taxonID	type	description	language	source
FD7487B1FF90FFDCFF205EA73978FDA7.taxon	type_taxon	Type species: Chthonius terribilis With, 1906, by original designation.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF90FFD6FF205D5138E1FDAB.taxon	description	Figs 2 – 6	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF90FFD6FF205D5138E1FDAB.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-GZKL- 19 - 23 - 01): China, Guizhou Province, Kaili City, Zhouxi Town, Gazuida Cave, under stones within 20 – 200 m from the cave entrance (Temperature: 16 ° C, Humidity: 70 %) [26 ° 28 ′ 58.77 ″ N, 107 ° 55 ′ 38.24 ″ E], 716 m a. s. l., 12 October 2019, Zegang Feng & Lingchen Zhao leg. Paratypes: 2 ♂ (Ps. - MHBU-GZKL- 19 - 23 - 02 & GZKL- 19 - 23 - 03), 4 ♀ (Ps. - MSWU-GZKL- 19 - 23 - 04 – GZKL- 19 - 23 - 07), all with the same data as the holotype.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF90FFD6FF205D5138E1FDAB.taxon	etymology	Etymology. The specific name is derived from the Latin word “ acutus ”, meaning acute, which refers to the distinctly sharp epistome.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF90FFD6FF205D5138E1FDAB.taxon	diagnosis	Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome distinctly pointed, triangular; posterior margin of carapace with 2 setae; tergites II – VI each with 4 setae. Pedipalps slender, femur 6.58 – 6.67 (♂), 6.63 – 6.94 (♀) times longer than broad; chela 7.29 – 7.65 (♂), 7.38 – 7.79 (♀) times longer than broad; both chelal fingers with intercalary teeth, movable chelal finger teeth distinctly more retrorse than those of the fixed chelal finger.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF90FFD6FF205D5138E1FDAB.taxon	description	Description. Males (holotype and paratypes) (Figs 3 A, 4 A – F, 5, 6). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 4 C, 5 A, C): carapace 0.96 – 1.00 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome distinctly pointed, triangular, with 2 setae flanking base; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 10 – 11 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 5 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 4 D, 5 B): large, about as long as carapace, 2.42 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 20 – 23 teeth, fourth one largest; movable finger with 24 – 26 retrorse contiguous small teeth; galea completely vestigial. Serrula exterior with 24 – 26 and serrula interior with 14 – 15 blades. Rallum with 7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 5 E). Pedipalp (Figs 4 A, B, E, 5 D, 6 A, B): long and slender, trochanter 1.10 – 1.20, femur 6.58 – 6.67, patella 2.27 – 2.35, chela 7.29 – 7.65, hand 2.46 – 2.57 times longer than broad; femur 2.50 – 2.55 times longer than patella; movable chelal finger 1.92 – 1.93 times longer than hand and 0.65 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissures present on patella (Figs 4 E, 5 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming an oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated at same level as est and distal to b (Fig. 6 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 44 – 46 macrodenticles, long and pointed, plus 13 – 14 intercalary microdenticles, 57 – 60 in total; movable chelal finger with 43 – 46 macrodenticles (markedly smaller than teeth on fixed chelal finger), markedly retrorse and pointed, plus 15 – 16 intercalary microdenticles, 58 – 62 in total (Fig. 6 A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 4 B, 6 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4 – 5: 4: 4: 4: 4: 4: 4: 4 – 5: 4 – 5: 4: T 2 T: 0, tergites VII – IX each with an unpaired median seta. Sternal chaetotaxy III – XII: 12 – 13: 12 – 13: 8 – 9: 7 – 8: 7: 7: 7 – 8: 7 – 8: 0: 2. Anterior genital operculum with 9 – 12 setae, genital opening slit-like, with 18 – 20 marginal setae on each side, 47 – 50 in total (Fig. 4 F). Legs (Fig. 6 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.85 – 1.92 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.26 – 2.59 times longer than tibia. Femoropatella of leg IV 3.53 – 3.81 times longer than deep; tibia 6.08 – 6.45 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.67 – 4.11 times longer than deep (TS = 0.30), tarsus 14.17 – 15.50 times longer than deep and 2.51 – 2.58 times longer than metatarsus (TS = 0.33 – 0.36). Arolium slightly shorter than the claws, not divided; claws simple. Adult females (paratypes) (Figs 3 B, 4 G). Mostly same as males, but a little larger; with same coxal chaetotaxy as males; tergal chaetotaxy I – XII: 4: 4: 4: 4: 4: 4: 4 – 5: 5: 5: 4: T 2 T: 0; sternal chaetotaxy IV – XII: 11 – 12: 8: 7 – 8: 7: 7 – 8: 8 – 9: 7: 0: 2; anterior genital operculum with 10 – 11 setae, posterior margin with 11 – 12 marginal setae, 22 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.56 – 3.78 times longer than deep (TS = 0.31 – 0.32), tarsus 13.67 – 14.67 times longer than deep and 2.56 – 2.59 times longer than metatarsus (TS = 0.33 – 0.35). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Males: body length 2.07 – 2.13. Pedipalps: trochanter 0.22 – 0.24 / 0.20 (1.10 – 1.20), femur 1.20 – 1.25 / 0.18 – 0.19 (6.58 – 6.67), patella 0.47 – 0.50 / 0.20 – 0.22 (2.27 – 2.35), chela 1.75 – 1.76 / 0.23 – 0.24 (7.29 – 7.65), hand 0.59 / 0.23 – 0.24 (2.46 – 2.57), movable chelal finger length 1.13 – 1.14. Chelicera 0.75 – 0.80 / 0.31 – 0.33 (2.42), movable finger length 0.40 – 0.42. Carapace 0.65 – 0.68 / 0.68 (0.96 – 1.00). Leg I: trochanter 0.20 – 0.21 / 0.18 (1.11 – 1.17), femur 0.72 – 0.75 / 0.10 (7.20 – 7.50), patella 0.39 / 0.09 (4.33), tibia 0.32 – 0.35 / 0.07 (4.57 – 5.00), tarsus 0.79 – 0.83 / 0.06 (13.17 – 13.83). Leg IV: trochanter 0.28 – 0.32 / 0.18 – 0.20 (1.56 – 1.60), femoropatella 1.03 – 1.13 / 0.27 – 0.32 (3.53 – 3.81), tibia 0.71 – 0.79 / 0.11 – 0.13 (6.08 – 6.45), metatarsus 0.33 – 0.37 / 0.09 (3.67 – 4.11), tarsus 0.85 – 0.93 / 0.06 (14.17 – 15.50). Females: body length 2.16 – 2.29. Pedipalps: trochanter 0.27 / 0.20 – 0.22 (1.23 – 1.35), femur 1.25 – 1.26 / 0.18 – 0.19 (6.63 – 6.94), patella 0.50 / 0.22 – 0.23 (2.17 – 2.27), chela 1.77 – 1.87 / 0.24 (7.38 – 7.79), hand 0.59 – 0.62 / 0.24 (2.46 – 2.58), movable chelal finger length 1.16 – 1.23. Chelicera 0.82 – 0.86 / 0.35 – 0.36 (2.34 – 2.39), movable finger length 0.44 – 0.46. Carapace 0.67 – 0.69 / 0.72 – 0.77 (0.90 – 0.93). Leg I: trochanter 0.17 – 0.20 / 0.16 – 0.18 (1.06 – 1.11), femur 0.69 – 0.72 / 0.09 – 0.10 (7.20 – 7.67), patella 0.37 – 0.39 / 0.08 – 0.09 (4.33 – 4.63), tibia 0.33 – 0.34 / 0.07 (4.71 – 4.86), tarsus 0.78 – 0.81 / 0.06 (13.00 – 13.50). Leg IV: trochanter 0.26 – 0.28 / 0.17 (1.53 – 1.65), femoropatella 0.99 – 1.03 / 0.27 (3.67 – 3.81), tibia 0.70 – 0.72 / 0.11 – 0.12 (6.00 – 6.36), metatarsus 0.32 – 0.34 / 0.09 (3.56 – 3.78), tarsus 0.82 – 0.88 / 0.06 (13.67 – 14.67).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF90FFD6FF205D5138E1FDAB.taxon	discussion	Remarks. Tyrannochthonius acutus sp. nov. is similar to T. hispidus sp. nov. in having intercalary teeth on both chelal fingers and a pointed, triangular epistome, but differs by a larger body size (body length 2.07 – 2.13 (J), 2.16 – 2.29 (♀) vs. 1.56 – 1.78 (J), 1.77 – 1.99 (♀) mm, chela 7.29 – 7.65 (J), 7.38 – 7.79 (♀) vs. 6.68 – 6.72 (J), 6.20 – 6.79 (♀) times longer than board), the number of blades of rallum (7 vs. 6) and the presence of more fixed chelal finger teeth (57 – 60 vs. 42 – 43). Tyrannochthonius acutus sp. nov. can be distinguished from T. chixing, T. harveyi and T. zhai by the number of setae at the anterior margin of the carapace (6 vs. 4) and the presence of intercalary teeth on both chelal fingers, from T. akaelus and T. ganshuanensis by the number of setae on tergite II (4 vs. 2) and from T. antridraconis by the presence of more chelal teeth (57 – 60 vs. 33 – 36 teeth on the fixed chelal finger; 58 – 62 vs. 30 – 36 teeth on the movable chelal finger) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF90FFD6FF205D5138E1FDAB.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF9AFFC8FF205D7A38E1FCEE.taxon	description	Figs 7 – 11	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF9AFFC8FF205D7A38E1FCEE.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-HBUARA # 2022 - 493 - 01): China, Guizhou Province, Hezhang County, Shuitangbao Township, Caoziping Village, Shenren Cave, under stones and detritus in the deep zone (Temperature: 13 ° C, Humidity: 90 %) [27 ° 0 ′ 55.82 ″ N, 104 ° 40 ′ 36.98 ″ E], 2042 m a. s. l., 31 July 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun & Wenlong Fan leg. Paratypes: 4 ♂ (Ps. - MHBU-HBUARA # 2022 - 493 - 02 – HBUARA # 2022 - 493 - 05), 7 ♀ (Ps. - MSWU-HBUARA # 2022 - 493 - 06 – HBUARA # 2022 - 493 - 12), all with the same data as the holotype.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF9AFFC8FF205D7A38E1FCEE.taxon	etymology	Etymology. The specific name is derived from the Latin word “ altus ”, meaning high, which refers to the species that live at high altitudes.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF9AFFC8FF205D7A38E1FCEE.taxon	diagnosis	Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome pointed and triangular; posterior margin of carapace with 2 setae; tergites I – II each with 4 setae. Pedipalps slender, femur 8.50 – 8.79 (♂), 8.06 – 8.44 (♀) times longer than broad; chela 8.25 – 8.57 (♂), 7.41 – 7.42 (♀) times longer than broad; both chelal fingers with intercalary teeth.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF9AFFC8FF205D7A38E1FCEE.taxon	description	Description. Males (holotype and paratypes) (Figs 7 F, 8 A, 9 A – F, 10, 11). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 9 C, 10 A): carapace 0.95 – 0.97 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome pointed and triangular, with 2 setae flanking base; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 14 – 15 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 10 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 9 D, 10 B): large, about as long as carapace, 2.48 – 2.63 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 18 – 20 teeth, distal one largest; movable finger with 18 – 19 retrorse contiguous small teeth; galea completely vestigial. Serrula exterior with 21 – 22 and serrula interior with 14 – 15 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 10 E). Pedipalp (Figs 9 A, B, E, 10 D, 11 A, B): long and slender, trochanter 1.21 – 1.22, femur 8.50 – 8.79, patella 2.14 – 2.30, chela 8.25 – 8.57, hand 2.95 – 2.96 times longer than broad; femur 2.67 – 2.89 times longer than patella; movable chelal finger 1.69 – 1.79 times longer than hand and 0.61 – 0.62 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 9 E, 10 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb midway between st and b; b and t situated subdistally, t slightly situated distal to b; est situated distal to t (Fig. 11 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 24 – 26 macrodenticles, long and pointed, plus 15 – 16 intercalary microdenticles, 39 – 42 in total; movable chelal finger with 34 – 35 macrodenticles (markedly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 10 – 11 intercalary microdenticles, 44 – 46 in total (Fig. 11 A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 9 B, 11 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 2 – 4: 4: 3 – 4: 4: 4: 4: 5: 4 – 5: T 2 T: 0; tergites IX and Ⅹ each with an unpaired median seta. Sternal chaetotaxy III – XII: 16: 12 – 14: 8 – 9: 7 – 9: 8: 8 – 9: 8 – 9: 7 – 8: 0: 2. Anterior genital operculum with 11 – 12 setae, genital opening slit-like, with 7 – 12 marginal setae on each side, 27 – 35 in total (Fig. 9 F). Legs (Fig. 11 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.90 – 1.92 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.32 – 2.47 times longer than tibia. Femoropatella of leg IV 5.18 – 5.20 times longer than deep; tibia 6.80 – 6.91 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.50 – 4.38 times longer than deep (TS = 0.29 – 0.32), tarsus 13.83 – 16.33 times longer than deep and 2.80 – 2.96 times longer than metatarsus (TS = 0.24 – 0.27). Arolium slightly shorter than the claws, not divided; claws simple. Adult females (paratypes) (Figs 8 B, 9 G). Mostly same as males, but a little larger; with same coxal chaetotaxy as males; tergal chaetotaxy I – XII: 4: 4: 3 – 4: 4: 4: 4: 4: 4: 4 – 5: 4: T 2 T: 0; sternal chaetotaxy IV – XII: 13: 9 – 10: 9: 7 – 10: 8 – 9: 8 – 9: 6 – 7: 0: 2; anterior genital operculum with 10 – 11 setae, posterior margin with 14 – 15 marginal setae, 25 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.88 – 4.13 times longer than deep (TS = 0.29 – 0.30), tarsus 14.67 – 15.67 times longer than deep and 2.67 – 3.03 times longer than metatarsus (TS = 0.25 – 0.30). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Males: body length 2.36 – 2.37. Pedipalps: trochanter 0.22 – 0.23 / 0.18 – 0.19 (1.21 – 1.22), femur 1.23 – 1.36 / 0.14 – 0.16 (8.50 – 8.79), patella 0.46 – 0.47 / 0.20 – 0.22 (2.14 – 2.30), chela 1.80 – 1.98 / 0.21 – 0.24 (8.25 – 8.57), hand 0.62 – 0.71 / 0.21 – 0.24 (2.95 – 2.96), movable chelal finger length 1.11 – 1.20. Chelicera 0.72 – 0.79 / 0.29 – 0.30 (2.48 – 2.63), movable finger length 0.40 – 0.43. Carapace 0.59 – 0.65 / 0.62 – 0.67 (0.95 – 0.97). Leg I: trochanter 0.17 – 0.19 / 0.15 – 0.18 (1.06 – 1.13), femur 0.71 – 0.80 / 0.09 (7.89 – 8.89), patella 0.37 – 0.42 / 0.08 (4.63 – 5.25), tibia 0.32 – 0.37 / 0.06 – 0.07 (5.29 – 5.33), tarsus 0.79 – 0.86 / 0.06 (13.17 – 14.33). Leg IV: trochanter 0.27 – 0.30 / 0.16 – 0.18 (1.67 – 1.69), femoropatella 1.04 – 1.14 / 0.20 – 0.22 (5.18 – 5.20), tibia 0.68 – 0.76 / 0.10 – 0.11 (6.80 – 6.91), metatarsus 0.28 – 0.35 / 0.08 (3.50 – 4.38), tarsus 0.83 – 0.98 / 0.06 (13.83 – 16.33). Females: body length 2.61 – 2.95. Pedipalps: trochanter 0.22 – 0.23 / 0.18 – 0.19 (1.21 – 1.22), femur 1.35 – 1.37 / 0.16 – 0.17 (8.06 – 8.44), patella 0.51 / 0.21 (2.43), chela 1.93 – 2.00 / 0.26 – 0.27 (7.41 – 7.42), hand 0.69 – 0.71 / 0.26 – 0.27 (2.63 – 2.65), movable chelal finger length 1.19 – 1.22. Chelicera 0.79 – 0.82 / 0.31 – 0.32 (2.55 – 2.56), movable finger length 0.44. Carapace 0.67 – 0.68 / 0.68 – 0.70 (0.97 – 0.99). Leg I: trochanter 0.19 / 0.16 – 0.17 (1.12 – 1.19), femur 0.78 – 0.79 / 0.09 – 0.10 (7.90 – 8.67), patella 0.41 – 0.43 / 0.08 – 0.09 (4.78 – 5.13), tibia 0.34 – 0.36 / 0.06 (5.67 – 6.00), tarsus 0.83 – 0.84 / 0.06 – 0.07 (12.00 – 13.83). Leg IV: trochanter 0.29 / 0.16 (1.81), femoropatella 1.10 – 1.14 / 0.21 – 0.22 (5.18 – 5.24), tibia 0.74 – 0.76 / 0.11 – 0.12 (6.33 – 6.73), metatarsus 0.31 – 0.33 / 0.08 (3.88 – 4.13), tarsus 0.88 – 0.94 / 0.06 (14.67 – 15.67).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF9AFFC8FF205D7A38E1FCEE.taxon	discussion	Remarks. Tyrannochthonius altus sp. nov. is similar to T. babaowanensis sp. nov. in having intercalary teeth on both chelal fingers and a pointed, triangular epistome, but differs by the presence of a slender chela (chela 8.25 – 8.57 (♂) vs. 7.27 (♂) times longer than board), the number of setae on tergite Ⅹ (4 – 5 vs. 2) and the number of blades of coxal spines (14 – 15 vs. 8 – 9). Tyrannochthonius altus sp. nov. can be distinguished from T. akaelus, T. ganshuanensis and T. harveyi by the number of setae on the tergites I – II (4 vs. 2), from T. chixing and T. zhai by the number of setae at the anterior margin of the carapace (6 vs. 4), a larger body size (body length min. 2.36 mm vs. max. 1.64 mm) and the presence of intercalary teeth on both chelal fingers, from T. antridraconis by the relative position of trichobothrium sb (sb midway between b and st vs. nearer b than st) and the presence of a slender chela (chela 8.25 – 8.57 (♂) vs. 7.90 – 8.00 (♂) times longer than board) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF9AFFC8FF205D7A38E1FCEE.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF84FFC2FF205CBE38E1FBEA.taxon	description	Figs 12 – 16	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF84FFC2FF205CBE38E1FBEA.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-HBUARA # 2022 - 506 - 01): China, Guizhou Province, Anlong County, Dushan Town, Polao Village, Hei Cave, under stones and clods in the deep zone (Temperature: 16 ° C, Humidity: 65 %) [25 ° 18 ′ 46.56 ″ N, 105 ° 36 ′ 2.69 ″ E], 1067 m a. s. l., 6 August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun & Wenlong Fan leg. Paratypes: 3 ♂ (Ps. - MHBU-HBUARA # 2022 - 506 - 02 – HBUARA # 2022 - 506 - 04), 3 ♀ (Ps. - MSWU-HBUARA # 2022 - 506 - 05 – HBUARA # 2022 - 506 - 07), all with the same data as the holotype.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF84FFC2FF205CBE38E1FBEA.taxon	etymology	Etymology. The specific name is derived from the Latin word “ arificus ”, meaning arid, which refers to the species that lives in a dry environment.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF84FFC2FF205CBE38E1FBEA.taxon	diagnosis	Diagnosis (J ♀). Small-sized troglomorphic species with slightly elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, rounded and obtuse; posterior margin of carapace with 2 setae; tergites I – Ⅴ each with 4 setae. Palpal femur 5.27 – 5.55 (♂), 5.36 – 5.50 (♀) times longer than broad; chela 6.29 – 6.62 (♂), 6.07 – 6.20 (♀) times longer than broad; both chelal fingers with intercalary teeth.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF84FFC2FF205CBE38E1FBEA.taxon	description	Description. Males (holotype and paratypes) (Figs 13 A, 14 A – F, H, 15, 16). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 14 C, D, 15 A): carapace 0.93 – 0.95 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small, rounded and obtuse; with 17 – 18 setae arranged s 3 – 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 3 – 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 10 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 15 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 14 E, 15 B): large, about as long as carapace, 2.05 – 2.20 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on dorsal side. Both fingers well provided with teeth, fixed finger with 11 – 12 teeth, distal one largest; movable finger with 11 – 13 retrorse contiguous small teeth; galea completely vestigial (Fig. 15 B). Serrula exterior with 18 – 20 and serrula interior with 11 – 13 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 15 E). Pedipalp (Figs 14 A, B, H, 16 A, B): long and slender, trochanter 1.15 – 1.25, femur 5.27 – 5.55, patella 2.15 – 2.17, chela 6.29 – 6.62, hand 2.07 – 2.15 times longer than broad; femur 2.18 – 2.23 times longer than patella; movable chelal finger 2.10 – 2.14 times longer than hand and 0.69 – 0.70 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 14 H, 15 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated distal to b and at same level as est (Fig. 16 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 30 – 32 macrodenticles, slightly retrorse and pointed, plus 25 – 26 intercalary microdenticles, 56 – 57 in total; movable chelal finger with 12 – 13 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 10 – 11 intercalary microdenticles and 17 – 19 vestigial, rounded and contiguous basal teeth, 41 in total (Fig. 16 A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 14 B, 16 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 4: 4: 4: 5: 5: 5: 5: 3: T 2 T: 0, tergites VI – IX each with an unpaired median seta. Sternal chaetotaxy III – XII: 13 – 15: 13 – 14: 10: 9: 9: 9: 9 – 10: 9: 0: 2. Anterior genital operculum with 10 – 11 setae, genital opening slit-like, with 11 – 14 marginal setae on each side, 35 – 36 in total (Fig. 14 F). Legs (Fig. 16 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.76 – 1.89 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.22 – 2.37 times longer than tibia. Femoropatella of leg IV 2.84 – 3.06 times longer than deep; tibia 4.78 – 5.00 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.17 times longer than deep (TS = 0.26 – 0.37), tarsus 11.75 – 14.67 times longer than deep and 2.32 – 2.47 times longer than metatarsus (TS = 0.25 – 0.34). Arolium slightly shorter than the claws, not divided; claws simple. Adult females (paratypes) (Figs 13 B, 14 G). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; with same tergal chaetotaxy as males; sternal chaetotaxy IV – XII: 14 – 15: 10: 9 – 10: 9: 9 – 10: 9 – 11: 8 – 9: 0: 2; anterior genital operculum with 10 – 11 setae, posterior margin with 14 marginal setae, 24 – 25 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.00 – 3.50 times longer than deep (TS = 0.33), tarsus 12.00 – 14.67 times longer than deep and 2.29 – 2.44 times longer than metatarsus (TS = 0.27 – 0.29). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Males: body length 1.31 – 1.40. Pedipalps: trochanter 0.15 / 0.12 – 0.13 (1.15 – 1.25), femur 0.58 – 0.61 / 0.11 (5.27 – 5.55), patella 0.26 – 0.28 / 0.12 – 0.13 (2.15 – 2.17), chela 0.86 – 0.88 / 0.13 – 0.14 (6.29 – 6.62), hand 0.28 – 0.29 / 0.13 – 0.14 (2.07 – 2.15), movable chelal finger length 0.60 – 0.61. Chelicera 0.41 – 0.44 / 0.20 (2.05 – 2.20), movable finger length 0.23 – 0.24. Carapace 0.41 – 0.43 / 0.43 – 0.46 (0.93 – 0.95). Leg I: trochanter 0.12 / 0.10 (1.20), femur 0.36 – 0.37 / 0.06 – 0.07 (5.14 – 6.17), patella 0.19 – 0.21 / 0.05 (3.80 – 4.20), tibia 0.18 – 0.19 / 0.04 (4.50 – 4.75), tarsus 0.40 – 0.45 / 0.03 – 0.04 (11.25 – 13.33). Leg IV: trochanter 0.16 – 0.17 / 0.11 – 0.12 (1.33 – 1.55), femoropatella 0.52 – 0.54 / 0.17 – 0.19 (2.84 – 3.06), tibia 0.40 – 0.43 / 0.08 – 0.09 (4.78 – 5.00), metatarsus 0.19 / 0.06 (3.17), tarsus 0.44 – 0.47 / 0.03 – 0.04 (11.75 – 14.67). Females: body length 1.62 – 1.69. Pedipalps: trochanter 0.16 – 0.17 / 0.12 – 0.14 (1.21 – 1.33), femur 0.59 – 0.66 / 0.11 – 0.12 (5.36 – 5.50), patella 0.30 – 0.32 / 0.13 – 0.15 (2.13 – 2.31), chela 0.91 – 0.93 / 0.15 (6.07 – 6.20), hand 0.31 / 0.15 (2.07), movable chelal finger length 0.62 – 0.63. Chelicera 0.45 – 0.47 / 0.21 – 0.23 (2.04 – 2.14), movable finger length 0.24 – 0.25. Carapace 0.45 – 0.48 / 0.46 – 0.49 (0.98). Leg I: trochanter 0.12 – 0.13 / 0.10 – 0.11 (1.18 – 1.20), femur 0.37 – 0.39 / 0.06 – 0.07 (5.57 – 6.17), patella 0.19 – 0.22 / 0.06 (3.17 – 3.67), tibia 0.19 – 0.20 / 0.04 – 0.05 (4.00 – 4.75), tarsus 0.44 – 0.45 / 0.04 (11.00 – 11.25). Leg IV: trochanter 0.18 / 0.11 (1.64), femoropatella 0.50 – 0.56 / 0.17 – 0.19 (2.94 – 2.95), tibia 0.39 – 0.45 / 0.08 – 0.09 (4.88 – 5.00), metatarsus 0.18 – 0.21 / 0.06 (3.00 – 3.50), tarsus 0.44 – 0.48 / 0.03 – 0.04 (12.00 – 14.67).