distal one largest; movable finger with 11–13 retrorse contiguous small teeth; galea represented by a very slight bump on movable finger (Fig. 105B). Serrula exterior with 17–19 and serrula interior with 13–14 blades. Rallum with 7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 105E).</p><p>Pedipalp (Figs 104A, B, E, 105D, 106A, B): long and slender, trochanter 1.47–1.67, femur 7.36–7.38, patella 2.29–2.33, chela 8.11–8.38, hand 2.89–3.00 times longer than broad; femur 2.94–3.00 times longer than patella; movable chelal finger 1.75 times longer than hand and 0.62–0.63 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 104E, 105D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated distal to b and proximal to est (Fig. 106A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 20–21 macrodenticles, slightly retrorse, long and pointed; movable chelal finger with 25–26 macrodenticles (smaller and denser than teeth on fixed chelal finger), retrorse and pointed (Fig. 106A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 104B, 106B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 2–3: 4: 4: 4: 4: 4: 4: 4: 4: 4: T2T: 0. Sternal chaetotaxy III–XII: 12–13: 11–12: 8: 7: 8: 9: 9: 9: 0: 2. Anterior genital operculum with 10–11 setae, genital opening slit-like, with 15–18 marginal setae on each side, 42–45 in total (Fig. 104F).</p><p>Legs (Fig. 106C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.81–1.94 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.21–2.24 times longer than tibia. Femoropatella of leg IV 4.05–4.32 times longer than deep; tibia 6.20–6.33 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.86–4.29 times longer than deep (TS= 0.26–0.27), tarsus 11.50–14.40 times longer than deep and 2.40–2.56 times longer than metatarsus (TS= 0.29). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 103B, 104G). Mostly same as males, but a little larger; with the same chaetotaxy of coxae as males; tergal chaetotaxy I–XII: 2–4: 4: 4: 4: 4: 4: 4: 4: 4–5: 4: T2T: 0; sternal chaetotaxy IV–XII: 11: 8–10: 9: 9: 9: 8–10: 8–9: 0: 2; anterior genital operculum with 9–10 setae, posterior margin with 12 marginal setae, 21–22 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.63–4.00 times longer than deep (TS= 0.25–0.31), tarsus 11.00–12.17 times longer than deep and 2.36–2.52 times longer than metatarsus (TS= 0.29).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 1.81–1.94. Pedipalps: trochanter 0.25/0.15–0.17 (1.47–1.67), femur 0.96–1.03/0.13–0.14 (7.36–7.38), patella 0.32–0.35/0.14–0.15 (2.29–2.33), chela 1.34–1.46/0.16–0.18 (8.11–8.38), hand 0.48–0.52/0.16–0.18 (2.89– 3.00), movable chelal finger length 0.84–0.91. Chelicera 0.49–0.52/0.20–0.21 (2.45–2.48), movable finger length 0.27–0.29. Carapace 0.46–0.49/0.47–0.49 (0.98–1.00). Leg I: trochanter 0.16/0.12–0.14 (1.14–1.33), femur 0.56– 0.62/0.07–0.08 (7.75–8.00), patella 0.31–0.32/0.06–0.07 (4.57–5.17), tibia 0.25–0.29/0.06 (4.17–4.83), tarsus 0.56– 0.64/0.05 (11.20–12.80). Leg IV: trochanter 0.21–0.24/0.13–0.14 (1.62–1.71), femoropatella 0.82–0.89/0.19–0.22 (4.05–4.32), tibia 0.57–0.62/0.09–0.10 (6.20–6.33), metatarsus 0.27–0.30/0.07 (3.86–4.29), tarsus 0.69–0.72/0.05– 0.06 (11.50–14.40).</p><p>Females: body length 1.94–2.01. Pedipalps: trochanter 0.22–0.27/0.14–0.15 (1.57–1.80), femur 0.95–1.02/0.14 (6.79–7.29), patella 0.33–0.37/0.15 (2.20–2.47), chela 1.33–1.46/0.17–0.19 (7.68–7.82), hand 0.48–0.53/0.17– 0.19 (2.79–2.82), movable chelal finger length 0.82–0.89. Chelicera 0.50–0.53/0.21–0.23 (2.30–2.38), movable finger length 0.27–0.30. Carapace 0.49–0.53/0.47–0.49 (1.04–1.08). Leg I: trochanter 0.16/0.14 (1.14), femur 0.58–0.61/0.07–0.08 (7.63–8.29), patella 0.31–0.32/0.07 (4.43–4.57), tibia 0.27–0.28/0.05–0.06 (4.50–5.60), tarsus 0.59–0.66/0.05–0.06 (11.00–11.80). Leg IV: trochanter 0.22–0.25/0.14–0.15 (1.57–1.67), femoropatella 0.82– 0.88/0.20–0.21 (4.10–4.19), tibia 0.59–0.62/0.09 (6.56–6.89), metatarsus 0.28–0.29/0.07–0.08 (3.63–4.00), tarsus 0.66–0.73/0.06 (11.00–12.17).</p><p>Remarks. Tyrannochthonius planus sp. nov. is similar to T. pictus sp. nov. in lacking an epistome on the carapace as well as intercalary teeth on both chelal fingers, but differs by having fewer teeth on both chelal fingers (fixed chelal finger with 20–21 vs. 30–32 teeth; movable chelal finger with 25–26 vs. 40–43 teeth), the trait of the teeth at movable chelal fingertip (distinctly retrorse vs. slightly retrorse) and the number of blades of rallum (7 vs. 8).</p><p>Tyrannochthonius planus sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on the fixed chelal finger, from T. harveyi by the number of setae at the anterior margin of the carapace (6 vs. 4) and on tergites II–III (4 vs. 2), the presence of a slender palp (chela 8.11–8.38 (♂), 7.68–7.82 (♀) times vs. 7.50 (♂), 7.22 (♀) times longer than board, length min. 1.33 mm vs. max. 1.30 mm; palpal femur 7.36–7.38 (♂), 6.79–7.29 (♀) times vs. 6.29 (♂), 5.80 (♀) times longer than board, length min. 0.95 mm vs. max. 0.88 mm) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FFEDFF5BFF2059FA38E1F9B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF17FF5EFF20594238E1F80A.text	FD7487B1FF17FF5EFF20594238E1F80A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius qilinensis Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius qilinensis sp. nov. ‹ḆĦƟṁae›</p><p>Figs 107–110</p><p>Type material. Holotype ♀ (Ps.-MHBU-GZJH-19-26-35): China, Guizhou Province, Jianhe County, Yangasha Sub-district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.46168&amp;materialsCitation.latitude=26.76613" title="Search Plazi for locations around (long 108.46168/lat 26.76613)">Chuandong Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.46168&amp;materialsCitation.latitude=26.76613" title="Search Plazi for locations around (long 108.46168/lat 26.76613)">Qilin Cave</a>, under a stone in the deep zone (Temperature: 15°C, Humidity: 80%) [26°45′58.07″N, 108°27′42.06″E], 743 m a.s.l., 13 October 2019, Zegang Feng &amp; Lingchen Zhao leg.