Cyclophorus phongnhakebangensis P. V. von Oheimb, sp. nov.

urn:lsid:zoobank.org:act:FD4FE74D-974A-4C6F-B968-15A09D2B1BB0

Etymology. The name refers to the type locality of this species in Phong Nha-Ke Bang National Park.

Conchological description. Medium-sized Cyclophorus species, overall shell shape rounded, aperture typically circular. Shell height 20.5 to 26.2 mm (mean = 24.1 mm), shell breadth 26.0 to 32.8 mm (mean = 29.8 mm). Number of whorls 4.75 to 5.25. Early whorls without or with bluish colouration. Shape of body whorl slightly compressed or compressed; usually rounded. Lip reflected and simple. Lip colour ivory, white or peach. Wing-shaped projection at columellar margin absent, or present and not pronounced. Peripheral band usually dark brown. Colour pattern generally variable. Typical elements of pattern above position of peripheral band: zig-zag stripes, spiral lines and spiral bands in different shades of brown. Typical elements of pattern below position of peripheral band: spiral lines, spiral bands and transverse lines in darker shades of brown.

Type specimens. VNM032 (Fig 11a; holotype), specimen in ethanol; NHMUK 20130904/ 4; GenBank accession numbers MN153314 (COI), MN153363 (16S), MN153410 (28S); female individual; shell height 24.5 mm, shell breadth 29.6 mm, number of whorls 5. VNM035 (Fig 11b; paratype), specimen in ethanol; NHMUK 20130894/3; GenBank accession numbers MN153315 (COI), MN153364 (16S), MN153411 (28S); female individual; shell height 24.0 mm, shell breadth 29.4 mm, number of whorls 5. VNM031 (Fig 11c; paratype), specimen in ethanol; NHMUK 20130904/3; GenBank accession numbers MG720883 (COI), MG720933 (16S), MG720983 (28S); male individual; shell height 23.2 mm, shell breadth 27.3 mm, number of whorls 5.

Locality and sampling data. VNM032, VNM031: Vietnam, Quang Binh, Phong Nha-Ke Bang National Park (17.50317˚, 106.26045˚); 03 March 2012; leg. Jonathan Ablett, Hao Van Luong, Fred Naggs & Sang Van Pham. VNM035: Vietnam, Quang Binh, Phong Nha-Ke Bang National Park (17.54147˚, 106.23532˚); 03 March 2012; leg. Jonathan Ablett, Hao Van Luong, Fred Naggs & Sang Van Pham.

Diagnosis. Cyclophorus phongnhakebangensis sp. nov. can be differentiated from all herein characterised Cyclophorus species based on DNA sequence data. Diagnostic nucleotide substitutions of the cytochrome c oxidase subunit I (COI) gene: 46:G, 95:C, 133:A, 178:C, 242: C, 259:C, 277:C, 388:G, 391:A/G, 409:G, 412:C, 415:C, 433:T, 439:A, 473:A, 474:G, 475:C, 544: G, 565:C, 595:C, 637:C (numbers indicate the position in the studied COI fragment).

Remarks. This species corresponds to lineage 06 in Oheimb et al. [6]. The studied material of C. phongnhakebangensis has been found in Phong Nha-Ke Bang National Park in central Quang Binh province (Fig 4). None of the other herein characterised species is known to occur sympatrically with this species.