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF84FFC2FF205CBE38E1FBEA.taxon	discussion	Remarks. Tyrannochthonius arificus sp. nov. is similar to T. yanshanensis sp. nov. in having intercalary teeth on both chelal fingers and a small, rounded and obtuse epistome, but differs by the larger body size (body length 1.62 – 1.69 (♀) vs. 1.29 – 1.33 (♀) mm; chela 6.07 – 6.20 (♀) vs. 5.69 – 5.75 (♀) times longer than board), the number of setae on tergite Ⅴ (4 vs. 5) and tergite Ⅹ (3 vs. 5), more intercalary teeth on the fixed chelal finger (25 – 26 vs. 14), the number of blades of coxal spines (10 vs. 7 – 8) and the number of blades of rallum (8 vs. 7). Tyrannochthonius arificus sp. nov. can be distinguished from T. akaelus, T. ganshuanensis and T. harveyi by the number of setae on the tergites I – II (4 vs. 2), from T. chixing and T. zhai by the number of setae at the anterior margin of the carapace (6 vs. 4), a shorter palp (chela length max. 0.93 vs. min. 1.21 mm; palpal femur length max. 0.66 vs. min. 0.84 mm) and the presence of intercalary teeth on both chelal fingers, from T. antridraconis by a smaller size (body length max. 1.40 vs. min. 1.80 mm; chela length max. 0.88 vs. min. 1.70 mm) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF84FFC2FF205CBE38E1FBEA.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF8EFFC4FF205BBA38E1FB92.taxon	description	Figs 17 – 21	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF8EFFC4FF205BBA38E1FB92.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-GZYJ- 20 - 14 - 01): China, Guizhou Province, Yinjiang County, Zhongxing Sub-district, Tianchi Village, Babaowan Cave, on a stone wall in the deep zone (Temperature: 14 ° C, Humidity: 85 %) [28 ° 3 ′ 53.07 ″ N, 108 ° 21 ′ 52.89 ″ E], 655 m a. s. l., 23 August 2020, Zegang Feng, Hongru Xu & Yanmeng Hou leg.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF8EFFC4FF205BBA38E1FB92.taxon	etymology	Etymology. Named after the type locality, Babaowan Cave.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF8EFFC4FF205BBA38E1FB92.taxon	diagnosis	Diagnosis (♂). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, pointed, triangular; posterior margin of carapace with 2 setae; tergites I – VI each with 4 setae. Pedipalps slender, femur 7.67 times longer than broad; chela 7.27 times longer than broad; both chelal fingers with intercalary teeth (present in the distal 1 / 3 of the fingers); chemosensory setae present on dorsum of chelal hand.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF8EFFC4FF205BBA38E1FB92.taxon	description	Description. Male (holotype), female unknown (Figs 17 F, 18 – 21). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 19 C, 20 A): carapace 1.04 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome pointed and small, triangular, with 2 setae flanking base; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 8 – 9 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 20 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 19 D, 20 B): large, about as long as carapace, 2.37 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 14 teeth, distal one largest; movable finger with 13 retrorse contiguous small teeth; galea completely vestigial (Fig. 20 B). Serrula exterior with 21 and serrula interior with 14 blades. Rallum with 7 blades, the distal one longer and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 20 E). Pedipalp (Figs 19 A, B, F, 21 A, B): long and slender, trochanter 1.22, femur 7.67, patella 2.72, chela 7.27, hand 2.41 times longer than broad; femur 2.35 times longer than patella; movable chelal finger 2.06 times longer than hand and 0.68 times longer than chela. Setae generally long and acuminate; one distal lyrifissures present on patella (Figs 19 F, 20 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, b situated at same level as est, t slightly distal to it (Fig. 21 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 42 macrodenticles, slightly retrorse and pointed, plus 15 intercalary microdenticles (present in the distal 1 / 3 of the fingers, till est), 57 in total; movable chelal finger with 31 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 11 intercalary microdenticles (present in the distal 1 / 3 of the fingers, till b) and 14 vestigial, rounded and contiguous basal teeth, 56 in total (Fig. 21 A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 19 B, 21 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 4: 4: 4: 4: 5: 5: 5: 2: T 2 T: 0. Sternal chaetotaxy III – XII: 14: 12: 7: 7: 8: 9: 9: 7: 0: 2. Anterior genital operculum with 13 setae, genital opening slit-like, with 14 – 15 marginal setae on each side, 42 in total (Fig. 19 E). Legs (Fig. 21 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.97 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.43 times longer than tibia. Femoropatella of leg IV 4.21 times longer than deep; tibia 7.00 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.75 times longer than deep (TS = 0.33), tarsus 14.00 times longer than deep and 2.80 times longer than metatarsus (TS = 0.32). Arolium slightly shorter than the claws, not divided; claws simple. Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Male: body length 2.35. Pedipalps: trochanter 0.22 / 0.18 (1.22), femur 1.15 / 0.15 (7.67), patella 0.49 / 0.18 (2.72), chela 1.60 / 0.22 (7.27), hand 0.53 / 0.22 (2.41), movable chelal finger length 1.09. Chelicera 0.64 / 0.27 (2.37), movable finger length 0.35. Carapace 0.59 / 0.57 (1.04). Leg I: trochanter 0.19 / 0.15 (1.27), femur 0.69 / 0.08 (8.63), patella 0.35 / 0.07 (5.00), tibia 0.30 / 0.06 (5.00), tarsus 0.73 / 0.06 (12.17). Leg IV: trochanter 0.27 / 0.16 (1.69), femoropatella 1.01 / 0.24 (4.21), tibia 0.70 / 0.10 (7.00), metatarsus 0.30 / 0.08 (3.75), tarsus 0.84 / 0.06 (14.00).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF8EFFC4FF205BBA38E1FB92.taxon	discussion	Remarks. Female unknown. Tyrannochthonius babaowanensis sp. nov. is similar to T. antridraconis in having intercalary teeth on both chelal fingers and a small, rounded and obtuse epistome, but differs by a larger body size (body length 1.62 – 1.69 (♂) vs. 1.29 – 1.33 (♂) mm), the number of setae on tergite VII (5 vs. 4) and tergite Ⅹ (2 vs. 4 – 5), the proportion of intercalary teeth on the fixed finger (1 / 3 vs. 2 / 3), the number of blades of coxal spines (8 – 9 vs. 12 – 15) and the number of blades of rallum (7 vs. 8). Tyrannochthonius babaowanensis sp. nov. can be distinguished from T. chixing and T. zhai by the larger size (body length 2.35 vs. max. 1.64 mm; chela length 1.60 vs. max. 1.24 mm), the number of setae at the anterior margin of the carapace (6 vs. 4) and the presence of intercalary teeth on both chelal fingers, from T. akaelus, T. ganshuanensis and T. harveyi by the number of setae on the tergites I – II (4 vs. 2) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF8EFFC4FF205BBA38E1FB92.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF88FFFEFF205B6038E1FEF2.taxon	description	Figs 22 – 26	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF88FFFEFF205B6038E1FEF2.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-HBUARA # 2021 - 202 - 01): China, Guizhou Province, Libo County, Maolan Town, Yaochao Village, Yanjia Cave, under stones in the deep zone (Temperature: 13 ° C, Humidity: 90 %) [25 ° 22 ′ 41.97 ″ N, 108 ° 3 ′ 40.08 ″ E], 609.5 m a. s. l., 1 August 2021, Zegang Feng, Hongru Xu, Liu Fu & Nana Zhan leg. Paratypes: 2 ♂ (Ps. - MHBU-HBUARA # 2021 - 202 - 02 & HBUARA # 2021 - 202 - 03), 2 ♀ (Ps. - MSWU-HBUARA # 2021 - 202 - 04 & HBUARA # 2021 - 202 - 05), all with the same data as the holotype.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF88FFFEFF205B6038E1FEF2.taxon	etymology	Etymology. The specific name is derived from the Latin word “ breviculus ”, meaning short, which refers to the short pedipalps.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF88FFFEFF205B6038E1FEF2.taxon	diagnosis	Diagnosis (J ♀). Small-sized troglomorphic species with slightly elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, rounded and obtuse; posterior margin of carapace with 2 setae; tergites I – Ⅴ each with 4 setae (more variation in females). Palpal femur 5.10 – 5.33 (♂), 5.00 – 5.30 (♀) times longer than broad; chela 5.69 – 5.85 (♂), 5.57 – 5.64 (♀) times longer than broad; both chelal fingers with intercalary teeth; chemosensory setae present on dorsum of chelal hand.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF88FFFEFF205B6038E1FEF2.taxon	description	Description. Males (holotype and paratypes) (Figs 23 A, 24 A – C, E, F, H). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 24 C, 25 A): carapace 0.95 – 0.98 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small, rounded and obtuse; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 8 – 9 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 25 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 24 E, 25 B): large, about as long as carapace, 2.05 – 2.11 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation both dorsal and ventral side. Both fingers well provided with teeth, fixed finger with 10 – 11 teeth, distal one largest; movable finger with 13 – 15 retrorse contiguous small teeth; galea completely vestigial (Fig. 25 B). Serrula exterior with 17 – 18 and serrula interior with 12 – 14 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 25 E). Pedipalp (Figs 24 A, B, H, 25 D, 26 A, B): long and slender, trochanter 1.00 – 1.18, femur 5.10 – 5.33, patella 2.00 – 2.09, chela 5.69 – 5.85, hand 1.85 – 1.92 times longer than broad; femur 2.09 – 2.13 times longer than patella; movable chelal finger 2.12 – 2.13 times longer than hand and 0.69 – 0.70 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 24 H, 25 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb midway between st and b; b and t situated subdistally, t situated distal to b and midway between it and est; est situated distal to b (Fig. 26 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 29 – 30 macrodenticles, slightly retrorse and pointed, plus 13 – 15 intercalary microdenticles, 42 – 45 in total; movable chelal finger with 9 – 10 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 8 intercalary microdenticles and 14 – 16 vestigial, rounded and contiguous basal teeth, 32 – 33 in total (Fig. 26 A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 24 B, 26 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 4: 4: 4: 5: 5: 5: 5: 4: T 2 T: 0, tergites VI – IX each with an unpaired median seta. Sternal chaetotaxy III – XII: 13 – 14: 11 – 13: 8 – 9: 9: 9: 8 – 9: 9: 9 – 10: 0: 2. Anterior genital operculum with 10 – 11 setae, genital opening slit-like, with 10 – 12 marginal setae on each side, 32 – 33 in total (Fig. 24 F). Legs (Fig. 26 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.87 – 2.00 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.00 – 2.27 times longer than tibia. Femoropatella of leg IV 2.67 times longer than deep; tibia 4.00 – 4.43 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 2.80 times longer than deep (TS = 0.36), tarsus 11.33 – 11.67 times longer than deep and 2.43 – 2.50 times longer than metatarsus (TS = 0.24 – 0.29). Arolium slightly shorter than the claws, not divided; claws simple. Adult females (paratypes) (Figs 23 B, 24 D, G). Mostly same as males, but a little larger; with same chaetotaxy of coxae; tergal chaetotaxy I – XII: 3 – 4: 2 – 4: 4 – 5: 4 – 5: 5: 5: 5: 5: 5: 4: T 2 T: 0; sternal chaetotaxy IV – XII: 12 – 13: 7 – 9: 8 – 9: 9: 9: 10: 9: 0: 2; anterior genital operculum with 10 setae, posterior margin with 14 – 16 marginal setae, 24 – 26 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 2.33 – 2.80 times longer than deep (TS = 0.36 – 0.43), tarsus 11.00 – 12.00 times longer than deep and 2.36 – 2.57 times longer than metatarsus (TS = 0.24 – 0.25). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Males: body length 1.24 – 1.26. Pedipalps: trochanter 0.11 – 0.13 / 0.11 (1.00 – 1.18), femur 0.48 – 0.51 / 0.09 – 0.10 (5.10 – 5.33), patella 0.23 – 0.24 / 0.11 – 0.12 (2.00 – 2.09), chela 0.74 – 0.76 / 0.13 (5.69 – 5.85), hand 0.24 – 0.25 / 0.13 (1.85 – 1.92), movable chelal finger length 0.51 – 0.53. Chelicera 0.38 – 0.39 / 0.18 – 0.19 (2.05 – 2.11), movable finger length 0.20. Carapace 0.38 – 0.40 / 0.40 – 0.41 (0.95 – 0.98). Leg I: trochanter 0.10 – 0.11 / 0.09 (1.11 – 1.22), femur 0.28 / 0.05 – 0.06 (4.67 – 5.60), patella 0.14 – 0.15 / 0.05 (2.80 – 3.00), tibia 0.15 – 0.16 / 0.04 (3.75 – 4.00), tarsus 0.32 – 0.34 / 0.03 (10.67 – 11.33). Leg IV: trochanter 0.13 – 0.14 / 0.09 – 0.10 (1.40 – 1.44), femoropatella 0.40 / 0.15 (2.67), tibia 0.28 – 0.31 / 0.07 (4.00 – 4.43), metatarsus 0.14 / 0.05 (2.80), tarsus 0.34 – 0.35 / 0.03 (11.33 – 11.67). Females: body length 1.33 – 1.44. Pedipalps: trochanter 0.12 – 0.14 / 0.11 – 0.12 (1.08 – 1.09), femur 0.53 – 0.55 / 0.10 – 0.11 (5.00 – 5.30), patella 0.25 – 0.26 / 0.13 (1.92 – 2.00), chela 0.78 – 0.79 / 0.14 (5.57 – 5.64), hand 0.26 – 0.27 / 0.14 (1.86 – 1.93), movable chelal finger length 0.53 – 0.55. Chelicera 0.42 – 0.43 / 0.20 – 0.21 (2.05 – 2.10), movable finger length 0.22 – 0.23. Carapace 0.40 – 0.42 / 0.42 – 0.45 (0.93 – 0.95). Leg I: trochanter 0.11 / 0.09 – 0.10 (1.10 – 1.22), femur 0.27 – 0.30 / 0.06 (4.50 – 5.00), patella 0.14 – 0.16 / 0.05 (2.80 – 3.20), tibia 0.17 / 0.04 (4.25), tarsus 0.33 – 0.35 / 0.04 (8.25 – 8.75). Leg IV: trochanter 0.12 – 0.14 / 0.09 – 0.10 (1.33 – 1.40), femoropatella 0.42 – 0.46 / 0.14 – 0.15 (2.80 – 3.29), tibia 0.32 – 0.33 / 0.07 (4.57 – 4.71), metatarsus 0.14 / 0.05 – 0.06 (2.33 – 2.80), tarsus 0.33 – 0.36 / 0.03 (11.00 – 12.00).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF88FFFEFF205B6038E1FEF2.taxon	discussion	Remarks. Tyrannochthonius breviculus sp. nov. is similar to T. yanshanensis sp. nov. in having intercalary teeth on both chelal fingers and a small, rounded and obtuse epistome, but differs by a shorter palp (chela length 0.78 – 0.79 (♀) vs. 0.91 – 0.92 (♀) mm; palpal femur 5.00 – 5.30 (♀) vs. 4.92 – 5.08 (♀) times longer than board, length 0.53 – 0.55 (♀) vs. 0.59 – 0.61 (♀) mm; movable chelal finger 2.04 (♀) vs. 2.17 – 2.21 (♀) times longer than hand), the number of setae on the tergites VIII ‒ IX (5 vs. 6), the relative position of trichobothrium sb (sb midway between st and b vs. nearer st than b) and the number of blades of rallum (8 vs. 7). Tyrannochthonius breviculus sp. nov. can be distinguished from T. chixing, T. harveyi and T. zhai by the presence of intercalary teeth on both chelal fingers, from T. akaelus, T. antridraconis and T. ganshuanensis by a shorter palp (chela length max. 0.79 vs. min. 1.21 mm; palpal femur length max. 0.55 vs. min. 0.90 mm) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF88FFFEFF205B6038E1FEF2.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFB2FFFCFF205E9D38E1F826.taxon	description	Figs 27 – 31	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFB2FFFCFF205E9D38E1F826.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-HBUARA # 2022 - 518 - 01): China, Guizhou Province, Danzhai County, Longquan Town, Jinguadong Village, Jingua Cave, under stones about 20 m from the cave entrance (Temperature: 12 ° C, Humidity: 90 %) [26 ° 8 ′ 15.94 ″ N, 107 ° 48 ′ 25.09 ″ E], 749 m a. s. l., 12 August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun & Wenlong Fan leg. Paratype: 1 ♀ (Ps. - MSWU-HBUARA # 2022 - 518 - 02), with the same data as the holotype.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFB2FFFCFF205E9D38E1F826.taxon	etymology	Etymology. The specific name is derived from a combination of the Latin word “ brevis ” and “ spineus ”, meaning short and spined, respectively, which refers to the short coxal spines.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFB2FFFCFF205E9D38E1F826.taxon	diagnosis	Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin and finely denticulated, epistome small and represented by a slight bump; posterior margin of the carapace with 2 setae; tergites I – II each with 2 setae. Pedipalps slender, femur 6.64 (♂), 6.67 (♀) times longer than broad; chela 7.39 (♂), 7.53 (♀) times longer than broad; both chelal fingers without intercalary teeth, movable chelal finger teeth distinctly smaller and denser than fixed chelal finger teeth; chemosensory setae present on dorsum of chelal hand; coxal spines short.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFB2FFFCFF205E9D38E1F826.taxon	description	Description. Male (holotype) (Figs 28 A, 29 A – F, 30, 31). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 29 C, 30 A): carapace 1.02 times longer than broad, slightly narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and represented by a slight bump, with 2 setae flanking base; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 8 – 9 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 30 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 29 D, 30 B): large, about as long as carapace, 2.36 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 12 teeth, distal one largest; movable finger with 16 retrorse contiguous small teeth; galea represented by a very slight bump on movable finger (Fig. 30 B). Serrula exterior with 17 and serrula interior with 13 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 30 E). Pedipalp (Figs 29 A, B, E, 31 A, B): long and slender, trochanter 1.33, femur 6.64, patella 2.13, chela 7.39, hand 2.39 times longer than broad; femur 2.74 times longer than patella; movable chelal finger 2.05 times longer than hand and 0.66 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 29 E, 30 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb midway between st and b; b and t situated subdistally, t situated distal to b and proximal to est (Fig. 31 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 26 macrodenticles, slightly retrorse and pointed; movable chelal finger with 18 macrodenticles (markedly smaller than teeth on fixed chelal finger), continuous, retrorse and pointed, plus 18 small, rounded and contiguous basal teeth, 36 in total (Fig. 31 A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 29 B, 31 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 2: 2: 4: 4: 4: 4: 4: 5: 5: 4: T 2 T: 0. Sternal chaetotaxy III – XII: 12: 12: 7: 8: 9: 9: 9: 9: 0: 2. Anterior genital operculum with 10 setae, genital opening slit-like, with 12 – 14 marginal setae on each side, 36 in total (Fig. 29 F). Legs (Fig. 31 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.76 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.32 times longer than tibia. Femoropatella of leg IV 3.43 times longer than deep; tibia 5.67 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.29 times longer than deep (TS = 0.26), tarsus 12.80 times longer than deep and 2.78 times longer than metatarsus (TS = 0.33). Arolium slightly shorter than the claws, not divided; claws simple. Adult female (paratype) (Figs 28 B, 29 G). Mostly same as male, but a little larger; chaetotaxy of coxae: P 3, I 3, II 3 – 4, III 5, IV 5; with same tergal chaetotaxy as male; sternal chaetotaxy IV – XII: 12: 8: 9: 9: 9: 9: 9: 0: 2; anterior genital operculum with 10 setae, posterior margin with 12 marginal setae, 22 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 4.00 times longer than deep (TS = 0.25), tarsus 12.40 times longer than deep and 2.58 times longer than metatarsus (TS = 0.31). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Male: body length 1.73. Pedipalps: trochanter 0.20 / 0.15 (1.33), femur 0.93 / 0.14 (6.64), patella 0.34 / 0.16 (2.13), chela 1.33 / 0.18 (7.39), hand 0.43 / 0.18 (2.39), movable chelal finger length 0.88. Chelicera 0.52 / 0.22 (2.36), movable finger length 0.28. Carapace 0.49 / 0.48 (1.02). Leg I: trochanter 0.14 / 0.13 (1.08), femur 0.51 / 0.07 (7.29), patella 0.29 / 0.07 (4.14), tibia 0.25 / 0.05 (5.00), tarsus 0.58 / 0.05 (11.60). Leg IV: trochanter 0.22 / 0.14 (1.57), femoropatella 0.72 / 0.21 (3.43), tibia 0.51 / 0.09 (5.67), metatarsus 0.23 / 0.07 (3.29), tarsus 0.64 / 0.05 (12.80). Female: body length 2.12. Pedipalps: trochanter 0.21 / 0.16 (1.31), femur 1.00 / 0.15 (6.67), patella 0.36 / 0.17 (2.12), chela 1.43 / 0.19 (7.53), hand 0.47 / 0.19 (2.47), movable chelal finger length 0.94. Chelicera 0.58 / 0.24 (2.42), movable finger length 0.30. Carapace 0.55 / 0.54 (1.02). Leg I: trochanter 0.14 / 0.14 (1.00), femur 0.56 / 0.08 (7.00), patella 0.28 / 0.07 (4.00), tibia 0.26 / 0.05 (5.20), tarsus 0.62 / 0.05 (12.40). Leg IV: trochanter 0.23 / 0.13 (1.77), femoropatella 0.73 / 0.21 (3.48), tibia 0.51 / 0.09 (5.67), metatarsus 0.24 / 0.06 (4.00), tarsus 0.62 / 0.05 (12.40).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFB2FFFCFF205E9D38E1F826.taxon	discussion	Remarks. Tyrannochthonius brevispinus sp. nov. is similar to T. oblongus sp. nov. in having a small and slight bump epistome and the same number of setae on tergites I – II (2), but differs by a smaller size (body length 1.73 (♂) vs. 2.18 – 2.26 (♂) mm; chela 7.39 (♂), 7.53 (♀) vs. 7.27 (♂), 6.78 – 7.09 (♀) times longer than board, length 1.33 (♂), 1.43 (♀) vs. 1.60 (♂) mm, 1.56 (♀) mm; palpal femur length 0.93 (♂), 1.00 (♀) vs. 1.11 – 1.17 (♂), 1.09 – 1.17 (♀) mm), the proportion of movable chelal finger and chelal hand (movable chelal finger 2.05 (♂), 2.00 (♀) vs. 1.74 – 1.79 (♂), 1.74 – 1.75 (♀) times longer than hand), the relative position of trichobothrium sb (sb midway between b and st vs. nearer b than st) and the number of blades of rallum (8 vs. 7). Tyrannochthonius brevispinus sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on both chelal fingers, from T. harveyi by the number of setae at the anterior margin of the carapace (6 vs. 4), a larger body size (body length min. 1.73 mm vs. max. 1.56 mm; chela length min. 1.33 mm vs. max. 1.30 mm; palpal femur length min. 0.93 mm vs. max. 0.88 mm), the number of setae on tergite III (4 vs. 2) and the number of blades of rallum (8 vs. 6) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFB2FFFCFF205E9D38E1F826.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFBCFFF5FF205F8838E1FA56.taxon	description	Figs 32 – 36	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFBCFFF5FF205F8838E1FA56.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-HBUARA # 2022 - 305 - 01): China, Guizhou Province, Jiangkou County, Nuxi Town, Mengjiatun Village, Liangfeng Cave, under stones and detritus in the deep zone (Temperature: 13 ° C, Humidity: 80 %) [27 ° 49 ′ 37.84 ″ N, 108 ° 51 ′ 43.34 ″ E], 408 m a. s. l., 4 July 2022, Yanmeng Hou, Nana Zhan, Long Lin & Jianzhou Sun leg. Paratypes: 2 ♀ (Ps. - MSWU-HBUARA # 2022 - 305 - 02 & HBUARA # 2022 - 305 - 03), all with the same data as the holotype.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFBCFFF5FF205F8838E1FA56.taxon	etymology	Etymology. The specific name is derived from the Latin word “ duo ”, meaning dual, which refers to the presence of two well-developed anterior eyes.