</p><p>Etymology. Named after the type locality, Qilin Cave.</p><p>Diagnosis (♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, represented by a bump; posterior margin of carapace with 2 setae; tergites I–VI each with 2 setae. Pedipalps slender, femur 7.72 times longer than broad; chela 7.62 times longer than broad; both chelal fingers with intercalary teeth, but present at fingertips only; chelal fingers curved in dorsal view; chemosensory setae present on dorsum of chelal hand.</p><p>Description. Female (holotype), male unknown (Figs 107–110).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 108C, 109A): carapace 0.93 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, represented by a bump, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 11–12 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 109C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 108D, 109B): large, about as long as carapace, 2.38 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 17 teeth, distal one largest; movable finger with 13 retrorse contiguous small teeth; galea completely vestigial (Fig. 109B). Serrula exterior with 18 and serrula interior with 13 blades. Rallum with 8 blades, the distal one longer and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 109E).</p><p>Pedipalp (Figs 108A, B, F, 109D, 110A, B): long and slender, trochanter 1.45, femur 7.72, patella 2.65, chela 7.62, hand 2.88 times longer than broad; femur 2.28 times longer than patella; movable chelal finger 1.67 times longer than hand and 0.63 times longer than chela. Setae generally long and acuminate; one distal lyrifissures present on patella (Figs 108F, 109D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, b situated at same level as it and proximal to t (Fig. 110A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 43 macrodenticles, slightly retrorse and pointed, plus 4 intercalary microdenticles (present at fingertip only, till et), 47 in total; movable chelal finger with 35 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 4 intercalary microdenticles (present at fingertip only, till et on fixed chelal finger) and 10 vestigial, rounded and contiguous basal teeth, 49 in total (Fig. 110A). Chelal fingers markedly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 108B, 110B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 2: 2: 2: 2: 4: 4: 5: 4: 5: 4: T2T: 0. Sternal chaetotaxy IV–XII: 13: 10: 7: 8: 7: 9: 7: 0: 2. Anterior genital operculum with 11 setae, genital opening slit-like, with 14 marginal setae on each side, 25 in total (Fig. 108E).</p><p>Legs (Fig. 110C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 2.00 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.39 times longer than tibia. Femoropatella of leg IV 5.08 times longer than deep; tibia 6.23 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 4.30 times longer than deep (TS= 0.30), tarsus 14.00 times longer than deep and 2.28 times longer than metatarsus (TS= 0.27). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Female: body length 2.41. Pedipalps: trochanter 0.32/0.22 (1.45), femur 1.39/0.18 (7.72), patella 0.61/0.23 (2.65), chela 1.98/0.26 (7.62), hand 0.75/0.26 (2.88), movable chelal finger length 1.25. Chelicera 0.76/0.32 (2.38), movable finger length 0.40. Carapace 0.68/0.73 (0.93). Leg I: trochanter 0.25/0.18 (1.39), femur 0.86/0.10 (8.60), patella 0.43/0.09 (4.78), tibia 0.38/0.07 (5.43), tarsus 0.91/0.07 (13.00). Leg IV: trochanter 0.34/0.19 (1.79), femoropatella 1.27/0.25 (5.08), tibia 0.81/0.13 (6.23), metatarsus 0.43/0.10 (4.30), tarsus 0.98/0.07 (14.00).</p><p>Remarks. Male unknown. Tyrannochthonius qilinensis sp. nov. is similar to T. akaelus in having intercalary teeth on both chelal fingers and a small epistome, but differs by a larger body size (body length 2.41 vs. 2.10 mm; chela length 1.98 vs. 1.28 mm; palpal femur 7.72 vs. 6.60 times longer than board, length 1.39 vs. 0.90 mm; movable chelal finger 1.67 vs. 2.10 times longer than hand; femoropatella of leg IV length 1.27 vs. 0.79 mm), the number of setae on tergite IV (2 vs. 4) and the position of trichobothrium t (t situated distal to est vs. at same level as it / est).</p><p>It is worth mentioning that T. qilinensis sp. nov. is similar to T. pandus in having a pair of curved chelal fingers, but differs by the presence of intercalary teeth on both chelal fingers and a larger body size (body length 2.41 vs. 1.67 mm; chela 7.62 vs. 7.06 times longer than board, length 1.98 vs. 1.20 mm; palpal femur 7.72 vs. 6.77 times longer than board, length 1.39 vs. 0.88 mm) (Hou et al. 2022a).</p><p>Tyrannochthonius qilinensis sp. nov. can be distinguished from T. chixing, T. zhai and T. harveyi by the presence of intercalary teeth on both chelal fingers and the number of setae at the anterior margin of the carapace (6 vs. 4), from T. antridraconis by the number of setae on the tergites I–IV (2 vs. 4), from T. ganshuanensis by the number of setae on tergite IV (2 vs. 4), a larger size (body length 2.41 (♀) vs. 1.40–1.80 (♀) mm; chela 7.62 (♀) vs. 6.90–7.30 (♀) times longer than board, length 1.98 (♀) vs. 1.21–1.44 (♀) mm; palpal femur 7.72 (♀) vs. 5.90–6.70 (♀) times longer than board, length 1.39 (♀) vs. 0.95–0.97 (♀) mm; movable chelal finger 1.67 (♀) vs. 2.00–2.20 (♀) times longer than hand) and the relative position of trichobothrium sb (sb closer to st than to b vs. closer to b) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FF17FF5EFF20594238E1F80A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF11FF53FF205F8838E1F8FE.text	FD7487B1FF11FF53FF205F8838E1F8FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius quattuor Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius quattuor sp. nov. ‹四ƭƟṁae›</p><p>Figs 111–115</p><p>Type material. Holotype ♂ (Ps.