KEY TO THE SPECIES- GROUPS OF SAGOLA SHARP

The key is based on male specimens because most female specimens are indistinguishable based on external morphology.

1. Hind-tarsal claw enlarged relative to fore and mid-claw (Fig. 56k)................ ........................ notabilis species-group

1′. Hind-tarsal claw simple, similar to fore and mid-claw.......................................2

2(1′). Abdominal tergite V distinctly depressed, bearing processes from anterior and posterior edges and long, dense setae between processes (Fig. 54f)............... ...................... excavata species-group

2′. Abdominal tergite V simple, unmodified ............................................................... 3

3(2′). Frontal sulcus reversed Y- or V-shaped, divided with each posterior branch reaching vertexal fovea (Fig. 52p, r: arrows) ......................... furcata species-group

3′. Frontal sulcus not divided...................4

4(3′). Fore femur with semicircular depression (Fig. 4u: arrow)...................................5

4′. Fore femur without semicircular depression....................................................... 13

5(4). Head transverse (Figs. 25g, 27e) ........6

5′. Head round or bluntly triangular ........7

6(5). Anterior frontal fovea small, round, but not externally observable owing to concealment by connected frontal rostrum (Fig. 27e: arrow) ................................... .......................... flavipes species-group

6′. Anterior frontal fovea elongate, partially covered by frontal rostrum (Fig. 25g) ............................ incisa species-group

7(5′). Eye relatively more prominent due to depressed frontal gena (Fig. 23c) ......... ............................ valida species-group

7′. Eye normal and frontal gena not depressed .................................................................... 8

8(7′). Antennomeres 5–6 enlarged (Fig. 21e) ............................ parva species-group

8′. Antennomeres not enlarged.................9

9(8′). Ventral surface of head with pair of processes laterally (Fig. 19j: arrow)....... ........................... pulchra species-group

9′. Ventral surface of head without processes laterally...................................10

10(9′). Gular area of head with transverse depression and medial process (Fig. 16i: arrow).............. baiknami species-group

10′. Gular area of head with small, round depression without medial process......11

11(10′). Gular area of head with bean-shaped opening (Fig. 8j: arrow); ventral surface of head with pair of short carinae from posterior point of eyes, as long as eye (Fig. 8j).......... rugifrons species-group

11′. Gular area of head with round opening; ventral surface of head without carinae .........................................................12

12(11′). Ventral surface of head with large round depression (Fig. 12m: arrow) ................ ............................. bifida species-group

12′. Ventral surface of head with transverse depression (Fig. 4s: arrow) ................... ........................... hirtalis species-group

13(4′). Head with dense setae around posterior frontal fovea (Fig. 29a–b) ..................... ........................ helmorei species-group

13′. Head with sparse setae throughout...14

14(13′). Ventral surface of head distinctly convex, with heart-shaped depression (Fig. 35d) ......... sharpi species-group

14′. Ventral surface of head slightly convex or flat, without heart-shaped depression ................................................... 15

15(14′). Frontal sulcus deep and long, reaching between behind eye to occiput (Fig. 38j–k)........ auripila species-group

15′. Frontal sulcus not exceeding posterior margin of eye ....................................16

16(15′). Ventral surface of head with a pair of cone-shaped temporal depressions and small triangular medial processes (Fig. 50c: short arrow) .......................... ........................ owensae species-group

16′. Ventral surface of head without coneshaped temporal depressions and medial processes............................................17

17(16′). Head triangular, ventral temple of head depressed and projecting (Figs. 64f, 66n, 70y–z, 75d).......................................... 18

17′. Head round, ventral temple of head simple.................................................21

18(17). Posterior frontal fovea large and rounded, as large as eye (Fig. 75d) ........................ ............................. genalis species-group

18′. Posterior frontal fovea small.............19

19(18′). Ventral surface of neck with patch of long, dense setae anteriorly (Fig. 66o–t, 69)..... ................................................................ 20

19′. Ventral surface of neck without patch of setae anteriorly.................................. ................. moehauensis species-group

20(19). Genitalia robust and convex (Fig. 66g –l) ............................... major species-group

20′. Genitalia broad and flat (Fig. 70a–x) ...................... insignis species-group

21(17′). Ventral surface of head with process behind mouthparts bearing dense setae along posterior margin (Figs. 41f, 43d, 77b)............................................... 22

21′. Ventral surface of head without process behind mouthparts.............................24

22(21). Ventral surface of head with W-shaped process (Fig. 77b: arrow) ...................... .................... opercularis species-group

22′. Ventral surface of head with round process (Figs. 41f, 43d)..........................23

23(22′). Ventral surface of head with distinct triangular process; eye one-third length of temple (Fig. 43d)............................. ........................... strialis species-group

23′. Ventral surface of head with round process; eye large, as long as temple (Fig. 41f)..... arboricola species-group

24(21′). Anterior frontal fovea open anteriorly (Fig. 34c: arrow) ................................... ........................ gourlayi species-group

24′. Anterior frontal fovea closed anteriorly ......................................................... 25

25(24′). Ventral surface of head convex with setose gula.........................................26

25′. Ventral surface of head simple..........27

26(25). Larger, 2.3–2.9 mm total length; posterior frontal fovea elongate (Fig. 31j) .......................... rustica species-group

26′. Smaller, 1.7–1.9 mm total length; posterior frontal fovea round (Fig. 33a)..... ......................... lescheni species-group

27(25′). Antennomere 1 approximately 2.0 times longer than wide, with dull surface (Fig. 4q) ............... gilae species-group

27′. Antennomere 1 approximately 1.5– 2.0 times longer than wide, with surface at least slightly shining (Fig. n) ....................................................... 28

28(27′). Frontal rostrum rectangular, lobes continuous, reaching posterior frontal fovea (Fig. 59o)........... misella species-group

28′. Frontal rostrum round, lobes not continuous, not reaching eye if continuous ........................................................ 29

29(28′). Elytra subquadrate; hind wings reduced to small pads (Fig. 76a–b) .................... .................. flemingensis species-group

29′. Elytra rectangular; hind wings fully developed...........................................30

30(29′). Median lobe of genitalia divided vertically, major lobe partially covers minor lobe (Fig. 47f–j)..... socia species-group

30′. Median lobe of genitalia not divided (Fig. 45f–k)..... anisarthra species-group