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFBCFFF5FF205F8838E1FA56.taxon	diagnosis	Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace with two anterior corneate eyes, posterior eyes reduced into eyespots; anterior margin of carapace thin, finely denticulated, epistome small, pointed, triangular; posterior margin of carapace with 2 setae; tergites I – II each with 4 setae. Pedipalps slender, femur 5.85 (♂), 5.79 – 6.08 (♀) times longer than broad; chela 6.94 (♂), 5.85 – 6.56 (♀) times longer than broad; both chelal fingers with intercalary teeth; chemosensory setae present on dorsum of chelal hand.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFBCFFF5FF205F8838E1FA56.taxon	description	Description. Male (holotype) (Figs 32 D, 33 A, 34 A – F, 35, 36). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 34 C, 35 A): carapace 1.00 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; anterior pair of eyes well developed, posterior pair of eyes reduced into eyespots; anterior margin slightly serrate; epistome small and pointed, triangular; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 9 – 11 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 35 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 34 D, 35 B): large, about as long as carapace, 2.08 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation both dorsal and ventral side. Both fingers well provided with teeth, fixed finger with 15 teeth, distal one largest; movable finger with 15 retrorse contiguous small teeth; galea completely vestigial (Fig. 35 B). Serrula exterior with 17 and serrula interior with 13 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 35 E). Pedipalp (Figs 34 A, B, E, 35 D, 36 A, B): long and slender, trochanter 1.36, femur 5.85, patella 2.27, chela 6.94, hand 2.25 times longer than broad; femur 2.24 times longer than patella; movable chelal finger 2.14 times longer than hand and 0.69 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 34 E, 35 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, t situated at same level as est and distal to b (Fig. 36 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 32 macrodenticles, slightly retrorse and pointed, plus 21 intercalary microdenticles, 53 in total; movable chelal finger with 16 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 10 intercalary microdenticles and 14 vestigial, rounded and contiguous basal teeth, 40 in total (Fig. 36 A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 34 B, 36 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 3: 4: 6: 7: 8: 8: 6: 4: T 2 T: 0, tergites VI – IX each with an unpaired median seta. Sternal chaetotaxy III – XII: 12: 12: 9: 9: 9: 10: 10: 8: 0: 2. Anterior genital operculum with 12 setae, genital opening slit-like, with 17 – 18 marginal setae on each side, 47 in total (Fig. 34 F). Legs (Fig. 36 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 2.09 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.13 times longer than tibia. Femoropatella of leg IV 2.91 times longer than deep; tibia 4.80 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.29 times longer than deep (TS = 0.35), tarsus 11.60 times longer than deep and 2.52 times longer than metatarsus (TS = 0.36). Arolium slightly shorter than the claws, not divided; claws simple. Adult females (paratypes) (Figs 33 B, 34 G). Mostly same as male, but a little larger; with same chaetotaxy of coxae; tergal chaetotaxy I – XII: 4: 4: 4: 4 – 6: 7 – 8: 8: 8: 8 – 9: 8 – 9: 4: T 2 T: 0; sternal chaetotaxy IV – XII: 11 – 13: 12: 11 – 12: 12: 12: 11 – 12: 9: 0: 2; anterior genital operculum with 11 setae, posterior margin with 12 – 14 marginal setae, 23 – 25 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.00 – 3.43 times longer than deep (TS = 0.33), tarsus 11.60 – 11.80 times longer than deep and 2.42 – 2.46 times longer than metatarsus (TS = 0.31 – 0.34). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Male: body length 1.73. Pedipalps: trochanter 0.19 / 0.14 (1.36), femur 0.76 / 0.13 (5.85), patella 0.34 / 0.15 (2.27), chela 1.11 / 0.16 (6.94), hand 0.36 / 0.16 (2.25), movable chelal finger length 0.77. Chelicera 0.52 / 0.25 (2.08), movable finger length 0.28. Carapace 0.52 / 0.52 (1.00). Leg I: trochanter 0.15 / 0.12 (1.25), femur 0.46 / 0.07 (6.57), patella 0.22 / 0.07 (3.14), tibia 0.24 / 0.05 (4.80), tarsus 0.51 / 0.04 (12.75). Leg IV: trochanter 0.19 / 0.14 (1.36), femoropatella 0.67 / 0.23 (2.91), tibia 0.48 / 0.10 (4.80), metatarsus 0.23 / 0.07 (3.29), tarsus 0.58 / 0.05 (11.60). Females: body length 1.93 – 2.12. Pedipalps: trochanter 0.19 – 0.21 / 0.14 – 0.16 (1.19 – 1.50), femur 0.79 – 0.81 / 0.13 – 0.14 (5.79 – 6.08), patella 0.36 / 0.15 – 0.16 (2.25 – 2.40), chela 1.17 – 1.18 / 0.18 – 0.20 (5.85 – 6.56), hand 0.38 – 0.39 / 0.18 – 0.20 (1.95 – 2.11), movable chelal finger length 0.80 – 0.82. Chelicera 0.57 – 0.58 / 0.28 – 0.29 (2.00 – 2.04), movable finger length 0.31 – 0.32. Carapace 0.56 / 0.61 – 0.63 (0.89 – 0.92). Leg I: trochanter 0.13 – 0.14 / 0.12 – 0.13 (1.08), femur 0.47 – 0.49 / 0.07 – 0.08 (5.88 – 7.00), patella 0.22 – 0.24 / 0.07 (3.14 – 3.43), tibia 0.24 / 0.05 (4.80), tarsus 0.48 – 0.54 / 0.05 – 0.06 (9.00 – 9.60). Leg IV: trochanter 0.20 – 0.23 / 0.14 – 0.15 (1.33 – 1.64), femoropatella 0.70 / 0.23 (3.04), tibia 0.52 – 0.53 / 0.10 (5.20 – 5.30), metatarsus 0.24 / 0.07 – 0.08 (3.00 – 3.43), tarsus 0.58 – 0.59 / 0.05 (11.60 – 11.80).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFBCFFF5FF205F8838E1FA56.taxon	discussion	Remarks. Tyrannochthonius duo sp. nov. can be distinguished from other described Chinese troglomorphic Tyrannochthonius species by the presence of a pair of well-developed anterior eyes and more setae on tergites V ‒ IX (6 – 9).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFBCFFF5FF205F8838E1FA56.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFB9FFEFFF20592138E1FA22.taxon	description	Figs 37 – 41	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFB9FFEFFF20592138E1FA22.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-HBUARA # 2022 - 490 - 01): China, Guizhou Province, Hezhang County, Pingshan Town, Jiangnan Village, Xianren Cave, under stones, detritus and on the cave walls within 20 – 200 m from the cave entrance (Temperature: 14 ° C, Humidity: 85 %) [27 ° 7 ′ 51.46 ″ N, 104 ° 57 ′ 20.45 ″ E], 1446 m a. s. l., 30 July 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun & Wenlong Fan leg. Paratypes: 13 ♂ (Ps. - MHBU-HBUARA # 2022 - 490 - 02 – HBUARA # 2022 - 490 - 14), 15 ♀ (Ps. - MHBU-HBUARA # 2022 - 490 - 15 – HBUARA # 2022 - 490 - 29), all with the same data as the holotype; 4 ♂ (Ps. - MSWU- GZC 190808 - 01 – GZC 190808 - 04), 8 ♀ (Ps. - MSWU- GZC 190808 - 05 – GZC 190808 - 12), 8 August 2019, Zegang Feng, Zhaoyi Li & Chen Zhang leg., all with the same location as the holotype.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFB9FFEFFF20592138E1FA22.taxon	etymology	Etymology. The specific name is derived from the Latin word “ gracilis ”, meaning gracile, which refers to the elongated coxal spines.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFB9FFEFFF20592138E1FA22.taxon	diagnosis	Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with 2 setae; tergites I – IV each with 2 setae. Pedipalps slender, femur 8.50 – 8.88 (♂), 8.05 – 8.53 (♀) times longer than broad; chela 8.79 – 9.04 (♂), 8.41 – 8.81 (♀) times longer than broad; only fixed chelal finger with intercalary teeth.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFB9FFEFFF20592138E1FA22.taxon	description	Description. Males (holotype and paratypes) (Figs 38 A, 39 A – F, 40, 41). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 39 C, 40 A): carapace 0.99 – 1.00 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, triangular, with 2 setae flanking base; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exter ior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 11 – 12 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 40 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 39 D, 40 B): large, about as long as carapace, 2.50 – 2.53 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 15 – 18 teeth, distal one largest; movable finger with 18 – 21 retrorse contiguous small teeth; galea completely vestigial (Fig. 40 B). Serrula exterior with 21 – 22 and serrula interior with 14 – 15 blades. Rallum with 6 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 40 E). Pedipalp (Figs 39 A, B, E, 40 D, 41 A, B): long and slender, trochanter 1.24 – 1.40, femur 8.50 – 8.88, patella 2.70 – 3.05, chela 8.79 – 9.04, hand 3.08 – 3.30 times longer than broad; femur 2.47 – 2.48 times longer than patella; movable chelal finger 1.63 – 1.76 times longer than hand and 0.60 – 0.62 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 39 E, 40 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, t situated distal to b and slightly proximal to est (Fig. 41 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 28 – 30 macrodenticles, long and pointed, plus 14 – 15 intercalary microdenticles, 42 – 45 in total; movable chelal finger with 34 – 37 macrodenticles (smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 5 – 6 vestigial, rounded and contiguous basal teeth, 40 – 42 in total, without intercalary teeth (Fig. 41 A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 39 B, 41 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 2: 2: 2: 2: 4: 4: 4: 4: 5: 4: T 2 T: 0; tergite IX with an unpaired median seta. Sternal chaetotaxy III – XII: 11 – 12: 10: 8: 7 – 8: 6 – 7: 7: 7: 7: 0: 2. Anterior genital operculum with 10 – 12 setae, genital opening slit-like, with 9 – 11 marginal setae on each side, 31 – 33 in total (Fig. 39 F). Legs (Fig. 41 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.76 – 1.77 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.37 – 2.41 times longer than tibia. Femoropatella of leg IV 3.93 – 4.41 times longer than deep; tibia 6.75 – 6.83 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.80 – 4.00 times longer than deep (TS = 0.23 – 0.26), tarsus 14.14 – 14.57 times longer than deep and 2.48 – 2.68 times longer than metatarsus (TS = 0.27). Arolium slightly shorter than the claws, not divided; claws simple. Adult females (paratypes) (Figs 37 D, 38 B, 39 G). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 3, II 3 – 4, III 5, IV 5; with same tergal chaetotaxy as males; sternal chaetotaxy IV – XII: 12 – 14: 8: 7 – 8: 7: 7: 7: 7: 0: 2; anterior genital operculum with 10 setae, posterior margin with 10 – 12 marginal setae, 20 – 22 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 4.56 times longer than deep (TS = 0.22 – 0.27), tarsus 13.25 – 14.57 times longer than deep and 2.49 – 2.59 times longer than metatarsus (TS = 0.21 – 0.26). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Males: body length 1.96 – 2.06. Pedipalps: trochanter 0.26 – 0.28 / 0.20 – 0.21 (1.24 – 1.40), femur 1.51 – 1.53 / 0.17 – 0.18 (8.50 – 8.88), patella 0.61 – 0.62 / 0.20 – 0.23 (2.70 – 3.05), chela 2.08 – 2.11 / 0.23 – 0.24 (8.79 – 9.04), hand 0.74 – 0.76 / 0.23 – 0.24 (3.08 – 3.30), movable chelal finger length 1.24 – 1.30. Chelicera 0.80 – 0.86 / 0.32 – 0.34 (2.50 – 2.53), movable finger length 0.43 – 0.45. Carapace 0.68 – 0.71 / 0.68 – 0.72 (0.99 – 1.00). Leg I: trochanter 0.23 / 0.18 – 0.19 (1.21 – 1.28), femur 0.85 – 0.88 / 0.09 – 0.10 (8.80 – 9.44), patella 0.48 – 0.50 / 0.09 (5.33 – 5.56), tibia 0.38 – 0.39 / 0.06 – 0.08 (4.88 – 6.33), tarsus 0.90 – 0.94 / 0.06 – 0.07 (13.43 – 15.00). Leg IV: trochanter 0.32 – 0.33 / 0.18 – 0.20 (1.65 – 1.78), femoropatella 1.18 – 1.19 / 0.27 – 0.30 (3.93 – 4.41), tibia 0.81 – 0.82 / 0.12 (6.75 – 6.83), metatarsus 0.38 – 0.40 / 0.10 (3.80 – 4.00), tarsus 0.99 – 1.02 / 0.07 (14.14 – 14.57). Females: body length 2.12 – 2.58. Pedipalps: trochanter 0.27 – 0.30 / 0.21 (1.29 – 1.43), femur 1.61 – 1.62 / 0.19 – 0.20 (8.05 – 8.53), patella 0.64 – 0.67 / 0.23 – 0.24 (2.67 – 2.91), chela 2.27 – 2.29 / 0.26 – 0.27 (8.41 – 8.81), hand 0.78 – 0.83 / 0.26 – 0.27 (3.00 – 3.07), movable chelal finger length 1.39 – 1.43. Chelicera 0.89 – 0.91 / 0.36 – 0.38 (2.39 – 2.47), movable finger length 0.48 – 0.49. Carapace 0.74 – 0.76 / 0.74 – 0.76 (0.97 – 1.03). Leg I: trochanter 0.23 – 0.24 / 0.18 – 0.19 (1.21 – 1.33), femur 0.89 – 0.92 / 0.09 – 0.10 (9.20 – 9.89), patella 0.47 / 0.09 (5.22), tibia 0.39 – 0.41 / 0.07 – 0.08 (5.13 – 5.57), tarsus 0.95 – 0.98 / 0.07 (13.57 – 14.00). Leg IV: trochanter 0.35 / 0.19 (1.84), femoropatella 1.23 – 1.25 / 0.27 – 0.28 (4.39 – 4.63), tibia 0.82 – 0.86 / 0.12 (6.83 – 7.17), metatarsus 0.41 / 0.09 (4.56), tarsus 1.02 – 1.06 / 0.07 – 0.08 (13.25 – 14.57).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFB9FFEFFF20592138E1FA22.taxon	discussion	Remarks. Tyrannochthonius gracilis sp. nov. is similar to T. parcidentatus sp. nov. in having intercalary teeth on the fixed chelal finger only and a pointed, triangular epistome, but differs by a smaller body size but longer appendages (body length 1.96 – 2.06 (♂), 2.12 – 2.58 (♀) vs. 2.29 – 2.62 (♂), 2.64 – 2.66 (♀) mm; chela length 2.08 – 2.11 (♂), 2.27 – 2.29 (♀) vs. 1.89 – 1.93 (♂), 2.02 – 2.05 (♀) mm) and more fixed chelal finger teeth (42 ‒ 45 vs. 30 ‒ 31). Tyrannochthonius gracilis sp. nov. can be distinguished from T. akaelus, T. antridraconis and T. ganshuanensis by the presence of intercalary teeth on the fixed chelal finger only, from T. harveyi by the presence of intercalary teeth on the fixed chelal finger, from T. chixing and T. zhai by the number of setae on tergites I – IV (2 vs. 4) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFB9FFEFFF20592138E1FA22.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFA3FFE2FF205AC738E1F8FE.taxon	description	Figs 42 – 46	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFA3FFE2FF205AC738E1F8FE.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-GZJH- 19 - 26 - 01): China, Guizhou Province, Jianhe County, Yangasha Sub-district, Chuandong Village, Qilin Cave, under stones within 100 – 300 m from the cave entrance (Temperature: 15 ° C, Humidity: 80 %) [26 ° 45 ′ 58.07 ″ N, 108 ° 27 ′ 42.06 ″ E], 743 m a. s. l., 13 October 2019, Zegang Feng & Lingchen Zhao leg. Paratypes: 12 ♂ (Ps. - MHBU-GZJH- 19 - 26 - 02 – GZJH- 19 - 26 - 13), 21 ♀ (Ps. - MHBU-GZJH- 19 - 26 - 14 – GZJH- 19 - 26 - 34), all with the same data as the holotype; 15 ♂ (Ps. - MSWU-HBUARA # 2022 - 521 - 01 – HBUARA # 2022 - 521 - 15), 12 ♀ (Ps. - MSWU-HBUARA # 2022 - 521 - 16 – HBUARA # 2022 - 521 - 27), 13 August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun & Wenlong Fan leg., all with the same location as the holotype.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFA3FFE2FF205AC738E1F8FE.taxon	etymology	Etymology. The specific name is derived from the Latin word “ hispidus ” (= hirsute), which refers to the increased number of setae at the posterior margin of the carapace.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFA3FFE2FF205AC738E1F8FE.taxon	diagnosis	Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with 2 – 4 setae; tergites I – VI each with 4 setae. Pedipalps slender, femur 5.80 – 5.93 (♂), 5.93 – 6.14 (♀) times longer than broad; chela 6.68 – 6.72 (♂), 6.20 – 6.79 (♀) times longer than broad; both chelal fingers with intercalary teeth.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFA3FFE2FF205AC738E1F8FE.taxon	description	Description. Males (holotype and paratypes) (Figs 43 A, 44 A – F, 45, 46). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 44 C, 45 A): carapace 0.94 – 0.98 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, triangular, with 2 setae flanking base; with 18 – 20 setae arranged s 3 – 4 s: 4: 4: 2 – 3: 2 – 4, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 7 – 10 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 45 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 44 D, 45 B): large, about as long as carapace, 2.29 – 2.30 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 10 – 12 teeth, distal one largest; movable finger with 10 – 13 retrorse contiguous small teeth; galea completely vestigial (Fig. 45 B). Serrula exterior with 18 – 20 and serrula interior with 14 – 15 blades. Rallum with 6 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 45 E). Pedipalp (Figs 44 A, B, E, 45 D, 46 A, B): long and slender, trochanter 1.20 – 1.31, femur 5.80 – 5.93, patella 2.00 – 2.19, chela 6.68 – 6.72, hand 2.32 – 2.33 times longer than broad; femur 2.49 – 2.59 times longer than patella; movable chelal finger 1.77 – 1.83 times longer than hand and 0.61 – 0.64 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 44 E, 45 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated distal to b and slightly proximal to est (Fig. 46 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 26 – 28 macrodenticles, long and pointed, plus 15 – 16 intercalary microdenticles, 42 – 43 in total; movable chelal finger with 22 – 24 macrodenticles (markedly smaller than teeth on fixed chelal finger), retrorse and pointed, plus 9 – 10 intercalary microdenticles and 7 – 8 vestigial, rounded and contiguous basal teeth, 38 – 42 in total (Fig. 46 A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 44 B, 46 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 4: 4: 4: 4: 4 – 5: 5: 5: 4: T 2 T: 0; tergites VIII – IX each with an unpaired median seta. Sternal chaetotaxy III – XII: 13 – 15: 13 – 14: 10: 9: 9: 9: 9 – 11: 9: 0: 2. Anterior genital operculum with 10 setae, genital opening slit-like, with 10 – 13 marginal setae on each side, 31 – 34 in total (Fig. 44 F). Legs (Fig. 46 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.79 – 1.88 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.27 – 2.43 times longer than tibia. Femoropatella of leg IV 2.68 – 2.70 times longer than deep; tibia 5.00 – 5.38 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.14 times longer than deep (TS = 0.32), tarsus 12.50 – 12.75 times longer than deep and 2.27 – 2.32 times longer than metatarsus (TS = 0.26 – 0.29). Arolium slightly shorter than the claws, not divided; claws simple. Adult females (paratypes) (Figs 43 B, 44 G). Mostly same as males, but a little larger; with same chaetotaxy of coxae as males; tergal chaetotaxy I – XII: 4: 4: 4: 4: 4: 4: 4: 5: 5: 4: T 2 T: 0; sternal chaetotaxy IV – XII: 14: 10: 9: 9: 10: 9 – 10: 8 – 9: 0: 2; anterior genital operculum with 9 – 10 setae, posterior margin with 13 – 14 marginal setae, 23 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.50 – 3.67 times longer than deep (TS = 0.33 – 0.36), tarsus 10.00 times longer than deep and 2.27 – 2.38 times longer than metatarsus (TS = 0.32). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Males: body length 1.56 – 1.78. Pedipalps: trochanter 0.17 – 0.18 / 0.13 – 0.15 (1.20 – 1.31), femur 0.83 – 0.87 / 0.14 – 0.15 (5.80 – 5.93), patella 0.32 – 0.35 / 0.16 (2.00 – 2.19), chela 1.21 – 1.27 / 0.18 – 0.19 (6.68 – 6.72), hand 0.42 – 0.44 / 0.18 – 0.19 (2.32 – 2.33), movable chelal finger length 0.77 – 0.78. Chelicera 0.53 – 0.55 / 0.23 – 0.24 (2.29 – 2.30), movable finger length 0.28 – 0.29. Carapace 0.48 – 0.50 / 0.49 – 0.53 (0.94 – 0.98). Leg I: trochanter 0.14 – 0.15 / 0.12 (1.17 – 1.25), femur 0.43 – 0.45 / 0.07 – 0.08 (5.38 – 6.43), patella 0.24 / 0.07 (3.43), tibia 0.21 – 0.22 / 0.05 – 0.06 (3.50 – 4.40), tarsus 0.50 – 0.51 / 0.04 – 0.05 (10.20 – 12.50). Leg IV: trochanter 0.20 – 0.21 / 0.12 – 0.13 (1.62 – 1.67), femoropatella 0.59 – 0.62 / 0.22 – 0.23 (2.68 – 2.70), tibia 0.43 – 0.45 / 0.08 – 0.09 (5.00 – 5.38), metatarsus 0.22 / 0.07 (3.14), tarsus 0.50 – 0.51 / 0.07 (12.50 – 12.75). Females: body length 1.77 – 1.99. Pedipalps: trochanter 0.18 – 0.20 / 0.14 (1.29 – 1.43), femur 0.86 – 0.89 / 0.14 – 0.15 (5.93 – 6.14), patella 0.35 – 0.36 / 0.16 (2.19 – 2.25), chela 1.24 – 1.29 / 0.19 – 0.20 (6.20 – 6.79), hand 0.44 – 0.45 / 0.19 – 0.20 (2.25 – 2.32), movable chelal finger length 0.77 – 0.82. Chelicera 0.55 – 0.58 / 0.24 – 0.25 (2.29 – 2.32), movable finger length 0.29. Carapace 0.49 / 0.53 – 0.54 (0.91 – 0.92). Leg I: trochanter 0.12 – 0.15 / 0.12 – 0.13 (1.00 – 1.15), femur 0.44 – 0.45 / 0.07 – 0.08 (5.50 – 6.43), patella 0.24 – 0.26 / 0.07 (3.43 – 3.71), tibia 0.21 / 0.05 – 0.06 (2.63 – 3.50), tarsus 0.49 – 0.51 / 0.05 (9.80 – 10.20). Leg IV: trochanter 0.20 / 0.13 (1.54), femoropatella 0.60 – 0.62 / 0.20 – 0.21 (2.95 – 3.00), tibia 0.44 – 0.45 / 0.08 – 0.09 (5.00 – 5.50), metatarsus 0.21 – 0.22 / 0.06 (3.50 – 3.67), tarsus 0.50 / 0.05 (10.00).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFA3FFE2FF205AC738E1F8FE.taxon	discussion	Remarks. Tyrannochthonius hispidus sp. nov. is similar to T. acutus sp. nov. in having intercalary teeth on both chelal fingers and a pointed, triangular epistome, but differs by a smaller body size (body length 1.56 – 1.78 (J), 1.77 – 1.99 (♀) vs. 2.07 – 2.13 (J), 2.16 – 2.29 (♀) mm; chela 6.68 – 6.72 (J), 6.20 – 6.79 (♀) vs. 7.29 – 7.65 (J), 7.38 – 7.79 (♀) times longer than board), fewer fixed chelal finger teeth (42 ‒ 43 vs. 57 ‒ 60) and the number of blades of rallum (6 vs. 7). Tyrannochthonius hispidus sp. nov. can be distinguished from T. chixing, T. harveyi and T. zhai by the presence of intercalary teeth on both chelal fingers, from T. akaelus and T. ganshuanensis by the number of setae on tergites I – II (4 vs. 2), from T. antridraconis by a smaller size (body length max. 1.78 mm vs. min. 1.8 mm; chela 6.68 – 6.72 times vs. 7.90 – 8.00 times longer than board, length 1.21 – 1.27 (J) vs. 1.70 – 1.76 (J) mm), more fixed chelal finger teeth (42 ‒ 43 vs. 33 ‒ 36) and the relative position of trichobothrium sb (sb closer to st than to b vs. closer to b) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFA3FFE2FF205AC738E1F8FE.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFAAFF98FF205F8838E1FF4B.taxon	description	Figs 47 – 51	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFAAFF98FF205F8838E1FF4B.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-GZC 190804 - 01): China, Guizhou Province, Pu’an County, Qingshan Town, Xiajiehe Village, Liujiadadong Cave, under stones and clods in the deep zone (Temperature: 13 ° C, Humidity: 90 %) [25 ° 28 ′ 6.88 ″ N, 105 ° 0 ′ 19.57 ″ E], 1559 m a. s. l., 4 August 2019, Zegang Feng, Zhaoyi Li & Chen Zhang leg. Paratypes: 3 ♂ (Ps. - MHBU-GZC 190804 - 02 – GZC 190804 - 04), 5 ♀ (Ps. - MSWU-GZC 190804 - 05 – GZC 190804 - 09), all with the same data as the holotype.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFAAFF98FF205F8838E1FF4B.taxon	etymology	Etymology. The specific name is derived from the Latin word “ infirmus ”, meaning weak, which refers to the presence of a pair of weak eyespots.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFAAFF98FF205F8838E1FF4B.taxon	diagnosis	Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace with a pair of anterior eyespots only; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with 2 setae; tergites I – Ⅴ each with 4 setae. Pedipalps slender, femur 6.72 – 6.88 (♂), 6.47 – 6.94 (♀) times longer than broad; chela 7.27 – 7.42 (♂), 6.28 – 6.83 (♀) times longer than broad; both chelal fingers with intercalary teeth.