- MHBU-HBUARA#2022-484 - 01): China, Guizhou Province, Dafang County, Yangchang Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.6528&amp;materialsCitation.latitude=27.078781" title="Search Plazi for locations around (long 105.6528/lat 27.078781)">Guanjiadadong Cave</a>, under stones within 50–100 m from the cave entrance (Temperature: 17°C, Humidity: 85%) [27°4′43.61″N, 105°39′10.09″E], 1555 m a.s.l., 28 July 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun &amp; Wenlong Fan leg. Paratypes: 2 ♂ (Ps.- MHBU-HBUARA#2022-484 - 02 &amp; HBUARA#2022-484-03), 9 ♀ (Ps.- MSWU-HBUARA#2022-484 - 04 –HBUARA#2022-484-12), all with the same data as the holotype .</p><p>Etymology. The specific name is derived from the Latin word “ quattuor ”, meaning four, which refers to the tergites (except tergite XII) each with four setae.</p><p>Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with 2 setae; tergites I–XI each with 4 setae (except the females). Pedipalps slender, femur 7.80–8.40 (♂), 7.33–7.75 (♀) times longer than broad; chela 7.91–8.25 (♂), 6.89–7.25 (♀) times longer than broad; both chelal fingers without intercalary teeth, movable chelal finger teeth markedly smaller than fixed chelal finger teeth and distinctly retrorse.</p><p>Description. Males (holotype and paratypes) (Figs 112A, 113A–F, 114, 115).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 113C, 114A): carapace 1.02–1.04 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, triangular, with 2 setae flanking base; with 18 setae arranged s4–5s: 4–5: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 13–15 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 114C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 113D, 114B): large, about as long as carapace, 2.36–2.48 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 18–21 teeth, distal one largest; movable finger with 17–18 retrorse contiguous small teeth; galea completely vestigial (Fig. 114B). Serrula exterior with 22–24 and serrula interior with 14–15 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 114E).</p><p>Pedipalp (Figs 113A, B, E, 114D, 115A, B): long and slender, trochanter 1.18–1.28, femur 7.80–8.40, patella 2.44–2.50, chela 7.91–8.25, hand 2.91–2.95 times longer than broad; femur 2.66–2.80 times longer than patella; movable chelal finger 1.64–1.69 times longer than hand and 0.60–0.61 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 113E, 114D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated distal to b and proximal to est (Fig. 115A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 31–35 macrodenticles, slightly retrorse, long and pointed; movable chelal finger with 49–51 macrodenticles (markedly smaller than teeth on fixed chelal finger), continuous and markedly retrorse (Fig. 115A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 113B, 115B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4: 4: 4: 4: 4: T2T: 0. Sternal chaetotaxy III–XII: 14–15: 13: 9–10: 8–9: 7–9: 8–9: 9: 9: 0: 2. Anterior genital operculum with 9–11 setae, genital opening slit-like, with 14–16 marginal setae on each side, 38–42 in total (Fig. 113F).</p><p>Legs (Fig. 115C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 2.06–2.13 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.23–2.35 times longer than tibia. Femoropatella of leg IV 4.17–4.79 times longer than deep; tibia 6.10–6.30 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.44–3.50 times longer than deep (TS= 0.36–0.39), tarsus 12.33–13.67 times longer than deep and 2.64–2.65 times longer than metatarsus (TS= 0.26–0.32). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 112B, 113G). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5–6; tergal chaetotaxy I–XII: 3–4: 4: 4: 4: 4: 4: 4: 4: 4–5: 4: T2T: 0; sternal chaetotaxy IV–XII: 12–14: 9: 7–8: 9: 8–9: 9–10: 9: 0: 2; anterior genital operculum with 10 setae, posterior margin with 14–16 marginal setae, 24–26 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.20–3.33 times longer than deep (TS= 0.27–0.41), tarsus 13.00–13.50 times longer than deep and 2.53–2.60 times longer than metatarsus (TS= 0.33–0.37).</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 2.28–2.48. Pedipalps: trochanter 0.20–0.23/0.17–0.18 (1.18–1.28), femur 1.17–1.26/0.15 (7.80–8.40), patella 0.44–0.45/0.18 (2.44–2.50), chela 1.65–1.74/0.20–0.22 (7.91–8.25), hand 0.59–0.64/0.20–0.22 (2.91–2.95), movable chelal finger length 1.00–1.05. Chelicera 0.59–0.62/0.25 (2.36–2.48), movable finger length 0.33–0.34. Carapace 0.56–0.59/0.55–0.57 (1.02–1.04). Leg I: trochanter 0.16/0.15–0.16 (1.00–1.07), femur 0.64–0.68/0.08–0.09 (7.56–8.00), patella 0.30–0.33/0.07 (4.29–4.71), tibia 0.30–0.31/0.06 (5.00–5.17), tarsus 0.67–0.73/0.06 (11.17– 12.17). Leg IV: trochanter 0.26–0.28/0.16–0.17 (1.63–1.65), femoropatella 0.91–0.96/0.19–0.23 (4.17–4.79), tibia 0.61–0.63/0.10 (6.10–6.30), metatarsus 0.28–0.31/0.08–0.09 (3.44–3.50), tarsus 0.74–0.82/0.06 (12.33–13.67).</p><p>Females: body length 2.48–2.79. Pedipalps: trochanter 0.23–0.24/0.18–0.19 (1.26–1.28), femur 1.24– 1.32/0.16–0.18 (7.33–7.75), patella 0.46–0.50/0.18–0.21 (2.38–2.56), chela 1.74–1.86/0.24–0.27 (6.89–7.25), hand 0.63–0.70/0.24–0.27 (2.59–2.63), movable chelal finger length 1.05–1.02. Chelicera 0.62–0.71/0.27–0.29 (2.30–2.45), movable finger length 0.35–0.38. Carapace 0.58–0.63/0.58–0.62 (1.00–1.02). Leg I: trochanter 0.17– 0.19/0.16–0.17 (1.06–1.12), femur 0.66–0.75/0.09–0.10 (7.33–7.50), patella 0.35–0.38/0.07–0.09 (4.22–5.00), tibia 0.31–0.34/0.06–0.07 (4.86–5.17), tarsus 0.72–0.75/0.06 (12.00–12.50). Leg IV: trochanter 0.28–0.30/0.17–0.18 (1.65–1.67), femoropatella 0.94–1.01/0.22–0.25 (4.04–4.27), tibia 0.60–0.63/0.10–0.11 (5.45–6.30), metatarsus 0.30–0.32/0.09–0.10 (3.20–3.33), tarsus 0.78–0.81/0.06 (13.00–13.50).