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFAAFF98FF205F8838E1FF4B.taxon	description	Description. Males (holotype and paratypes) (Figs 48 A, 49 A – F, H, 50, 51). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 49 C, D, 50 A): carapace 0.97 – 0.99 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; anterior eyes reduced into eyespots, posterior eyes reduced completely; anterior margin slightly serrate; epistome small and pointed, triangular, with 2 setae flanking base; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 11 – 13 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 50 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 49 E, 50 B): large, about as long as carapace, 2.29 – 2.30 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 12 – 14 teeth, distal one largest; movable finger with 12 – 13 retrorse contiguous small teeth; galea completely vestigial (Fig. 50 B). Serrula exterior with 18 – 21 and serrula interior with 13 – 15 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 50 E). Pedipalp (Figs 49 A, B, H, 50 D, 51 A, B): long and slender, trochanter 1.42 – 1751, femur 6.72 – 6.88, patella 2.09 – 2.32, chela 7.27 – 7.42, hand 2.64 – 2.67 times longer than broad; femur 2.50 – 2.63 times longer than patella; movable chelal finger 1.74 – 1.78 times longer than hand and 0.63 – 0.64 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 49 H, 50 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, t situated midway between it and est; est situated distal to b (Fig. 51 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 28 – 31 macrodenticles, slightly retrorse and pointed, plus 20 – 21 intercalary microdenticles, 48 – 52 in total; movable chelal finger with 25 – 26 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 18 – 19 intercalary microdenticles and 8 – 10 vestigial, rounded and contiguous basal teeth, 52 – 54 in total (Fig. 51 A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 49 B, 51 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 4: 4: 4: 5: 5 – 6: 5: 5: 4: T 2 T: 0; tergites VII – IX each with an unpaired median seta. Sternal chaetotaxy III – XII: 12 – 13: 12 – 13: 8 – 9: 10: 9 – 10: 8 – 10: 9 – 10: 9 – 10: 0: 2. Anterior genital operculum with 10 – 11 setae, genital opening slit-like, with 20 – 23 marginal setae on each side, 53 in total (Fig. 49 F). Legs (Fig. 51 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.92 – 2.09 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.44 – 2.59 times longer than tibia. Femoropatella of leg IV 3.59 – 3.85 times longer than deep; tibia 6.25 – 6.58 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.56 – 3.60 times longer than deep (TS = 0.28), tarsus 12.43 – 13.83 times longer than deep and 2.42 – 2.59 times longer than metatarsus (TS = 0.36 – 0.37). Arolium slightly shorter than the claws, not divided; claws simple. Adult females (paratypes) (Figs 48 B, 49 G). Mostly same as males, but a little larger; with same chaetotaxy of coxae as males; tergal chaetotaxy I – XII: 4: 4: 4: 4: 4: 4 – 5: 5 – 6: 5 – 7: 6: 4: T 2 T: 0; sternal chaetotaxy IV – XII: 11 – 13: 9: 10: 9 – 10: 9 – 10: 10 – 11: 10: 0: 2; anterior genital operculum with 10 setae, posterior margin with 11 – 13 marginal setae, 21 – 23 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.44 – 3.56 times longer than deep (TS = 0.28 – 0.29), tarsus 12.29 – 13.50 times longer than deep and 2.53 – 2.77 times longer than metatarsus (TS = 0.35 – 0.38). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Males: body length 2.41 – 2.54. Pedipalps: trochanter 0.27 – 0.35 / 0.19 – 0.20 (1.42 – 1.75), femur 1.10 – 1.21 / 0.16 – 0.18 (6.72 – 6.88), patella 0.44 – 0.46 / 0.19 – 0.22 (2.09 – 2.32), chela 1.60 – 1.78 / 0.22 – 0.24 (7.27 – 7.42), hand 0.58 – 0.64 / 0.22 – 0.24 (2.64 – 2.67), movable chelal finger length 1.01 – 1.14. Chelicera 0.69 – 0.71 / 0.30 – 0.31 (2.29 – 2.30), movable finger length 0.37 – 0.40. Carapace 0.62 – 0.68 / 0.64 – 0.69 (0.97 – 0.99). Leg I: trochanter 0.19 – 0.21 / 0.15 – 0.17 (1.24 – 1.27), femur 0.67 – 0.75 / 0.09 – 0.11 (6.82 – 7.44), patella 0.32 – 0.39 / 0.09 (3.56 – 4.33), tibia 0.29 – 0.34 / 0.06 – 0.08 (4.25 – 4.83), tarsus 0.75 – 0.83 / 0.06 – 0.07 (11.86 – 12.50). Leg IV: trochanter 0.31 – 0.33 / 0.17 – 0.19 (1.74 – 1.82), femoropatella 0.97 – 1.04 / 0.27 (3.59 – 3.85), tibia 0.75 – 0.79 / 0.12 (6.25 – 6.58), metatarsus 0.32 – 0.36 / 0.09 – 0.10 (3.56 – 3.60), tarsus 0.83 – 0.87 / 0.06 – 0.07 (12.43 – 13.83). Females: body length 2.50 – 2.67. Pedipalps: trochanter 0.21 – 0.24 / 0.18 (1.17 – 1.33), femur 1.10 – 1.18 / 0.17 (6.47 – 6.94), patella 0.45 / 0.20 (2.25), chela 1.57 – 1.64 / 0.24 – 0.25 (6.28 – 6.83), hand 0.59 – 0.61 / 0.24 – 0.25 (2.36 – 2.54), movable chelal finger length 1.00 – 1.04. Chelicera 0.69 – 0.71 / 0.32 – 0.33 (2.15 – 2.16), movable finger length 0.37 – 0.39. Carapace 0.66 – 0.67 / 0.69 – 0.70 (0.94 – 0.97). Leg I: trochanter 0.17 – 0.19 / 0.15 – 0.16 (1.06 – 1.27), femur 0.64 – 0.68 / 0.10 (6.40 – 6.80), patella 0.35 / 0.08 – 0.09 (3.89 – 4.38), tibia 0.28 – 0.32 / 0.07 – 0.08 (4.00), tarsus 0.70 – 0.77 / 0.07 (10.00 – 11.00). Leg IV: trochanter 0.29 / 0.17 (1.71), femoropatella 0.95 – 0.97 / 0.25 – 0.26 (3.73 – 3.80), tibia 0.69 – 0.72 / 0.11 – 0.12 (6.00 – 6.27), metatarsus 0.31 – 0.32 / 0.09 (3.44 – 3.56), tarsus 0.81 – 0.86 / 0.06 – 0.07 (12.29 – 13.50).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFAAFF98FF205F8838E1FF4B.taxon	discussion	Remarks. Tyrannochthonius infirmus sp. nov. can be distinguished from other described Chinese troglomorphic Tyrannochthonius species by the presence of a pair of anterior eyespots.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFAAFF98FF205F8838E1FF4B.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFD4FF9EFF205E2F38E1F826.taxon	description	Figs 52 – 56	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFD4FF9EFF205E2F38E1F826.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-HBUARA # 2021 - 168 - 01): China, Guizhou Province, Panzhou City, Danxia Town, Zimu Cave, under stones in the deep zone (Temperature: 14 ° C, Humidity: 90 %) [25 ° 40 ′ 29.59 ″ N, 104 ° 37 ′ 36.64 ″ E], 1787 m a. s. l., 15 July 2021, Zegang Feng, Hongru Xu, Liu Fu & Nana Zhan leg. Paratypes: 3 ♂ (Ps. - MHBU-HBUARA # 2021 - 168 - 02 – HBUARA # 2021 - 168 - 04), 5 ♀ (Ps. - MSWU-HBUARA # 2021 - 168 - 05 – HBUARA # 2021 - 168 - 09), all with the same data as the holotype.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFD4FF9EFF205E2F38E1F826.taxon	etymology	Etymology. The specific name is derived from the Latin word “ latus ”, meaning wide, which refers to the widened chelal hands.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFD4FF9EFF205E2F38E1F826.taxon	diagnosis	Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, represented by a bump; posterior margin of carapace with 2 setae; tergites I – VII each with 4 setae. Pedipalps slender, femur 9.29 – 9.33 (♂), 8.75 – 8.79 (♀) times longer than broad; chela 6.69 – 7.13 (♂), 6.15 – 6.58 (♀) times longer than broad; chelal hand wider, hand 2.80 – 2.90 (♂), 2.53 – 2.72 (♀) times longer than broad (hand wide 0.30 – 0.35 (♂), 0.36 – 0.40 (♀) mm); both chelal fingers without intercalary teeth.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFD4FF9EFF205E2F38E1F826.taxon	description	Description. Males (holotype and paratypes) (Figs 53 A, 54 A – F, 55, 56). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 54 C, 55 A): carapace 0.92 – 0.96 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, represented by a bump, with 2 setae flanking base; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 11 – 12 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 55 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 54 D, 55 B): large, about as long as carapace, 2.39 – 2.45 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 18 – 20 teeth, distal one largest; movable finger with 14 – 17 retrorse contiguous small teeth; galea completely vestigial (Fig. 55 B). Serrula exterior with 20 – 24 and serrula interior with 14 – 15 blades. Rallum with 7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 55 E). Pedipalp (Figs 54 A, B, E, 55 D, 56 A, B): long and slender, trochanter 1.22 – 1.29, femur 9.29 – 9.33, patella 2.52 – 2.58, chela 6.69 – 7.13, hand 2.80 – 2.90 times longer than broad; femur 2.71 – 2.72 times longer than patella; movable chelal finger 1.41 – 1.46 times longer than hand and 0.59 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 54 E, 55 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated at same level as it; est situated distal to b (Fig. 56 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 29 – 31 macrodenticles, long and pointed, slightly retrorse; movable chelal finger with 30 – 32 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed (Fig. 56 A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 54 B, 56 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 4: 4: 4: 4: 4: 4 – 5: 5: 4: T 2 T: 0; tergite IX with an unpaired median seta. Sternal chaetotaxy III – XII: 13 – 15: 12 – 14: 8: 8 – 9: 9: 9 – 11: 9 – 11: 9: 0: 2. Anterior genital operculum with 14 setae, genital opening slit-like, with 13 – 15 marginal setae on each side, 41 – 44 in total (Fig. 54 F). Legs (Fig. 56 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.78 – 1.87 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.44 – 2.65 times longer than tibia. Femoropatella of leg IV 4.96 – 5.32 times longer than deep; tibia 7.00 – 7.75 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.60 – 4.10 times longer than deep (TS = 0.27 – 0.28), tarsus 14.43 – 16.00 times longer than deep and 2.73 – 2.81 times longer than metatarsus (TS = 0.26 – 0.30). Arolium slightly shorter than the claws, not divided; claws simple. Adult females (paratypes) (Figs 53 B, 54 G). Mostly same as males, but a little larger; with same chaetotaxy of coxae as males; tergal chaetotaxy I – XII: 4: 4: 4: 4: 4: 4: 4: 4 – 5: 5: 4: T 2 T: 0; sternal chaetotaxy IV – XII: 13: 9: 9 – 10: 10 – 11: 10 – 11: 10 – 11: 9: 0: 2; anterior genital operculum with 11 setae, posterior margin with 12 marginal setae, 23 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.70 – 4.00 times longer than deep (TS = 0.28 – 0.35), tarsus 12.75 – 13.88 times longer than deep and 2.76 – 2.78 times longer than metatarsus (TS = 0.25 – 0.30). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Males: body length 2.59 – 2.80. Pedipalps: trochanter 0.27 – 0.28 / 0.21 – 0.23 (1.22 – 1.29), femur 1.58 – 1.68 / 0.17 – 0.18 (9.29 – 9.33), patella 0.58 – 0.62 / 0.23 – 0.24 (2.52 – 2.58), chela 2.14 – 2.34 / 0.30 – 0.35 (6.69 – 7.13), hand 0.87 – 0.98 / 0.30 – 0.35 (2.80 – 2.90), movable chelal finger length 1.27 – 1.38. Chelicera 0.81 – 0.86 / 0.33 – 0.36 (2.39 – 2.45), movable finger length 0.45 – 0.48. Carapace 0.65 – 0.71 / 0.71 – 0.74 (0.92 – 0.96). Leg I: trochanter 0.20 – 0.26 / 0.18 – 0.19 (1.11 – 1.37), femur 0.84 – 0.91 / 0.11 (7.64 – 8.27), patella 0.45 – 0.51 / 0.10 (4.50 – 5.10), tibia 0.37 – 0.41 / 0.08 (4.63 – 5.13), tarsus 0.98 – 1.00 / 0.07 (14.00 – 14.29). Leg IV: trochanter 0.33 / 0.19 (1.74), femoropatella 1.24 – 1.33 / 0.25 (4.96 – 5.32), tibia 0.84 – 0.93 / 0.12 (7.00 – 7.75), metatarsus 0.36 – 0.41 / 0.10 (3.60 – 4.10), tarsus 1.01 – 1.02 / 0.07 (14.43 – 16.00). Females: body length 2.91 – 3.11. Pedipalps: trochanter 0.29 – 0.32 / 0.23 – 0.26 (1.23 – 1.26), femur 1.67 – 1.75 / 0.19 – 0.20 (8.75 – 8.79), patella 0.68 – 0.70 / 0.25 – 0.27 (2.59 – 2.72), chela 2.37 – 2.46 / 0.36 – 0.40 (6.15 – 6.58), hand 0.98 – 1.01 / 0.36 – 0.40 (2.53 – 2.72), movable chelal finger length 1.39 – 1.45. Chelicera 0.94 – 0.97 / 0.38 – 0.40 (2.43 – 2.47), movable finger length 0.52 – 0.55. Carapace 0.72 – 0.75 / 0.77 – 0.84 (0.89 – 0.94). Leg I: trochanter 0.23 – 0.26 / 0.19 – 0.20 (1.21 – 1.30), femur 0.94 – 0.99 / 0.11 – 0.12 (8.25 – 8.55), patella 0.51 – 0.53 / 0.10 – 0.11 (4.82 – 5.10), tibia 0.39 – 0.41 / 0.07 (5.57 – 5.86), tarsus 1.02 – 1.05 / 0.08 (12.75 – 13.13). Leg IV: trochanter 0.34 – 0.36 / 0.19 – 0.21 (1.71 – 1.79), femoropatella 1.32 – 1.38 / 0.25 – 0.26 (5.28 – 5.31), tibia 0.92 – 0.93 / 0.12 (7.67 – 7.75), metatarsus 0.37 – 0.40 / 0.10 (3.70 – 4.00), tarsus 1.02 – 1.11 / 0.08 (12.75 – 13.88).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFD4FF9EFF205E2F38E1F826.taxon	discussion	Remarks. Tyrannochthonius latus sp. nov. is similar to T. nanxingensis sp. nov. in lacking intercalary teeth on both chelal fingers, but differs by a longer palp (chela length 2.14 – 2.34 (♂), 2.37 – 2.46 (♀) vs. 1.92 – 1.97 (♂), 1.85 (♀) mm; palpal femur length 1.58 – 1.68 (♂), 1.67 – 1.75 (♀) vs. 1.34 – 1.35 (♂), 1.24 (♀) mm; movable chelal finger 1.41 – 1.46 vs. 1.78 – 1.86 times longer than hand), the relative position of trichobothrium sb (sb closer to st than to b vs. closer to b), the number of setae on tergite IV (4 vs. 5) and the number of blades of rallum (7 vs. 8). Tyrannochthonius latus sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on fixed chelal fingers, from T. harveyi by the number of setae on tergites I – III (4 vs. 2) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFD4FF9EFF205E2F38E1F826.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFDEFF94FF205F8838E1FE3A.taxon	description	Figs 57 – 61	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFDEFF94FF205F8838E1FE3A.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-HBUARA # 2022 - 504 - 01): China, Guizhou Province, Xingren City, Xinlongchang Town, Lianzhuang Village, Daxiao Cave, under stones, detritus and on the cave walls within 200 – 400 m from the cave entrance (Temperature: 15 ° C, Humidity: 85 %) [25 ° 26 ′ 16.92 ″ N, 105 ° 6 ′ 58.31 ″ E], 1473 m a. s. l., 5 August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun & Wenlong Fan leg. Paratypes: 14 ♂ (Ps. - MHBU-HBUARA # 2022 - 504 - 02 – HBUARA # 2022 - 504 - 15), 6 ♀ (Ps. - MSWU- HBUARA # 2022 - 504 - 16 – HBUARA # 2022 - 504 - 21), all with the same data as the holotype.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFDEFF94FF205F8838E1FE3A.taxon	etymology	Etymology. The specific name is derived from the Latin word “ maculosus ”, meaning speckled, which refers to the speckled tergites.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFDEFF94FF205F8838E1FE3A.taxon	diagnosis	Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with 2 setae; tergites I – III each with 4 setae. Pedipalps slender, femur 6.47 – 6.80 (♂), 6.67 – 6.80 (♀) times longer than broad; chela 6.81 – 7.45 (♂), 6.55 (♀) times longer than broad; both chelal fingers with intercalary teeth.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFDEFF94FF205F8838E1FE3A.taxon	description	Description. Males (holotype and paratypes) (Figs 57 F, 58 A, 59 A – F, 60, 61). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 59 C, 60 A): carapace 1.02 – 1.04 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, triangular, with 2 setae flanking base; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 3 – 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 9 – 10 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 60 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 59 D, 60 B): large, about as long as carapace, 2.22 – 2.33 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 12 – 14 teeth, distal one largest; movable finger with 14 – 17 retrorse contiguous small teeth; galea completely vestigial (Fig. 60 B). Serrula exterior with 18 – 21 and serrula interior with 14 – 15 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 60 E). Pedipalp (Figs 59 A, B, E, 60 D, 61 A, B): long and slender, trochanter 1.22 – 1.35, femur 6.47 – 6.80, patella 2.22 – 2.28, chela 6.81 – 7.45, hand 2.38 – 2.65 times longer than broad; femur 2.43 – 2.49 times longer than patella; movable chelal finger 1.83 – 1.90 times longer than hand and 0.65 – 0.66 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 59 E, 60 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb midway between st and b; b and t situated subdistally, t situated at same level as it; est situated distal to b (Fig. 61 C). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 34 – 35 macrodenticles, slightly retrorse and pointed, plus 24 – 25 intercalary microdenticles, 58 – 60 in total; movable chelal finger with 20 – 22 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 18 – 19 intercalary microdenticles and 12 – 14 vestigial, rounded and contiguous basal teeth, 52 – 53 in total (Fig. 61 C). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 59 B, 61 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 4: 4 – 5: 4 – 5: 5: 5: 5 – 6: 5: 4: T 2 T: 0; tergite IX with an unpaired median seta. Sternal chaetotaxy III – XII: 14 – 15: 13 – 14: 9 – 10: 8 – 10: 9 – 10: 9: 9: 9: 0: 2. Anterior genital operculum with 10 setae, genital opening slit-like, with 17 – 22 marginal setae on each side, 45 – 50 in total (Fig. 59 F). Legs (Fig. 61 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.94 – 1.97 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.33 – 2.36 times longer than tibia. Femoropatella of leg IV 3.63 – 3.79 times longer than deep; tibia 5.82 – 6.00 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.11 – 3.63 times longer than deep (TS = 0.21 – 0.24), tarsus 11.83 – 12.67 times longer than deep and 2.54 – 2.62 times longer than metatarsus (TS = 0.28 – 0.32). Arolium slightly shorter than the claws, not divided; claws simple. Adult females (paratypes) (Figs 58 B, 59 G). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; tergal chaetotaxy I – XII: 4: 3 – 4: 4: 4 – 5: 5 – 6: 5 – 7: 5 – 6: 5 – 6: 5 – 6: 4: T 2 T: 0; sternal chaetotaxy IV – XII: 14: 10: 9 – 10: 9 – 10: 9 – 10: 10: 9 – 10: 0: 2; anterior genital operculum with 12 setae, posterior margin with 14 – 15 marginal setae, 26 – 27 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.00 – 3.22 times longer than deep (TS = 0.24 – 0.25), tarsus 11.33 – 12.33 times longer than deep and 2.55 – 2.83 times longer than metatarsus (TS = 0.34 – 0.35). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Males: body length 2.22 – 2.31. Pedipalps: trochanter 0.22 – 0.23 / 0.17 – 0.18 (1.22 – 1.35), femur 0.97 – 1.02 / 0.15 (6.47 – 6.80), patella 0.40 – 0.41 / 0.18 (2.22 – 2.28), chela 1.43 – 1.49 / 0.20 – 0.21 (6.81 – 7.45), hand 0.50 – 0.53 / 0.20 – 0.21 (2.38 – 2.65), movable chelal finger length 0.95 – 0.97. Chelicera 0.60 – 0.63 / 0.27 (2.22 – 2.33), movable finger length 0.33 – 0.35. Carapace 0.59 / 0.57 – 0.58 (1.02 – 1.04). Leg I: trochanter 0.17 – 0.19 / 0.14 – 0.15 (1.21 – 1.27), femur 0.60 – 0.61 / 0.08 (7.50 – 7.63), patella 0.31 / 0.07 – 0.08 (3.88 – 4.43), tibia 0.27 – 0.28 / 0.06 (4.50 – 4.67), tarsus 0.63 – 0.66 / 0.06 (10.50 – 11.00). Leg IV: trochanter 0.25 – 0.27 / 0.15 – 0.16 (1.67 – 1.69), femoropatella 0.87 – 0.91 / 0.24 (3.63 – 3.79), tibia 0.64 – 0.66 / 0.11 (5.82 – 6.00), metatarsus 0.28 – 0.29 / 0.08 – 0.09 (3.11 – 3.63), tarsus 0.71 – 0.76 / 0.06 (11.83 – 12.67). Females: body length 2.50 – 2.59. Pedipalps: trochanter 0.22 / 0.18 – 0.19 (1.16 – 1.22), femur 1.00 – 1.02 / 0.15 (6.67 – 6.80), patella 0.40 / 0.18 – 0.19 (2.11 – 2.22), chela 1.44 / 0.22 (6.55), hand 0.51 / 0.22 (2.32), movable chelal finger length 0.94. Chelicera 0.64 – 0.65 / 0.29 – 0.30 (2.17 – 2.21), movable finger length 0.35. Carapace 0.61 – 0.62 / 0.62 – 0.63 (0.98). Leg I: trochanter 0.18 / 0.15 – 0.16 (1.13 – 1.20), femur 0.58 – 0.61 / 0.08 – 0.09 (6.44 – 7.63), patella 0.30 – 0.32 / 0.08 (3.75 – 4.00), tibia 0.27 – 0.28 / 0.06 – 0.07 (3.86 – 4.67), tarsus 0.60 – 0.64 / 0.06 (10.00 – 10.67). Leg IV: trochanter 0.27 – 0.28 / 0.15 (1.80 – 1.87), femoropatella 0.90 – 0.92 / 0.23 (3.91 – 4.00), tibia 0.65 – 0.70 / 0.11 (5.91 – 6.36), metatarsus 0.24 – 0.29 / 0.08 – 0.09 (3.00 – 3.22), tarsus 0.68 – 0.74 / 0.06 (11.33 – 12.33).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFDEFF94FF205F8838E1FE3A.taxon	discussion	Remarks. Tyrannochthonius maculosus sp. nov. is similar to T. multidentatus sp. nov. in having intercalary teeth on both chelal fingers and a pointed epistome, but differs by the trait of teeth on movable chelal finger (slightly retrorse and well-spaced vs. markedly retrorse and continuous), the relative position of trichobothrium sb (sb midway between st and b vs. closer to b) and the number of blades of rallum (8 vs. 6). Tyrannochthonius maculosus sp. nov. can be distinguished from T. chixing, T. harveyi and T. zhai by the presence of intercalary teeth on both chelal fingers, from T. akaelus and T. ganshuanensis by the number of setae on tergites I – II (4 vs. 2), from T. antridraconis by a shorter palp (chela 6.81 – 7.45 (♂), 6.55 (♀) vs. 7.90 – 8.00 (♂), 6.90 – 7.55 (♀) times longer than broad; palpal femur 6.47 – 6.80 (♂), 6.67 – 6.80 (♀) vs. 7.30 – 8.30 (♂), 7.10 – 7.50 (♀) times longer than broad) and the relative position of trichobothrium sb (sb midway between st and b vs. closer to b) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFDEFF94FF205F8838E1FE3A.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFD8FF8CFF205ECA3E10FE1E.taxon	description	Figs 62 – 69	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFD8FF8CFF205ECA3E10FE1E.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-HBUARA # 2021 - 204 - 01): China, Guizhou Province, Pingtang County, Tangbian Town, Xinhong Village, Xiaohui Cave, under stones and on the cave walls in the deep zone (Temperature: 15 ° C, Humidity: 80 %) [25 ° 38 ′ 43.65 ″ N, 106 ° 45 ′ 46.81 ″ E], 888.6 m a. s. l., 3 August 2021, Zegang Feng, Hongru Xu, Liu Fu & Nana Zhan leg. Paratypes: 1 ♂ (Ps. - MHBU-HBUARA # 2021 - 204 - 02), 2 ♀ (Ps. - MSWU-HBUARA # 2021 - 204 - 03 & HBUARA # 2021 - 204 - 04), all with the same data as the holotype; 2 ♂ (Ps. - MHBU-HBUARA # 2021 - 205 - 01 & HBUARA # 2021 - 205 - 02), 5 ♀ (Ps. - MSWU-HBUARA # 2021 - 205 - 03 – HBUARA # 2021 - 205 - 07) Tangbian Town, Bailu Village, Weier Cave, under stones in the deep zone [25 ° 38 ′ 8.52 ″ N, 106 ° 43 ′ 8.16 ″ E], 919.3 m a. s. l., all with the same collection date and collectors as the holotype; 9 ♂ (Ps. - MHBU-HBUARA # 2022 - 513 - 01 – HBUARA # 2022 - 513 - 09), 5 ♀ (Ps. - MSWU-HBUARA # 2022 - 513 - 10 – HBUARA # 2022 - 513 - 14), Luodian County, Bianyan Town, Dahao Village, Da Cave, under stones in the deep zone [25 ° 40 ′ 5.91 ″ N, 106 ° 41 ′ 23.07 ″ E], 930 m a. s. l., 09 August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun & Wenlong Fan leg.; 5 ♂ (Ps. - MHBU-HBUARA # 2022 - 514 - 01 – HBUARA # 2022 - 514 - 05), 2 ♀ (Ps. - MSWU-HBUARA # 2022 - 514 - 06 & HBUARA # 2022 - 514 - 07), Luodian County, Bianyan Town, Dashang Village, Qixia Cave, under stones in the deep zone [25 ° 37 ′ 55.58 ″ N, 106 ° 42 ′ 23.24 ″ E], 956 m a. s. l., 10 August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun & Wenlong Fan leg.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFD8FF8CFF205ECA3E10FE1E.taxon	etymology	Etymology. The specific name is derived from a combination of the Latin word “ multus ” and “ cavus ”, meaning numerous and cavernous, respectively, which refers to the species found in several caves.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFD8FF8CFF205ECA3E10FE1E.taxon	diagnosis	Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and represented by a slight bump; posterior margin of carapace with 2 setae; tergites I – IV each with 2 setae. Pedipalps slender, femur 6.79 – 7.08 (♂), 6.53 – 7.21 (♀) times longer than broad; chela 7.53 – 8.20 (♂), 7.05 – 8.38 (♀) times longer than broad; both chelal fingers without intercalary teeth.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFD8FF8CFF205ECA3E10FE1E.taxon	description	Description. Males (holotype and paratypes) (Figs 64 F, 66 A, 67 A – F, 68, 69). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 67 C, 68 A): carapace 1.02 – 1.14 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and represented by a slight bump, with 2 setae flanking base; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 3 – 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 8 – 10 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 68 C); intercoxal tubercle absent; without sub-oral seta Chelicera (Figs 67 D, 68 B): large, about as long as carapace, 2.27 – 2.42 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 11 – 13 teeth, distal one largest; movable finger with 12 – 13 retrorse contiguous small teeth; galea represented by a very slight bump on movable finger (Fig. 68 B). Serrula exterior with 18 – 21 and serrula interior with 13 – 15 blades. Rallum with 7 – 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 68 E). Pedipalp (Figs 67 A, B, E, 68 D, 69 A, B): long and slender, trochanter 1.20 – 1.47, femur 6.79 – 7.08, patella 2.40 – 2.86, chela 7.53 – 8.20, hand 2.65 – 2.94 times longer than broad; femur 2.39 – 2.68 times longer than patella; movable chelal finger 1.65 – 1.80 times longer than hand and 0.62 – 0.64 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 67 E, 68 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, t situated at same level as est and distal to b (Fig. 69 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 24 – 27 macrodenticles, slightly retrorse, long and pointed; movable chelal finger with 30 – 33 macrodenticles (markedly smaller than teeth on fixed chelal finger), continuous and retrorse, almost prostrate (Fig. 69 A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 67 B, 69 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 2: 2: 2: 2: 2 – 4: 4: 4: 4: 4: 2: T 2 T: 0. Sternal chaetotaxy III – XII: 12 – 15: 11 – 13: 7 – 10: 7 – 10: 7 – 10: 7 – 9: 8 – 10: 7 – 10: 0: 2. Anterior genital operculum with 9 – 10 setae, genital opening slit-like, with 13 – 18 marginal setae on each side, 38 – 45 in total (Fig. 67 F). Legs (Fig. 69 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.78 – 2.07 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.56 – 2.88 times longer than tibia. Femoropatella of leg IV 3.86 – 4.10 times longer than deep; tibia 5.90 – 7.38 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.86 – 4.43 times longer than deep (TS = 0.23 – 0.30), tarsus 13.60 – 15.60 times longer than deep and 2.39 – 2.71 times longer than metatarsus (TS = 0.23 – 0.25). Arolium slightly shorter than the claws, not divided; claws simple. Adult females (paratypes) (Figs 66 B, 67 G). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 3, II 3 – 4, III 5, IV 4 – 5; tergal chaetotaxy I – XII: 2: 2: 2: 2: 3 – 4: 4: 4: 4: 4: 2: T 2 T: 0; sternal chaetotaxy IV – XII: 11 – 13: 7 – 9: 7 – 10: 7 – 9: 7 – 9: 9: 7 – 9: 0: 2; anterior genital operculum with 9 – 10 setae, posterior margin with 12 – 14 marginal setae, 21 – 24 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.25 – 4.29 times longer than deep (TS = 0.25 – 0.32), tarsus 14.20 – 15.40 times longer than deep and 2.45 – 2.73 times longer than metatarsus (TS = 0.20 – 0.27). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Males: body length 1.71 – 2.01. Pedipalps: trochanter 0.18 – 0.22 / 0.14 – 0.16 (1.20 – 1.47), femur 0.92 – 1.02 / 0.13 – 0.15 (6.79 – 7.08), patella 0.36 – 0.41 / 0.13 – 0.15 (2.40 – 2.86), chela 1.23 – 1.35 / 0.13 – 0.17 (7.53 – 8.20), hand 0.44 – 0.50 / 0.13 – 0.17 (2.65 – 2.94), movable chelal finger length 0.79 – 0.84. Chelicera 0.46 – 0.53 / 0.19 – 0.22 (2.27 – 2.42), movable finger length 0.24 – 0.29. Carapace 0.49 – 0.53 / 0.44 – 0.49 (1.02 – 1.14). Leg I: trochanter 0.15 – 0.18 / 0.13 – 0.14 (1.07 – 1.29), femur 0.57 – 0.63 / 0.07 – 0.08 (7.13 – 8.43), patella 0.29 – 0.34 / 0.06 – 0.07 (4.14 – 5.33), tibia 0.23 – 0.26 / 0.05 – 0.06 (4.00 – 5.20), tarsus 0.61 – 0.70 / 0.05 – 0.06 (11.67 – 13.80). Leg IV: trochanter 0.22 – 0.26 / 0.14 – 0.16 (1.56 – 1.79), femoropatella 0.82 – 0.90 / 0.20 – 0.22 (3.86 – 4.10), tibia 0.59 – 0.63 / 0.08 – 0.10 (5.90 – 7.38), metatarsus 0.27 – 0.31 / 0.07 (3.86 – 4.43), tarsus 0.68 – 0.78 / 0.05 (13.60 – 15.60). Females: body length 1.81 – 2.10. Pedipalps: trochanter 0.17 – 0.20 / 0.15 – 0.16 (1.13 – 1.27), femur 0.96 – 1.05 / 0.14 – 0.16 (6.53 – 7.21), patella 0.37 – 0.41 / 0.13 – 0.17 (2.29 – 2.92), chela 1.30 – 1.42 / 0.16 – 0.20 (7.05 – 8.38), hand 0.47 – 0.53 / 0.16 – 0.20 (2.58 – 2.94), movable chelal finger length 0.81 – 0.86. Chelicera 0.50 – 0.53 / 0.22 – 0.24 (2.17 – 2.32), movable finger length 0.27 – 0.29. Carapace 0.51 – 0.56 / 0.46 – 0.53 (1.04 – 1.13). Leg I: trochanter 0.14 – 0.17 / 0.13 – 0.15 (1.00 – 1.21), femur 0.55 – 0.61 / 0.07 – 0.09 (6.78 – 8.43), patella 0.28 – 0.34 / 0.07 (4.00 – 4.86), tibia 0.23 – 0.25 / 0.05 – 0.06 (3.83 – 5.00), tarsus 0.62 – 0.70 / 0.05 – 0.06 (10.67 – 14.00). Leg IV: trochanter 0.20 – 0.25 / 0.13 – 0.15 (1.43 – 1.71), femoropatella 0.80 – 0.88 / 0.19 – 0.23 (3.82 – 4.42), tibia 0.54 – 0.61 / 0.08 – 0.10 (6.00 – 7.00), metatarsus 0.26 – 0.30 / 0.07 – 0.08 (3.25 – 4.29), tarsus 0.71 – 0.77 / 0.05 (14.20 – 15.40).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFD8FF8CFF205ECA3E10FE1E.taxon	discussion	Remarks. Tyrannochthonius multicavus sp. nov. is similar to T. harveyi in having the same number of setae on tergites I – III (2) and lacking intercalary teeth on both chelal fingers, but differs by the number of setae at the anterior margin of the carapace (6 vs. 4), a larger body size (body length min. 1.71 mm vs. max. 1.56 mm), the relative position of trichobothrium sb (sb closer to b than to st vs. midway between st and b), the number of setae on tergites IX and Ⅹ (2 vs. 4) and the proportion of movable chelal finger and chelal hand (1.65 – 1.80 vs. 2.29 times). Tyrannochthonius multicavus sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on the fixed chelal finger (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFD8FF8CFF205ECA3E10FE1E.taxon	distribution	Distribution. Known only from the Xiaohui, Weier, Da, and Qixia caves.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFC0FF86FF205EEE398EF9C6.taxon	description	Figs 70 – 75	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFC0FF86FF205EEE398EF9C6.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-GZFG- 20 - 10 - 01): China, Guizhou Province, Fenggang County, Longquan Sub-district, Guanyin Cave, under the stones in the deep zone (Temperature: 16 ° C, Humidity: 85 %) [27 ° 56 ′ 40.91 ″ N, 107 ° 44 ′ 1.99 ″ E], 684 m a. s. l., 20 August 2020, Zegang Feng, Hongru Xu & Yanmeng Hou leg. Paratypes: 2 ♂ (Ps. - MHBU-GZFG- 20 - 10 - 02 & GZFG- 20 - 10 - 03), 4 ♀ (Ps. - MHBU-GZFG- 20 - 10 - 04 – GZFG- 20 - 10 - 07), all with the same data as the holotype; 2 ♀ (Ps. - MSWU-GZSN- 20 - 11 - 01 & GZSN- 20 - 11 - 02), Sinan County, Yangjiaao Township, Guanyinshan Village, Anjialin Cave [27 ° 52 ′ 36.09 ″ N, 107 ° 54 ′ 49.02 ″ E], 641 m a. s. l., 21 August 2020, with the same collectors as the holotype; 2 ♀ (Ps. - MSWU-HBUARA # 2022 - 475 - 01 & HBUARA # 2022 - 475 - 02), Anjialin Cave, 23 July 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun & Wenlong Fan leg.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFC0FF86FF205EEE398EF9C6.taxon	etymology	Etymology. The specific name is derived from a combination of the Latin word “ multus ” and “ dentatus ”, meaning numerous and toothed, respectively, which refers to the numerous teeth on the chelal fingers.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFC0FF86FF205EEE398EF9C6.taxon	diagnosis	Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with 2 setae; tergites I – VI each with 4 setae. Pedipalps slender, femur 7.29 – 7.69 (♂), 6.59 – 6.94 (♀) times longer than broad; chela 7.40 – 7.84 (♂), 6.78 – 7.23 (♀) times longer than broad; both chelal fingers with intercalary teeth and teeth closely arranged.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFC0FF86FF205EEE398EF9C6.taxon	description	Description. Males (holotype and paratypes) (Figs 72 A, 73 A – F, 74, 75). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 73 C, 74 A): carapace 1.02 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, triangular, with 2 setae flanking base; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 9 – 12 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 74 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 73 D, 74 B): large, about as long as carapace, 2.28 – 2.32 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on dorsal side. Both fingers well provided with teeth, fixed finger with 18 – 21 teeth, distal one largest; movable finger with 18 – 21 retrorse contiguous small teeth; galea completely vestigial (Fig. 74 B). Serrula exterior with 17 – 18 and serrula interior with 11 – 12 blades. Rallum with 6 – 7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 74 E). Pedipalp (Figs 73 A, B, E, 74 D, 75 A, B): long and slender, trochanter 1.13 – 1.19, femur 7.29 – 7.69, patella 2.12 – 2.31, chela 7.40 – 7.84, hand 2.55 – 2.63 times longer than broad; femur 2.70 – 2.83 times longer than patella; movable chelal finger 1.82 – 1.94 times longer than hand and 0.63 – 0.65 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 73 E, 74 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, t situated at same level as it; est situated distal to b (Fig. 75 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced closely along the margin, larger teeth present in the middle of the row, becoming smaller distally and proximally: fixed chelal finger with 34 – 38 macrodenticles, slightly retrorse and pointed, plus 28 – 31 intercalary microdenticles, 62 – 69 in total; movable chelal finger with 36 – 40 macrodenticles (slightly smaller than teeth on fixed chelal finger), continuous, markedly retrorse and pointed, plus 17 – 18 intercalary microdenticles, 53 – 58 in total (Fig. 75 A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 73 B, 75 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 4: 4: 4: 4: 4 – 5: 5 – 6: 5: 4 – 5: T 2 T: 0. Sternal chaetotaxy III – XII: 12: 12 – 13: 8: 7 – 8: 7 – 9: 7 – 9: 7 – 9: 7: 0: 2. Anterior genital operculum with 10 setae, genital opening slit-like, with 10 – 13 marginal setae on each side, 32 – 34 in total (Fig. 73 F). Legs (Fig. 75 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.84 – 1.93 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.58 – 2.64 times longer than tibia. Femoropatella of leg IV 3.90 – 3.95 times longer than deep; tibia 6.33 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.57 – 3.71 times longer than deep (TS = 0.38 – 0.40), tarsus 14.20 times longer than deep and 2.73 – 2.84 times longer than metatarsus (TS = 0.41 – 0.42). Arolium slightly shorter than the claws, not divided; claws simple. Adult females (paratypes) (Figs 71 D, 72 B, 73 G). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5 – 6; tergal chaetotaxy I – XII: 4: 4: 4: 4: 4: 4: 4: 5 – 6: 5 – 6: 4: T 2 T: 0; sternal chaetotaxy IV – XII: 12 – 13: 8 – 10: 7 – 9: 7 – 8: 7 – 9: 7 – 9: 7: 0: 2; anterior genital operculum with 9 – 10 setae, posterior margin with 11 – 12 marginal setae, 20 – 22 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.50 – 4.00 times longer than deep (TS = 0.36 – 0.41), tarsus 12.50 – 13.17 times longer than deep and 2.68 – 2.75 times longer than metatarsus (TS = 0.38 – 0.46). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Males: body length 1.97 – 1.99. Pedipalps: trochanter 0.18 – 0.19 / 0.16 (1.13 – 1.19), femur 1.00 – 1.02 / 0.13 – 0.14 (7.29 – 7.69), patella 0.36 – 0.37 / 0.16 – 0.17 (2.12 – 2.31), chela 1.48 – 1.49 / 0.19 – 0.20 (7.40 – 7.84), hand 0.50 – 0.51 / 0.19 – 0.20 (2.55 – 2.63), movable chelal finger length 0.93 – 0.97. Chelicera 0.57 – 0.58 / 0.25 (2.28 – 2.32), movable finger length 0.30 – 0.31. Carapace 0.51 – 0.53 / 0.50 – 0.52 (1.02). Leg I: trochanter 0.15 – 0.16 / 0.14 (1.07 – 1.14), femur 0.57 – 0.58 / 0.08 (7.13 – 7.25), patella 0.30 – 0.31 / 0.07 (4.29 – 4.43), tibia 0.24 – 0.25 / 0.06 (4.00 – 4.17), tarsus 0.62 – 0.66 / 0.05 (12.40 – 13.20). Leg IV: trochanter 0.23 / 0.14 – 0.15 (1.53 – 1.64), femoropatella 0.82 – 0.83 / 0.21 (3.90 – 3.95), tibia 0.57 / 0.09 (6.33), metatarsus 0.25 – 0.26 / 0.07 (3.57 – 3.71), tarsus 0.71 / 0.05 (14.20). Females: body length 1.91 – 2.16. Pedipalps: trochanter 0.21 – 0.22 / 0.16 – 0.18 (1.17 – 1.31), femur 1.10 – 1.12 / 0.16 – 0.17 (6.59 – 6.94), patella 0.40 – 0.45 / 0.18 – 0.20 (2.22 – 2.28), chela 1.56 – 1.59 / 0.22 – 0.23 (6.78 – 7.23), hand 0.55 – 0.57 / 0.22 – 0.23 (2.39 – 2.55), movable chelal finger length 0.98 – 1.00. Chelicera 0.63 – 0.64 / 0.26 – 0.28 (2.29 – 2.42), movable finger length 0.33 – 0.34. Carapace 0.57 – 0.59 / 0.57 – 0.58 (0.98 – 1.04). Leg I: trochanter 0.17 – 0.18 / 0.14 – 0.16 (1.06 – 1.29), femur 0.61 – 0.63 / 0.08 – 0.09 (6.78 – 7.75), patella 0.32 – 0.34 / 0.08 (4.00 – 4.25), tibia 0.26 – 0.27 / 0.06 (4.33 – 4.50), tarsus 0.65 – 0.70 / 0.06 (10.83 – 11.67). Leg IV: trochanter 0.23 – 0.28 / 0.15 – 0.16 (1.53 – 1.75), femoropatella 0.91 – 0.93 / 0.22 – 0.23 (4.00 – 4.23), tibia 0.60 – 0.63 / 0.10 (6.00 – 6.30), metatarsus 0.28 – 0.29 / 0.07 – 0.09 (3.22 – 4.00), tarsus 0.75 – 0.79 / 0.06 (12.50 – 13.17).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFC0FF86FF205EEE398EF9C6.taxon	discussion	Remarks. Tyrannochthonius multidentatus sp. nov. is similar to T. maculosus sp. nov. in having intercalary teeth on both chelal fingers and a small, pointed epistome, but differs by a smaller body size (body length max. 2.16 mm vs. min. 2.22 mm), the relative position of trichobothrium sb (sb closer to b than to st vs. midway between st and b), the trait of teeth on movable chelal finger (markedly retrorse and continuous, without vestigial proximal half teeth vs. slightly retrorse and well-spaced, with vestigial proximal half teeth) and the number of blades of rallum (6 – 7 vs. 8). Tyrannochthonius multidentatus sp. nov. can be distinguished from T. chixing, T. harveyi and T. zhai by having intercalary teeth on both chelal fingers and the number of setae at the anterior margin of the carapace (6 vs. 4), from T. akaelus and T. ganshuanensis by the number of setae on tergites I – II (4 vs. 2), from T. antridraconis by a shorter palp (chela length max. 1.59 mm vs. min. 1.68 mm; palpal femur length max. 1.12 mm vs. min. 1.18 mm), more chelal finger teeth (fixed chelal finger with 62 – 69 vs. 33 – 36 teeth; movable chelal finger with 53 – 58 vs. 30 – 36 teeth), the number of blades of rallum (6 – 7 vs. 8) and coxal spines (9 – 12 vs. 12 – 15) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFC0FF86FF205EEE398EF9C6.taxon	distribution	Distribution. Known only from the Guanyin and Anjialin caves.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFCAFFB8FF2059D138E1FA82.taxon	description	Figs 76 – 80	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFCAFFB8FF2059D138E1FA82.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-HBUARA # 2022 - 502 - 01): China, Guizhou Province, Panzhou City, Zhuhai Town, Nanxing Village, Feng Cave, on the cave walls (Temperature: 16 ° C, Humidity: 80 %) [25 ° 35 ′ 45.36 ″ N, 104 ° 52 ′ 20.05 ″ E], 1615 m a. s. l., 4 August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun & Wenlong Fan leg. Paratypes: 2 ♂ (Ps. - MHBU-HBUARA # 2022 - 502 - 02 & HBUARA # 2022 - 502 - 03), 1 ♀ (Ps. - MSWU-HBUARA # 2022 - 502 - 04), all with the same data as the holotype.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFCAFFB8FF2059D138E1FA82.taxon	etymology	Etymology. Named after the village of Nanxing, near the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFCAFFB8FF2059D138E1FA82.taxon	diagnosis	Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, represented by a bump; posterior margin of carapace with 2 setae; tergites I – III each with 4 setae. Pedipalps slender, femur 8.38 – 8.44 (♂), 7.75 (♀) times longer than broad; chela 7.04 – 7.01 (♂), 7.12 (♀) times longer than broad; both chelal fingers without intercalary teeth, movable chelal finger teeth slightly closer than fixed chelal finger teeth.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFCAFFB8FF2059D138E1FA82.taxon	description	Description. Males (holotype and paratypes) (Figs 77 A, 78 A – F, 79, 80). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 78 C, 79 A): carapace 1.02 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, represented by a bump, with 2 setae flanking base; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 10 – 12 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 79 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 78 D, 79 B): large, about as long as carapace, 2.23 – 2.37 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on dorsal side. Both fingers well provided with teeth, fixed finger with 14 – 16 teeth, distal one largest; movable finger with 15 – 18 retrorse contiguous small teeth; galea completely vestigial (Fig. 79 B). Serrula exterior with 20 – 21 and serrula interior with 14 – 15 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 79 E). Pedipalp (Figs 78 A, B, E, 79 D, 80 A, B): long and slender, trochanter 1.30 – 1.35, femur 8.38 – 8.44, patella 2.24 – 2.50, chela 7.04 – 7.11, hand 2.46 – 2.56 times longer than broad; femur 2.70 – 2.85 times longer than patella; movable chelal finger 1.78 – 1.86 times longer than hand and 0.64 – 0.65 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 78 E, 79 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, t situated at same level as est and distal to b (Fig. 80 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 27 – 28 macrodenticles, long and pointed, slightly retrorse; movable chelal finger with 30 – 32 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed (Fig. 80 A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 78 B, 80 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 4: 5: 4 – 6: 5: 5 – 6: 5: 5: 4: T 2 T: 0; tergites VIII and IX each with an unpaired median seta. Sternal chaetotaxy III – XII: 14: 12 – 13: 9 – 10: 9: 9 – 10: 9: 10 – 11: 9: 0: 2. Anterior genital operculum with 10 – 12 setae, genital opening slit-like, with 13 – 16 marginal setae on each side, 38 – 41 in total (Fig. 78 F). Legs (Fig. 80 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.80 – 1.81 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.62 – 2.71 times longer than tibia. Femoropatella of leg IV 4.52 – 5.00 times longer than deep; tibia 7.08 – 8.09 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 4.33 – 4.67 times longer than deep (TS = 0.24 – 0.26), tarsus 13.86 – 14.57 times longer than deep and 2.43 – 2.49 times longer than metatarsus (TS = 0.30 – 0.31). Arolium slightly shorter than the claws, not divided; claws simple. Adult female (paratype) (Figs 76 E, 77 B, 78 G). Mostly same as males, but a little larger; with same chaetotaxy of coxae as males; tergal chaetotaxy I – XII: 4: 4: 4: 5: 6: 6: 5: 5: 6: 4: T 2 T: 0; sternal chaetotaxy IV – XII: 14: 9: 9: 10: 10: 10: 11: 0: 2; anterior genital operculum with 10 setae, posterior margin with 12 marginal setae, 22 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 4.00 times longer than deep (TS = 0.28), tarsus 15.83 times longer than deep and 2.64 times longer than metatarsus (TS = 0.41). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Males: body length 2.59 – 2.75. Pedipalps: trochanter 0.26 – 0.27 / 0.20 (1.30 – 1.35), femur 1.34 – 1.35 / 0.16 (8.38 – 8.44), patella 0.47 – 0.50 / 0.20 – 0.21 (2.24 – 2.50), chela 1.92 – 1.97 / 0.27 – 0.28 (7.04 – 7.11), hand 0.69 / 0.27 – 0.28 (2.46 – 2.56), movable chelal finger length 1.23 – 1.28. Chelicera 0.69 – 0.71 / 0.30 – 0.31 (2.23 – 2.37), movable finger length 0.39. Carapace 0.64 – 0.67 / 0.63 – 0.66 (1.02). Leg I: trochanter 0.19 – 0.22 / 0.18 (1.06 – 1.22), femur 0.83 – 0.85 / 0.10 (8.30 – 8.50), patella 0.46 – 0.47 / 0.09 – 0.10 (4.70 – 5.11), tibia 0.34 / 0.08 (4.25), tarsus 0.89 – 0.92 / 0.06 – 0.07 (13.14 – 14.83). Leg IV: trochanter 0.27 – 0.34 / 0.16 – 0.19 (1.69 – 1.79), femoropatella 1.22 – 1.25 / 0.25 – 0.27 (4.52 – 5.00), tibia 0.85 – 0.89 / 0.11 – 0.12 (7.08 – 8.09), metatarsus 0.39 – 0.42 / 0.09 (4.33 – 4.67), tarsus 0.97 – 1.02 / 0.07 (13.86 – 14.57). Female: body length 2.83. Pedipalps: trochanter 0.27 / 0.20 (1.35), femur 1.24 / 0.16 (7.75), patella 0.47 / 0.22 (2.14), chela 1.85 / 0.26 (7.12), hand 0.64 / 0.26 (2.46), movable chelal finger length 1.18. Chelicera 0.65 / 0.30 (2.17), movable finger length 0.39. Carapace 0.61 / 0.66 (0.92). Leg I: trochanter 0.19 / 0.17 (1.12), femur 0.76 / 0.10 (7.60), patella 0.41 / 0.09 (4.56), tibia 0.32 / 0.07 (4.57), tarsus 0.81 / 0.06 (13.50). Leg IV: trochanter 0.32 / 0.20 (1.60), femoropatella 1.12 / 0.25 (4.48), tibia 0.81 / 0.11 (7.36), metatarsus 0.36 / 0.09 (4.00), tarsus 0.95 / 0.06 (15.83).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFCAFFB8FF2059D138E1FA82.taxon	discussion	Remarks. Tyrannochthonius nanxingensis sp. nov. is similar to T. latus sp. nov. in lacking intercalary teeth on both chelal fingers, but differs by a shorter palp (chela length max. 1.97 mm vs. min. 2.14 mm; palpal femur length max. 1.35 mm vs. min. 1.58 mm; movable chelal finger 1.78 – 1.86 vs. 1.41 – 1.46 times longer than hand), the relative position of trichobothrium sb (sb closer to b than to st vs. closer to st), the number of setae on tergite IV (5 vs. 4) and the number of blades of rallum (8 vs. 7). Tyrannochthonius nanxingensis sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on fixed chelal fingers, from T. harveyi by the number of setae on tergites I – III (4 vs. 2) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFCAFFB8FF2059D138E1FA82.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFF4FFBFFF205A9238E1F8FE.taxon	description	Figs 81 – 85	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFF4FFBFFF205A9238E1F8FE.