</p><p>Remarks. Tyrannochthonius quattuor sp. nov. is similar to T. latus sp. nov. in having the same number of setae on tergites I–VII (4) and lacking intercalary teeth on both chelal fingers, but differs by having a smaller body size (body length 2.28–2.45 (♂), 2.48–2.79 (♀) vs. 2.59–2.80 (♂), 2.91–3.11 (♀) mm; chela 7.91–8.25 (♂), 6.89–7.25 (♀) times vs. 6.69–7.13 (♂), 6.15–6.58 (♀) times longer than board, length 1.65–1.74 (♂), 1.74–1.86 (♀) vs. 2.14– 2.34 (♂), 2.37–2.46 (♀) mm; palpal femur 7.80–8.40 (♂), 7.33–7.75 (♀) times vs. 9.29–9.33 (♂), 8.75–8.79 (♀) times longer than board, length 1.17–1.26 (♂), 1.24–1.32 (♀) vs. 1.58–1.68 (♂), 1.67–1.75 (♀) mm), the trait of teeth on the movable chelal finger (continuous and markedly retrorse vs. well-spaced and slightly retrorse) and the number of blades of rallum (8 vs. 7) and coxal spines (13–15 vs. 11–12).</p><p>Tyrannochthonius quattuor sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on the fixed chelal finger, from T. harveyi by the number of setae at the anterior margin of the carapace (6 vs. 4) and on tergites I–III (4 vs. 2) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FF11FF53FF205F8838E1F8FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF1BFF55FF205F8838E1F840.text	FD7487B1FF1BFF55FF205F8838E1F840.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius umidus Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius umidus sp. nov. ‹AEÎƟṁae›</p><p>Figs 116–120</p><p>Type material. Holotype ♂ (Ps.-MHBU-GZGY-20-06-01): China, Guizhou Province, Wudang District, Shuitian Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.81707&amp;materialsCitation.latitude=26.715212" title="Search Plazi for locations around (long 106.81707/lat 26.715212)">Lizi Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.81707&amp;materialsCitation.latitude=26.715212" title="Search Plazi for locations around (long 106.81707/lat 26.715212)">Da Cave</a>, under the stones in the deep zone (Temperature: 14°C, Humidity: 95%) [26°42′54.76″N, 106°49′1.46″E], 1334 m a.s.l., 17 August 2020, Zegang Feng, Hongru Xu &amp; Yanmeng Hou leg. Paratypes: 13 ♂ (Ps.-MHBU-GZGY-20-06-02–GZGY-20-06-14), 7 ♀ (Ps.-MSWU-GZGY-20-06-15–GZGY-20-06-21), all with the same data as the holotype .</p><p>Etymology. The specific name is derived from the Latin word “ umidus ”, meaning humid, which refers to the species that live in an extremely humid environment.</p><p>Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, without epistome; posterior margin of carapace with 2 setae; tergites I–III each with 2 setae. Pedipalps slender, femur 7.86–7.93 (♂), 7.60–8.23 (♀) times longer than broad; chela 8.44–9.06 (♂), 7.75–8.33 (♀) times longer than broad; both chelal fingers without intercalary teeth, movable chelal finger teeth markedly smaller than fixed chelal finger teeth.</p><p>Description. Males (holotype and paratypes) (Figs 116E, 117A, 118A–F, 119, 120).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 118C, 119A): carapace 1.13–1.14 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; without epistome; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 12–13 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 119C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 118D, 119B): large, about as long as carapace, 2.26–2.41 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 9–11 teeth, distal one largest; movable finger with 13–16 retrorse contiguous small teeth; galea completely vestigial (Fig. 119B). Serrula exterior with 20–22 and serrula interior with 12–14 blades. Rallum with 7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 119E).</p><p>Pedipalp (Figs 118A, B, E, 119D, 120A, B): long and slender, trochanter 1.25–1.31, femur 7.86–7.93, patella 2.80, chela 8.44–9.06, hand 2.94–3.12 times longer than broad; femur 2.62–2.64 times longer than patella; movable chelal finger 1.79–1.83 times longer than hand and 0.63 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 118E, 119D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, t situated at same level as est and distal to b (Fig. 120A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 26–29 macrodenticles, slightly retrorse, long and pointed; movable chelal finger with 39–42 macrodenticles (markedly smaller than teeth on fixed chelal finger), continuous and retrorse, almost prostrate (Fig. 120A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 118B, 120B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 2: 2: 2: 1–2: 4: 3–4: 4: 4: 4: 2: T2T: 0. Sternal chaetotaxy III–XII: 11–12: 11–12: 8: 7: 7–8: 7: 9: 8–9: 0: 2. Anterior genital operculum with 9 setae, genital opening slit-like, with 13–16 marginal setae on each side, 36–39 in total (Fig. 118F).</p><p>Legs (Fig. 120C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.88–2.06 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.41–2.52 times longer than tibia. Femoropatella of leg IV 4.36–4.60 times longer than deep; tibia 6.00–6.30 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 4.29 times longer than deep (TS= 0.30), tarsus 16.00–16.40 times longer than deep and 2.67–2.73 times longer than metatarsus (TS= 0.26–0.29). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Adult females (paratypes) (Figs 117B, 118G). Mostly same as males, but a little larger; with same chaetotaxy of coxae as males; tergal chaetotaxy I–XII: 2: 2: 2: 2: 4: 4: 4: 4: 4: 2: T2 T: 0; sternal chaetotaxy IV–XII: 10–12: 8–9: 7: 7–9: 8–9: 9: 8: 0: 2; anterior genital operculum with 10–11 setae, posterior margin with 10–12 marginal setae, 21–22 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.75–3.86 times longer than deep (TS= 0.30–0.33), tarsus 14.80–15.40 times longer than deep and 2.57–2.74 times longer than metatarsus (TS= 0.25–0.31) .