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-GZGY- 20 - 04 - 01): China, Guizhou Province, Huaxi District, Gaopo Township, Houzi Cave, under stones in the deep zone (Temperature: 16 ° C, Humidity: 75 %) [26 ° 16 ′ 50.73 ″ N, 106 ° 49 ′ 10.46 ″ E], 1485 m a. s. l., 16 August 2020, Zegang Feng, Hongru Xu & Yanmeng Hou leg. Paratypes: 1 ♂ (Ps. - MHBU-GZGY- 20 - 04 - 02), with the same data as the holotype; 8 ♂ (Ps. - MHBU-HBUARA # 2022 - 481 - 01 – HBUARA # 2022 - 481 - 08), 2 ♀ (Ps. - MSWU-HBUARA # 2022 - 481 - 09 & HBUARA # 2022 - 481 - 10), with the same location as the holotype, 26 July 2022, Yanmeng Hou, Lu Zhang & Jianzhou Sun leg.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFF4FFBFFF205A9238E1F8FE.taxon	etymology	Etymology. The specific name is derived from the Latin word “ oblongus ” (= rectangular), which refers to the presence of a nearly rectangular carapace.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFF4FFBFFF205A9238E1F8FE.taxon	diagnosis	Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and obtuse, represented by a bump; posterior margin of carapace with 2 setae; tergites I – II each with 2 setae. Pedipalps slender, femur 7.80 – 7.93 (♂), 7.27 – 7.80 (♀) times longer than broad; chela 7.27 (♂), 6.78 – 7.09 (♀) times longer than broad; both chelal fingers without intercalary teeth.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFF4FFBFFF205A9238E1F8FE.taxon	description	Description. Males (holotype and paratypes) (Figs 81 F, 82 A, 83 A – F, 84, 85). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 83 C, 84 A): carapace 1.05 – 1.09 times longer than broad, only slightly narrowed posteriorly resulting in rectangular appearance; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and obtuse, represented by a bump, with 2 setae flanking base; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 8 – 9 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 84 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 83 D, 84 B): large, about as long as carapace, 2.32 – 2.36 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 15 – 17 teeth, distal one largest; movable finger with 17 – 18 retrorse contiguous small teeth; galea completely vestigial (Fig. 84 B). Serrula exterior with 17 – 19 and serrula interior with 13 – 15 blades. Rallum with 7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 84 E). Pedipalp (Figs 83 A, B, E, 84 D, 85 A, B): long and slender, trochanter 1.28 – 1.33, femur 7.80 – 7.93, patella 2.41 – 2.67, chela 7.27, hand 2.59 – 2.64 times longer than broad; femur 2.78 – 2.85 times longer than patella; movable chelal finger 1.74 – 1.79 times longer than hand and 0.63 – 0.64 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 83 E, 84 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, t situated at same level as est and distal to b (Fig. 85 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 23 – 25 macrodenticles, slightly retrorse, long and pointed; movable chelal finger with 22 – 23 macrodenticles (markedly smaller than teeth on fixed chelal finger), continuous and retrorse, almost prostrate, plus 8 – 10 vestigial, rounded and contiguous basal teeth, 30 – 33 in total (Fig. 85 A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 83 B, 85 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 2: 2: 3 – 4: 4: 4: 4: 4: 4: 4 – 6: 4: T 2 T: 0. Sternal chaetotaxy III – XII: 14 – 15: 10 – 13: 8 – 9: 8 – 9: 8 – 9: 9 – 10: 10: 7 – 8: 0: 2. Anterior genital operculum with 10 – 11 setae, genital opening slit-like, with 12 – 14 marginal setae on each side, 35 – 37 in total (Fig. 83 F). Legs (Fig. 85 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.85 – 1.97 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.63 – 2.66 times longer than tibia. Femoropatella of leg IV 4.90 – 4.95 times longer than deep; tibia 7.20 – 8.00 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 4.57 – 5.14 times longer than deep (TS = 0.28), tarsus 14.83 – 15.00 times longer than deep and 2.47 – 2.81 times longer than metatarsus (TS = 0.31 – 0.32). Arolium slightly shorter than the claws, not divided; claws simple. Adult females (paratypes) (Figs 82 B, 83 G). Mostly same as males, but a little larger; with same chaetotaxy of coxae as males; tergal chaetotaxy I – XII: 2: 2: 4: 4: 4: 4: 4: 4: 4: 4: T 2 T: 0; sternal chaetotaxy IV – XII: 10 – 11: 8 – 9: 7 – 8: 7 – 9: 8 – 9: 9: 7: 0: 2; anterior genital operculum with 10 – 12 setae, posterior margin with 14 – 15 marginal setae, 24 – 27 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.88 – 4.57 times longer than deep (TS = 0.28 – 0.29), tarsus 13.83 – 17.00 times longer than deep and 2.66 – 2.68 times longer than metatarsus (TS = 0.29 – 0.30). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Males: body length 2.18 – 2.26. Pedipalps: trochanter 0.20 – 0.23 / 0.15 – 0.18 (1.28 – 1.33), femur 1.11 – 1.17 / 0.14 – 0.15 (7.80 – 7.93), patella 0.40 – 0.41 / 0.15 – 0.17 (2.41 – 2.67), chela 1.60 / 0.22 (7.27), hand 0.57 – 0.58 / 0.22 (2.59 – 2.64), movable chelal finger length 1.01 – 1.02. Chelicera 0.58 – 0.59 / 0.25 (2.32 – 2.36), movable finger length 0.32. Carapace 0.59 – 0.62 / 0.56 – 0.57 (1.05 – 1.09). Leg I: trochanter 0.17 / 0.15 – 0.16 (1.06 – 1.13), femur 0.67 – 0.72 / 0.08 – 0.09 (8.00 – 8.38), patella 0.34 – 0.39 / 0.08 (4.25 – 4.88), tibia 0.29 – 0.30 / 0.06 (4.83 – 5.00), tarsus 0.77 – 0.79 / 0.06 (12.83 – 13.17). Leg IV: trochanter 0.25 – 0.30 / 0.15 – 0.17 (1.67 – 1.76), femoropatella 0.99 – 1.03 / 0.20 – 0.21 (4.90 – 4.95), tibia 0.72 / 0.09 – 0.10 (7.20 – 8.00), metatarsus 0.32 – 0.36 / 0.07 (4.57 – 5.14), tarsus 0.89 – 0.90 / 0.06 (14.83 – 15.00). Females: body length 2.33 – 2.53. Pedipalps: trochanter 0.21 – 0.23 / 0.16 – 0.17 (1.24 – 1.44), femur 1.09 – 1.17 / 0.15 (7.27 – 7.80), patella 0.36 – 0.42 / 0.17 – 0.18 (2.12 – 2.33), chela 1.56 / 0.22 – 0.23 (6.78 – 7.09), hand 0.56 – 0.57 / 0.22 – 0.23 (2.48 – 2.55), movable chelal finger length 0.98 – 0.99. Chelicera 0.59 / 0.25 – 0.26 (2.27 – 2.36), movable finger length 0.31. Carapace 0.57 – 0.59 / 0.56 (1.02 – 1.05). Leg I: trochanter 0.17 / 0.14 – 0.15 (1.13 – 1.21), femur 0.67 / 0.08 (8.38), patella 0.37 / 0.07 (5.29), tibia 0.28 / 0.05 – 0.07 (4.00 – 5.60), tarsus 0.70 – 0.72 / 0.05 – 0.06 (12.00 – 14.00). Leg IV: trochanter 0.27 – 0.28 / 0.15 – 0.16 (1.75 – 1.80), femoropatella 0.96 – 0.97 / 0.20 (4.80 – 4.85), tibia 0.65 – 0.68 / 0.09 (7.22 – 7.56), metatarsus 0.31 – 0.32 / 0.07 – 0.08 (3.88 – 4.57), tarsus 0.83 – 0.85 / 0.05 – 0.06 (13.83 – 17.00).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFF4FFBFFF205A9238E1F8FE.taxon	discussion	Remarks. Tyrannochthonius oblongus sp. nov. is similar to T. brevispinus sp. nov. in having the same number of setae on tergites I – II (2) and lacking intercalary teeth on both chelal fingers, but differs by a larger body size (body length min. 2.18 mm vs. max. 2.12 mm; chela min. 1.56 mm vs. max. 1.43 mm; palpal femur length min. 1.09 mm vs. max. 1.00 mm; movable chelal finger 1.74 – 1.79 (♂), 1.74 – 1.75 (♀) vs. 2.05 (♂), 2.00 (♀) times longer than hand), the relative position of trichobothrium sb (sb nearer b than st vs. midway between b and st) and the number of blades of rallum (7 vs. 8). Tyrannochthonius oblongus sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on the fixed chelal finger, from T. harveyi by the number of setae at the anterior margin of the carapace (6 vs. 4) and a slender palp (chela 7.27 (♂), 6.78 – 7.09 (♀) vs. 7.50 (♂), 7.22 (♀) times longer than board, length min. 1.56 mm vs. max. 1.30 mm; palpal femur 7.80 – 7.93 (♂), 7.27 – 7.80 (♀) vs. 6.29 (♂), 5.80 (♀) times longer than board, length min. 1.09 mm vs. max. 0.88 mm) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFF4FFBFFF205A9238E1F8FE.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFFFFFB1FF205F8838E1F86E.taxon	description	Figs 86 – 90	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFFFFFB1FF205F8838E1F86E.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-HBUARA # 2022 - 489 - 01): China, Guizhou Province, Dafang County, Lvtang Township, Fengchaner Village, Daniu Cave, under the clods and on the cave walls in the deep zone (Temperature: 15 ° C, Humidity: 85 %) [27 ° 3 ′ 16.72 ″ N, 105 ° 29 ′ 9.31 ″ E], 1574 m a. s. l., 29 July 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun & Wenlong Fan leg. Paratypes: 1 ♂ (Ps. - MHBU-HBUARA # 2022 - 489 - 02), 3 ♀ (Ps. - MSWU-HBUARA # 2022 - 489 - 03 – HBUARA # 2022 - 489 - 05), all with the same data as the holotype.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFFFFFB1FF205F8838E1F86E.taxon	etymology	Etymology. The specific name is derived from a combination of the Latin word “ parcus ” and “ dentatus ”, meaning few and toothed, respectively, which refers to the low number of intercalary teeth on the fixed chelal finger.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFFFFFB1FF205F8838E1F86E.taxon	diagnosis	Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with 2 setae; tergites I – III each with 2 setae. Pedipalps slender, femur 9.07 – 9.13 (♂), 8.41 – 8.81 (♀) times longer than broad; chela 8.77 – 9.00 (♂), 7.88 – 8.42 (♀) times longer than broad; only fixed chelal finger with a few intercalary teeth.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFFFFFB1FF205F8838E1F86E.taxon	description	Description. Males (holotype and paratype) (Figs 87 A, 88 A – C, E, F, H, 89, 90). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 88 C, 89 A): carapace 0.98 – 1.02 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, triangular, with 2 setae flanking base; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 3 – 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 11 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 89 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 88 E, 89 B): large, about as long as carapace, 2.34 – 2.45 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 20 – 21 teeth, distal one largest; movable finger with 15 retrorse contiguous small teeth; galea completely vestigial (Fig. 89 B). Serrula exterior with 17 – 18 and serrula interior with 13 – 14 blades. Rallum with 7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 89 E). Pedipalp (Figs 88 A, B, H, 89 D, 90 A, B): long and slender, trochanter 1.28 – 1.41, femur 9.07 – 9.13, patella 2.50, chela 8.77 – 9.00, hand 3.32 – 3.38 times longer than broad; femur 2.72 – 2.74 times longer than patella; movable chelal finger 1.53 – 1.55 times longer than hand and 0.58 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 88 H, 89 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb midway between b and st; b and t situated subdistally, t situated distal to b and proximal to est (Fig. 90 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 26 – 28 macrodenticles, long and pointed, slightly retrorse, plus 3 – 4 intercalary microdenticles, 30 – 31 in total; movable chelal finger with 44 – 46 macrodenticles (smaller and denser than teeth on fixed chelal finger), slightly retrorse and pointed, without intercalary teeth (Fig. 90 A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 88 B, 90 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 2: 2: 2: 2 – 3: 4: 4: 4: 5: 5: 3 – 4: T 2 T: 0; tergites VIII and IX each with an unpaired median seta. Sternal chaetotaxy III – XII: 14 – 16: 12 – 13: 8: 8: 7 – 8: 7 – 8: 7 – 8: 7: 0: 2. Anterior genital operculum with 11 setae, genital opening slit-like, with 12 – 15 marginal setae on each side, 36 – 39 in total (Fig. 88 F). Legs (Fig. 90 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.90 – 1.93 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.34 – 2.44 times longer than tibia. Femoropatella of leg IV 4.48 – 4.63 times longer than deep; tibia 6.64 – 6.73 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.56 – 3.67 times longer than deep (TS = 0.28 – 0.30), tarsus 15.50 – 15.67 times longer than deep and 2.85 – 2.91 times longer than metatarsus (TS = 0.32 – 0.33). Arolium slightly shorter than the claws, not divided; claws simple. Adult females (paratypes) (Figs 87 B, 88 D, G). Mostly same as males, but a little larger; with 17 – 18 setae of carapace arranged s 4 s: 4: 4: 2: 2; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; tergal chaetotaxy I – XII: 2: 2: 2: 2: 4: 4: 4: 5: 5: 4: T 2 T: 0; sternal chaetotaxy IV – XII: 12: 8 – 9: 8: 7: 7: 7: 7: 0: 2; anterior genital operculum with 10 – 11 setae, posterior margin with 13 – 14 marginal setae, 23 – 25 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.67 – 4.00 times longer than deep (TS = 0.27 – 0.28), tarsus 15.00 – 16.33 times longer than deep and 2.72 – 2.73 times longer than metatarsus (TS = 0.31 – 0.37). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Males: body length 2.29 – 2.62. Pedipalps: trochanter 0.23 – 0.24 / 0.17 – 0.18 (1.28 – 1.41), femur 1.36 – 1.37 / 0.15 (9.07 – 9.13), patella 0.50 / 0.20 (2.50), chela 1.89 – 1.93 / 0.21 – 0.22 (8.77 – 9.00), hand 0.71 – 0.73 / 0.21 – 0.22 (3.32 – 3.38), movable chelal finger length 1.10 – 1.12. Chelicera 0.68 – 0.71 / 0.29 (2.34 – 2.45), movable finger length 0.37 – 0.39. Carapace 0.61 – 0.62 / 0.61 – 0.62 (0.98 – 1.02). Leg I: trochanter 0.19 / 0.15 – 0.17 (1.12 – 1.27), femur 0.74 – 0.79 / 0.09 (8.22 – 8.78), patella 0.39 – 0.41 / 0.08 (4.88 – 5.13), tibia 0.32 – 0.35 / 0.06 – 0.07 (5.00 – 5.33), tarsus 0.78 – 0.82 / 0.06 (13.00 – 13.67). Leg IV: trochanter 0.27 – 0.30 / 0.15 – 0.16 (1.69 – 2.00), femoropatella 1.11 – 1.12 / 0.24 – 0.25 (4.48 – 4.63), tibia 0.73 – 0.74 / 0.11 (6.64 – 6.73), metatarsus 0.32 – 0.33 / 0.09 (3.56 – 3.67), tarsus 0.93 – 0.94 / 0.06 (15.50 – 15.67). Females: body length 2.64 – 2.66. Pedipalps: trochanter 0.23 – 0.25 / 0.20 (1.15 – 1.25), femur 1.41 – 1.43 / 0.16 – 0.17 (8.41 – 8.81), patella 0.51 – 0.52 / 0.20 – 0.22 (2.36 – 2.55), chela 2.02 – 2.05 / 0.24 – 0.26 (7.88 – 8.42), hand 0.75 – 0.77 / 0.24 – 0.26 (2.96 – 3.13), movable chelal finger length 1.19. Chelicera 0.75 – 0.77 / 0.30 – 0.31 (2.48 – 2.50), movable finger length 0.41 – 0.42. Carapace 0.63 – 0.65 / 0.63 – 0.64 (0.98 – 1.03). Leg I: trochanter 0.18 – 0.21 / 0.16 – 0.17 (1.13 – 1.24), femur 0.78 – 0.80 / 0.10 (7.80 – 8.00), patella 0.42 / 0.08 – 0.09 (4.67 – 5.25), tibia 0.35 – 0.37 / 0.06 (5.83 – 6.17), tarsus 0.79 – 0.84 / 0.06 (13.17 – 14.00). Leg IV: trochanter 0.28 – 0.31 / 0.17 – 0.18 (1.65 – 1.72), femoropatella 1.08 – 1.16 / 0.24 – 0.25 (4.50 – 4.64), tibia 0.72 – 0.75 / 0.11 (6.55 – 6.82), metatarsus 0.33 – 0.36 / 0.09 (3.67 – 4.00), tarsus 0.90 – 0.98 / 0.06 (15.00 – 16.33).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFFFFFB1FF205F8838E1F86E.taxon	discussion	Remarks. Tyrannochthonius parcidentatus sp. nov. is similar to T. gracilis sp. nov. in having intercalary teeth on the fixed chelal finger only and a pointed, triangular epistome, but differs by a larger body size but shorter appendages (body length 2.29 – 2.62 (♂), 2.64 – 2.66 (♀) vs. 1.96 – 2.06 (♂), 2.12 – 2.58 (♀) mm; chela length 1.89 – 1.93 (♂), 2.02 – 2.05 (♀) vs. 2.08 – 2.11 (♂), 2.27 – 2.29 (♀) mm; movable chelal finger 1.53 – 1.55 (♂), 1.55 – 1.59 (♀) times vs. 1.63 – 1.76 (♂), 1.67 – 1.83 (♀) times longer than board), fewer fixed chelal finger teeth (30 ‒ 31 vs. 42 ‒ 45) and the number of blades of rallum (7 vs. 6). Tyrannochthonius parcidentatus sp. nov. can be distinguished from T. akaelus, T. antridraconis and T. ganshuanensis by the presence of intercalary teeth on the fixed chelal finger only, from T. harveyi by the presence of intercalary teeth on the fixed chelal finger, from T. chixing and T. zhai by the number of setae on tergites I – III (2 vs. 4) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFFFFFB1FF205F8838E1F86E.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFF9FFA8FF205F883985F86E.taxon	description	Figs 91 – 96	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFF9FFA8FF205F883985F86E.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-HBUARA # 2022 - 515 - 01): China, Guizhou Province, Dushan County, Baiquan Town, Jiabao Village, Gaoluo Cave, under stones near the cave entrance and in the deep zone (Temperature: 13 ° C, Humidity: 90 %) [25 ° 44 ′ 27.78 ″ N, 107 ° 33 ′ 47.64 ″ E], 967 m a. s. l., 11 August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun & Wenlong Fan leg. Paratypes: 5 ♂ (Ps. - MHBU-HBUARA # 2022 - 515 - 02 – HBUARA # 2022 - 515 - 06), 7 ♀ (Ps. - MSWU-HBUARA # 2022 - 515 - 07 – HBUARA # 2022 - 515 - 13), all with the same data as the holotype; 16 ♂ (Ps. - MHBU-HBUARA # 2022 - 516 - 01 – HBUARA # 2022 - 516 - 16), 13 ♀ (Ps. - MSWU-HBUARA # 2022 - 516 - 17 – HBUARA # 2022 - 516 - 29), Jiabao Village, Cangjun Cave [25 ° 44 ′ 31.99 ″ N, 107 ° 34 ′ 3.76 ″ E], 948 m a. s. l., with the same collection date and collectors as the holotype.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFF9FFA8FF205F883985F86E.taxon	etymology	Etymology. The specific name is derived from the Latin word “ pictus ” (= figured), which refers to the dorsal surface of the cheliceral palm possessing rounded whitish patches.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFF9FFA8FF205F883985F86E.taxon	diagnosis	Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, without epistome; posterior margin of carapace with 2 setae; tergite I with 2 setae, tergites III – VIII each with 4 setae. Pedipalps slender, femur 7.29 – 7.71 (♂), 7.07 – 7.29 (♀) times longer than broad; chela 8.06 – 8.50 (♂), 7.24 – 8.00 (♀) times longer than broad; both chelal fingers without intercalary teeth, movable chelal finger teeth smaller and denser than fixed chelal finger teeth.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFF9FFA8FF205F883985F86E.taxon	description	Description. Males (holotype and paratypes) (Figs 93 A, 94 A – F, 95, 96). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 94 C, 95 A): carapace 1.02 – 1.08 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; without epistome; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 8 – 9 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 95 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 94 D, 95 B): large, about as long as carapace, 2.36 – 2.52 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 13 – 16 teeth, distal one largest; movable finger with 15 – 17 retrorse contiguous small teeth; galea represented by a very slight bump on movable finger (Fig. 95 B). Serrula exterior with 17 – 20 and serrula interior with 13 – 15 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 95 E). Pedipalp (Figs 94 A, B, E, 95 D, 96 A, B): long and slender, trochanter 1.24 – 1.31, femur 7.29 – 7.71, patella 2.27 – 2.40, chela 8.06 – 8.50, hand 2.83 – 3.00 times longer than broad; femur 2.83 – 2.97 times longer than patella; movable chelal finger 1.76 – 1.79 times longer than hand and 0.62 – 0.63 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 94 E, 95 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated distal to b; est situated distal to t (Fig. 96 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 30 – 32 macrodenticles, slightly retrorse, long and pointed; movable chelal finger with 40 – 43 macrodenticles (smaller and denser than teeth on fixed chelal finger), slightly retrorse and pointed (Fig. 96 A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 94 B, 96 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 2: 2 – 4: 4: 4: 4: 4: 4: 4: 4 – 5: 4: T 2 T: 0. Sternal chaetotaxy III – XII: 12 – 15: 12 – 13: 8 – 9: 8 – 9: 9: 9: 9 – 10: 9: 0: 2. Anterior genital operculum with 9 – 11 setae, genital opening slit-like, with 15 – 17 marginal setae on each side, 41 – 46 in total (Fig. 94 F). Legs (Fig. 96 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.81 – 2.00 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.31 – 2.37 times longer than tibia. Femoropatella of leg IV 3.87 – 4.00 times longer than deep; tibia 6.50 – 7.33 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.50 – 3.63 times longer than deep (TS = 0.25 – 0.34), tarsus 13.17 – 15.00 times longer than deep and 2.59 – 2.72 times longer than metatarsus (TS = 0.24 – 0.31). Arolium slightly shorter than the claws, not divided; claws simple. Adult females (paratypes) (Figs 91 F, 92 F, 93 B, 94 G). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3 – 4, I 3, II 4, III 5, IV 5; tergal chaetotaxy I – XII: 2: 3 – 4: 4: 4: 4: 4: 4: 4: 4 – 5: 4: T 2 T: 0; sternal chaetotaxy IV – XII: 11 – 12: 8 – 10: 9: 8 – 9: 8 – 9: 8 – 10: 9 – 10: 0: 2; anterior genital operculum with 10 setae, posterior margin with 12 – 15 marginal setae, 22 – 25 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.13 – 3.86 times longer than deep (TS = 0.30 – 0.36), tarsus 11.83 – 15.40 times longer than deep and 2.72 – 2.88 times longer than metatarsus (TS = 0.25 – 0.30). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Males: body length 1.80 – 2.09. Pedipalps: trochanter 0.20 – 0.21 / 0.16 – 0.17 (1.24 – 1.31), femur 0.98 – 1.08 / 0.13 – 0.14 (7.29 – 7.71), patella 0.34 – 0.38 / 0.15 – 0.16 (2.27 – 2.40), chela 1.45 – 1.57 / 0.18 – 0.19 (8.06 – 8.50), hand 0.51 – 0.55 / 0.18 – 0.19 (2.83 – 3.00), movable chelal finger length 0.91 – 0.98. Chelicera 0.52 – 0.55 / 0.21 – 0.23 (2.36 – 2.52), movable finger length 0.28 – 0.30. Carapace 0.47 – 0.54 / 0.46 – 0.51 (1.02 – 1.08). Leg I: trochanter 0.16 – 0.17 / 0.14 – 0.15 (1.07 – 1.21), femur 0.58 – 0.65 / 0.08 (7.25 – 8.13), patella 0.32 – 0.33 / 0.07 (4.57 – 4.71), tibia 0.27 – 0.29 / 0.05 – 0.06 (3.50 – 5.40), tarsus 0.64 – 0.68 / 0.05 – 0.06 (11.17 – 13.60). Leg IV: trochanter 0.24 – 0.26 / 0.14 – 0.15 (1.67 – 1.86), femoropatella 0.87 – 0.95 / 0.22 – 0.24 (3.87 – 4.00), tibia 0.61 – 0.66 / 0.09 – 0.10 (6.50 – 7.33), metatarsus 0.28 – 0.29 / 0.08 (3.50 – 3.63), tarsus 0.73 – 0.79 / 0.05 – 0.06 (13.17 – 15.00). Females: body length 2.18 – 2.25. Pedipalps: trochanter 0.20 – 0.23 / 0.16 – 0.17 (1.25 – 1.44), femur 1.02 – 1.08 / 0.14 – 0.15 (7.07 – 7.29), patella 0.35 – 0.39 / 0.16 – 0.17 (2.19 – 2.44), chela 1.46 – 1.56 / 0.19 – 0.21 (7.24 – 8.00), hand 0.54 – 0.56 / 0.19 – 0.21 (2.67 – 2.84), movable chelal finger length 0.88 – 0.99. Chelicera 0.55 – 0.57 / 0.24 (2.29 – 2.38), movable finger length 0.29 – 0.32. Carapace 0.53 – 0.55 / 0.51 – 0.54 (1.00 – 1.04). Leg I: trochanter 0.16 / 0.14 – 0.15 (1.07 – 1.14), femur 0.58 – 0.66 / 0.08 (7.25 – 8.25), patella 0.31 – 0.33 / 0.07 – 0.08 (4.00 – 4.71), tibia 0.27 – 0.29 / 0.05 – 0.06 (4.67 – 5.80), tarsus 0.64 – 0.69 / 0.05 – 0.06 (10.67 – 13.60). Leg IV: trochanter 0.24 – 0.27 / 0.14 – 0.16 (1.56 – 1.93), femoropatella 0.86 – 0.92 / 0.22 – 0.23 (3.74 – 4.00), tibia 0.58 – 0.63 / 0.09 – 0.10 (5.90 – 7.00), metatarsus 0.25 – 0.29 / 0.07 – 0.08 (3.13 – 3.86), tarsus 0.71 – 0.79 / 0.05 – 0.06 (11.83 – 15.40).