</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 2.00–2.04. Pedipalps: trochanter 0.20–0.21/0.16 (1.25–1.31), femur 1.10–1.11/0.14 (7.86–7.93), patella 0.42/0.15 (2.80), chela 1.52–1.54/0.17–0.18 (8.44–9.06), hand 0.53/0.17–0.18 (2.94–3.12), movable chelal finger length 0.95–0.97. Chelicera 0.52–0.53/0.22–0.23 (2.26–2.41), movable finger length 0.27–0.29. Carapace 0.54– 0.57/0.48–0.50 (1.13–1.14). Leg I: trochanter 0.15–0.17/0.15 (1.00–1.13), femur 0.64–0.66/0.08 (8.00–8.25), patella 0.32–0.34/0.07 (4.57–4.86), tibia 0.27–0.29/0.05–0.06 (4.83–5.40), tarsus 0.68–0.70/0.05 (13.60–14.00). Leg IV: trochanter 0.23–0.25/0.15–0.16 (1.44–1.67), femoropatella 0.92–0.96/0.20–0.22 (4.36–4.60), tibia 0.60–0.63/0.10 (6.00–6.30), metatarsus 0.30/0.07 (4.29), tarsus 0.80–0.82/0.05 (16.00–16.40).</p><p>Females: body length 1.82–2.12. Pedipalps: trochanter 0.20/0.16–0.17 (1.18–1.25), femur 1.07–1.14/0.13–0.15 (7.60–8.23), patella 0.40–0.44/0.15–0.16 (2.67–2.75), chela 1.50–1.55/0.18–0.20 (7.75–8.33), hand 0.51–0.56/0.18– 0.20 (2.80–2.83), movable chelal finger length 0.97–0.98. Chelicera 0.51–0.54/0.21–0.23 (2.35–2.43), movable finger length 0.28–0.30. Carapace 0.52–0.55/0.48–0.51 (1.08). Leg I: trochanter 0.14–0.16/0.13–0.14 (1.08–1.14), femur 0.61–0.67/0.08 (7.63–8.38), patella 0.32–0.33/0.07 (4.57–4.71), tibia 0.26–0.29/0.06 (4.33–4.83), tarsus 0.69–0.70/0.06 (11.50–11.67). Leg IV: trochanter 0.20–0.22/0.14–0.15 (1.43–1.47), femoropatella 0.90–0.95/0.17– 0.18 (5.00–5.59), tibia 0.59–0.64/0.09–0.10 (6.40–6.56), metatarsus 0.27–0.30/0.07–0.08 (3.75–3.86), tarsus 0.74– 0.77/0.05 (14.80–15.40).</p><p>Remarks. Tyrannochthonius umidus sp. nov. is similar to T. pinguis sp. nov. in lacking an epistome on the carapace as well as intercalary teeth on both chelal fingers, but differs by the presence of more slender chela (chela 8.44–9.06 (♂), 7.75–8.33 (♀) vs. 7.45–7.95 (♂), 7.52 (♀) times longer than board; movable chelal finger 1.79–1.83 (♂), 1.73–1.92 (♀) times vs. 1.61–1.68 (♂), 1.57–1.69 (♀) times longer than hand), the relative position of trichobothrium sb (sb 1.20–1.22 times vs. 1.67–1.70 times as far from st as from b) and the number of blades of coxal spines (12–13 vs. 8–9).</p><p>Tyrannochthonius umidus sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on the fixed chelal finger, from T. harveyi by the number of setae at the anterior margin of the carapace (6 vs. 4), the number of setae on sternites VI–Ⅹ (7–9 vs. 6) and a larger body size (body length min. 1.82 mm vs. max. 1.56 mm; chelal length min. 1.50 mm vs. max. 1.30 mm; palpal femur length min. 1.07 mm vs. max. 0.88 mm) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FF1BFF55FF205F8838E1F840	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF05FF42FF205F8838E1FA56.text	FD7487B1FF05FF42FF205F8838E1FA56.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius yanshanensis Hou & Feng & Zhang 2023	<div><p>Tyrannochthonius yanshanensis sp. nov. ‹ȒƜƟṁae›</p><p>Figs 121–125</p><p>Type material. Holotype ♀ (Ps.-MHBU-GZWC-20-17-01): China, Guizhou Province, Wuchuan County, Yanshan Town, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.89641&amp;materialsCitation.latitude=28.803144" title="Search Plazi for locations around (long 107.89641/lat 28.803144)">Yanshan Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.89641&amp;materialsCitation.latitude=28.803144" title="Search Plazi for locations around (long 107.89641/lat 28.803144)">Yanshan Cave</a>, on the cave walls in the deep zone (Temperature: 17°C, Humidity: 70%) [28°48′11.32″N, 107°53′47.07″E], 951 m a.s.l., 24 August 2020, Zegang Feng, Hongru Xu &amp; Yanmeng Hou leg. Paratype: 1 ♀ (Ps.-MSWU-GZWC-20-17-02), with the same data as the holotype .</p><p>Etymology. Named after the type locality, Yanshan Cave.</p><p>Diagnosis (♀). Small-sized troglomorphic species with slightly elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, rounded and obtuse, represented a slight bump; posterior margin of carapace with 2 setae; tergites I–III each with 4 setae. Palpal femur 4.92–5.08 times longer than broad; chela 5.69–5.75 times longer than broad; both chelal fingers with intercalary teeth; chemosensory setae present on dorsum of chelal hand.</p><p>Description. Female (holotype and paratype), male unknown (Figs 122–125).</p><p>Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</p><p>Cephalothorax (Figs 123C, 124A): carapace 0.83–0.87 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small, rounded and obtuse, represented by a slight bump; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 7–8 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 124C); intercoxal tubercle absent; without sub-oral seta.</p><p>Chelicera (Figs 123D, 124B): large, about as long as carapace, 1.96–2.04 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation both dorsal and ventral side. Both fingers well provided with teeth, fixed finger with 13 teeth, distal one largest; movable finger with 12–13 retrorse contiguous small teeth; galea completely vestigial (Fig. 124B). Serrula exterior with 18 and serrula interior with 13–14 blades. Rallum with 7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 124E).</p><p>Pedipalp (Figs 123A, B, F, 124D, 125A, B): long and slender, trochanter 1.17–1.33, femur 4.92–5.08, patella 1.86–1.93, chela 5.69–5.75, hand 1.81 times longer than broad; femur 2.19–2.35 times longer than patella; movable chelal finger 2.17–2.21 times longer than hand and 0.69–0.70 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 123F, 124D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated distal to b and at same level as est (Fig. 125A).A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 29–30 macrodenticles, slightly retrorse and pointed, plus 14– 15 intercalary microdenticles, 44 in total; movable chelal finger with 9 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 8 intercalary microdenticles and 20–22 vestigial, rounded and contiguous basal teeth, 37–39 in total (Fig. 125A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 123B, 125B).</p><p>Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 4–5: 5: 5: 5–6: 6: 6: 5: T2T: 0, tergites VI–IX each with an unpaired median seta. Sternal chaetotaxy IV–XII: 12–13: 9–10: 9–10: 9: 10–11: 9–11: 9: 0: 2. Anterior genital operculum with 10 setae, genital opening slit-like, with 12–13 marginal setae on each side, 22–23 in total (Fig. 123E).</p><p>Legs (Fig. 125C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.87–2.13 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.06–2.19 times longer than tibia. Femoropatella of leg IV 2.59–2.65 times longer than deep; tibia 3.88–4.13 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 2.50 times longer than deep (TS= 0.33), tarsus 7.75–8.00 times longer than deep and 2.07–2.13 times longer than metatarsus (TS= 0.23–0.25). Arolium slightly shorter than the claws, not divided; claws simple.</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Females: body length 1.29–1.33. Pedipalps: trochanter 0.14–0.16/0.12 (1.17–1.33), femur 0.59–0.61/0.12 (4.92–5.08), patella 0.26–0.27/0.14 (1.86–1.93), chela 0.91–0.92/0.16 (5.69–5.75), hand 0.29/0.16 (1.81), movable chelal finger length 0.63–0.64. Chelicera 0.49–0.51/0.25 (1.96–2.04), movable finger length 0.27. Carapace 0.43–0.45/0.52 (0.83–0.87). Leg I: trochanter 0.12/0.10–0.11 (1.09–1.20), femur 0.28–0.32/0.06 (4.67–5.33), patella 0.15/0.06 (2.50), tibia 0.16–0.17/0.05 (3.20–3.40), tarsus 0.35/0.04–0.05 (7.00–8.75). Leg IV: trochanter 0.15–0.16/0.10 (1.50–1.60), femoropatella 0.44–0.45/0.17 (2.59–2.65), tibia 0.31–0.33/0.08 (3.88–4.13), metatarsus 0.15/0.06 (2.50), tarsus 0.31–0.32/0.04 (7.75–8.00).</p><p>Remarks. Male unknown. Tyrannochthonius yanshanensis sp. nov. is similar to T. breviculus sp. nov. in having intercalary teeth on both chelal fingers and a small, rounded and obtuse epistome, but differs by a longer palp (chela length 0.91–0.92 (♀) vs. 0.78–0.79 (♀) mm; palpal femur 4.92–5.08 (♀) vs. 5.00–5.30 (♀) times longer than board, length 0.59–0.61 (♀) vs. 0.53–0.55 (♀) mm; movable chelal finger 2.17–2.21 (♀) vs. 2.04 (♀) times longer than hand), the number of setae on the tergites VIII‒IX (6 vs. 5), the relative position of trichobothrium sb (sb nearer st than b vs. midway between st and b) and the number of blades of rallum (7 vs. 8).</p><p>Tyrannochthonius yanshanensis sp. nov. can be distinguished from T. chixing, T. harveyi and T. zhai by the presence of intercalary teeth on the movable chelal finger, from T. akaelus and T. ganshuanensis by the number of setae on tergites Ⅰ–II (4 vs. 2) and from T. antridraconis by a smaller size (body length max. 1.33 mm vs. min. 1.80 mm; chela 5.69–5.75 vs. 6.90–7.55 times longer than board, length 0.91–0.92 vs. 1.68–1.70 mm; palpal femur 4.92–5.08 vs. 7.10–7.50 times longer than board) (Mahnert 2009; Gao et al. 2018, 2020).</p><p>Distribution. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/FD7487B1FF05FF42FF205F8838E1FA56	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF0EFF40FF2059273F8CFAEC.text	FD7487B1FF0EFF40FF2059273F8CFAEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius Chamberlin 1929	<div><p>Key to the species of Tyrannochthonius from China (subspecies not included)</p><p>1 Carapace with eyes or eyespots.......................................................................... 2</p><p>- Carapace without eyes or eyespots....................................................................... 6</p><p>2 Without corneate eyes, only anterior pair of eyespots present................................... T. infirmus sp. nov.</p><p>- With at least a pair of well-developed corneate eyes......................................................... 3</p><p>3 Chelal finger with intercalary teeth....................................................................... 4</p><p>- Chelal finger without intercalary teeth.................................................................... 5</p><p>4 Troglomorphic species with larger body size and elongate appendages: e.g. body length min. 1.73 (♂, ♀) mm, palpal femur length min. 0.76 (♂, ♀) mm................................................................. T. duo sp. nov.</p><p>- Epigean species with smaller body size and without elongated appendages: e.g. body length 1.20 (♀) mm, palpal femur length 0.47 (♀) mm...................................................................... T. robustus Beier, 1951</p><p>5 Carapace with 18 setae and a dentate epistome; tergites VIII‒IX each with 8 setae......... T. japonicus (Ellingsen, 1907)</p><p>- Carapace with 16 setae and a small triangular epistome; tergites VIII‒IX each with 6 setae..................................................................................................... T. pachythorax Redikorzev, 1938</p><p>6 At least one finger of chela with intercalary teeth............................................................ 7</p><p>- Both chelal fingers without intercalary teeth............................................................... 23</p><p>7 Only fixed chelal finger with intercalary teeth.............................................................. 8</p><p>- Both chelal fingers with intercalary teeth................................................................. 11</p><p>8 Tergites I–III each with 2 setae; carapace anterior margin with 6 setae........................................... 9</p><p>- Tergites I–III each with 4 setae; carapace anterior margin with 4 setae......................................... ... 10</p><p>9 More chelal finger teeth: e.g. fixed chelal finger with 28‒30 macrodenticles, plus 14‒ 15 intercalary microdenticles, 42‒45 in total; smaller body size but longer appendages: e.g. body length 1.96–2.06 (♂), 2.12–2.58 (♀) mm; chela length 2.08–2.11 (♂), 2.27–2.29 (♀) mm; movable chelal finger 1.63–1.76 times longer than hand and 0.60–0.62 times longer than chela............................................................................................. T. gracilis sp. nov.