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFF9FFA8FF205F883985F86E.taxon	discussion	Remarks. Tyrannochthonius pictus sp. nov. is similar to T. planus sp. nov. in lacking an epistome on the carapace as well as intercalary teeth on both chelal fingers, but differs by having a greater number of teeth on the chelal fingers (fixed chelal finger with 30 – 32 vs. 20 – 21 teeth; movable chelal finger with 40 – 43 vs. 25 – 26 teeth) and the number of blades of rallum (8 vs. 7). Tyrannochthonius pictus sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on the fixed chelal finger, from T. harveyi by the number of setae at the anterior margin of the carapace (6 vs. 4), a slender palp (chela 8.06 – 8.50 (♂), 7.24 – 8.00 (♀) times vs. 7.50 (♂), 7.22 (♀) times longer than board, length min. 1.45 mm vs. max. 1.30 mm; palpal femur 7.29 – 7.71 (♂), 7.07 – 7.29 (♀) times vs. 6.29 (♂), 5.80 (♀) times longer than board, length min. 0.98 mm vs. max. 0.88 mm) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFF9FFA8FF205F883985F86E.taxon	distribution	Distribution. Known only from the Gaoluo and Cangjun caves.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFE0FFA1FF205F8838E1F92A.taxon	description	Figs 97 – 101	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFE0FFA1FF205F8838E1F92A.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-HBUARA # 2022 - 510 - 01): China, Guizhou Province, Ziyun County, Daying Town, Daying Village, Zharou Cave, under the stones and clods in the deep zone (Temperature: 15 ° C, Humidity: 90 %) [25 ° 29 ′ 11.07 ″ N, 106 ° 18 ′ 40.83 ″ E], 1102 m a. s. l., 8 August 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun & Wenlong Fan leg. Paratypes: 1 ♂ (Ps. - MHBU-HBUARA # 2022 - 510 - 02), 3 ♀ (Ps. - MSWU-HBUARA # 2022 - 510 - 03 – HBUARA # 2022 - 510 - 05), all with the same data as the holotype.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFE0FFA1FF205F8838E1F92A.taxon	etymology	Etymology. The specific name is derived from the Latin word “ pinguis ”, meaning fat, which refers to the enlarged abdomen present in this species.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFE0FFA1FF205F8838E1F92A.taxon	diagnosis	Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, without epistome; posterior margin of carapace with 2 setae; tergites I – IV each with 2 setae. Pedipalps slender, femur 8.07 – 8.14 (♂), 7.79 – 8.33 (♀) times longer than broad; chela 7.45 – 7.95 (♂), 7.52 (♀) times longer than broad; both chelal fingers without intercalary teeth, movable chelal finger teeth markedly smaller than fixed chelal finger teeth.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFE0FFA1FF205F8838E1F92A.taxon	description	Description. Males (holotype and paratype) (Figs 98 A, 99 A – F, 100, 101). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 99 C, 100 A): carapace 1.06 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; without epistome; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 3 – 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 8 – 9 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 100 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 99 D, 100 B): large, about as long as carapace, 2.39 – 2.41 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 10 – 11 teeth, distal one largest; movable finger with 13 – 15 retrorse contiguous small teeth; galea completely vestigial (Fig. 100 B). Serrula exterior with 19 – 22 and serrula interior with 14 – 15 blades. Rallum with 7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 100 E). Pedipalp (Figs 99 A, B, E, 100 D, 101 A, B): long and slender, trochanter 1.18 – 1.25, femur 8.07 – 8.14, patella 2.71 – 2.88, chela 7.45 – 7.95, hand 2.80 – 2.95 times longer than broad; femur 2.46 – 2.48 times longer than patella; movable chelal finger 1.61 – 1.68 times longer than hand and 0.60 – 0.62 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 99 E, 100 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, t situated at same level as est and distal to b (Fig. 101 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 22 – 23 macrodenticles, slightly retrorse, long and pointed; movable chelal finger with 31 – 33 macrodenticles (markedly smaller than teeth on fixed chelal finger), continuous and retrorse, almost prostrate (Fig. 101 A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 99 B, 101 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 2: 2: 2: 2: 4: 4: 4: 4: 4: 2: T 2 T: 0. Sternal chaetotaxy III – XII: 12 – 13: 10 – 12: 8: 7 – 8: 7: 7 – 8: 7: 7: 0: 2. Anterior genital operculum with 10 setae, genital opening slit-like, with 15 – 16 marginal setae on each side, 41 in total (Fig. 99 F). Legs (Fig. 101 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.76 – 1.92 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.82 – 2.85 times longer than tibia. Femoropatella of leg IV 4.42 – 4.64 times longer than deep; tibia 6.90 – 7.11 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 4.13 times longer than deep (TS = 0.24 – 0.30), tarsus 14.67 – 14.83 times longer than deep and 2.67 – 2.70 times longer than metatarsus (TS = 0.25 – 0.28). Arolium slightly shorter than the claws, not divided; claws simple. Adult females (paratypes) (Figs 98 B, 99 G). Mostly same as males, but a little larger; with same chaetotaxy of coxae as males; tergal chaetotaxy I – XII: 2: 2: 2: 2: 4: 4: 4: 4: 4 – 5: 2: T 2 T: 0; sternal chaetotaxy IV – XII: 11 – 12: 8: 7: 7: 7: 7: 7: 0: 2; anterior genital operculum with 10 setae, posterior margin with 10 – 12 marginal setae, 20 – 22 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.88 – 4.38 times longer than deep (TS = 0.29), tarsus 15.50 – 16.20 times longer than deep and 2.61 – 2.66 times longer than metatarsus (TS = 0.21 – 0.28). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Males: body length 1.94 – 2.13. Pedipalps: trochanter 0.20 / 0.16 – 0.17 (1.18 – 1.25), femur 1.13 – 1.14 / 0.14 (8.07 – 8.14), patella 0.46 / 0.16 – 0.17 (2.71 – 2.88), chela 1.49 – 1.51 / 0.19 – 0.20 (7.45 – 7.95), hand 0.56 / 0.19 – 0.20 (2.80 – 2.95), movable chelal finger length 0.90 – 0.94. Chelicera 0.53 – 0.55 / 0.22 – 0.23 (2.39 – 2.41), movable finger length 0.29 – 0.30. Carapace 0.53 – 0.55 / 0.50 – 0.52 (1.06). Leg I: trochanter 0.18 / 0.15 – 0.16 (1.13 – 1.20), femur 0.67 – 0.71 / 0.08 (8.38 – 8.88), patella 0.37 – 0.38 / 0.07 (5.29 – 5.43), tibia 0.27 – 0.28 / 0.06 (4.50 – 4.67), tarsus 0.77 – 0.79 / 0.06 (12.83 – 13.17). Leg IV: trochanter 0.27 – 0.28 / 0.16 – 0.17 (1.59 – 1.75), femoropatella 1.02 – 1.06 / 0.22 – 0.24 (4.42 – 4.64), tibia 0.64 – 0.69 / 0.09 – 0.10 (6.90 – 7.11), metatarsus 0.33 / 0.08 (4.13), tarsus 0.88 – 0.89 / 0.06 (14.67 – 14.83). Females: body length 1.84 – 2.24. Pedipalps: trochanter 0.19 – 0.23 / 0.15 – 0.17 (1.27 – 1.35), femur 1.09 – 1.25 / 0.14 – 0.15 (7.79 – 8.33), patella 0.45 – 0.50 / 0.15 – 0.17 (2.94 – 3.00), chela 1.43 – 1.58 / 0.19 – 0.21 (7.52), hand 0.52 – 0.61 / 0.19 – 0.21 (2.74 – 2.90), movable chelal finger length 0.88 – 0.96. Chelicera 0.52 – 0.55 / 0.22 – 0.24 (2.29 – 2.36), movable finger length 0.28 – 0.30. Carapace 0.52 – 0.54 / 0.50 – 0.51 (1.04 – 1.06). Leg I: trochanter 0.14 – 0.19 / 0.15 – 0.16 (0.93 – 1.19), femur 0.68 – 0.75 / 0.08 (8.50 – 9.38), patella 0.34 – 0.40 / 0.07 – 0.08 (4.86 – 5.00), tibia 0.27 – 0.29 / 0.05 – 0.06 (4.83 – 5.40), tarsus 0.75 – 0.81 / 0.06 (12.50 – 13.50). Leg IV: trochanter 0.27 – 0.28 / 0.14 – 0.16 (1.75 – 1.93), femoropatella 0.95 – 1.10 / 0.21 (4.52 – 5.24), tibia 0.63 – 0.70 / 0.09 – 0.10 (7.00), metatarsus 0.31 – 0.35 / 0.08 (3.88 – 4.38), tarsus 0.81 – 0.93 / 0.05 – 0.06 (15.50 – 16.20).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFE0FFA1FF205F8838E1F92A.taxon	discussion	Remarks. Tyrannochthonius pinguis sp. nov. is similar to T. umidus sp. nov. in lacking an epistome on the carapace as well as intercalary teeth on both chelal fingers, but differs by the presence of more robust chela (chela 7.45 – 7.95 (♂), 7.52 (♀) vs. 8.44 – 9.06 (♂), 7.75 – 8.33 (♀) times longer than board; movable chelal finger 1.61 – 1.68 (♂), 1.57 – 1.69 (♀) vs. 1.79 – 1.83 (♂), 1.73 – 1.92 (♀) times longer than hand), the relative position of trichobothrium sb (sb 1.67 – 1.70 vs. 1.20 – 1.22 times as far from st as from b) and the number of blades of coxal spines (8 – 9 vs. 12 – 13). Tyrannochthonius pinguis sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on the fixed chelal finger, from T. harveyi by the number of setae at the anterior margin of the carapace (6 vs. 4) and a larger body size (body length min. 1.84 mm vs. max. 1.56 mm; chelal length min. 1.43 mm vs. max. 1.30 mm; palpal femur length min. 1.09 mm vs. max. 0.88 mm) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFE0FFA1FF205F8838E1F92A.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFEDFF5BFF2059FA38E1F9B2.taxon	description	Figs 102 – 106	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFEDFF5BFF2059FA38E1F9B2.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-GZC 190801 - 01 - 01): China, Guizhou Province, Sandu County, Zhonghe Town, Sandong Village, Yanggong Cave, under the stones in the deep zone (Temperature: 13 ° C, Humidity: 90 %) [25 ° 42 ′ 48.06 ″ N, 107 ° 56 ′ 00.66 ″ E], 800 m a. s. l., 1 August 2019, Zegang Feng, Zhaoyi Li & Chen Zhang leg. Paratypes: 1 ♂ (Ps. - MHBU-GZC 190801 - 01 - 02), 2 ♀ (Ps. - MSWU-GZC 190801 - 01 - 03 & GZC 190801 - 01 - 04), all with the same data as the holotype.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFEDFF5BFF2059FA38E1F9B2.taxon	etymology	Etymology. The specific name is derived from the Latin word “ planus ”, meaning flat, which refers to the relatively straight anterior margin of the carapace.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFEDFF5BFF2059FA38E1F9B2.taxon	diagnosis	Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated and relatively straight, without epistome; posterior margin of carapace with 2 setae; tergites II – VIII each with 4 setae. Pedipalps slender, femur 7.36 – 7.38 (♂), 6.79 – 7.29 (♀) times longer than broad; chela 8.11 – 8.38 (♂), 7.68 – 7.82 (♀) times longer than broad; both chelal fingers without intercalary teeth, movable chelal finger teeth smaller and denser than fixed chelal finger teeth.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFEDFF5BFF2059FA38E1F9B2.taxon	description	Description. Males (holotype and paratype) (Figs 103 A, 104 A – F, 105, 106). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 104 C, 105 A): carapace 0.98 – 1.00 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; without epistome; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 8 – 9 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 105 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 104 D, 105 B): large, about as long as carapace, 2.45 – 2.48 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 12 – 13 teeth, distal one largest; movable finger with 11 – 13 retrorse contiguous small teeth; galea represented by a very slight bump on movable finger (Fig. 105 B). Serrula exterior with 17 – 19 and serrula interior with 13 – 14 blades. Rallum with 7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 105 E). Pedipalp (Figs 104 A, B, E, 105 D, 106 A, B): long and slender, trochanter 1.47 – 1.67, femur 7.36 – 7.38, patella 2.29 – 2.33, chela 8.11 – 8.38, hand 2.89 – 3.00 times longer than broad; femur 2.94 – 3.00 times longer than patella; movable chelal finger 1.75 times longer than hand and 0.62 – 0.63 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 104 E, 105 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated distal to b and proximal to est (Fig. 106 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 20 – 21 macrodenticles, slightly retrorse, long and pointed; movable chelal finger with 25 – 26 macrodenticles (smaller and denser than teeth on fixed chelal finger), retrorse and pointed (Fig. 106 A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 104 B, 106 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 2 – 3: 4: 4: 4: 4: 4: 4: 4: 4: 4: T 2 T: 0. Sternal chaetotaxy III – XII: 12 – 13: 11 – 12: 8: 7: 8: 9: 9: 9: 0: 2. Anterior genital operculum with 10 – 11 setae, genital opening slit-like, with 15 – 18 marginal setae on each side, 42 – 45 in total (Fig. 104 F). Legs (Fig. 106 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.81 – 1.94 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.21 – 2.24 times longer than tibia. Femoropatella of leg IV 4.05 – 4.32 times longer than deep; tibia 6.20 – 6.33 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.86 – 4.29 times longer than deep (TS = 0.26 – 0.27), tarsus 11.50 – 14.40 times longer than deep and 2.40 – 2.56 times longer than metatarsus (TS = 0.29). Arolium slightly shorter than the claws, not divided; claws simple. Adult females (paratypes) (Figs 103 B, 104 G). Mostly same as males, but a little larger; with the same chaetotaxy of coxae as males; tergal chaetotaxy I – XII: 2 – 4: 4: 4: 4: 4: 4: 4: 4: 4 – 5: 4: T 2 T: 0; sternal chaetotaxy IV – XII: 11: 8 – 10: 9: 9: 9: 8 – 10: 8 – 9: 0: 2; anterior genital operculum with 9 – 10 setae, posterior margin with 12 marginal setae, 21 – 22 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.63 – 4.00 times longer than deep (TS = 0.25 – 0.31), tarsus 11.00 – 12.17 times longer than deep and 2.36 – 2.52 times longer than metatarsus (TS = 0.29). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Males: body length 1.81 – 1.94. Pedipalps: trochanter 0.25 / 0.15 – 0.17 (1.47 – 1.67), femur 0.96 – 1.03 / 0.13 – 0.14 (7.36 – 7.38), patella 0.32 – 0.35 / 0.14 – 0.15 (2.29 – 2.33), chela 1.34 – 1.46 / 0.16 – 0.18 (8.11 – 8.38), hand 0.48 – 0.52 / 0.16 – 0.18 (2.89 – 3.00), movable chelal finger length 0.84 – 0.91. Chelicera 0.49 – 0.52 / 0.20 – 0.21 (2.45 – 2.48), movable finger length 0.27 – 0.29. Carapace 0.46 – 0.49 / 0.47 – 0.49 (0.98 – 1.00). Leg I: trochanter 0.16 / 0.12 – 0.14 (1.14 – 1.33), femur 0.56 – 0.62 / 0.07 – 0.08 (7.75 – 8.00), patella 0.31 – 0.32 / 0.06 – 0.07 (4.57 – 5.17), tibia 0.25 – 0.29 / 0.06 (4.17 – 4.83), tarsus 0.56 – 0.64 / 0.05 (11.20 – 12.80). Leg IV: trochanter 0.21 – 0.24 / 0.13 – 0.14 (1.62 – 1.71), femoropatella 0.82 – 0.89 / 0.19 – 0.22 (4.05 – 4.32), tibia 0.57 – 0.62 / 0.09 – 0.10 (6.20 – 6.33), metatarsus 0.27 – 0.30 / 0.07 (3.86 – 4.29), tarsus 0.69 – 0.72 / 0.05 – 0.06 (11.50 – 14.40). Females: body length 1.94 – 2.01. Pedipalps: trochanter 0.22 – 0.27 / 0.14 – 0.15 (1.57 – 1.80), femur 0.95 – 1.02 / 0.14 (6.79 – 7.29), patella 0.33 – 0.37 / 0.15 (2.20 – 2.47), chela 1.33 – 1.46 / 0.17 – 0.19 (7.68 – 7.82), hand 0.48 – 0.53 / 0.17 – 0.19 (2.79 – 2.82), movable chelal finger length 0.82 – 0.89. Chelicera 0.50 – 0.53 / 0.21 – 0.23 (2.30 – 2.38), movable finger length 0.27 – 0.30. Carapace 0.49 – 0.53 / 0.47 – 0.49 (1.04 – 1.08). Leg I: trochanter 0.16 / 0.14 (1.14), femur 0.58 – 0.61 / 0.07 – 0.08 (7.63 – 8.29), patella 0.31 – 0.32 / 0.07 (4.43 – 4.57), tibia 0.27 – 0.28 / 0.05 – 0.06 (4.50 – 5.60), tarsus 0.59 – 0.66 / 0.05 – 0.06 (11.00 – 11.80). Leg IV: trochanter 0.22 – 0.25 / 0.14 – 0.15 (1.57 – 1.67), femoropatella 0.82 – 0.88 / 0.20 – 0.21 (4.10 – 4.19), tibia 0.59 – 0.62 / 0.09 (6.56 – 6.89), metatarsus 0.28 – 0.29 / 0.07 – 0.08 (3.63 – 4.00), tarsus 0.66 – 0.73 / 0.06 (11.00 – 12.17).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFEDFF5BFF2059FA38E1F9B2.taxon	discussion	Remarks. Tyrannochthonius planus sp. nov. is similar to T. pictus sp. nov. in lacking an epistome on the carapace as well as intercalary teeth on both chelal fingers, but differs by having fewer teeth on both chelal fingers (fixed chelal finger with 20 – 21 vs. 30 – 32 teeth; movable chelal finger with 25 – 26 vs. 40 – 43 teeth), the trait of the teeth at movable chelal fingertip (distinctly retrorse vs. slightly retrorse) and the number of blades of rallum (7 vs. 8). Tyrannochthonius planus sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on the fixed chelal finger, from T. harveyi by the number of setae at the anterior margin of the carapace (6 vs. 4) and on tergites II – III (4 vs. 2), the presence of a slender palp (chela 8.11 – 8.38 (♂), 7.68 – 7.82 (♀) times vs. 7.50 (♂), 7.22 (♀) times longer than board, length min. 1.33 mm vs. max. 1.30 mm; palpal femur 7.36 – 7.38 (♂), 6.79 – 7.29 (♀) times vs. 6.29 (♂), 5.80 (♀) times longer than board, length min. 0.95 mm vs. max. 0.88 mm) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FFEDFF5BFF2059FA38E1F9B2.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF17FF5EFF20594238E1F80A.taxon	description	Figs 107 – 110	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF17FF5EFF20594238E1F80A.taxon	materials_examined	Type material. Holotype ♀ (Ps. - MHBU-GZJH- 19 - 26 - 35): China, Guizhou Province, Jianhe County, Yangasha Sub-district, Chuandong Village, Qilin Cave, under a stone in the deep zone (Temperature: 15 ° C, Humidity: 80 %) [26 ° 45 ′ 58.07 ″ N, 108 ° 27 ′ 42.06 ″ E], 743 m a. s. l., 13 October 2019, Zegang Feng & Lingchen Zhao leg.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF17FF5EFF20594238E1F80A.taxon	etymology	Etymology. Named after the type locality, Qilin Cave.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF17FF5EFF20594238E1F80A.taxon	diagnosis	Diagnosis (♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, represented by a bump; posterior margin of carapace with 2 setae; tergites I – VI each with 2 setae. Pedipalps slender, femur 7.72 times longer than broad; chela 7.62 times longer than broad; both chelal fingers with intercalary teeth, but present at fingertips only; chelal fingers curved in dorsal view; chemosensory setae present on dorsum of chelal hand.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF17FF5EFF20594238E1F80A.taxon	description	Description. Female (holotype), male unknown (Figs 107 – 110). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 108 C, 109 A): carapace 0.93 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, represented by a bump, with 2 setae flanking base; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 11 – 12 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 109 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 108 D, 109 B): large, about as long as carapace, 2.38 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 17 teeth, distal one largest; movable finger with 13 retrorse contiguous small teeth; galea completely vestigial (Fig. 109 B). Serrula exterior with 18 and serrula interior with 13 blades. Rallum with 8 blades, the distal one longer and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 109 E). Pedipalp (Figs 108 A, B, F, 109 D, 110 A, B): long and slender, trochanter 1.45, femur 7.72, patella 2.65, chela 7.62, hand 2.88 times longer than broad; femur 2.28 times longer than patella; movable chelal finger 1.67 times longer than hand and 0.63 times longer than chela. Setae generally long and acuminate; one distal lyrifissures present on patella (Figs 108 F, 109 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, b situated at same level as it and proximal to t (Fig. 110 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 43 macrodenticles, slightly retrorse and pointed, plus 4 intercalary microdenticles (present at fingertip only, till et), 47 in total; movable chelal finger with 35 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 4 intercalary microdenticles (present at fingertip only, till et on fixed chelal finger) and 10 vestigial, rounded and contiguous basal teeth, 49 in total (Fig. 110 A). Chelal fingers markedly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 108 B, 110 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 2: 2: 2: 2: 4: 4: 5: 4: 5: 4: T 2 T: 0. Sternal chaetotaxy IV – XII: 13: 10: 7: 8: 7: 9: 7: 0: 2. Anterior genital operculum with 11 setae, genital opening slit-like, with 14 marginal setae on each side, 25 in total (Fig. 108 E). Legs (Fig. 110 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 2.00 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.39 times longer than tibia. Femoropatella of leg IV 5.08 times longer than deep; tibia 6.23 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 4.30 times longer than deep (TS = 0.30), tarsus 14.00 times longer than deep and 2.28 times longer than metatarsus (TS = 0.27). Arolium slightly shorter than the claws, not divided; claws simple. Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Female: body length 2.41. Pedipalps: trochanter 0.32 / 0.22 (1.45), femur 1.39 / 0.18 (7.72), patella 0.61 / 0.23 (2.65), chela 1.98 / 0.26 (7.62), hand 0.75 / 0.26 (2.88), movable chelal finger length 1.25. Chelicera 0.76 / 0.32 (2.38), movable finger length 0.40. Carapace 0.68 / 0.73 (0.93). Leg I: trochanter 0.25 / 0.18 (1.39), femur 0.86 / 0.10 (8.60), patella 0.43 / 0.09 (4.78), tibia 0.38 / 0.07 (5.43), tarsus 0.91 / 0.07 (13.00). Leg IV: trochanter 0.34 / 0.19 (1.79), femoropatella 1.27 / 0.25 (5.08), tibia 0.81 / 0.13 (6.23), metatarsus 0.43 / 0.10 (4.30), tarsus 0.98 / 0.07 (14.00).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF17FF5EFF20594238E1F80A.taxon	discussion	Remarks. Male unknown. Tyrannochthonius qilinensis sp. nov. is similar to T. akaelus in having intercalary teeth on both chelal fingers and a small epistome, but differs by a larger body size (body length 2.41 vs. 2.10 mm; chela length 1.98 vs. 1.28 mm; palpal femur 7.72 vs. 6.60 times longer than board, length 1.39 vs. 0.90 mm; movable chelal finger 1.67 vs. 2.10 times longer than hand; femoropatella of leg IV length 1.27 vs. 0.79 mm), the number of setae on tergite IV (2 vs. 4) and the position of trichobothrium t (t situated distal to est vs. at same level as it / est). It is worth mentioning that T. qilinensis sp. nov. is similar to T. pandus in having a pair of curved chelal fingers, but differs by the presence of intercalary teeth on both chelal fingers and a larger body size (body length 2.41 vs. 1.67 mm; chela 7.62 vs. 7.06 times longer than board, length 1.98 vs. 1.20 mm; palpal femur 7.72 vs. 6.77 times longer than board, length 1.39 vs. 0.88 mm) (Hou et al. 2022 a). Tyrannochthonius qilinensis sp. nov. can be distinguished from T. chixing, T. zhai and T. harveyi by the presence of intercalary teeth on both chelal fingers and the number of setae at the anterior margin of the carapace (6 vs. 4), from T. antridraconis by the number of setae on the tergites I – IV (2 vs. 4), from T. ganshuanensis by the number of setae on tergite IV (2 vs. 4), a larger size (body length 2.41 (♀) vs. 1.40 – 1.80 (♀) mm; chela 7.62 (♀) vs. 6.90 – 7.30 (♀) times longer than board, length 1.98 (♀) vs. 1.21 – 1.44 (♀) mm; palpal femur 7.72 (♀) vs. 5.90 – 6.70 (♀) times longer than board, length 1.39 (♀) vs. 0.95 – 0.97 (♀) mm; movable chelal finger 1.67 (♀) vs. 2.00 – 2.20 (♀) times longer than hand) and the relative position of trichobothrium sb (sb closer to st than to b vs. closer to b) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF17FF5EFF20594238E1F80A.