</p><p>- Fewer chelal finger teeth: e.g. fixed chelal finger with 26‒28 macrodenticles, plus 3‒ 4 intercalary microdenticles, 30‒31 in total; larger body size but shorter appendages: e.g. body length 2.29–2.62 (♂), 2.64–2.66 (♀) mm; chela length 1.89–1.93 (♂), 2.02–2.05 (♀) mm; movable chelal finger 1.53–1.55 times longer than hand and 0.58 times longer than chela............................................................................................. T. parcidentatus sp. nov.</p><p>10 Epistome present; body length 1.64 (♀) mm, chela 5.39 (♀) times longer than board (length 1.24 (♀) mm); chelal fingers with fewer teeth: fixed chelal finger with 31–32 teeth, movable chelal finger with 6–7 teeth, smooth and well-spaced.............................................................................. T. chixingi Gao, Wynne &amp; Zhang, 2018</p><p>- Epistome absent; body length 1.40 (♀) mm, chela 6.37 (♀) times longer than board (length 1.21 (♀) mm); chelal fingers with more teeth: fixed chelal finger with 37 teeth, movable chelal finger with 26 teeth, retrorse, pointed and continuous............................................................................... T. zhai Gao, Zhang and Chen, 2020</p><p>11 Tergites I–II each with 2 setae.......................................................................... 12</p><p>- Tergites I–II each with 4 setae........................................................................ ... 14</p><p>12 Smaller: body length max. 2.10 (♀) mm, palpal femur length max. 0.97 (♀) mm, chela length max. 1.44 (♀) mm; tergite IV with 4 setae........................................................................................ 13</p><p>- Larger: body length 2.41 (♀) mm, palpal femur length 1.39 (♀) mm, chela length 1.98 (♀) mm; tergite IV with 2 setae........................................................................................ T. qilinensis sp. nov.</p><p>13 With a slightly slender chela: hand 2.50 (♀) times, femur 6.60 (♀) times, chela 7.70 (♀) times longer than broad; fewer numerous intercalary teeth; trichobothrium sb indistinctly nearer st than b ......................... T. akaelus Mahnert, 2009</p><p>- With a slightly stouter chela: hand 2.30 (♀) times, femur 5.90–6.70 (♀) times, chela 6.90–7.30 (♀) times longer than broad; more numerous intercalary teeth; trichobothrium sb slightly nearer b than st ............. T. ganshuanensis Mahnert, 2009</p><p>14 Smaller: body length max. 1.40 (♂), 1.69 (♀) mm; chela length max. 0.88 (♂), 0.93 (♀) mm; epistome small, obtuse.... 15</p><p>- Larger: body length min. 1.56 (♂), 1.77 (♀) mm; chela length min. 1.21 (♂), 1.24 (♀) mm; epistome large, pointed...... 17</p><p>15 Coxae II with 7–9 terminally indented coxal spines on each side; fewer fixed chelal finger teeth (40–44); chela 5.69–5.85 (♂), 5.57–5.75 (♀) times longer than board................................................................... 16</p><p>- Coxae II with 10 terminally indented coxal spines on each side; more fixed chelal finger teeth (56–57); chela 6.29–6.62 (♂), 6.07–6.20 (♀) times longer than board...................................................... T. arificus sp. nov.</p><p>16 Tergites VIII‒IX each with 6 setae; rallum with 7 pinnate blades; chela length 0.91–0.92 (♀) mm; palpal femur 4.92–5.08 (♀) times longer than board (length 0.59–0.61 (♀) mm); movable chelal finger 2.17–2.21 (♀) times longer than hand; trichobothrium sb nearer st than b ............................................................. T. yanshanensis sp. nov.</p><p>- Tergites VIII‒IX each with 5 setae; rallum with 8 pinnate blades; chela length 0.74–0.76 (♂), 0.78–0.79 (♀) mm; palpal femur 5.10–5.33 (♂), 5.00–5.30 (♀) times longer than board (length 0.48–0.51 (♂), 0.53–0.55 (♀) mm); movable chelal finger 2.12–2.13 (♂), 2.04 (♀) times longer than hand; trichobothrium sb midway between st and b ........ T. breviculus sp. nov.</p><p>17 Body length generally min. 2.00 (♂, ♀) mm, chela length min. 1.43 (♂, ♀) mm; palpal femur length min. 0.97 (♂, ♀) mm.. .................................................................................................. 18</p><p>- Body length max. 1.99 (♂) mm, chela length max. 1.29 (♂) mm; palpal femur length max. 0.89 (♂) mm T. hispidus sp. nov.</p><p>18 Trichobothrium sb nearer b than st or midway between st and b ............................................... 19</p><p>- Trichobothrium sb nearer st than b .......................................................... T. acutus sp. nov.</p><p>19 Shorter appendages: chela length max. 1.59 (♂, ♀) mm; palpal femur length max. 1.10 (♂, ♀) mm; femoropatella of leg IV length max. 0.93 (♂, ♀) mm........................................................................... 20</p><p>- Longer appendages: chela length min. 1.60 (♂, ♀) mm; palpal femur length min. 1.15 (♂, ♀) mm; femoropatella of leg IV length min. 1.01 (♂, ♀) mm........................................................................... 21</p><p>20 Movable chelal finger teeth markedly retrorse and continuous, without vestigial proximal half teeth; trichobothrium sb nearer b than st; rallum with 6–7 pinnate blades............................................... T. multidentatus sp. nov.</p><p>- Movable chelal finger teeth slightly retrorse and well-spaced, with vestigial proximal half teeth; trichobothrium sb midway between st and b; rallum with 8 pinnate blades............................................. T. maculosus sp. nov.</p><p>21 Trichobothrium sb nearer b than st; body length max. 2.35 (♂, ♀) mm, chela 7.27–8.00 times longer than board (length max. 1.76 (♂, ♀) mm), palpal femur 7.30–8.30 times longer than board; tergite VIII with 5–6 setae....................... 22</p><p>- Trichobothrium sb midway between st and b; body length min. 2.36 (♂, ♀) mm, chela 8.25–8.57 times longer than board (length min. 1.80 (♂, ♀) mm), palpal femur 8.50–8.79 times longer than board; tergite VIII with 4 setae.... T. altus sp. nov.</p><p>22 Intercalary teeth present in the distal 2/3 of fixed chelal finger and till trichobothrium sb of movable chelal finger; tergite VII with 4 setae, tergite X with 4–5 setae; chela 7.9–8.0 (♂), 6.9–7.55 (♀) times longer than broad; coxae II with 12–15 terminally indented coxal spines on each side; rallum with 8 pinnate blades...................... T. antridraconis Mahnert, 2009</p><p>- Intercalary teeth present in tips (nearly 1/3) of both chelal fingers only; tergite VII with 5 setae, tergite X with 2 setae; chela 7.27 (♂) times longer than broad; coxae II with 8–9 terminally indented coxal spines on each side; rallum with 7 pinnate blades................................................................................ T. babaowanensis sp. nov.</p><p>23 Chelal fingers straight or slightly curved in dorsal view...................................................... 24</p><p>- Chelal fingers markedly curved in dorsal view.................................. T. pandus Hou, Gao &amp; Zhang, 2022</p><p>24 Epistome absent..................................................................................... 25</p><p>- Epistome present.................................................................................... 28</p><p>25 Tergites II–IV with 1–2 setae; trichobothrium sb nearer b than st .............................................. 26</p><p>- Tergites II–IV with 3–4 setae; trichobothrium sb nearer st than b .............................................. 27</p><p>26 Trichobothrium sb of movable chelal finger 1.20–1.22 times as far from st as from b; chela 8.44–9.06 (♂), 7.75–8.33 (♀) times longer than board; coxae II with 12–13 terminally indented coxal spines on each side................. T. umidus sp. nov.</p><p>- Trichobothrium sb of movable chelal finger 1.67–1.70 times as far from st as from b; chela 7.45–7.95 (♂), 7.52 (♀) times longer than board; coxae II with 8–9 terminally indented coxal spines on each side........................ T. pinguis sp. nov.</p><p>27 Fewer chelal fingers teeth: fixed chelal finger with 20–21 teeth, movable chelal finger with 25–26 teeth; slightly smaller body size (body length 1.81–1.94 (♂), 1.94–2.01 (♀) mm; chela length 1.34–1.46 (♂), 1.33–1.46 (♀) mm); rallum with 7 pinnate blades................................................................................ T. planus sp. nov.</p><p>- More chelal fingers teeth: fixed chelal finger with 30–32 teeth, movable chelal finger with 40–43 teeth; slightly larger body size (body length 1.80–2.09 (♂), 2.18–2.25 (♀) mm; chela length 1.45–1.57 (♂), 1.46–1.56 (♀) mm); rallum with 8 pinnate blades................................................................................. T. pictus sp. nov.</p><p>28 Tergites I–II each with 2 setae.......................................................................... 29</p><p>- Tergites I–II each with 3–4 setae........................................................................ 32</p><p>29 Tergites IV and X each with 4 setae; chela 7.27–7.50 times longer than board.................................... 30</p><p>- Tergites IV and X each with 2 setae; chela 7.53–8.20 times longer than board.................... T. multicavus sp. nov.</p><p>30 Carapace anterior margin with 6 setae and posterior margin with 2 setae; body length min. 1.73 (♂, ♀) mm, chela length min. 1.33 (♂, ♀) mm, palpal femur length min. 0.93 (♂, ♀) mm; rallum with 7–8 pinnate blades......................... 31</p><p>- Carapace anterior margin with 4 setae and posterior margin with 4 setae; body length max. 1.56 (♂, ♀) mm, chela length max. 1.30 (♂, ♀) mm, palpal femur length max. 0.88 (♂, ♀) mm; rallum with 6 pinnate blades................................................................................................. T. harveyi Gao, Zhang and Chen, 2020</p><p>31 Larger: body length 2.18–2.26 (♂), 2.33–2.53 (♀) mm, chela 7.27 (♂), 6.78–7.09 (♀) times longer than board (length 1.60 (♂), 1.56 (♀) mm), palpal femur length 1.11–1.17 (♂), 1.09–1.17 (♀) mm; movable chelal finger 1.74–1.79 (♂), 1.74–1.75 (♀) times longer than hand; trichobothrium sb nearer b than st; rallum with 7 pinnate blades............. T. oblongus sp. nov.</p><p>- Smaller: body length 1.73 (♂), 2.12 (♀) mm, chela 7.39 (♂), 7.53 (♀) times longer than board (length 1.33 (♂), 1.43 (♀) mm), palpal femur length 0.93 (♂), 1.00 (♀) mm; movable chelal finger 2.05 (♂), 2.00 (♀) times longer than hand; trichobothrium sb midway between st and b; rallum with 8 pinnate blades................................... T. brevispinus sp. nov.</p><p>32 Movable chelal finger teeth slightly retrorse and well-spaced; coxae II with 10–12 terminally indented coxal spines on each side............................................................................................... 33</p><p>- Movable chelal finger teeth markedly retrorse and continuous; coxae II with 13–15 terminally indented coxal spines on each side................................................................................. T. quattuor sp. nov.</p><p>33 Larger: body length 2.59–2.80 (♂), 2.91–3.11 (♀) mm, chela 2.14–2.34 (♂), 2.37–2.46 (♀) mm, movable chelal finger 1.41– 1.46 times longer than hand, palpal femur length 1.58–1.68 (♂), 1.67–1.75 (♀) mm; trichobothrium sb nearer st than b; rallum with 7 pinnate blades; tergite IV with 4 setae................................................... T. latus sp. nov.</p><p>- Smaller: body length 2.59–2.75 (♂), 2.83 (♀) mm, chela 1.92–1.97 (♂), 1.85 (♀) mm, movable chelal finger 1.78–1.86 times longer than hand, palpal femur length 1.34–1.35 (♂), 1.24 (♀) mm; trichobothrium sb nearer b than st; rallum with 8 pinnate blades; tergite IV with 5 setae........................................................ T. nanxingensis sp. nov.</p></div>	https://treatment.plazi.org/id/FD7487B1FF0EFF40FF2059273F8CFAEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hou, Yanmeng;Feng, Zegang;Zhang, Feng	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