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF11FF53FF205F8838E1F8FE.taxon	description	Figs 111 – 115	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF11FF53FF205F8838E1F8FE.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-HBUARA # 2022 - 484 - 01): China, Guizhou Province, Dafang County, Yangchang Town, Guanjiadadong Cave, under stones within 50 – 100 m from the cave entrance (Temperature: 17 ° C, Humidity: 85 %) [27 ° 4 ′ 43.61 ″ N, 105 ° 39 ′ 10.09 ″ E], 1555 m a. s. l., 28 July 2022, Yanmeng Hou, Lu Zhang, Jianzhou Sun & Wenlong Fan leg. Paratypes: 2 ♂ (Ps. - MHBU-HBUARA # 2022 - 484 - 02 & HBUARA # 2022 - 484 - 03), 9 ♀ (Ps. - MSWU-HBUARA # 2022 - 484 - 04 – HBUARA # 2022 - 484 - 12), all with the same data as the holotype.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF11FF53FF205F8838E1F8FE.taxon	etymology	Etymology. The specific name is derived from the Latin word “ quattuor ”, meaning four, which refers to the tergites (except tergite XII) each with four setae.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF11FF53FF205F8838E1F8FE.taxon	diagnosis	Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with 2 setae; tergites I – XI each with 4 setae (except the females). Pedipalps slender, femur 7.80 – 8.40 (♂), 7.33 – 7.75 (♀) times longer than broad; chela 7.91 – 8.25 (♂), 6.89 – 7.25 (♀) times longer than broad; both chelal fingers without intercalary teeth, movable chelal finger teeth markedly smaller than fixed chelal finger teeth and distinctly retrorse.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF11FF53FF205F8838E1F8FE.taxon	description	Description. Males (holotype and paratypes) (Figs 112 A, 113 A – F, 114, 115). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 113 C, 114 A): carapace 1.02 – 1.04 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, triangular, with 2 setae flanking base; with 18 setae arranged s 4 – 5 s: 4 – 5: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 13 – 15 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 114 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 113 D, 114 B): large, about as long as carapace, 2.36 – 2.48 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 18 – 21 teeth, distal one largest; movable finger with 17 – 18 retrorse contiguous small teeth; galea completely vestigial (Fig. 114 B). Serrula exterior with 22 – 24 and serrula interior with 14 – 15 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 114 E). Pedipalp (Figs 113 A, B, E, 114 D, 115 A, B): long and slender, trochanter 1.18 – 1.28, femur 7.80 – 8.40, patella 2.44 – 2.50, chela 7.91 – 8.25, hand 2.91 – 2.95 times longer than broad; femur 2.66 – 2.80 times longer than patella; movable chelal finger 1.64 – 1.69 times longer than hand and 0.60 – 0.61 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 113 E, 114 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated distal to b and proximal to est (Fig. 115 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 31 – 35 macrodenticles, slightly retrorse, long and pointed; movable chelal finger with 49 – 51 macrodenticles (markedly smaller than teeth on fixed chelal finger), continuous and markedly retrorse (Fig. 115 A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 113 B, 115 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 4: 4: 4: 4: 4: 4: 4: 4: T 2 T: 0. Sternal chaetotaxy III – XII: 14 – 15: 13: 9 – 10: 8 – 9: 7 – 9: 8 – 9: 9: 9: 0: 2. Anterior genital operculum with 9 – 11 setae, genital opening slit-like, with 14 – 16 marginal setae on each side, 38 – 42 in total (Fig. 113 F). Legs (Fig. 115 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 2.06 – 2.13 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.23 – 2.35 times longer than tibia. Femoropatella of leg IV 4.17 – 4.79 times longer than deep; tibia 6.10 – 6.30 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.44 – 3.50 times longer than deep (TS = 0.36 – 0.39), tarsus 12.33 – 13.67 times longer than deep and 2.64 – 2.65 times longer than metatarsus (TS = 0.26 – 0.32). Arolium slightly shorter than the claws, not divided; claws simple. Adult females (paratypes) (Figs 112 B, 113 G). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5 – 6; tergal chaetotaxy I – XII: 3 – 4: 4: 4: 4: 4: 4: 4: 4: 4 – 5: 4: T 2 T: 0; sternal chaetotaxy IV – XII: 12 – 14: 9: 7 – 8: 9: 8 – 9: 9 – 10: 9: 0: 2; anterior genital operculum with 10 setae, posterior margin with 14 – 16 marginal setae, 24 – 26 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.20 – 3.33 times longer than deep (TS = 0.27 – 0.41), tarsus 13.00 – 13.50 times longer than deep and 2.53 – 2.60 times longer than metatarsus (TS = 0.33 – 0.37). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Males: body length 2.28 – 2.48. Pedipalps: trochanter 0.20 – 0.23 / 0.17 – 0.18 (1.18 – 1.28), femur 1.17 – 1.26 / 0.15 (7.80 – 8.40), patella 0.44 – 0.45 / 0.18 (2.44 – 2.50), chela 1.65 – 1.74 / 0.20 – 0.22 (7.91 – 8.25), hand 0.59 – 0.64 / 0.20 – 0.22 (2.91 – 2.95), movable chelal finger length 1.00 – 1.05. Chelicera 0.59 – 0.62 / 0.25 (2.36 – 2.48), movable finger length 0.33 – 0.34. Carapace 0.56 – 0.59 / 0.55 – 0.57 (1.02 – 1.04). Leg I: trochanter 0.16 / 0.15 – 0.16 (1.00 – 1.07), femur 0.64 – 0.68 / 0.08 – 0.09 (7.56 – 8.00), patella 0.30 – 0.33 / 0.07 (4.29 – 4.71), tibia 0.30 – 0.31 / 0.06 (5.00 – 5.17), tarsus 0.67 – 0.73 / 0.06 (11.17 – 12.17). Leg IV: trochanter 0.26 – 0.28 / 0.16 – 0.17 (1.63 – 1.65), femoropatella 0.91 – 0.96 / 0.19 – 0.23 (4.17 – 4.79), tibia 0.61 – 0.63 / 0.10 (6.10 – 6.30), metatarsus 0.28 – 0.31 / 0.08 – 0.09 (3.44 – 3.50), tarsus 0.74 – 0.82 / 0.06 (12.33 – 13.67). Females: body length 2.48 – 2.79. Pedipalps: trochanter 0.23 – 0.24 / 0.18 – 0.19 (1.26 – 1.28), femur 1.24 – 1.32 / 0.16 – 0.18 (7.33 – 7.75), patella 0.46 – 0.50 / 0.18 – 0.21 (2.38 – 2.56), chela 1.74 – 1.86 / 0.24 – 0.27 (6.89 – 7.25), hand 0.63 – 0.70 / 0.24 – 0.27 (2.59 – 2.63), movable chelal finger length 1.05 – 1.02. Chelicera 0.62 – 0.71 / 0.27 – 0.29 (2.30 – 2.45), movable finger length 0.35 – 0.38. Carapace 0.58 – 0.63 / 0.58 – 0.62 (1.00 – 1.02). Leg I: trochanter 0.17 – 0.19 / 0.16 – 0.17 (1.06 – 1.12), femur 0.66 – 0.75 / 0.09 – 0.10 (7.33 – 7.50), patella 0.35 – 0.38 / 0.07 – 0.09 (4.22 – 5.00), tibia 0.31 – 0.34 / 0.06 – 0.07 (4.86 – 5.17), tarsus 0.72 – 0.75 / 0.06 (12.00 – 12.50). Leg IV: trochanter 0.28 – 0.30 / 0.17 – 0.18 (1.65 – 1.67), femoropatella 0.94 – 1.01 / 0.22 – 0.25 (4.04 – 4.27), tibia 0.60 – 0.63 / 0.10 – 0.11 (5.45 – 6.30), metatarsus 0.30 – 0.32 / 0.09 – 0.10 (3.20 – 3.33), tarsus 0.78 – 0.81 / 0.06 (13.00 – 13.50).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF11FF53FF205F8838E1F8FE.taxon	discussion	Remarks. Tyrannochthonius quattuor sp. nov. is similar to T. latus sp. nov. in having the same number of setae on tergites I – VII (4) and lacking intercalary teeth on both chelal fingers, but differs by having a smaller body size (body length 2.28 – 2.45 (♂), 2.48 – 2.79 (♀) vs. 2.59 – 2.80 (♂), 2.91 – 3.11 (♀) mm; chela 7.91 – 8.25 (♂), 6.89 – 7.25 (♀) times vs. 6.69 – 7.13 (♂), 6.15 – 6.58 (♀) times longer than board, length 1.65 – 1.74 (♂), 1.74 – 1.86 (♀) vs. 2.14 – 2.34 (♂), 2.37 – 2.46 (♀) mm; palpal femur 7.80 – 8.40 (♂), 7.33 – 7.75 (♀) times vs. 9.29 – 9.33 (♂), 8.75 – 8.79 (♀) times longer than board, length 1.17 – 1.26 (♂), 1.24 – 1.32 (♀) vs. 1.58 – 1.68 (♂), 1.67 – 1.75 (♀) mm), the trait of teeth on the movable chelal finger (continuous and markedly retrorse vs. well-spaced and slightly retrorse) and the number of blades of rallum (8 vs. 7) and coxal spines (13 – 15 vs. 11 – 12). Tyrannochthonius quattuor sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on the fixed chelal finger, from T. harveyi by the number of setae at the anterior margin of the carapace (6 vs. 4) and on tergites I – III (4 vs. 2) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF11FF53FF205F8838E1F8FE.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF1BFF55FF205F8838E1F840.taxon	description	Figs 116 – 120	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF1BFF55FF205F8838E1F840.taxon	materials_examined	Type material. Holotype ♂ (Ps. - MHBU-GZGY- 20 - 06 - 01): China, Guizhou Province, Wudang District, Shuitian Town, Lizi Village, Da Cave, under the stones in the deep zone (Temperature: 14 ° C, Humidity: 95 %) [26 ° 42 ′ 54.76 ″ N, 106 ° 49 ′ 1.46 ″ E], 1334 m a. s. l., 17 August 2020, Zegang Feng, Hongru Xu & Yanmeng Hou leg. Paratypes: 13 ♂ (Ps. - MHBU-GZGY- 20 - 06 - 02 – GZGY- 20 - 06 - 14), 7 ♀ (Ps. - MSWU-GZGY- 20 - 06 - 15 – GZGY- 20 - 06 - 21), all with the same data as the holotype.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF1BFF55FF205F8838E1F840.taxon	etymology	Etymology. The specific name is derived from the Latin word “ umidus ”, meaning humid, which refers to the species that live in an extremely humid environment.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF1BFF55FF205F8838E1F840.taxon	diagnosis	Diagnosis (J ♀). Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, without epistome; posterior margin of carapace with 2 setae; tergites I – III each with 2 setae. Pedipalps slender, femur 7.86 – 7.93 (♂), 7.60 – 8.23 (♀) times longer than broad; chela 8.44 – 9.06 (♂), 7.75 – 8.33 (♀) times longer than broad; both chelal fingers without intercalary teeth, movable chelal finger teeth markedly smaller than fixed chelal finger teeth.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF1BFF55FF205F8838E1F840.taxon	description	Description. Males (holotype and paratypes) (Figs 116 E, 117 A, 118 A – F, 119, 120). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 118 C, 119 A): carapace 1.13 – 1.14 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; without epistome; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 12 – 13 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 119 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 118 D, 119 B): large, about as long as carapace, 2.26 – 2.41 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 9 – 11 teeth, distal one largest; movable finger with 13 – 16 retrorse contiguous small teeth; galea completely vestigial (Fig. 119 B). Serrula exterior with 20 – 22 and serrula interior with 12 – 14 blades. Rallum with 7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 119 E). Pedipalp (Figs 118 A, B, E, 119 D, 120 A, B): long and slender, trochanter 1.25 – 1.31, femur 7.86 – 7.93, patella 2.80, chela 8.44 – 9.06, hand 2.94 – 3.12 times longer than broad; femur 2.62 – 2.64 times longer than patella; movable chelal finger 1.79 – 1.83 times longer than hand and 0.63 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 118 E, 119 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to b than to st; b and t situated subdistally, t situated at same level as est and distal to b (Fig. 120 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 26 – 29 macrodenticles, slightly retrorse, long and pointed; movable chelal finger with 39 – 42 macrodenticles (markedly smaller than teeth on fixed chelal finger), continuous and retrorse, almost prostrate (Fig. 120 A). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 118 B, 120 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 2: 2: 2: 1 – 2: 4: 3 – 4: 4: 4: 4: 2: T 2 T: 0. Sternal chaetotaxy III – XII: 11 – 12: 11 – 12: 8: 7: 7 – 8: 7: 9: 8 – 9: 0: 2. Anterior genital operculum with 9 setae, genital opening slit-like, with 13 – 16 marginal setae on each side, 36 – 39 in total (Fig. 118 F). Legs (Fig. 120 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.88 – 2.06 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.41 – 2.52 times longer than tibia. Femoropatella of leg IV 4.36 – 4.60 times longer than deep; tibia 6.00 – 6.30 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 4.29 times longer than deep (TS = 0.30), tarsus 16.00 – 16.40 times longer than deep and 2.67 – 2.73 times longer than metatarsus (TS = 0.26 – 0.29). Arolium slightly shorter than the claws, not divided; claws simple. Adult females (paratypes) (Figs 117 B, 118 G). Mostly same as males, but a little larger; with same chaetotaxy of coxae as males; tergal chaetotaxy I – XII: 2: 2: 2: 2: 4: 4: 4: 4: 4: 2: T 2 T: 0; sternal chaetotaxy IV – XII: 10 – 12: 8 – 9: 7: 7 – 9: 8 – 9: 9: 8: 0: 2; anterior genital operculum with 10 – 11 setae, posterior margin with 10 – 12 marginal setae, 21 – 22 in total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.75 – 3.86 times longer than deep (TS = 0.30 – 0.33), tarsus 14.80 – 15.40 times longer than deep and 2.57 – 2.74 times longer than metatarsus (TS = 0.25 – 0.31). Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Males: body length 2.00 – 2.04. Pedipalps: trochanter 0.20 – 0.21 / 0.16 (1.25 – 1.31), femur 1.10 – 1.11 / 0.14 (7.86 – 7.93), patella 0.42 / 0.15 (2.80), chela 1.52 – 1.54 / 0.17 – 0.18 (8.44 – 9.06), hand 0.53 / 0.17 – 0.18 (2.94 – 3.12), movable chelal finger length 0.95 – 0.97. Chelicera 0.52 – 0.53 / 0.22 – 0.23 (2.26 – 2.41), movable finger length 0.27 – 0.29. Carapace 0.54 – 0.57 / 0.48 – 0.50 (1.13 – 1.14). Leg I: trochanter 0.15 – 0.17 / 0.15 (1.00 – 1.13), femur 0.64 – 0.66 / 0.08 (8.00 – 8.25), patella 0.32 – 0.34 / 0.07 (4.57 – 4.86), tibia 0.27 – 0.29 / 0.05 – 0.06 (4.83 – 5.40), tarsus 0.68 – 0.70 / 0.05 (13.60 – 14.00). Leg IV: trochanter 0.23 – 0.25 / 0.15 – 0.16 (1.44 – 1.67), femoropatella 0.92 – 0.96 / 0.20 – 0.22 (4.36 – 4.60), tibia 0.60 – 0.63 / 0.10 (6.00 – 6.30), metatarsus 0.30 / 0.07 (4.29), tarsus 0.80 – 0.82 / 0.05 (16.00 – 16.40). Females: body length 1.82 – 2.12. Pedipalps: trochanter 0.20 / 0.16 – 0.17 (1.18 – 1.25), femur 1.07 – 1.14 / 0.13 – 0.15 (7.60 – 8.23), patella 0.40 – 0.44 / 0.15 – 0.16 (2.67 – 2.75), chela 1.50 – 1.55 / 0.18 – 0.20 (7.75 – 8.33), hand 0.51 – 0.56 / 0.18 – 0.20 (2.80 – 2.83), movable chelal finger length 0.97 – 0.98. Chelicera 0.51 – 0.54 / 0.21 – 0.23 (2.35 – 2.43), movable finger length 0.28 – 0.30. Carapace 0.52 – 0.55 / 0.48 – 0.51 (1.08). Leg I: trochanter 0.14 – 0.16 / 0.13 – 0.14 (1.08 – 1.14), femur 0.61 – 0.67 / 0.08 (7.63 – 8.38), patella 0.32 – 0.33 / 0.07 (4.57 – 4.71), tibia 0.26 – 0.29 / 0.06 (4.33 – 4.83), tarsus 0.69 – 0.70 / 0.06 (11.50 – 11.67). Leg IV: trochanter 0.20 – 0.22 / 0.14 – 0.15 (1.43 – 1.47), femoropatella 0.90 – 0.95 / 0.17 – 0.18 (5.00 – 5.59), tibia 0.59 – 0.64 / 0.09 – 0.10 (6.40 – 6.56), metatarsus 0.27 – 0.30 / 0.07 – 0.08 (3.75 – 3.86), tarsus 0.74 – 0.77 / 0.05 (14.80 – 15.40).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF1BFF55FF205F8838E1F840.taxon	discussion	Remarks. Tyrannochthonius umidus sp. nov. is similar to T. pinguis sp. nov. in lacking an epistome on the carapace as well as intercalary teeth on both chelal fingers, but differs by the presence of more slender chela (chela 8.44 – 9.06 (♂), 7.75 – 8.33 (♀) vs. 7.45 – 7.95 (♂), 7.52 (♀) times longer than board; movable chelal finger 1.79 – 1.83 (♂), 1.73 – 1.92 (♀) times vs. 1.61 – 1.68 (♂), 1.57 – 1.69 (♀) times longer than hand), the relative position of trichobothrium sb (sb 1.20 – 1.22 times vs. 1.67 – 1.70 times as far from st as from b) and the number of blades of coxal spines (12 – 13 vs. 8 – 9). Tyrannochthonius umidus sp. nov. can be distinguished from T. akaelus, T. antridraconis, T. chixing, T. ganshuanensis and T. zhai by lacking intercalary teeth on the fixed chelal finger, from T. harveyi by the number of setae at the anterior margin of the carapace (6 vs. 4), the number of setae on sternites VI – Ⅹ (7 – 9 vs. 6) and a larger body size (body length min. 1.82 mm vs. max. 1.56 mm; chelal length min. 1.50 mm vs. max. 1.30 mm; palpal femur length min. 1.07 mm vs. max. 0.88 mm) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF1BFF55FF205F8838E1F840.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF05FF42FF205F8838E1FA56.taxon	description	Figs 121 – 125	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF05FF42FF205F8838E1FA56.taxon	materials_examined	Type material. Holotype ♀ (Ps. - MHBU-GZWC- 20 - 17 - 01): China, Guizhou Province, Wuchuan County, Yanshan Town, Yanshan Village, Yanshan Cave, on the cave walls in the deep zone (Temperature: 17 ° C, Humidity: 70 %) [28 ° 48 ′ 11.32 ″ N, 107 ° 53 ′ 47.07 ″ E], 951 m a. s. l., 24 August 2020, Zegang Feng, Hongru Xu & Yanmeng Hou leg. Paratype: 1 ♀ (Ps. - MSWU-GZWC- 20 - 17 - 02), with the same data as the holotype.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF05FF42FF205F8838E1FA56.taxon	etymology	Etymology. Named after the type locality, Yanshan Cave.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF05FF42FF205F8838E1FA56.taxon	diagnosis	Diagnosis (♀). Small-sized troglomorphic species with slightly elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, rounded and obtuse, represented a slight bump; posterior margin of carapace with 2 setae; tergites I – III each with 4 setae. Palpal femur 4.92 – 5.08 times longer than broad; chela 5.69 – 5.75 times longer than broad; both chelal fingers with intercalary teeth; chemosensory setae present on dorsum of chelal hand.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF05FF42FF205F8838E1FA56.taxon	description	Description. Female (holotype and paratype), male unknown (Figs 122 – 125). Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 123 C, 124 A): carapace 0.83 – 0.87 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small, rounded and obtuse, represented by a slight bump; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1 / 2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 7 – 8 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig. 124 C); intercoxal tubercle absent; without sub-oral seta. Chelicera (Figs 123 D, 124 B): large, about as long as carapace, 1.96 – 2.04 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation both dorsal and ventral side. Both fingers well provided with teeth, fixed finger with 13 teeth, distal one largest; movable finger with 12 – 13 retrorse contiguous small teeth; galea completely vestigial (Fig. 124 B). Serrula exterior with 18 and serrula interior with 13 – 14 blades. Rallum with 7 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig. 124 E). Pedipalp (Figs 123 A, B, F, 124 D, 125 A, B): long and slender, trochanter 1.17 – 1.33, femur 4.92 – 5.08, patella 1.86 – 1.93, chela 5.69 – 5.75, hand 1.81 times longer than broad; femur 2.19 – 2.35 times longer than patella; movable chelal finger 2.17 – 2.21 times longer than hand and 0.69 – 0.70 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (Figs 123 F, 124 D). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist forming a nearly straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, very close to chelal teeth; dx situated distal to et; sb closer to st than to b; b and t situated subdistally, t situated distal to b and at same level as est (Fig. 125 A). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to ist). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 29 – 30 macrodenticles, slightly retrorse and pointed, plus 14 – 15 intercalary microdenticles, 44 in total; movable chelal finger with 9 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 8 intercalary microdenticles and 20 – 22 vestigial, rounded and contiguous basal teeth, 37 – 39 in total (Fig. 125 A). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 123 B, 125 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 4: 4 – 5: 5: 5: 5 – 6: 6: 6: 5: T 2 T: 0, tergites VI – IX each with an unpaired median seta. Sternal chaetotaxy IV – XII: 12 – 13: 9 – 10: 9 – 10: 9: 10 – 11: 9 – 11: 9: 0: 2. Anterior genital operculum with 10 setae, genital opening slit-like, with 12 – 13 marginal setae on each side, 22 – 23 in total (Fig. 123 E). Legs (Fig. 125 C, D): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.87 – 2.13 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.06 – 2.19 times longer than tibia. Femoropatella of leg IV 2.59 – 2.65 times longer than deep; tibia 3.88 – 4.13 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 2.50 times longer than deep (TS = 0.33), tarsus 7.75 – 8.00 times longer than deep and 2.07 – 2.13 times longer than metatarsus (TS = 0.23 – 0.25). Arolium slightly shorter than the claws, not divided; claws simple. Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses). Females: body length 1.29 – 1.33. Pedipalps: trochanter 0.14 – 0.16 / 0.12 (1.17 – 1.33), femur 0.59 – 0.61 / 0.12 (4.92 – 5.08), patella 0.26 – 0.27 / 0.14 (1.86 – 1.93), chela 0.91 – 0.92 / 0.16 (5.69 – 5.75), hand 0.29 / 0.16 (1.81), movable chelal finger length 0.63 – 0.64. Chelicera 0.49 – 0.51 / 0.25 (1.96 – 2.04), movable finger length 0.27. Carapace 0.43 – 0.45 / 0.52 (0.83 – 0.87). Leg I: trochanter 0.12 / 0.10 – 0.11 (1.09 – 1.20), femur 0.28 – 0.32 / 0.06 (4.67 – 5.33), patella 0.15 / 0.06 (2.50), tibia 0.16 – 0.17 / 0.05 (3.20 – 3.40), tarsus 0.35 / 0.04 – 0.05 (7.00 – 8.75). Leg IV: trochanter 0.15 – 0.16 / 0.10 (1.50 – 1.60), femoropatella 0.44 – 0.45 / 0.17 (2.59 – 2.65), tibia 0.31 – 0.33 / 0.08 (3.88 – 4.13), metatarsus 0.15 / 0.06 (2.50), tarsus 0.31 – 0.32 / 0.04 (7.75 – 8.00).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF05FF42FF205F8838E1FA56.taxon	discussion	Remarks. Male unknown. Tyrannochthonius yanshanensis sp. nov. is similar to T. breviculus sp. nov. in having intercalary teeth on both chelal fingers and a small, rounded and obtuse epistome, but differs by a longer palp (chela length 0.91 – 0.92 (♀) vs. 0.78 – 0.79 (♀) mm; palpal femur 4.92 – 5.08 (♀) vs. 5.00 – 5.30 (♀) times longer than board, length 0.59 – 0.61 (♀) vs. 0.53 – 0.55 (♀) mm; movable chelal finger 2.17 – 2.21 (♀) vs. 2.04 (♀) times longer than hand), the number of setae on the tergites VIII ‒ IX (6 vs. 5), the relative position of trichobothrium sb (sb nearer st than b vs. midway between st and b) and the number of blades of rallum (7 vs. 8). Tyrannochthonius yanshanensis sp. nov. can be distinguished from T. chixing, T. harveyi and T. zhai by the presence of intercalary teeth on the movable chelal finger, from T. akaelus and T. ganshuanensis by the number of setae on tergites Ⅰ – II (4 vs. 2) and from T. antridraconis by a smaller size (body length max. 1.33 mm vs. min. 1.80 mm; chela 5.69 – 5.75 vs. 6.90 – 7.55 times longer than board, length 0.91 – 0.92 vs. 1.68 – 1.70 mm; palpal femur 4.92 – 5.08 vs. 7.10 – 7.50 times longer than board) (Mahnert 2009; Gao et al. 2018, 2020).	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
FD7487B1FF05FF42FF205F8838E1FA56.taxon	distribution	Distribution. Known only from the type locality.	en	Hou, Yanmeng, Feng, Zegang, Zhang, Feng (2023): Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5262 (1): 1-158, DOI: 10.11646/zootaxa.5262.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5262.1.1
